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130 lines
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<document id="C4598C03E248BE7B8C89A87D73FEAC73" ID-CLB-Dataset="5016" ID-DOI="10.1649/0010-065X-74.1.45" ID-GBIF-Dataset="9e4b8160-11b2-4bcd-bcbf-2604a74c7d7f" ID-ISSN="1938-4394" ID-Zenodo-Dep="4790442" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621947045496" checkinUser="carolina" docAuthor="McPeak, Ron H., Lago, Paul K. & Hanley, Guy A." docDate="2020" docId="463687B7FF9BFFE9F0CCFF2FDBFBB389" docLanguage="en" docName="TheColeopBull.74.1.45-59.pdf" docOrigin="The Coleopterists Bulletin 74 (1)" docSource="http://dx.doi.org/10.1649/0010-065x-74.1.45" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Dichelonyx pusillus" docType="treatment" docVersion="8" lastPageNumber="57" masterDocId="BA0FFFCFFF97FFE5F02BFFEED96FB04C" masterDocTitle="Two New Sulcate Species of Dichelonyx Harris (Coleoptera: Scarabaeidae: Melolonthinae: Dichelonychini) from Southern California, USA and Baja California, Mexico" masterLastPageNumber="59" masterPageNumber="45" pageNumber="57" updateTime="1698989724174" updateUser="ExternalLinkService">
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<mods:title id="5F753A0365FECAB6F53100632F650A56">Two New Sulcate Species of Dichelonyx Harris (Coleoptera: Scarabaeidae: Melolonthinae: Dichelonychini) from Southern California, USA and Baja California, Mexico</mods:title>
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<mods:namePart id="19856EB7F8FC54EE1F067FAFEC7E5A32">McPeak, Ron H.</mods:namePart>
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<mods:namePart id="D0CA631317C14B0D0E9B85D0A5080AE2">Lago, Paul K.</mods:namePart>
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<mods:namePart id="C4181A5A5B17B8136A1C5EBBAB3C1B1E">Hanley, Guy A.</mods:namePart>
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<mods:title id="1014E826EB3475224F00BBB5A4CBB8E8">The Coleopterists Bulletin</mods:title>
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<mods:date id="02A5A652E949EC215C878C18FAD8D1B9">2020</mods:date>
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<treatment id="463687B7FF9BFFE9F0CCFF2FDBFBB389" ID-DOI="http://doi.org/10.5281/zenodo.5190754" ID-GBIF-Taxon="184092203" ID-Zenodo-Dep="5190754" LSID="urn:lsid:plazi:treatment:463687B7FF9BFFE9F0CCFF2FDBFBB389" httpUri="http://treatment.plazi.org/id/463687B7FF9BFFE9F0CCFF2FDBFBB389" lastPageNumber="57" pageId="12" pageNumber="57">
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<subSubSection id="8685652AFF9BFFE9F0CCFF2FDB36B09B" box="[231,601,193,215]" pageId="12" pageNumber="57" type="nomenclature">
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<paragraph id="CE2036A1FF9BFFE9F0CCFF2FDB36B09B" blockId="12.[231,601,193,242]" box="[231,601,193,215]" pageId="12" pageNumber="57">
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<heading id="956881CDFF9BFFE9F0CCFF2FDB36B09B" box="[231,601,193,215]" pageId="12" pageNumber="57">
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<taxonomicName id="099F4D22FF9BFFE9F0CCFF2FDB36B09B" authority="(LeConte, 1856)" baseAuthorityName="LeConte" baseAuthorityYear="1856" box="[231,601,193,215]" class="Insecta" family="Melolonthidae" genus="Dichelonyx" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="pusillus">
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<emphasis id="FCEBEAB3FF9BFFE9F0CCFF2FDB36B09B" bold="true" box="[231,601,193,215]" pageId="12" pageNumber="57">
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<emphasis id="FCEBEAB3FF9BFFE9F0CCFF2FD8C2B09B" bold="true" box="[231,429,193,215]" italics="true" pageId="12" pageNumber="57">Dichelonyx pusillus</emphasis>
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(
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<bibRefCitation id="AA0E4B50FF9BFFE9F190FF2CDB3DB09B" author="LeConte, J. L." box="[443,594,194,215]" pageId="12" pageNumber="57" pagination="225 - 288" refId="ref7717" refString="LeConte, J. L. 1856. Synopsis of the Melolonthidae of the United States. Journal of the Academy of Natural Sciences of Philadelphia 3 (2): 225 - 288." type="journal article" year="1856">LeConte, 1856</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8685652AFF9BFFE9F174FF32D88EB0BD" box="[351,481,220,242]" pageId="12" pageNumber="57" type="description">
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<paragraph id="CE2036A1FF9BFFE9F174FF32D88EB0BD" blockId="12.[231,601,193,242]" box="[351,481,220,242]" pageId="12" pageNumber="57">
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(
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<figureCitation id="56A42A24FF9BFFE9F14DFF32D8B5B0BD" box="[358,474,220,242]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="10.[197,236,1374,1393]" captionStartId-1="11.[175,214,1540,1559]" captionTargetBox-0="[189,1186,202,1346]" captionTargetBox-1="[157,1167,285,1519]" captionTargetId-0="figure-167@10.[173,1188,199,1358]" captionTargetId-1="figure-61@11.[154,1169,282,1523]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Figs. 20–22. Dichelonyx pusillus, male. 20) Habitus; 21–22) Head, dorsal and oblique views, respectively." captionText-1="Figs. 23–26. Dichelonyx pusillus. 23–24) Female head, dorsal and oblique views, respectively; 25) Female vittate elytra; 26) Parameres." figureDoi-0="http://doi.org/10.5281/zenodo.4790587" figureDoi-1="http://doi.org/10.5281/zenodo.4790602" httpUri-0="https://zenodo.org/record/4790587/files/figure.png" httpUri-1="https://zenodo.org/record/4790602/files/figure.png" pageId="12" pageNumber="57">Figs. 20–26</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="8685652AFF9BFFE9F084FEE9DB19B151" box="[175,630,263,285]" pageId="12" pageNumber="57" type="reference_group">
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<paragraph id="CE2036A1FF9BFFE9F084FEE9DB19B151" blockId="12.[175,630,263,285]" box="[175,630,263,285]" pageId="12" pageNumber="57">
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<treatmentCitationGroup id="EE8F118FFF9BFFE9F084FEE9DB19B151" box="[175,630,263,285]" pageId="12" pageNumber="57">
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<treatmentCitation id="4F3E10B0FF9BFFE9F084FEE9DB1DB151" author="LeConte, J. L." box="[175,626,263,285]" page="282" pageId="12" pageNumber="57" year="1856">
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<taxonomicName id="099F4D22FF9BFFE9F084FEE9DB1DB151" ID-CoL="35LHP" authority="LeConte 1856: 282" authorityName="LeConte" authorityPageNumber="282" authorityYear="1856" box="[175,626,263,285]" class="Insecta" family="Scarabaeidae" genus="Dichelonycha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="pusilla">
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<emphasis id="FCEBEAB3FF9BFFE9F084FEE9D85CB151" box="[175,307,263,285]" italics="true" pageId="12" pageNumber="57">Dichelonycha</emphasis>
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[sic]
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<emphasis id="FCEBEAB3FF9BFFE9F147FEE9D8C3B151" box="[364,428,263,285]" italics="true" pageId="12" pageNumber="57">pusilla</emphasis>
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<bibRefCitation id="AA0E4B50FF9BFFE9F19FFEE6DB1DB151" author="LeConte, J. L." box="[436,626,264,285]" pageId="12" pageNumber="57" pagination="225 - 288" refId="ref7717" refString="LeConte, J. L. 1856. Synopsis of the Melolonthidae of the United States. Journal of the Academy of Natural Sciences of Philadelphia 3 (2): 225 - 288." type="journal article" year="1856">LeConte 1856: 282</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="8685652AFF9BFFE9F0EEFEDCDBEFB30C" pageId="12" pageNumber="57" type="description">
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<paragraph id="CE2036A1FF9BFFE9F0EEFEDCDBEFB30C" blockId="12.[173,660,306,965]" pageId="12" pageNumber="57">
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Specimens of this species are typically uniformly dark brown, occasionally with the central elytral disc paler brown. The anterior clypeal angles of males (
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<figureCitation id="56A42A24FF9BFFE9F0D9FE6CD809B1DB" box="[242,358,386,408]" captionStart="Figs" captionStartId="10.[197,236,1374,1393]" captionTargetBox="[189,1186,202,1346]" captionTargetId="figure-167@10.[173,1188,199,1358]" captionTargetPageId="10" captionText="Figs. 20–22. Dichelonyx pusillus, male. 20) Habitus; 21–22) Head, dorsal and oblique views, respectively." figureDoi="http://doi.org/10.5281/zenodo.4790587" httpUri="https://zenodo.org/record/4790587/files/figure.png" pageId="12" pageNumber="57">Figs. 20–22</figureCitation>
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) and females (
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<figureCitation id="56A42A24FF9BFFE9F1DEFE6CDB07B1DB" box="[501,616,386,408]" captionStart="Figs" captionStartId="11.[175,214,1540,1559]" captionTargetBox="[157,1167,285,1519]" captionTargetId="figure-61@11.[154,1169,282,1523]" captionTargetPageId="11" captionText="Figs. 23–26. Dichelonyx pusillus. 23–24) Female head, dorsal and oblique views, respectively; 25) Female vittate elytra; 26) Parameres." figureDoi="http://doi.org/10.5281/zenodo.4790602" httpUri="https://zenodo.org/record/4790602/files/figure.png" pageId="12" pageNumber="57">Figs. 23–24</figureCitation>
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) are slightly rounded, with the lateral margins slightly divergent to the base. The pronotum is unicolored. The elytral setae are white or silvery white and generally distributed from margin to margin, but never densely spaced, often vittate, with third, fifth, seventh, and ninth costae glabrous (
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<figureCitation id="56A42A24FF9BFFE9F23BFDCCDB35B27B" box="[528,602,546,567]" captionStart="Figs" captionStartId="11.[175,214,1540,1559]" captionTargetBox="[157,1167,285,1519]" captionTargetId="figure-61@11.[154,1169,282,1523]" captionTargetPageId="11" captionText="Figs. 23–26. Dichelonyx pusillus. 23–24) Female head, dorsal and oblique views, respectively; 25) Female vittate elytra; 26) Parameres." figureDoi="http://doi.org/10.5281/zenodo.4790602" httpUri="https://zenodo.org/record/4790602/files/figure.png" pageId="12" pageNumber="57">Fig. 25</figureCitation>
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). The glabrous lines are generally more than half as wide as the setal bands. In some specimens, the setae are more evenly distributed, and the vittae are not at all apparent, but in these cases, the relatively sparse distribution of setae, along with the general dark coloration, is distinctive. The parameres are typical for genus, slightly angulate dorsally, with apically elongate lobes as robust in lateral view as those of
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<taxonomicName id="099F4D22FF9BFFE9F257FD18D863B36A" authorityName="McPeak & Lago & Hanley" authorityYear="2020" class="Insecta" family="Melolonthidae" genus="Dichelonyx" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="magnesae">
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<emphasis id="FCEBEAB3FF9BFFE9F257FD18D863B36A" italics="true" pageId="12" pageNumber="57">D. magnesae</emphasis>
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</taxonomicName>
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or more so, with down-curved acute apices (
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<figureCitation id="56A42A24FF9BFFE9F0DDFCC5D850B30C" box="[246,319,811,832]" captionStart="Figs" captionStartId="11.[175,214,1540,1559]" captionTargetBox="[157,1167,285,1519]" captionTargetId="figure-61@11.[154,1169,282,1523]" captionTargetPageId="11" captionText="Figs. 23–26. Dichelonyx pusillus. 23–24) Female head, dorsal and oblique views, respectively; 25) Female vittate elytra; 26) Parameres." figureDoi="http://doi.org/10.5281/zenodo.4790602" httpUri="https://zenodo.org/record/4790602/files/figure.png" pageId="12" pageNumber="57">Fig. 26</figureCitation>
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). The internal sac lacks sclerites.
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</paragraph>
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</subSubSection>
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<subSubSection id="8685652AFF9BFFE9F0EEFCABDBFBB389" pageId="12" pageNumber="57" type="description">
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<paragraph id="CE2036A1FF9BFFE9F0EEFCABDBFBB389" blockId="12.[173,660,306,965]" pageId="12" pageNumber="57">
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<taxonomicName id="099F4D22FF9BFFE9F0EEFCABD8EDB317" baseAuthorityName="LeConte" baseAuthorityYear="1856" box="[197,386,837,859]" class="Insecta" family="Melolonthidae" genus="Dichelonyx" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="pusillus">
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<emphasis id="FCEBEAB3FF9BFFE9F0EEFCABD8EDB317" box="[197,386,837,859]" italics="true" pageId="12" pageNumber="57">Dichelonyx pusillus</emphasis>
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</taxonomicName>
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occurs widely in the southern half of California. The species has also been taken in western Nevada and in
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<collectingRegion id="0C5BF843FF9BFFE9F1AFFC95DB61B3DC" box="[388,526,891,912]" country="Mexico" name="Baja California" pageId="12" pageNumber="57">Baja California</collectingRegion>
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,
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<collectingCountry id="B6887631FF9BFFE9F233FC95DB33B3DC" box="[536,604,891,912]" name="Mexico" pageId="12" pageNumber="57">Mexico</collectingCountry>
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(
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<bibRefCitation id="AA0E4B50FF9BFFE9F242FC95D869B3E7" author="Cornell, J. F." pageId="12" pageNumber="57" refId="ref7563" refString="Cornell, J. F. 1971. A taxonomic review of the beetle genus Dichelonyx Harris in North America (Scarabaeidae: Melolonthinae). PhD dissertation, Oregon State University, Corvallis, OR, 253 pp." type="book" year="1971">Cornell 1971</bibRefCitation>
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). The two new species occur sympatrically with
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<taxonomicName id="099F4D22FF9BFFE9F0F3FC5ED859B389" baseAuthorityName="LeConte" baseAuthorityYear="1856" box="[216,310,943,965]" class="Insecta" family="Melolonthidae" genus="Dichelonyx" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="pusillus">
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<emphasis id="FCEBEAB3FF9BFFE9F0F3FC5ED859B389" box="[216,310,943,965]" italics="true" pageId="12" pageNumber="57">D.pusillus</emphasis>
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</taxonomicName>
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only in the southern portion of its range.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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