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<document id="D7B1EF1D9B037F254E7A972C29449282" ID-DOI="10.5281/zenodo.170590" ID-GBIF-Dataset="22b7f18d-d80f-41c9-a21b-7d4114bd504a" ID-ISSN="1175­5326" ID-Zenodo-Dep="170590" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459713953354" checkinUser="plazi" docAuthor="Hamilton, Steven W. &amp; Holzenthal, Ralph W." docDate="2005" docId="1D3D879BFFB7A14EFEEDFD42B52B1751" docLanguage="en" docName="zt00810.pdf" docOrigin="Zootaxa 810" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Polyplectropus beccus Hamilton and Holzenthal, new species" docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="E104FFE3FFBDA145FFE5FF96B1311170" masterDocTitle="Five new species of Polycentropodidae (Trichoptera) from Ecuador and Venezuela" masterLastPageNumber="14" masterPageNumber="1" pageNumber="11" updateTime="1698207473436" updateUser="plazi">
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<mods:title id="3A9A4644D8AEA423C8BECFFA0318737D">Five new species of Polycentropodidae (Trichoptera) from Ecuador and Venezuela</mods:title>
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<mods:namePart id="DF5A08865D80181A6C2C9DDC849A007C">Hamilton, Steven W.</mods:namePart>
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<mods:namePart id="ED9DBFED907F39D96C18416F8BFFEA84">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="C036C8D7C73C24110234E345236666A8">text</mods:typeOfResource>
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<mods:date id="74D1E6AB45D11DDD505A34B3BD2CC6BB">2005</mods:date>
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<mods:number id="E47EC82EDC3FA868DCCA9C69DFEFF53C">810</mods:number>
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<mods:identifier id="8E581B66FD3F3482C07F891FD591DA44" type="DOI">10.5281/zenodo.170590</mods:identifier>
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<treatment id="1D3D879BFFB7A14EFEEDFD42B52B1751" ID-DOI="http://doi.org/10.5281/zenodo.6266234" ID-GBIF-Taxon="119334117" ID-Zenodo-Dep="6266234" LSID="urn:lsid:plazi:treatment:1D3D879BFFB7A14EFEEDFD42B52B1751" httpUri="http://treatment.plazi.org/id/1D3D879BFFB7A14EFEEDFD42B52B1751" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="DD8E6506FFB7A14FFEEDFD42B05B1261" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="952B368DFFB7A14FFEEDFD42B2C9139E" blockId="10.[264,1016,724,785]" box="[264,1016,724,750]" pageId="10" pageNumber="11">
<heading id="CE6381E1FFB7A14FFEEDFD42B2C9139E" bold="true" box="[264,1016,724,750]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="A7E0EA9FFFB7A14FFEEDFD42B2C9139E" bold="true" box="[264,1016,724,750]" pageId="10" pageNumber="11">
<taxonomicName id="52944D0EFFB7A14FFEEDFD42B26A139E" authority="Hamilton and Holzenthal" authorityName="Hamilton and Holzenthal" box="[264,859,724,750]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="beccus" status="sp. nov.">
<emphasis id="A7E0EA9FFFB7A14FFEEDFD42B325139E" bold="true" box="[264,532,724,750]" italics="true" pageId="10" pageNumber="11">Polyplectropus beccus</emphasis>
Hamilton and Holzenthal
</taxonomicName>
,
<taxonomicNameLabel id="BCD357E4FFB7A14FFC8DFD42B2C9139E" box="[872,1016,724,750]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="952B368DFFB7A14FFEEDFD61B05B1261" blockId="10.[264,1016,724,785]" box="[264,362,759,785]" pageId="10" pageNumber="11">
<figureCitation id="0DAF2A08FFB7A14FFEEDFD61B07C1261" box="[264,333,759,785]" captionStart="FIGURE 5" captionStartId="11.[264,368,1148,1172]" captionTargetBox="[269,1315,292,1114]" captionTargetId="figure@11.[269,1317,290,1124]" captionTargetPageId="11" captionText="FIGURE 5. Polycentropus beccus, new species. Wings: A—forewing; B—hind wing." httpUri="https://zenodo.org/record/170595/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
,
<figureCitation id="0DAF2A08FFB7A14FFEBEFD61B05B1261" box="[347,362,759,785]" captionStart="FIGURE 6" captionStartId="11.[264,367,1742,1766]" captionText="FIGURE 6. Polycentropus beccus, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—caudal; E—phallus, lateral; F—phallic sclerite, dorsal; G—phallobase, caudal; H—VIIIth tergite, dorsal." pageId="10" pageNumber="11">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DD8E6506FFB7A14FFEEDFCD1B2791601" pageId="10" pageNumber="11" type="description">
<paragraph id="952B368DFFB7A14FFEEDFCD1B20615B9" blockId="10.[264,1325,839,1905]" pageId="10" pageNumber="11">
While wing venation indicates affinities to the New World
<taxonomicName id="52944D0EFFB7A14FFC38FCD1B5BF1210" box="[989,1166,839,864]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="A7E0EA9FFFB7A14FFC38FCD1B5BF1210" box="[989,1166,839,864]" italics="true" pageId="10" pageNumber="11">Polyplectropus</emphasis>
</taxonomicName>
(
<figureCitation id="0DAF2A08FFB7A14FFB44FCD1B5DD1211" box="[1185,1260,839,865]" captionStart="FIGURE 5" captionStartId="11.[264,368,1148,1172]" captionTargetBox="[269,1315,292,1114]" captionTargetId="figure@11.[269,1317,290,1124]" captionTargetPageId="11" captionText="FIGURE 5. Polycentropus beccus, new species. Wings: A—forewing; B—hind wing." httpUri="https://zenodo.org/record/170595/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
), the male genitalia resemble no known New or Old World species of this genus (
<figureCitation id="0DAF2A08FFB7A14FFB71FCF9B5EC12F9" box="[1172,1245,879,905]" captionStart="FIGURE 6" captionStartId="11.[264,367,1742,1766]" captionText="FIGURE 6. Polycentropus beccus, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—caudal; E—phallus, lateral; F—phallic sclerite, dorsal; G—phallobase, caudal; H—VIIIth tergite, dorsal." pageId="10" pageNumber="11">Fig. 6</figureCitation>
). The shape of the preanal appendage (
<figureCitation id="0DAF2A08FFB7A14FFD6EFC01B3E012C1" box="[651,721,919,945]" captionStart="FIGURE 6" captionStartId="11.[264,367,1742,1766]" captionText="FIGURE 6. Polycentropus beccus, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—caudal; E—phallus, lateral; F—phallic sclerite, dorsal; G—phallobase, caudal; H—VIIIth tergite, dorsal." pageId="10" pageNumber="11">Fig. 6</figureCitation>
A, B &amp; D) is unique among
<taxonomicName id="52944D0EFFB7A14FFBC7FC01B5E312C0" box="[1058,1234,919,944]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="A7E0EA9FFFB7A14FFBC7FC01B5E312C0" box="[1058,1234,919,944]" italics="true" pageId="10" pageNumber="11">Polyplectropus</emphasis>
</taxonomicName>
, and to our knowledge all known polycentropodids. The 4, heavy and truncate setae on the dorsal lobe and the bifurcate posterior process of the preanal appendage (
<figureCitation id="0DAF2A08FFB7A14FFBFDFC71B56F1571" box="[1048,1118,999,1025]" captionStart="FIGURE 6" captionStartId="11.[264,367,1742,1766]" captionText="FIGURE 6. Polycentropus beccus, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—caudal; E—phallus, lateral; F—phallic sclerite, dorsal; G—phallobase, caudal; H—VIIIth tergite, dorsal." pageId="10" pageNumber="11">Fig. 6</figureCitation>
A &amp; B) are diagnostic. The phallus has a pair of large, symmetrical spines in the phallic membranes (
<figureCitation id="0DAF2A08FFB7A14FFB1CFB99B0261521" captionStart="FIGURE 6" captionStartId="11.[264,367,1742,1766]" captionText="FIGURE 6. Polycentropus beccus, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—caudal; E—phallus, lateral; F—phallic sclerite, dorsal; G—phallobase, caudal; H—VIIIth tergite, dorsal." pageId="10" pageNumber="11">Fig. 6</figureCitation>
E), an elongate phallic sclerite (
<figureCitation id="0DAF2A08FFB7A14FFD7DFBA1B3D11521" box="[664,736,1079,1105]" captionStart="FIGURE 6" captionStartId="11.[264,367,1742,1766]" captionText="FIGURE 6. Polycentropus beccus, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—caudal; E—phallus, lateral; F—phallic sclerite, dorsal; G—phallobase, caudal; H—VIIIth tergite, dorsal." pageId="10" pageNumber="11">Fig. 6</figureCitation>
E &amp; F), and a bluntly forked process in the apicodorsal phallic membranes (
<figureCitation id="0DAF2A08FFB7A14FFD86FBC9B39B1509" box="[611,682,1119,1145]" captionStart="FIGURE 6" captionStartId="11.[264,367,1742,1766]" captionText="FIGURE 6. Polycentropus beccus, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—caudal; E—phallus, lateral; F—phallic sclerite, dorsal; G—phallobase, caudal; H—VIIIth tergite, dorsal." pageId="10" pageNumber="11">Fig. 6</figureCitation>
G). The inferior appendages (
<figureCitation id="0DAF2A08FFB7A14FFBEBFBC9B5641509" box="[1038,1109,1119,1145]" captionStart="FIGURE 6" captionStartId="11.[264,367,1742,1766]" captionText="FIGURE 6. Polycentropus beccus, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—caudal; E—phallus, lateral; F—phallic sclerite, dorsal; G—phallobase, caudal; H—VIIIth tergite, dorsal." pageId="10" pageNumber="11">Fig. 6</figureCitation>
A, C &amp; D) resemble New World
<taxonomicName id="52944D0EFFB7A14FFE21FB11B35D15D0" box="[452,620,1159,1184]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="A7E0EA9FFFB7A14FFE21FB11B35D15D0" box="[452,620,1159,1184]" italics="true" pageId="10" pageNumber="11">Polycentropus</emphasis>
</taxonomicName>
in the medial connection more than they resemble those of
<taxonomicName id="52944D0EFFB7A14FFEEDFB39B08915B8" box="[264,440,1199,1224]" class="Insecta" family="Polycentropodidae" genus="Polyplectropus" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="A7E0EA9FFFB7A14FFEEDFB39B08915B8" box="[264,440,1199,1224]" italics="true" pageId="10" pageNumber="11">Polyplectropus</emphasis>
</taxonomicName>
, which are often broadly joined.
</paragraph>
<paragraph id="952B368DFFB7A14FFEDDFB41B22F1419" blockId="10.[264,1325,839,1905]" pageId="10" pageNumber="11">
Adult. Length of forewing: male
<quantity id="526C9B68FFB7A14FFD2DFB41B2671581" box="[712,854,1239,1265]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="10" pageNumber="11" unit="mm" value="6.0" valueMax="6.5" valueMin="5.5">5.5–6.5 mm</quantity>
(n=3). Body pale brown to yellow, legs stramineous, dorsum of head and thorax brown, clothed with long, dark setae; general vestiture of forewing with fine brown to dark brown setae, with many small patches of golden setae, base of forewing with long, erect setae.
</paragraph>
<paragraph id="952B368DFFB7A14FFEDDFAE1B2791601" blockId="10.[264,1325,839,1905]" pageId="10" pageNumber="11">
Male. Genitalia as in
<figureCitation id="0DAF2A08FFB7A14FFDDFFAE1B3B514E1" box="[570,644,1399,1425]" captionStart="FIGURE 6" captionStartId="11.[264,367,1742,1766]" captionText="FIGURE 6. Polycentropus beccus, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—caudal; E—phallus, lateral; F—phallic sclerite, dorsal; G—phallobase, caudal; H—VIIIth tergite, dorsal." pageId="10" pageNumber="11">Fig. 6</figureCitation>
. Posteromesal region of tergum VIII produced posteriad, shelf­like, with apical patch of microsetae. Sternum IX in lateral view subquadrate, in ventral view quadrate; anterior margin emarginate, posterior margin slightly produced medially. Tergum IX and X membranous, very reduced. Intermediate appendage absent. Preanal appendage very short, densely setose, with patch of stout, blunt setae apically, rounded, broad basally; mesoventral process triangular, directed ventrad, apex bifurcate, beak­like, with sclerotized dorsal branch and more slender, less sclerotized, closely appressed ventral branch. Inferior appendage densely setose, elongate, dorsoventral height less than length; dorsolateral flange an elongate lobe; ventrally with short, lobate projection. Phallobase moderately elongate; apicoventral projection absent; phallic membranes with single pair of long, curved spines; phallic sclerite oval, moderately elongate, apicolateral corners acute. Subphallic sclerite absent, but subphallic semisclerotized band connecting bases of preanal appendages beneath phallus.
</paragraph>
</subSubSection>
<caption id="C1EB6605FFB6A14EFEEDFBEAB58715E3" box="[264,1206,1148,1172]" httpUri="https://zenodo.org/record/170595/files/figure.png" pageId="11" pageNumber="12" targetBox="[269,1315,292,1114]" targetPageId="11">
<paragraph id="952B368DFFB6A14EFEEDFBEAB58715E3" blockId="11.[264,1206,1148,1172]" box="[264,1206,1148,1172]" pageId="11" pageNumber="12">
<emphasis id="A7E0EA9FFFB6A14EFEEDFBEAB0BB15E4" bold="true" box="[264,394,1148,1172]" pageId="11" pageNumber="12">FIGURE 5.</emphasis>
<taxonomicName id="52944D0EFFB6A14EFE7DFBEAB3B115E3" box="[408,640,1148,1171]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="beccus" status="sp. nov.">
<emphasis id="A7E0EA9FFFB6A14EFE7DFBEAB3B115E3" box="[408,640,1148,1171]" italics="true" pageId="11" pageNumber="12">Polycentropus beccus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCD357E4FFB6A14EFD68FBEAB22015E3" box="[653,785,1148,1171]" pageId="11" pageNumber="12" rank="species">new species</taxonomicNameLabel>
. Wings: A—forewing; B—hind wing.
</paragraph>
</caption>
<subSubSection id="DD8E6506FFB6A14EFEDDFB79B52B1751" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="952B368DFFB6A14EFEDDFB79B3EF1429" blockId="11.[264,1323,1263,1569]" pageId="11" pageNumber="12">
<emphasis id="A7E0EA9FFFB6A14EFEDDFB79B5F81479" bold="true" box="[312,1225,1263,1289]" pageId="11" pageNumber="12">
<typeStatus id="4A2F882FFFB6A14EFEDDFB79B09B1479" box="[312,426,1263,1289]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="ED83761DFFB6A14EFDF5FB79B3FA1479" box="[528,715,1263,1289]" name="Venezuela" pageId="11" pageNumber="12">VENEZUELA</collectingCountry>
: T. F. Amazonas [=Estado Amazonas]:
</emphasis>
Puerto Ayacucho (
<quantity id="526C9B68FFB6A14EFE68FA81B0EB1441" box="[397,474,1303,1329]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="12" unit="km" value="40.0">40 km</quantity>
S), El Tobogan, Caño Coromoto,
<date id="E12A104DFFB6A14EFC94FA81B5061441" box="[881,1079,1303,1329]" pageId="11" pageNumber="12" value="1989-01-24">24 January 1989</date>
, malaise, P.J. Spangler, R.A. Faitoute, C.B. Barr (
<collectionCode id="F385AE48FFB6A14EFD94FAA9B3E31429" box="[625,722,1343,1369]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12">NMNH</collectionCode>
).
</paragraph>
<paragraph id="952B368DFFB6A14EFEDDFAF1B29414A1" blockId="11.[264,1323,1263,1569]" pageId="11" pageNumber="12">
<emphasis id="A7E0EA9FFFB6A14EFEDDFAF1B55F14F1" bold="true" box="[312,1134,1383,1409]" pageId="11" pageNumber="12">
<typeStatus id="4A2F882FFFB6A14EFEDDFAF1B08814F1" box="[312,441,1383,1409]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="ED83761DFFB6A14EFE5BFAF1B34B14F1" box="[446,634,1383,1409]" name="Venezuela" pageId="11" pageNumber="12">VENEZUELA</collectingCountry>
: T. F. Amazonas [=Estado Amazonas]:
</emphasis>
Puerto Ayacucho (
<quantity id="526C9B68FFB6A14EFEA3FA19B0A714D9" box="[326,406,1423,1449]" metricMagnitude="4" metricUnit="m" metricValue="2.9" pageId="11" pageNumber="12" unit="km" value="29.0">29 km</quantity>
S), Río Agua Blanca,
<date id="E12A104DFFB6A14EFD57FA19B2AE14D9" box="[690,927,1423,1449]" pageId="11" pageNumber="12" value="1987-11-17">17 November 1987</date>
, P.J. Spangler, R.A. Faitoute—
<specimenCount id="8392FD04FFB6A14EFAC7FA19B07014A1" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
(
<collectionCode id="F385AE48FFB6A14EFEB5FA21B08014A1" box="[336,433,1463,1489]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12">NMNH</collectionCode>
); same data as holotype—
<specimenCount id="8392FD04FFB6A14EFD0FFA21B20214A1" box="[746,819,1463,1489]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
(
<collectionCode id="F385AE48FFB6A14EFCA6FA21B2A814A1" box="[835,921,1463,1489]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="11" pageNumber="12">UMSP</collectionCode>
).
</paragraph>
<paragraph id="952B368DFFB6A14EFEDDFA49B52B1751" blockId="11.[264,1323,1263,1569]" pageId="11" pageNumber="12">
Etymology. From
<taxonomicName id="52944D0EFFB6A14EFDFFFA49B3891489" authority="Latin" authorityName="Latin" box="[538,696,1503,1529]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="beccus">
<emphasis id="A7E0EA9FFFB6A14EFDFFFA49B35B1488" box="[538,618,1503,1528]" italics="true" pageId="11" pageNumber="12">beccus</emphasis>
, Latin
</taxonomicName>
for beak or bill, referring to the bifurcate process of the preanal appendage that resembles a birds beak in lateral aspect.
</paragraph>
</subSubSection>
</treatment>
</document>
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