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243 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.877.36698" ID-GBIF-Dataset="a4fd240c-caf7-41f1-b507-d9ffc15cdfbe" ID-PMC="PMC6775175" ID-Pensoft-Pub="1313-2970-877-31" ID-Pensoft-UUID="5BE094A8A1645289A9A3A154C4AF6293" ID-PubMed="31592220" ID-ZooBank="ABBC811D2A6D46159ECF0F44933BFBE1" ModsDocID="1313-2970-877-31" checkinTime="1570026182685" checkinUser="pensoft" docAuthor="Dayrat, Benoit, Goulding, Tricia C., Khalil, Munawar, Comendador, Joseph, Xuan, Qu ảng Ngo, Tan, Siong Kiat & Tan, Shau Hwai" docDate="2019" docId="D819240C8F695F69953D8549D3F5CC14" docLanguage="en" docName="ZooKeys 877: 31-80" docOrigin="ZooKeys 877" docSource="http://dx.doi.org/10.3897/zookeys.877.36698" docTitle="Laspionchis Dayrat & Goulding, gen. nov." docType="treatment" docUuid="47CA237B-3E0F-49BF-A866-C551E979A236" docUuidSource="ZooBank" docVersion="7" id="5BE094A8A1645289A9A3A154C4AF6293" lastPageNumber="42" masterDocId="5BE094A8A1645289A9A3A154C4AF6293" masterDocTitle="A new genus of air-breathing marine slugs from South-East Asia (Gastropoda, Pulmonata, Onchidiidae)" masterLastPageNumber="80" masterPageNumber="31" pageNumber="41" updateTime="1668167836593" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus of air-breathing marine slugs from South-East Asia (Gastropoda, Pulmonata, Onchidiidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dayrat, Benoit</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Goulding, Tricia C.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Khalil, Munawar</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Comendador, Joseph</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Xuan, Qu ảng Ngo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tan, Siong Kiat</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tan, Shau Hwai</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>877</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>31</mods:start>
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<mods:end>80</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.877.36698</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.877.36698</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-877-31</mods:identifier>
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<mods:identifier type="ZooBank">ABBC811D2A6D46159ECF0F44933BFBE1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">5BE094A8A1645289A9A3A154C4AF6293</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159619267" LSID="urn:lsid:zoobank.org:act:47CA237B-3E0F-49BF-A866-C551E979A236" httpUri="http://treatment.plazi.org/id/D819240C8F695F69953D8549D3F5CC14" lastPageId="11" lastPageNumber="42" pageId="10" pageNumber="41">
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<subSubSection pageId="10" pageNumber="41" type="nomenclature">
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<paragraph pageId="10" pageNumber="41">
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Genus
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<taxonomicName LSID="D819240C-8F69-5F69-953D-8549D3F5CC14" authority="Dayrat & Goulding" class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis" order="Systellommatophora" pageId="10" pageNumber="41" phylum="Mollusca" rank="genus">Laspionchis Dayrat & Goulding</taxonomicName>
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<taxonomicNameLabel pageId="10" pageNumber="41">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="41" type="type species">
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<paragraph pageId="10" pageNumber="41">Type species.</paragraph>
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<paragraph pageId="10" pageNumber="41">
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis boucheti" order="Systellommatophora" pageId="10" pageNumber="41" phylum="Mollusca" rank="species" species="boucheti">
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<emphasis italics="true" pageId="10" pageNumber="41">Laspionchis boucheti</emphasis>
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</taxonomicName>
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, designated here.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="41" type="etymology">
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<paragraph pageId="10" pageNumber="41">Etymology.</paragraph>
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<paragraph pageId="10" pageNumber="41">
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Combination of
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<emphasis italics="true" pageId="10" pageNumber="41">
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<normalizedToken originalValue="láspi">laspi</normalizedToken>
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</emphasis>
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, a Greek word meaning mud, and
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<emphasis italics="true" pageId="10" pageNumber="41">onchis</emphasis>
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, a word derived from the Greek
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<emphasis italics="true" pageId="10" pageNumber="41">
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<normalizedToken originalValue="ὁ">ὁ</normalizedToken>
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<normalizedToken originalValue="ὂγκος">ὂγκος</normalizedToken>
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</emphasis>
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(mass, tumor) and used in the past for onchidiid slugs.
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis" order="Systellommatophora" pageId="10" pageNumber="41" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="41">Laspionchis</emphasis>
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</taxonomicName>
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conveniently refers to those onchidiid species that always live on mud and are covered with a thin layer of mud.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="41" type="gender">
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<paragraph pageId="10" pageNumber="41">Gender.</paragraph>
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<paragraph pageId="10" pageNumber="41">
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Gender masculine of
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<emphasis italics="true" pageId="10" pageNumber="41">onchis</emphasis>
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(ICZN Art. 30.1.1), a word derived from the masculine Greek word
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<emphasis italics="true" pageId="10" pageNumber="41">
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<normalizedToken originalValue="ὁ">ὁ</normalizedToken>
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<normalizedToken originalValue="ὂγκος">ὂγκος</normalizedToken>
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</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="41" type="diagnosis">
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<paragraph pageId="10" pageNumber="41">Diagnosis.</paragraph>
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<paragraph pageId="10" pageNumber="41">
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Body not flattened. No dorsal gills. Dorsal eyes present on notum. Retractable, central papilla (usually with four dorsal eyes) present, often raised above dorsal surface. Eyes at tip of short ocular tentacles. Male opening below right ocular tentacle (or below it and very slightly to its left). No transversal protuberance on oral lobes. Foot wide. Pneumostome median, on ventral hyponotum. Intestinal loops exactly between types I and II (with a transitional loop on average descending at 6
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<normalizedToken originalValue="o’clock">o'clock</normalizedToken>
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). Rectal gland absent. Accessory penial gland present with a hollow spine and a muscular sac. Penis with hooks: numerous, densely arranged next to each other, and pointed.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="42" pageId="10" pageNumber="41" type="remarks">
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<paragraph pageId="10" pageNumber="41">Remarks.</paragraph>
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<paragraph lastPageId="11" lastPageNumber="42" pageId="10" pageNumber="41">
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No external diagnostic feature unambiguously distinguishes
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis" order="Systellommatophora" pageId="10" pageNumber="41" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="41">Laspionchis</emphasis>
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</taxonomicName>
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from other onchidiid genera. Externally,
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis" order="Systellommatophora" pageId="10" pageNumber="41" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="41">Laspionchis</emphasis>
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</taxonomicName>
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slugs are especially difficult to distinguish from
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paromoionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paromoionchis" order="Systellommatophora" pageId="10" pageNumber="41" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="41">Paromoionchis</emphasis>
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</taxonomicName>
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slugs, which live in the same habitat (mud surface) and are often found together at the exact same sites. Also, for a non-expert,
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis" order="Systellommatophora" pageId="10" pageNumber="41" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="41">Laspionchis</emphasis>
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</taxonomicName>
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slugs could easily be confused with
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronina" order="Systellommatophora" pageId="10" pageNumber="41" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="41">Peronina</emphasis>
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</taxonomicName>
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or
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="10" pageNumber="41" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="41">Onchidium</emphasis>
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</taxonomicName>
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slugs, although those are characterized by distinctive, external features. However,
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis" order="Systellommatophora" pageId="10" pageNumber="41" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="41">Laspionchis</emphasis>
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</taxonomicName>
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is characterized by a unique combination of internal and external characters: no dorsal gills, male opening
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<pageBreakToken pageId="11" pageNumber="42" start="start">below</pageBreakToken>
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the right eye tentacle (or below it and very slightly to its left), no rectal gland, intestinal loops between types I and II (i.e., with a transitional loop on average oriented at 6
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<normalizedToken originalValue="o’clock">o'clock</normalizedToken>
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), accessory penial gland present with a muscular sac, penis with numerous, pointed hooks densely arranged next to each other. According to our data, any onchidiid slug with this combination of characters belongs to
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis" order="Systellommatophora" pageId="11" pageNumber="42" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="11" pageNumber="42">Laspionchis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="11" pageNumber="42">
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Intestinal loops between types I and II, with a transitional loop on average oriented at 6
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<normalizedToken originalValue="o’clock">o'clock</normalizedToken>
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, could almost be regarded as diagnostic of
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis" order="Systellommatophora" pageId="11" pageNumber="42" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="11" pageNumber="42">Laspionchis</emphasis>
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</taxonomicName>
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slugs, acknowledging the existence of variation (both intra-specific and inter-specific). Indeed, in
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis" order="Systellommatophora" pageId="11" pageNumber="42" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="11" pageNumber="42">Laspionchis</emphasis>
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</taxonomicName>
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slugs, the transitional loop is normally oriented at 6
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<normalizedToken originalValue="o’clock">o'clock</normalizedToken>
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, even though, strictly speaking, its orientation actually varies between 5 and 7
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<normalizedToken originalValue="o’clock">o'clock</normalizedToken>
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(
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Intestinal loops, dorsal view A, B Type I C, D Type II E-H Between types I and II. Small black arrows show the direction of the intestinal transport. The clockwise loop is in blue. The counterclockwise loop is in yellow. The transitional loop (between clockwise and counterclockwise loops) is in green. Red arrows indicate the orientation of the transitional loop. A Type I, with a transitional loop oriented at 3 o'clock, redrawn from Plate (1893: pl. 8, fig. 29) B Type I, with a transitional loop oriented at 3 o'clock, Wallaconchis sinanui (from Goulding et al. 2018: fig. 8 D) C Type II, with a transitional loop oriented at 9 o'clock, redrawn from Plate (1893: pl. 8, fig. 31) D Type II, with a transitional loop oriented at 9 o'clock, Paromoionchis tumidus (from Dayrat et al. 2019: fig. 12 A) E Between types I and II, with a descending transitional loop at 7 o'clock, holotype, Laspionchis boucheti, Australia, Northern Territory, [1688 H] (NTM P. 57614) F Between types I and II, with a descending transitional loop at 6 o'clock, L. boucheti, Australia, Northern Territory, [1681] (NTM P. 57612) G Between types I and II, with a descending transitional loop at 6 o'clock, L. boucheti, Vietnam, [5610] (ITBZC IM 00017) H Between types I and II, with a descending transitional loop at 5 o'clock, L. boucheti, Australia, Queensland, [2612] (MTQ). Scale bars: 1 mm (B), 2 mm (D, E), 5 mm (F-H)." figureDoi="10.3897/zookeys.877.36698.figure1" httpUri="https://binary.pensoft.net/fig/343041" pageId="11" pageNumber="42" tableDoi="10.3897/zookeys.877.36698.figure1">Fig. 1</figureCitation>
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). The intestinal loops of some individuals of other species can sometimes be characterized by a transitional loop oriented within that same range (between 5 and 7
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<normalizedToken originalValue="o’clock">o'clock</normalizedToken>
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), such as in species with intestinal loops of type I and a transitional loop oriented from 3 to 6
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<normalizedToken originalValue="o’clock">o'clock</normalizedToken>
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(as in
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="11" pageNumber="42" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="11" pageNumber="42">Wallaconchis</emphasis>
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</taxonomicName>
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, see
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.763.21252" author="Goulding, TC" journalOrPublisher="ZooKeys" pageId="41" pageNumber="72" pagination="1 - 111" refId="B15" refString="Goulding, TC, Khalil, M, Tan, SH, Dayrat, B, 2018a. Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda: Pulmonata: Onchidiidae). ZooKeys 763: 1 - 111" title="Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda: Pulmonata: Onchidiidae)." url="https://doi.org/10.3897/zookeys.763.21252" volume="763" year="2018 a">Goulding et al. 2018a</bibRefCitation>
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), and in species with intestinal loops of type II and a transitional loop oriented from 6 to 9
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<normalizedToken originalValue="o’clock">o'clock</normalizedToken>
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(as in
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paromoionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paromoionchis" order="Systellommatophora" pageId="11" pageNumber="42" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="11" pageNumber="42">Paromoionchis</emphasis>
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</taxonomicName>
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, see
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<bibRefCitation DOI="https://doi.org/10.5852/ejt.2019.500" author="Dayrat, B" journalOrPublisher="European Journal of Taxonomy" pageId="41" pageNumber="72" pagination="1 - 77" refId="B13" refString="Dayrat, B, Goulding, TC, Khalil, M, Apte, D, Bourke, AJ, Comendador, J, Tan, SH, 2019. A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1 - 77" title="A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae)." url="https://doi.org/10.5852/ejt.2019.500" volume="500" year="2019">Dayrat et al. 2019</bibRefCitation>
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). However, the important difference here is that a transitional loop oriented at 6
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<normalizedToken originalValue="o’clock">o'clock</normalizedToken>
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is the norm in
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<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Laspionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laspionchis" order="Systellommatophora" pageId="11" pageNumber="42" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="11" pageNumber="42">Laspionchis</emphasis>
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</taxonomicName>
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, while it is not the norm in those species from other genera.
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</paragraph>
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<paragraph pageId="11" pageNumber="42">
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A new generic name is needed because no existing name applies to the clade described here. Based on the examination of all the type specimens available in
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<taxonomicName family="Onchidiidae" lsidName="" pageId="11" pageNumber="42" rank="family">Onchidiidae</taxonomicName>
|
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(especially those of all the type species), a careful study of all the original descriptions (especially when no type specimens were available), and our ongoing taxonomic revision of every genus of the family (
|
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.636.8879" author="Dayrat, B" journalOrPublisher="ZooKeys" pageId="40" pageNumber="71" pagination="1 - 40" refId="B10" refString="Dayrat, B, Goulding, TC, Apte, D, Bhave, V, Comendador, J, Ngo, XQ, Tan, SK, Tan, SH, 2016. Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1 - 40" title="Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae)." url="https://doi.org/10.3897/zookeys.636.8879" volume="636" year="2016">Dayrat et al. 2016</bibRefCitation>
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,
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<bibRefCitation DOI="https://doi.org/10.1080/00222933.2017.1347297" author="Dayrat, B" journalOrPublisher="Journal of Natural History" pageId="40" pageNumber="71" pagination="1851 - 1897" refId="B11" refString="Dayrat, B, Goulding, TC, Apte, D, Bhave, V, Ngo, XQ, 2017. A new genus and four new species of onchidiid slugs from South-East Asia (Mollusca: Gastropoda: Pulmonata: Onchidiidae). Journal of Natural History 51: 1851 - 1897" title="A new genus and four new species of onchidiid slugs from South-East Asia (Mollusca: Gastropoda: Pulmonata: Onchidiidae)." url="https://doi.org/10.1080/00222933.2017.1347297" volume="51" year="2017">2017</bibRefCitation>
|
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,
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<bibRefCitation author="Dayrat, B" journalOrPublisher="Raffles Bulletin of Zoology" pageId="41" pageNumber="72" pagination="814 - 837" refId="B12" refString="Dayrat, B, Goulding, TC, Khalil, M, Lozouet, P, Tan, SH, 2018. Systematic revision one clade at a time: A new genus of onchidiid slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae). Raffles Bulletin of Zoology 66: 814 - 837" title="Systematic revision one clade at a time: A new genus of onchidiid slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae)." volume="66" year="2018">2018</bibRefCitation>
|
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,
|
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|
|
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2019.500" author="Dayrat, B" journalOrPublisher="European Journal of Taxonomy" pageId="41" pageNumber="72" pagination="1 - 77" refId="B13" refString="Dayrat, B, Goulding, TC, Khalil, M, Apte, D, Bourke, AJ, Comendador, J, Tan, SH, 2019. A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1 - 77" title="A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae)." url="https://doi.org/10.5852/ejt.2019.500" volume="500" year="2019">2019</bibRefCitation>
|
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;
|
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|
<bibRefCitation DOI="https://doi.org/10.1127/arch.moll/146/121-133" author="Dayrat, B" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="40" pageNumber="71" pagination="121 - 133" refId="B9" refString="Dayrat, B, Goulding, TC, 2017. Systematics of the onchidiid slug Onchidinaaustralis (Mollusca: Gastropoda: Pulmonata). Archiv fuer Molluskenkunde 146: 121 - 133" title="Systematics of the onchidiid slug Onchidinaaustralis (Mollusca: Gastropoda: Pulmonata)." url="https://doi.org/10.1127/arch.moll/146/121-133" volume="146" year="2017">Dayrat and Goulding 2017</bibRefCitation>
|
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;
|
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|
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.763.21252" author="Goulding, TC" journalOrPublisher="ZooKeys" pageId="41" pageNumber="72" pagination="1 - 111" refId="B15" refString="Goulding, TC, Khalil, M, Tan, SH, Dayrat, B, 2018a. Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda: Pulmonata: Onchidiidae). ZooKeys 763: 1 - 111" title="Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda: Pulmonata: Onchidiidae)." url="https://doi.org/10.3897/zookeys.763.21252" volume="763" year="2018 a">Goulding et al. 2018a</bibRefCitation>
|
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,
|
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<bibRefCitation author="Goulding, TC" journalOrPublisher="Raffles Bulletin of Zoology" pageId="41" pageNumber="72" pagination="337 - 349" refId="B16" refString="Goulding, TC, Khalil, M, Tan, SH, Dayrat, B, 2018b. A new genus and a new species of onchidiid slugs from eastern Indonesia (Gastropoda: Euthyneura: Onchidiidae). Raffles Bulletin of Zoology 66: 337 - 349" title="A new genus and a new species of onchidiid slugs from eastern Indonesia (Gastropoda: Euthyneura: Onchidiidae)." volume="66" year="2018 b">b</bibRefCitation>
|
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,
|
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|
|
<bibRefCitation DOI="https://doi.org/10.1071/IS17094" author="Goulding, TC" journalOrPublisher="Invertebrate Systematics" pageId="41" pageNumber="72" pagination="803 - 826" refId="B17" refString="Goulding, TC, Tan, SH, Tan, SK, Apte, D, Bhave, V, Narayana, S, Salunkhe, R, Dayrat, B, 2018c. A revision of Peronina Plate, 1893 (Gastropoda: Euthyneura: Onchidiidae) based on mitochondrial and nuclear DNA sequences, morphology, and natural history. Invertebrate Systematics 32: 803 - 826" title="A revision of Peronina Plate, 1893 (Gastropoda: Euthyneura: Onchidiidae) based on mitochondrial and nuclear DNA sequences, morphology, and natural history." url="https://doi.org/10.1071/IS17094" volume="32" year="2018 c">c</bibRefCitation>
|
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|
|
), it appears that there is no generic name of which the type species matches the diagnosis of this genus. For a recent review of the application of all existing generic names of
|
|||
|
|
<taxonomicName family="Onchidiidae" lsidName="" pageId="11" pageNumber="42" rank="family">Onchidiidae</taxonomicName>
|
|||
|
|
, see
|
|||
|
|
<bibRefCitation DOI="https://doi.org/10.1080/00222933.2017.1347297" author="Dayrat, B" journalOrPublisher="Journal of Natural History" pageId="40" pageNumber="71" pagination="1851 - 1897" refId="B11" refString="Dayrat, B, Goulding, TC, Apte, D, Bhave, V, Ngo, XQ, 2017. A new genus and four new species of onchidiid slugs from South-East Asia (Mollusca: Gastropoda: Pulmonata: Onchidiidae). Journal of Natural History 51: 1851 - 1897" title="A new genus and four new species of onchidiid slugs from South-East Asia (Mollusca: Gastropoda: Pulmonata: Onchidiidae)." url="https://doi.org/10.1080/00222933.2017.1347297" volume="51" year="2017">Dayrat et al. (2017</bibRefCitation>
|
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|
|
: 1861).
|
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|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|