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<document id="822E7C0A3C724607FCAAC5F71F862FD9" ID-DOI="10.1111/zoj.12381" ID-GBIF-Dataset="d73105df-ed15-4fed-8798-0e9b473a114a" ID-Zenodo-Dep="270304" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1467563411394" checkinUser="plazi" docAuthor="Adán Ṕerez-Garćia, Francisco Ortega &amp; Emiliano Jiménez Fuentes" docDate="2016" docId="931F87FCFFFBDC5EFC0D3544D27F43EB" docLanguage="en" docName="zjls_177_648.pdf" docOrigin="Zoological Journal of the Linnean Society 177" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Pelorochelon soriana Ṕerez-Garćia, Ortega &amp; Fuentes, 2016, SP. NOV." docType="treatment" docVersion="7" lastPageNumber="660" masterDocId="6F26FF84FFF2DC52FF9B3039D6104052" masterDocTitle="Taxonomy, systematics, and diversity of the European oldest testudinids" masterLastPageNumber="675" masterPageNumber="648" pageNumber="657" updateTime="1698363157625" updateUser="plazi">
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<mods:title id="FF8072588F5D372417E9E414D57C9D33">Taxonomy, systematics, and diversity of the European oldest testudinids</mods:title>
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<mods:namePart id="A7AFFF2576B8AF418F598B111EFE4439">Adán Ṕerez-Garćia</mods:namePart>
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<mods:namePart id="D62BAA47484072B86484D0344E780EC0">Francisco Ortega</mods:namePart>
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<mods:namePart id="2143DACF83A431115B86C8AF1D501985">Emiliano Jiménez Fuentes</mods:namePart>
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<paragraph id="1B0936EAFFFBDC5BFC0D3544D30E45C6" blockId="9.[918,1310,1402,1457]" box="[918,1310,1402,1428]" pageId="9" pageNumber="657">
<heading id="40418186FFFBDC5BFC0D3544D30E45C6" allCaps="true" box="[918,1310,1402,1428]" centered="true" fontSize="9" level="6" pageId="9" pageNumber="657" reason="4">
<taxonomicName id="DCB64D69FFFBDC5BFC0D3544D2A345C3" box="[918,1203,1405,1426]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="675" phylum="Chordata" rank="species" species="soriana" status="sp. nov.">
<emphasis id="29C2EAF8FFFBDC5BFC0D3544D2A345C3" bold="true" box="[918,1203,1405,1426]" italics="true" pageId="9" pageNumber="657">PELOROCHELON SORIANA</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32F15783FFFBDC5BFB2635B9D30E45C6" box="[1213,1310,1402,1428]" pageId="9" pageNumber="657">SP. NOV.</taxonomicNameLabel>
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</paragraph>
<paragraph id="1B0936EAFFFBDC5BFC6335AED2AC45E3" blockId="9.[918,1310,1402,1457]" box="[1016,1212,1431,1457]" pageId="9" pageNumber="657">
<heading id="40418186FFFBDC5BFC6335AED2AC45E3" allCaps="true" box="[1016,1212,1431,1457]" centered="true" fontSize="11" level="1" pageId="9" pageNumber="657" reason="1">
(
<figureCitation id="838D2A6FFFFBDC5BFB9A35A0D24245E3" box="[1025,1106,1431,1457]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="9" pageNumber="657">FIGS 2</figureCitation>
, 
<figureCitation id="838D2A6FFFFBDC5BFBFA35AED26245E3" box="[1121,1138,1431,1457]" captionStart="Figure 3" captionStartId="3.[144,223,1803,1822]" captionTargetBox="[166,1401,198,1770]" captionTargetId="figure@3.[166,1401,198,1770]" captionTargetPageId="3" captionText="Figure 3. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, shells STUS 14227 and STUS 14228, in ventral view. B, shells STUS 14229 and STUS 14230, in dorsal view. C, STUS 14399, anterior half of a shell, in dorsal (C 1) and ventral (C 2) views. D, STUS 14398, anterior half of a shell, in dorsal (D 1) and ventral (D 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270307/files/figure.png" pageId="9" pageNumber="657">3</figureCitation>
, 
<figureCitation id="838D2A6FFFFBDC5BFB1935AED28445E3" box="[1154,1172,1431,1457]" captionStart="Figure 4" captionStartId="4.[166,245,1588,1607]" captionText="Figure 4. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, STUS 14224, partial shell, in dorsal view (A 1), and plastron, in ventral view (A 2). B, STUS 14226, left hemiplastron, in dorsal (B 1) and ventral (B 2) views. C, STUS 14225, posterior region of a carapace, in dorsal (C 1) and ventral (C 2) views. D, STUS 10840, first neural, in dorsal (D 1) and ventral (D 2) views; E, STUS 10862, first neural, in dorsal (E 1) and ventral (E 2) views; F, STUS 8711, first neural, in dorsal (F 1) and ventral (F 2) views; G, STUS 10786, pygal, in dorsal (G 1) and ventral (G 2) views; H, STUS 10841, pygal, in dorsal (H 1) and ventral (H 2) views; I, STUS 10785, pygal, in dorsal (I 1) and ventral (I 2) views; J, STUS 10761, right xiphiplastron, in ventral (J 1) and dorsal (J 2) views; K, STUS 10787, left xiphiplastron, in dorsal (K 1) and ventral (K 2) views; L, STUS 10719, left xiphiplastron, in dorsal (L 1) and ventral (L 2) views; M, STUS 10760, right xiphiplastron, in ventral (M 1) and dorsal (M 2) views; N, STUS 10851, left xiphiplastron, in dorsal (N 1) and ventral (N 2) views; O, STUS 11696, left xiphiplastron, in dorsal (O 1) and ventral (O 2) views. Scale bar = 10 cm." pageId="9" pageNumber="657">4</figureCitation>
, 
<figureCitation id="838D2A6FFFFBDC5BFB3835AED2A245E3" box="[1187,1202,1431,1457]" captionStart="Figure 5" captionStartId="6.[166,245,1411,1430]" captionTargetBox="[280,1324,198,1378]" captionTargetId="figure@6.[280,1325,198,1378]" captionTargetPageId="6" captionText="Figure 5. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, STUS 8707, right epiplastron, in ventral (A 1), dorsal (A 2), anterior (A 3), and lateral (A 4) views. B, STUS 13802, left epiplastron, in dorsal (B 1), ventral (B 2), anterior (B 3), and lateral (B 4) views. C, STUS 11698, right epiplastron, right half of the entoplastron and anterior region of the right hyoplastron, in ventral (C 1), dorsal (C 2), anterior (C 3), and lateral (C 4) views. D, STUS 10857, right epiplastron, in ventral (D 1) and dorsal (D 2) views. E, STUS 10849, left epiplastron, in dorsal (E 1) and ventral (E 2) views. F, STUS 11697, left epiplastron and left half of the entoplastron, in dorsal (F 1) and ventral (F 2) views. G, STUS 10843, right epiplastron, in ventral (G 1) and dorsal (G 2) views. H, STUS 13753, left epiplastron, in dorsal (H 1) and ventral (H 2) views. I, STUS 11693, left epiplastron, in dorsal (I 1) and ventral (I 2) views. J, STUS 10850, right epiplastron, in ventral (J 1) and dorsal (J 2) views. K, STUS 10852, left epiplastron, in dorsal (K 1) and ventral (K 2) views. L, STUS 11695, left epiplastron, in dorsal (L 1) and ventral (L 2) views. M, STUS 10853, left epiplastron, in dorsal (M 1) and ventral (M 2) views. N, STUS 13763, left epiplastron, in dorsal (N 1) and ventral (N 2) views. O, STUS 11694, left epiplastron, in dorsal (O 1) and ventral (O 2) views. P, STUS 13736, right epiplastron, in ventral (P 1) and dorsal (P 2) views. Q, STUS 13762, left epiplastron, in dorsal (Q 1) and ventral (Q 2) views. R, STUS 13754, left epiplastron, in dorsal (R 1) and ventral (R 2) views. S, STUS 10860, right epiplastron, in ventral (S 1) and dorsal (S 2) views. T, STUS 10864, right epiplastron, in ventral (T 1) and dorsal (T 2) views. U, STUS 10856, right epiplastron, in ventral (U 1) and dorsal (U 2) views. V, STUS 10861, left epiplastron, in dorsal (V 1) and ventral (V 2) views. W, STUS 10814, left epiplastron, in dorsal (W 1) and ventral (W 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270308/files/figure.png" pageId="9" pageNumber="657">5</figureCitation>
)
</heading>
</paragraph>
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<subSubSection id="53AC6561FFFBDC5BFCBC35F0D591462A" pageId="9" pageNumber="657" type="reference_group">
<paragraph id="1B0936EAFFFBDC5BFCBC35F0D23E4649" blockId="9.[807,1421,1481,1900]" pageId="9" pageNumber="657">
<taxonomicName id="DCB64D69FFFBDC5BFCBC35F0D5DC458D" box="[807,972,1481,1503]" class="Reptilia" family="Testudinidae" genus="Hadrianus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="657" phylum="Chordata" rank="species" species="sp">
<emphasis id="29C2EAF8FFFBDC5BFCBC35F0D5B3458D" bold="true" box="[807,931,1481,1503]" italics="true" pageId="9" pageNumber="657">Hadrianus</emphasis>
sp.
</taxonomicName>
Jiḿenez Fuentes 
<emphasis id="29C2EAF8FFFBDC5BFB3B35F0D2C9458D" bold="true" box="[1184,1241,1481,1503]" italics="true" pageId="9" pageNumber="657">et al.</emphasis>
, 1988; Jiḿenez Fuentes, 1992, 2003; Cuesta Ruiz-Colmenares &amp; Jiḿenez Fuentes, 1994
</paragraph>
<paragraph id="1B0936EAFFFBDC5BFCA4361CD591462A" blockId="9.[807,1421,1481,1900]" pageId="9" pageNumber="657">
? 
<taxonomicName id="DCB64D69FFFBDC5BFCD1361DD5FF4668" box="[842,1007,1572,1594]" class="Reptilia" family="Testudinidae" genus="Hadrianus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="657" phylum="Chordata" rank="species" species="sp">
<emphasis id="29C2EAF8FFFBDC5BFCD1361DD5D64668" bold="true" box="[842,966,1572,1594]" italics="true" pageId="9" pageNumber="657">Hadrianus</emphasis>
sp.
</taxonomicName>
Jiḿenez Fuentes 
<emphasis id="29C2EAF8FFFBDC5BFB5A361CD2EB4668" bold="true" box="[1217,1275,1572,1594]" italics="true" pageId="9" pageNumber="657">et al.</emphasis>
, 1989 
<emphasis id="29C2EAF8FFFBDC5BFCBC367AD581460B" bold="true" box="[807,913,1603,1625]" italics="true" pageId="9" pageNumber="657">
<typeStatus id="C40D8848FFFBDC5BFCBC367AD59D460B" box="[807,909,1603,1625]" pageId="9" pageNumber="657" type="holotype">Holotype</typeStatus>
.
</emphasis>
An almost complete shell, STUS 14223 (
<figureCitation id="838D2A6FFFFBDC5BFCB4365BD564462A" box="[815,884,1634,1656]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="9" pageNumber="657">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="53AC6561FFFBDC59FCBC36B8D44A416A" lastPageId="11" lastPageNumber="659" pageId="9" pageNumber="657" type="materials_examined">
<paragraph id="1B0936EAFFFBDC59FCBC36B8D44A416A" blockId="9.[807,1421,1481,1900]" lastBlockId="11.[144,759,199,1901]" lastPageId="11" lastPageNumber="659" pageId="9" pageNumber="657">
<emphasis id="29C2EAF8FFFBDC5BFCBC36B8D58E46C4" bold="true" box="[807,926,1665,1686]" italics="true" pageId="9" pageNumber="657">
<typeStatus id="C40D8848FFFBDC5BFCBC36B8D58A46C4" box="[807,922,1665,1686]" pageId="9" pageNumber="657" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Forty-four specimens including several almost complete and partial shells, and isolated plates (
<figureCitation id="838D2A6FFFFBDC5BFC1E3684D5DE4681" box="[901,974,1725,1747]" captionStart="Figure 3" captionStartId="3.[144,223,1803,1822]" captionTargetBox="[166,1401,198,1770]" captionTargetId="figure@3.[166,1401,198,1770]" captionTargetPageId="3" captionText="Figure 3. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, shells STUS 14227 and STUS 14228, in ventral view. B, shells STUS 14229 and STUS 14230, in dorsal view. C, STUS 14399, anterior half of a shell, in dorsal (C 1) and ventral (C 2) views. D, STUS 14398, anterior half of a shell, in dorsal (D 1) and ventral (D 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270307/files/figure.png" pageId="9" pageNumber="657">Figs 3</figureCitation>
, 
<figureCitation id="838D2A6FFFFBDC5BFC7D3687D5E64681" box="[998,1014,1726,1747]" captionStart="Figure 4" captionStartId="4.[166,245,1588,1607]" captionText="Figure 4. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, STUS 14224, partial shell, in dorsal view (A 1), and plastron, in ventral view (A 2). B, STUS 14226, left hemiplastron, in dorsal (B 1) and ventral (B 2) views. C, STUS 14225, posterior region of a carapace, in dorsal (C 1) and ventral (C 2) views. D, STUS 10840, first neural, in dorsal (D 1) and ventral (D 2) views; E, STUS 10862, first neural, in dorsal (E 1) and ventral (E 2) views; F, STUS 8711, first neural, in dorsal (F 1) and ventral (F 2) views; G, STUS 10786, pygal, in dorsal (G 1) and ventral (G 2) views; H, STUS 10841, pygal, in dorsal (H 1) and ventral (H 2) views; I, STUS 10785, pygal, in dorsal (I 1) and ventral (I 2) views; J, STUS 10761, right xiphiplastron, in ventral (J 1) and dorsal (J 2) views; K, STUS 10787, left xiphiplastron, in dorsal (K 1) and ventral (K 2) views; L, STUS 10719, left xiphiplastron, in dorsal (L 1) and ventral (L 2) views; M, STUS 10760, right xiphiplastron, in ventral (M 1) and dorsal (M 2) views; N, STUS 10851, left xiphiplastron, in dorsal (N 1) and ventral (N 2) views; O, STUS 11696, left xiphiplastron, in dorsal (O 1) and ventral (O 2) views. Scale bar = 10 cm." pageId="9" pageNumber="657">4</figureCitation>
, 
<figureCitation id="838D2A6FFFFBDC5BFB963684D20C4681" box="[1037,1052,1725,1747]" captionStart="Figure 5" captionStartId="6.[166,245,1411,1430]" captionTargetBox="[280,1324,198,1378]" captionTargetId="figure@6.[280,1325,198,1378]" captionTargetPageId="6" captionText="Figure 5. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, STUS 8707, right epiplastron, in ventral (A 1), dorsal (A 2), anterior (A 3), and lateral (A 4) views. B, STUS 13802, left epiplastron, in dorsal (B 1), ventral (B 2), anterior (B 3), and lateral (B 4) views. C, STUS 11698, right epiplastron, right half of the entoplastron and anterior region of the right hyoplastron, in ventral (C 1), dorsal (C 2), anterior (C 3), and lateral (C 4) views. D, STUS 10857, right epiplastron, in ventral (D 1) and dorsal (D 2) views. E, STUS 10849, left epiplastron, in dorsal (E 1) and ventral (E 2) views. F, STUS 11697, left epiplastron and left half of the entoplastron, in dorsal (F 1) and ventral (F 2) views. G, STUS 10843, right epiplastron, in ventral (G 1) and dorsal (G 2) views. H, STUS 13753, left epiplastron, in dorsal (H 1) and ventral (H 2) views. I, STUS 11693, left epiplastron, in dorsal (I 1) and ventral (I 2) views. J, STUS 10850, right epiplastron, in ventral (J 1) and dorsal (J 2) views. K, STUS 10852, left epiplastron, in dorsal (K 1) and ventral (K 2) views. L, STUS 11695, left epiplastron, in dorsal (L 1) and ventral (L 2) views. M, STUS 10853, left epiplastron, in dorsal (M 1) and ventral (M 2) views. N, STUS 13763, left epiplastron, in dorsal (N 1) and ventral (N 2) views. O, STUS 11694, left epiplastron, in dorsal (O 1) and ventral (O 2) views. P, STUS 13736, right epiplastron, in ventral (P 1) and dorsal (P 2) views. Q, STUS 13762, left epiplastron, in dorsal (Q 1) and ventral (Q 2) views. R, STUS 13754, left epiplastron, in dorsal (R 1) and ventral (R 2) views. S, STUS 10860, right epiplastron, in ventral (S 1) and dorsal (S 2) views. T, STUS 10864, right epiplastron, in ventral (T 1) and dorsal (T 2) views. U, STUS 10856, right epiplastron, in ventral (U 1) and dorsal (U 2) views. V, STUS 10861, left epiplastron, in dorsal (V 1) and ventral (V 2) views. W, STUS 10814, left epiplastron, in dorsal (W 1) and ventral (W 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270308/files/figure.png" pageId="9" pageNumber="657">5</figureCitation>
): shells 
<collectionCode id="7DA7AE2FFFFBDC5BFB0B3687D2C34681" box="[1168,1235,1726,1747]" pageId="9" pageNumber="657">STUS</collectionCode>
14224, 14227, 14228, 14229, and 14230; partial carapaces 
<collectionCode id="7DA7AE2FFFFBDC5BFAD136E5D39D46A3" box="[1354,1421,1756,1777]" pageId="9" pageNumber="657">STUS</collectionCode>
14225, 14398, and 14399; first neurals 
<collectionCode id="7DA7AE2FFFFBDC5BFA9A36C2D3544742" box="[1281,1348,1787,1808]" pageId="9" pageNumber="657">STUS</collectionCode>
8711, 10840, and 10862; pygals 
<collectionCode id="7DA7AE2FFFFBDC5BFBF83720D2B6477C" box="[1123,1190,1817,1838]" pageId="9" pageNumber="657">STUS</collectionCode>
10785, 10786, and 10841; partial plastra 
<collectionCode id="7DA7AE2FFFFBDC5BFBD73701D29F471F" box="[1100,1167,1848,1869]" pageId="9" pageNumber="657">STUS</collectionCode>
11697, 11698, and 14226; epiplastra 
<collectionCode id="7DA7AE2FFFFBDC5BFC67376FD22F4739" box="[1020,1087,1878,1899]" pageId="9" pageNumber="657">STUS</collectionCode>
8707, 10814, 10843, 10849, 10850, 10852, 10853, 10856, 10857, 10860, 10861, 10864, 11693, 11694, 11695, 13736, 13753, 13754, 13762, 13763, and 13802; xiphiplastra 
<collectionCode id="7DA7AE2FFFF9DC59FDC6313DD4B0414B" box="[605,672,260,281]" pageId="11" pageNumber="659">STUS</collectionCode>
10719, 10851, 10760, 10761, 10787, and 11696.
</paragraph>
</subSubSection>
<caption id="4FC96662FFF8DC58FF3D36B0D2494740" httpUri="https://zenodo.org/record/270309/files/figure.png" pageId="10" pageNumber="658" targetBox="[249,1365,198,1639]" targetPageId="10">
<paragraph id="1B0936EAFFF8DC58FF3D36B0D2494740" blockId="10.[166,1444,1673,1810]" pageId="10" pageNumber="658">
<emphasis id="29C2EAF8FFF8DC58FF3D36B0D70046CE" bold="true" box="[166,272,1673,1692]" pageId="10" pageNumber="658">Figure 7.</emphasis>
Holotypes and only known specimens of two European Eocene testudinids identified here as 
<emphasis id="29C2EAF8FFF8DC58FB6536B3D38E46CE" bold="true" box="[1278,1438,1673,1693]" italics="true" pageId="10" pageNumber="658">nomen dubium</emphasis>
. A, MNCN 73849, holotype of ‘ 
<emphasis id="29C2EAF8FFF8DC58FE78369FD4E946EB" bold="true" box="[483,761,1702,1721]" italics="true" pageId="10" pageNumber="658">Asturichelys multicostatus</emphasis>
’, from the middle-late Eocene of Llamaquique (Asturias, Spain), in dorsal (A1), ventral (A2), left lateral (A3), right lateral (A4), anterior (A5), and posterior (A6) views. B, MNHN.F CAT 1, holotype of ‘ 
<taxonomicName id="DCB64D69FFF8DC58FEDE36D9D41846A6" box="[325,520,1760,1780]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="658" phylum="Chordata" rank="species" species="castrensis">
<emphasis id="29C2EAF8FFF8DC58FEDE36D9D41846A6" bold="true" box="[325,520,1760,1780]" italics="true" pageId="10" pageNumber="658">Testudo castrensis</emphasis>
</taxonomicName>
’, from the middle Eocene of Castrais (Tarn, France), in dorsal (B1), ventral (B2), left lateral (B3), right lateral (B4), anterior (B5), and posterior (B6) views. Scale bar = 10 cm.
</paragraph>
</caption>
<subSubSection id="53AC6561FFF9DC59FF0B3179D48D41A2" pageId="11" pageNumber="659" type="etymology">
<paragraph id="1B0936EAFFF9DC59FF0B3179D48D41A2" blockId="11.[144,759,199,1901]" pageId="11" pageNumber="659">
<emphasis id="29C2EAF8FFF9DC59FF0B3179D71D4104" bold="true" box="[144,269,320,342]" italics="true" pageId="11" pageNumber="659">Etymology.</emphasis>
The specific name refers to 
<collectingCountry id="63A1767AFFF9DC59FDE43178D4AE4105" box="[639,702,321,343]" name="Syria" pageId="11" pageNumber="659">Soria</collectingCountry>
, the Spanish province where the 
<typeStatus id="C40D8848FFF9DC59FE433159D4194127" box="[472,521,352,373]" pageId="11" pageNumber="659">type</typeStatus>
locality is situated. 
<emphasis id="29C2EAF8FFF9DC59FF0B3147D7EE41C6" bold="true" box="[144,510,382,404]" italics="true" pageId="11" pageNumber="659">
<typeStatus id="C40D8848FFF9DC59FF0B3147D6D641C1" box="[144,198,382,403]" pageId="11" pageNumber="659">Type</typeStatus>
locality and horizon.
</emphasis>
Mazateŕon locality (
<figureCitation id="838D2A6FFFF9DC59FF0331A4D6C941E1" box="[152,217,413,435]" captionStart="Figure 1" captionStartId="1.[807,886,760,779]" captionTargetBox="[815,1412,198,726]" captionTargetId="figure@1.[815,1413,198,726]" captionTargetPageId="1" captionText="Figure 1. Geographical position of the type localities of the European Eocene testudinids discussed here. 1, Llamaquique (Asturias, Spain), middle-late Eocene, type locality of ‘ Asturichelys multicostatus ’. 2, Mazateŕon (Soria, Spain), middle Eocene, type locality of Pelorochelon soriana. 3, Falaise de Baby (Gironde, France), late Eocene, type locality of Cheirogaster maurini. 4, Castrais (Tarn, France), middle Eocene, type locality of ‘ Testudo castrensis ’. 5, Saint-Papoul (Aude, France), early Eocene, type locality of Fontainechelon cassouleti. 6, Salzburg (Austria), middle Eocene, type locality of ‘ Cheirogaster steinbacheri ’. 7, Mücheln (Germany), middle Eocene, type locality of Pelorochelon eocaenica." httpUri="https://zenodo.org/record/270305/files/figure.png" pageId="11" pageNumber="659">Fig. 1</figureCitation>
), 
<collectingCountry id="63A1767AFFF9DC59FF6A31A4D73D41E1" box="[241,301,413,435]" name="Syria" pageId="11" pageNumber="659">Soria</collectingCountry>
Province, Castilla y Léon, 
<collectingCountry id="63A1767AFFF9DC59FDC431A4D4B441E1" box="[607,676,413,435]" name="Spain" pageId="11" pageNumber="659">Spain</collectingCountry>
. Upper part of the Mazateŕon Formation, early Bartonian, MP 15–16, middle Eocene (
<bibRefCitation id="7F274B1BFFF9DC59FE2431E3D48141A2" author="Valero L &amp; Huerta P" box="[447,657,473,496]" journalOrPublisher="Geo-Temas" pageId="11" pageNumber="659" pagination="371 - 374" part="13" refString="Valero L, Huerta P, Garces´M, Beamud E, Gomez-Pac-´card M, Armenteros I. 2012. Magnetoestratigrafia del relleno Cenozoico de la Cuenca de Almazan. Geo-Temas 13: 371 - 374." title="Magnetoestratigrafia del relleno Cenozoico de la Cuenca de Almazan" type="journal article" year="2012">
Valero 
<emphasis id="29C2EAF8FFF9DC59FD8B31E3D45A41BD" bold="true" box="[528,586,473,495]" italics="true" pageId="11" pageNumber="659">et al.</emphasis>
, 2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="53AC6561FFF9DC59FF0B31C1D4B542B6" pageId="11" pageNumber="659" type="diagnosis">
<paragraph id="1B0936EAFFF9DC59FF0B31C1D4B542B6" blockId="11.[144,759,199,1901]" pageId="11" pageNumber="659">
<emphasis id="29C2EAF8FFF9DC59FF0B31C1D717425F" bold="true" box="[144,263,504,525]" italics="true" pageId="11" pageNumber="659">Diagnosis.</emphasis>
A species of 
<taxonomicName id="DCB64D69FFF9DC59FE5631C1D44D425C" box="[461,605,504,526]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="657" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="29C2EAF8FFF9DC59FE5631C1D44D425C" bold="true" box="[461,605,504,526]" italics="true" pageId="11" pageNumber="659">Pelorochelon</emphasis>
</taxonomicName>
with eight neurals, which differs from 
<taxonomicName id="DCB64D69FFF9DC59FDB1322ED4A6427E" box="[554,694,535,556]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="659" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFF9DC59FDB1322ED4A6427E" bold="true" box="[554,694,535,556]" italics="true" pageId="11" pageNumber="659">P. eocaenica</emphasis>
</taxonomicName>
by: rectangular morphology of the first or second neurals; short sides anteriorly positioned in the hexagonal second and third neurals; relatively short gular protrusion; relatively short epiplastral symphysis; very short dorsal epiplastral lip, in the sagittal plane; relatively short pectoral scutes.
</paragraph>
</subSubSection>
<subSubSection id="53AC6561FFF9DC5EFF0B32D4D7574466" lastPageId="12" lastPageNumber="660" pageId="11" pageNumber="659" type="description">
<paragraph id="1B0936EAFFF9DC59FF0B32D4D276416A" blockId="11.[144,759,199,1901]" lastBlockId="11.[807,1422,199,1901]" pageId="11" pageNumber="659">
<emphasis id="29C2EAF8FFF9DC59FF0B32D4D7BC4350" bold="true" box="[144,428,748,770]" italics="true" pageId="11" pageNumber="659">Description of the shell.</emphasis>
The length of the shell of the largest known specimens of 
<taxonomicName id="DCB64D69FFF9DC59FD9D3335D46C4372" box="[518,636,779,801]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="659" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFF9DC59FD9D3335D46C4372" bold="true" box="[518,636,779,801]" italics="true" pageId="11" pageNumber="659">P. soriana</emphasis>
</taxonomicName>
is close to 
<quantity id="DC4E9B0FFFF9DC59FF0B3313D6CA436D" box="[144,218,810,831]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="659" unit="cm" value="70.0">70 cm</quantity>
(
<figureCitation id="838D2A6FFFF9DC59FF6A3313D7234312" box="[241,307,810,832]" captionStart="Figure 3" captionStartId="3.[144,223,1803,1822]" captionTargetBox="[166,1401,198,1770]" captionTargetId="figure@3.[166,1401,198,1770]" captionTargetPageId="3" captionText="Figure 3. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, shells STUS 14227 and STUS 14228, in ventral view. B, shells STUS 14229 and STUS 14230, in dorsal view. C, STUS 14399, anterior half of a shell, in dorsal (C 1) and ventral (C 2) views. D, STUS 14398, anterior half of a shell, in dorsal (D 1) and ventral (D 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270307/files/figure.png" pageId="11" pageNumber="659">Fig. 3</figureCitation>
A). All preserved shells are crushed. However, the information available shows that its height was moderate. The outer surface of all the plates is granular. Well-developed carapace growth rings can be observed in all the specimens (
<figureCitation id="838D2A6FFFF9DC59FD12339DD4C143EB" box="[649,721,932,954]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 3B, 4A1). This taxon lacks medial or lateral carapace keels. A shallow notch is present in the anterior carapace margin (
<figureCitation id="838D2A6FFFF9DC59FEFB3439D7B94447" box="[352,425,1024,1046]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 3A, B, C, D). The nuchal plate is slightly wider than long. 
<taxonomicName id="DCB64D69FFF9DC59FDFD3427D6F64400" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="659" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFF9DC59FDFD3427D6F64400" bold="true" italics="true" pageId="11" pageNumber="659">Pelorochelon soriana</emphasis>
</taxonomicName>
has eight neural plates (
<figureCitation id="838D2A6FFFF9DC59FD8D3404D4714400" box="[534,609,1085,1107]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 3B). The neural series is irregular, considering its morphology, but it is continuous, from the contact with the nuchal plate to that with the first suprapygal. All neurals are longer than wide, except the sixth and seventh ones. A dorsal bump is present on the first neural (
<figureCitation id="838D2A6FFFF9DC59FEE034CDD7D6455B" box="[379,454,1268,1290]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 4D, E, F). The second neural of some specimens is rectangular, being longer than wide (
<figureCitation id="838D2A6FFFF9DC59FEF23508D7A44514" box="[361,436,1329,1351]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 3B). The first neural of these specimens is hexagonal, with their short sides positioned posteriorly. However, the rectangular neural is the first one in other shells (
<figureCitation id="838D2A6FFFF9DC59FDCB35B4D48745F0" box="[592,663,1421,1443]" captionStart="Figure 3" captionStartId="3.[144,223,1803,1822]" captionTargetBox="[166,1401,198,1770]" captionTargetId="figure@3.[166,1401,198,1770]" captionTargetPageId="3" captionText="Figure 3. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, shells STUS 14227 and STUS 14228, in ventral view. B, shells STUS 14229 and STUS 14230, in dorsal view. C, STUS 14399, anterior half of a shell, in dorsal (C 1) and ventral (C 2) views. D, STUS 14398, anterior half of a shell, in dorsal (D 1) and ventral (D 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270307/files/figure.png" pageId="11" pageNumber="659">Figs 3</figureCitation>
B, C, D, 4D, E, F). The other neurals are hexagonal, with the short sides anteriorly positioned. 
<taxonomicName id="DCB64D69FFF9DC59FDFD35F0D6F645AF" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="659" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFF9DC59FDFD35F0D6F645AF" bold="true" italics="true" pageId="11" pageNumber="659">Pelorochelon soriana</emphasis>
</taxonomicName>
has eight pairs of costals. All of them are noticeably wider than long. A well-developed alternating pattern of costals, composed by medially alternative long and short plates, versus laterally short and long ones, is present. 
<taxonomicName id="DCB64D69FFF9DC59FD9C365BD4E6462A" box="[519,758,1634,1656]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="659" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFF9DC59FD9C365BD4E6462A" bold="true" box="[519,758,1634,1656]" italics="true" pageId="11" pageNumber="659">Pelorochelon soriana</emphasis>
</taxonomicName>
has two suprapygals (
<figureCitation id="838D2A6FFFF9DC59FE0736B8D7F646C4" box="[412,486,1665,1687]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
, 
<figureCitation id="838D2A6FFFF9DC59FE6336B8D41446C4" box="[504,516,1665,1686]" captionStart="Figure 3" captionStartId="3.[144,223,1803,1822]" captionTargetBox="[166,1401,198,1770]" captionTargetId="figure@3.[166,1401,198,1770]" captionTargetPageId="3" captionText="Figure 3. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, shells STUS 14227 and STUS 14228, in ventral view. B, shells STUS 14229 and STUS 14230, in dorsal view. C, STUS 14399, anterior half of a shell, in dorsal (C 1) and ventral (C 2) views. D, STUS 14398, anterior half of a shell, in dorsal (D 1) and ventral (D 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270307/files/figure.png" pageId="11" pageNumber="659">3</figureCitation>
A, B, 4C). The first one is wider than the second. The posterior suprapygal is lenticular, and is embrassed by the first one. The morphology of the pygal could be rectangular or trapezoidal, being slightly wider than long or as wide as long (
<figureCitation id="838D2A6FFFF9DC59FE2E3723D7EF477D" box="[437,511,1818,1840]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
, 
<figureCitation id="838D2A6FFFF9DC59FD953723D40A477D" box="[526,538,1818,1839]" captionStart="Figure 3" captionStartId="3.[144,223,1803,1822]" captionTargetBox="[166,1401,198,1770]" captionTargetId="figure@3.[166,1401,198,1770]" captionTargetPageId="3" captionText="Figure 3. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, shells STUS 14227 and STUS 14228, in ventral view. B, shells STUS 14229 and STUS 14230, in dorsal view. C, STUS 14399, anterior half of a shell, in dorsal (C 1) and ventral (C 2) views. D, STUS 14398, anterior half of a shell, in dorsal (D 1) and ventral (D 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270307/files/figure.png" pageId="11" pageNumber="659">3</figureCitation>
A, B, 4C, G, H, I). This taxon has eleven pairs of peripherals. The distal margins of the anterior and posterior peripherals are dorsally directed. All of them lack pointed tips on the contact area of the sulci delimiting the marginal scutes with the carapacial rim (
<figureCitation id="838D2A6FFFF9DC59FCC4311BD5B64165" box="[863,934,290,312]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A, 3A, B, C, 4C).
</paragraph>
<paragraph id="1B0936EAFFF9DC59FCA43178D2CC449F" blockId="11.[807,1422,199,1901]" pageId="11" pageNumber="659">
A small cervical scute, that is almost as wide as long, is present (
<figureCitation id="838D2A6FFFF9DC59FC6F3166D2514127" box="[1012,1089,351,373]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 3B, C). This element is very short in visceral view (
<figureCitation id="838D2A6FFFF9DC59FBFC3147D2BE41C1" box="[1127,1198,382,404]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A2, 3A). This taxon has five relatively narrow vertebral scutes, the second to fourth being longer than wide. The third vertebral is slightly wider than the second, and both are wider than the fourth. The first one is wider that these vertebrals. The latero-posterior margins of this pentagonal vertebral are divergent towards the anterior region (
<figureCitation id="838D2A6FFFF9DC59FC29326DD5EB423B" box="[946,1019,596,618]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 3B, C, D, 4A1). The wider vertebral scute is the fifth, their latero-anterior margins being markedly divergent towards the posterior region. The first pair of pleurals is not in contact with the nuchal plate (
<figureCitation id="838D2A6FFFF9DC59FBAA32F7D26A42B1" box="[1073,1146,718,740]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 3B, C, D). This pair of scutes contacts with the first to fifth marginals. The second pair contacts with the fifth to seventh marginals. The third pair contact with the sixth to ninth marginal. The last pair of pleurals contact with the ninth to eleventh marginal. 
<taxonomicName id="DCB64D69FFF9DC59FB66335FD56D43C8" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="659" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFF9DC59FB66335FD56D43C8" bold="true" italics="true" pageId="11" pageNumber="659">Pelorochelon soriana</emphasis>
</taxonomicName>
lacks coincidence between the medial margins of the marginals and the costo-peripheral suture (
<figureCitation id="838D2A6FFFF9DC59FCB533FBD567438A" box="[814,887,962,984]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 3B, C). Twelve pairs of marginals are present. Thus, this taxon lacks a supracaudal scute. The first pair is significantly wider than long (
<figureCitation id="838D2A6FFFF9DC59FA863439D3784447" box="[1309,1384,1024,1046]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 3B, C, D). Therefore, this taxon lacks contact of the second marginals with the nuchal. The only marginals in contact with the first vertebral are those of the first pair. The anterior region of the last pair of marginals overlaps the posterior area of the second suprapygal (
<figureCitation id="838D2A6FFFF9DC59FB91348ED242449E" box="[1034,1106,1207,1229]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A1, 3B, 4C).
</paragraph>
<paragraph id="1B0936EAFFF9DC5EFCA434ECD4AB41A2" blockId="11.[807,1422,199,1901]" lastBlockId="12.[166,781,199,1901]" lastPageId="12" lastPageNumber="660" pageId="11" pageNumber="659">
The plastral lobes are relatively short and wide (
<figureCitation id="838D2A6FFFF9DC59FCB534CDD565455B" box="[814,885,1268,1290]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A2, 3A, 4A2, B). The anterior one is almost as long as the posterior. The anterior lobe is medially notched (
<figureCitation id="838D2A6FFFF9DC59FC093508D5CC4514" box="[914,988,1329,1351]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="11" pageNumber="659">Figs 2</figureCitation>
A2, 3A, C2, D2, 4A2, B, 5A–W). The morphology of the lateral margins of the epiplastra is recognized as subject to variability, this area being substraight in some specimens (
<figureCitation id="838D2A6FFFF9DC59FB3635B4D2E145F1" box="[1197,1265,1421,1443]" captionStart="Figure 5" captionStartId="6.[166,245,1411,1430]" captionTargetBox="[280,1324,198,1378]" captionTargetId="figure@6.[280,1325,198,1378]" captionTargetPageId="6" captionText="Figure 5. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, STUS 8707, right epiplastron, in ventral (A 1), dorsal (A 2), anterior (A 3), and lateral (A 4) views. B, STUS 13802, left epiplastron, in dorsal (B 1), ventral (B 2), anterior (B 3), and lateral (B 4) views. C, STUS 11698, right epiplastron, right half of the entoplastron and anterior region of the right hyoplastron, in ventral (C 1), dorsal (C 2), anterior (C 3), and lateral (C 4) views. D, STUS 10857, right epiplastron, in ventral (D 1) and dorsal (D 2) views. E, STUS 10849, left epiplastron, in dorsal (E 1) and ventral (E 2) views. F, STUS 11697, left epiplastron and left half of the entoplastron, in dorsal (F 1) and ventral (F 2) views. G, STUS 10843, right epiplastron, in ventral (G 1) and dorsal (G 2) views. H, STUS 13753, left epiplastron, in dorsal (H 1) and ventral (H 2) views. I, STUS 11693, left epiplastron, in dorsal (I 1) and ventral (I 2) views. J, STUS 10850, right epiplastron, in ventral (J 1) and dorsal (J 2) views. K, STUS 10852, left epiplastron, in dorsal (K 1) and ventral (K 2) views. L, STUS 11695, left epiplastron, in dorsal (L 1) and ventral (L 2) views. M, STUS 10853, left epiplastron, in dorsal (M 1) and ventral (M 2) views. N, STUS 13763, left epiplastron, in dorsal (N 1) and ventral (N 2) views. O, STUS 11694, left epiplastron, in dorsal (O 1) and ventral (O 2) views. P, STUS 13736, right epiplastron, in ventral (P 1) and dorsal (P 2) views. Q, STUS 13762, left epiplastron, in dorsal (Q 1) and ventral (Q 2) views. R, STUS 13754, left epiplastron, in dorsal (R 1) and ventral (R 2) views. S, STUS 10860, right epiplastron, in ventral (S 1) and dorsal (S 2) views. T, STUS 10864, right epiplastron, in ventral (T 1) and dorsal (T 2) views. U, STUS 10856, right epiplastron, in ventral (U 1) and dorsal (U 2) views. V, STUS 10861, left epiplastron, in dorsal (V 1) and ventral (V 2) views. W, STUS 10814, left epiplastron, in dorsal (W 1) and ventral (W 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270308/files/figure.png" pageId="11" pageNumber="659">Fig. 5</figureCitation>
J, M, Q) but subrounded in others (
<figureCitation id="838D2A6FFFF9DC59FBB73592D27E4593" box="[1068,1134,1451,1473]" captionStart="Figure 5" captionStartId="6.[166,245,1411,1430]" captionTargetBox="[280,1324,198,1378]" captionTargetId="figure@6.[280,1325,198,1378]" captionTargetPageId="6" captionText="Figure 5. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, STUS 8707, right epiplastron, in ventral (A 1), dorsal (A 2), anterior (A 3), and lateral (A 4) views. B, STUS 13802, left epiplastron, in dorsal (B 1), ventral (B 2), anterior (B 3), and lateral (B 4) views. C, STUS 11698, right epiplastron, right half of the entoplastron and anterior region of the right hyoplastron, in ventral (C 1), dorsal (C 2), anterior (C 3), and lateral (C 4) views. D, STUS 10857, right epiplastron, in ventral (D 1) and dorsal (D 2) views. E, STUS 10849, left epiplastron, in dorsal (E 1) and ventral (E 2) views. F, STUS 11697, left epiplastron and left half of the entoplastron, in dorsal (F 1) and ventral (F 2) views. G, STUS 10843, right epiplastron, in ventral (G 1) and dorsal (G 2) views. H, STUS 13753, left epiplastron, in dorsal (H 1) and ventral (H 2) views. I, STUS 11693, left epiplastron, in dorsal (I 1) and ventral (I 2) views. J, STUS 10850, right epiplastron, in ventral (J 1) and dorsal (J 2) views. K, STUS 10852, left epiplastron, in dorsal (K 1) and ventral (K 2) views. L, STUS 11695, left epiplastron, in dorsal (L 1) and ventral (L 2) views. M, STUS 10853, left epiplastron, in dorsal (M 1) and ventral (M 2) views. N, STUS 13763, left epiplastron, in dorsal (N 1) and ventral (N 2) views. O, STUS 11694, left epiplastron, in dorsal (O 1) and ventral (O 2) views. P, STUS 13736, right epiplastron, in ventral (P 1) and dorsal (P 2) views. Q, STUS 13762, left epiplastron, in dorsal (Q 1) and ventral (Q 2) views. R, STUS 13754, left epiplastron, in dorsal (R 1) and ventral (R 2) views. S, STUS 10860, right epiplastron, in ventral (S 1) and dorsal (S 2) views. T, STUS 10864, right epiplastron, in ventral (T 1) and dorsal (T 2) views. U, STUS 10856, right epiplastron, in ventral (U 1) and dorsal (U 2) views. V, STUS 10861, left epiplastron, in dorsal (V 1) and ventral (V 2) views. W, STUS 10814, left epiplastron, in dorsal (W 1) and ventral (W 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270308/files/figure.png" pageId="11" pageNumber="659">Fig. 5</figureCitation>
D, E, H). The well-developed dorsal epiplastral lip is concave. It shows a welldeveloped visceral relief in the latero-anterior region of the epiplastra. The epiplastral symphysis is relatively short, being shorter than the length of the entoplastron. In the sagittal plane, the dorsal epiplastral lip is very short, generally being much shorter than half of the length of the epiplastral symphysis. This taxon has relatively short gular protrusions. It lacks gular pocket, as well as a well-developed change of relief in the posterior margin of the epiplastral lip (
<figureCitation id="838D2A6FFFF9DC59FCB436C2D5604743" box="[815,880,1787,1809]" captionStart="Figure 5" captionStartId="6.[166,245,1411,1430]" captionTargetBox="[280,1324,198,1378]" captionTargetId="figure@6.[280,1325,198,1378]" captionTargetPageId="6" captionText="Figure 5. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, STUS 8707, right epiplastron, in ventral (A 1), dorsal (A 2), anterior (A 3), and lateral (A 4) views. B, STUS 13802, left epiplastron, in dorsal (B 1), ventral (B 2), anterior (B 3), and lateral (B 4) views. C, STUS 11698, right epiplastron, right half of the entoplastron and anterior region of the right hyoplastron, in ventral (C 1), dorsal (C 2), anterior (C 3), and lateral (C 4) views. D, STUS 10857, right epiplastron, in ventral (D 1) and dorsal (D 2) views. E, STUS 10849, left epiplastron, in dorsal (E 1) and ventral (E 2) views. F, STUS 11697, left epiplastron and left half of the entoplastron, in dorsal (F 1) and ventral (F 2) views. G, STUS 10843, right epiplastron, in ventral (G 1) and dorsal (G 2) views. H, STUS 13753, left epiplastron, in dorsal (H 1) and ventral (H 2) views. I, STUS 11693, left epiplastron, in dorsal (I 1) and ventral (I 2) views. J, STUS 10850, right epiplastron, in ventral (J 1) and dorsal (J 2) views. K, STUS 10852, left epiplastron, in dorsal (K 1) and ventral (K 2) views. L, STUS 11695, left epiplastron, in dorsal (L 1) and ventral (L 2) views. M, STUS 10853, left epiplastron, in dorsal (M 1) and ventral (M 2) views. N, STUS 13763, left epiplastron, in dorsal (N 1) and ventral (N 2) views. O, STUS 11694, left epiplastron, in dorsal (O 1) and ventral (O 2) views. P, STUS 13736, right epiplastron, in ventral (P 1) and dorsal (P 2) views. Q, STUS 13762, left epiplastron, in dorsal (Q 1) and ventral (Q 2) views. R, STUS 13754, left epiplastron, in dorsal (R 1) and ventral (R 2) views. S, STUS 10860, right epiplastron, in ventral (S 1) and dorsal (S 2) views. T, STUS 10864, right epiplastron, in ventral (T 1) and dorsal (T 2) views. U, STUS 10856, right epiplastron, in ventral (U 1) and dorsal (U 2) views. V, STUS 10861, left epiplastron, in dorsal (V 1) and ventral (V 2) views. W, STUS 10814, left epiplastron, in dorsal (W 1) and ventral (W 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270308/files/figure.png" pageId="11" pageNumber="659">Fig. 5</figureCitation>
A–W). The epiplastra are almost as wide as long. The entoplastron is subrhombic. The hyoplastra are longer that the hypoplastra. The axillary buttresses are in contact with the lateral region of the first pair of costals. The inguinal buttresses contact the lateral region of the fifth pair of costals. The xiphiplastra are shorter that the hypoplastra. 
<taxonomicName id="DCB64D69FFFEDC5EFF3D311BD7884165" box="[166,408,290,312]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFFEDC5EFF3D311BD7884165" bold="true" box="[166,408,290,312]" italics="true" pageId="12" pageNumber="660">Pelorochelon soriana</emphasis>
</taxonomicName>
lacks plastral hinges. It has a wider than long anal notch (
<figureCitation id="838D2A6FFFFEDC5EFE6B3178D4274104" box="[496,567,321,343]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="12" pageNumber="660">Figs 2</figureCitation>
A2, 3A, 4A2, B, J– O). The morphology of the lateral margins of the xiphiplastra, and that of the lateral margins of the anal notch, are also subject to variability (compare the two specimens in 
<figureCitation id="838D2A6FFFFEDC5EFEE23182D7AB4182" box="[377,443,443,465]" captionStart="Figure 3" captionStartId="3.[144,223,1803,1822]" captionTargetBox="[166,1401,198,1770]" captionTargetId="figure@3.[166,1401,198,1770]" captionTargetPageId="3" captionText="Figure 3. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, shells STUS 14227 and STUS 14228, in ventral view. B, shells STUS 14229 and STUS 14230, in dorsal view. C, STUS 14399, anterior half of a shell, in dorsal (C 1) and ventral (C 2) views. D, STUS 14398, anterior half of a shell, in dorsal (D 1) and ventral (D 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270307/files/figure.png" pageId="12" pageNumber="660">Fig. 3</figureCitation>
A, one of them with straight margins but subrounded margins in the other).
</paragraph>
<paragraph id="1B0936EAFFFEDC5EFF2431C1D7574466" blockId="12.[166,781,199,1901]" pageId="12" pageNumber="660">
The gular scutes of 
<taxonomicName id="DCB64D69FFFEDC5EFE2B31C0D437425F" box="[432,551,504,526]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFFEDC5EFE2B31C0D437425F" bold="true" box="[432,551,504,526]" italics="true" pageId="12" pageNumber="660">P. soriana</emphasis>
</taxonomicName>
do not overlap the entoplastron. These scutes contact the anterior margin of the entoplastron in some specimens (
<figureCitation id="838D2A6FFFFEDC5EFD04320CD4F64218" box="[671,742,565,587]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="12" pageNumber="660">Figs 2</figureCitation>
A2, 4B2) but other individuals lacks this contact (
<figureCitation id="838D2A6FFFFEDC5EFF35324BD6E442DA" box="[174,244,626,648]" captionStart="Figure 3" captionStartId="3.[144,223,1803,1822]" captionTargetBox="[166,1401,198,1770]" captionTargetId="figure@3.[166,1401,198,1770]" captionTargetPageId="3" captionText="Figure 3. Some of the paratypes of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province). A, shells STUS 14227 and STUS 14228, in ventral view. B, shells STUS 14229 and STUS 14230, in dorsal view. C, STUS 14399, anterior half of a shell, in dorsal (C 1) and ventral (C 2) views. D, STUS 14398, anterior half of a shell, in dorsal (D 1) and ventral (D 2) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270307/files/figure.png" pageId="12" pageNumber="660">Figs 3</figureCitation>
A, 3C2, 4A2). The anterior angle between the sagittal axis and the gularo-humeral sulcus is higher than 65°. The humero-pectoral sulcus is perpendicular to the axial plane in the medial area but shows a well-developed lateral change of curvature. The pectoral scutes are relatively short in the medial region, being considerably shorter than the entoplastron in most of the specimens. These scutes do not contact the entoplastron. The concavity of the sulcus between the abdominal and the femoral scutes is anteriorly directed, in the medial region, whereas that between the femoral and anal scutes is posteriorly directed (
<figureCitation id="838D2A6FFFFEDC5EFED333D8D79F43A4" box="[328,399,993,1015]" captionStart="Figure 2" captionStartId="2.[166,245,1850,1869]" captionTargetBox="[393,1217,826,1816]" captionTargetId="figure@2.[393,1218,826,1817]" captionTargetPageId="2" captionText="Figure 2. STUS 14223, holotype of Pelorochelon soriana, from the early Bartonian (middle Eocene) of the Spanish site of Mazateŕon (Soria Province), in dorsal (A) and ventral (B) views. Scale bar = 10 cm." httpUri="https://zenodo.org/record/270306/files/figure.png" pageId="12" pageNumber="660">Figs 2</figureCitation>
A2, 3A, 4A2, B2). The abdominal scutes are the longest. The second longer scutes are the humerals.
</paragraph>
</subSubSection>
<subSubSection id="53AC6561FFFEDC5EFF3D3405D27F43EB" pageId="12" pageNumber="660" type="discussion">
<paragraph id="1B0936EAFFFEDC5EFF3D3405D73945D6" blockId="12.[166,781,199,1901]" pageId="12" pageNumber="660">
<emphasis id="29C2EAF8FFFEDC5EFF3D3405D7014400" bold="true" box="[166,273,1084,1106]" italics="true" pageId="12" pageNumber="660">Remarks.</emphasis>
The comparison between the material of 
<taxonomicName id="DCB64D69FFFEDC5EFF3D3465D7234422" box="[166,307,1115,1137]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFF3D3465D7234422" bold="true" box="[166,307,1115,1137]" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
and the specimens attributable to the genus 
<taxonomicName id="DCB64D69FFFEDC5EFF653440D79E44DD" box="[254,398,1145,1167]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="657" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="29C2EAF8FFFEDC5EFF653440D79E44DD" bold="true" box="[254,398,1145,1167]" italics="true" pageId="12" pageNumber="660">Pelorochelon</emphasis>
</taxonomicName>
from Mazateŕon allows their assignment to two different species. Jiḿenez Fuentes (1992, 2003) indicated that the morphology of the dorsal region of the epiplastra of the taxon from Mazateŕon was different from that observed in the other European specimens of ‘ 
<taxonomicName id="DCB64D69FFFEDC5EFD95352BD49A457A" box="[526,650,1298,1320]" class="Reptilia" family="Testudinidae" genus="Hadrianus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="genus">
<emphasis id="29C2EAF8FFFEDC5EFD95352BD49A457A" bold="true" box="[526,650,1298,1320]" italics="true" pageId="12" pageNumber="660">Hadrianus</emphasis>
</taxonomicName>
’. However, Jiḿenez Fuentes (1992, 2003) did not describe those epiplastra, and he did not indicate what were these differences.
</paragraph>
<paragraph id="1B0936EAFFFEDC5EFF2435B4D2FA41BD" blockId="12.[166,781,199,1901]" lastBlockId="12.[829,1444,198,954]" pageId="12" pageNumber="660">
The neural formula of 
<taxonomicName id="DCB64D69FFFEDC5EFE5735B4D44845F0" box="[460,600,1421,1442]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFE5735B4D44845F0" bold="true" box="[460,600,1421,1442]" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
and 
<taxonomicName id="DCB64D69FFFEDC5EFD0C35B4D51D45F0" box="[663,781,1421,1442]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFFEDC5EFD0C35B4D51D45F0" bold="true" box="[663,781,1421,1442]" italics="true" pageId="12" pageNumber="660">P. soriana</emphasis>
</taxonomicName>
is different. Some shells of 
<taxonomicName id="DCB64D69FFFEDC5EFE713595D4664592" box="[490,630,1451,1473]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFE713595D4664592" bold="true" box="[490,630,1451,1473]" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
have all the neurals hexagonal, except the third one, which is rectangular, whereas, in other specimens, the nonhexagonal neurals are the fourth (rectangular) and the sixth (octagonal). In some specimens of 
<taxonomicName id="DCB64D69FFFEDC5EFD21361FD6C1460B" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFFEDC5EFD21361FD6C1460B" bold="true" italics="true" pageId="12" pageNumber="660">P. soriana</emphasis>
</taxonomicName>
, all neurals are hexagonal, except the second one (rectangular), although the nonhexagonal neural is the first one (rectangular) in other individuals. The second neural of 
<taxonomicName id="DCB64D69FFFEDC5EFE333699D42446E7" box="[424,564,1696,1717]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFE333699D42446E7" bold="true" box="[424,564,1696,1717]" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
is always hexagonal, their short sides being posteriorly positioned (as in 
<taxonomicName id="DCB64D69FFFEDC5EFF6736E4D76146A0" box="[252,369,1757,1778]" class="Reptilia" family="Testudinidae" genus="Hadrianus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="corsoni">
<emphasis id="29C2EAF8FFFEDC5EFF6736E4D76146A0" bold="true" box="[252,369,1757,1778]" italics="true" pageId="12" pageNumber="660">H. corsoni</emphasis>
</taxonomicName>
). The specimens of 
<taxonomicName id="DCB64D69FFFEDC5EFDEA36E4D4F746A0" box="[625,743,1757,1778]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFFEDC5EFDEA36E4D4F746A0" bold="true" box="[625,743,1757,1778]" italics="true" pageId="12" pageNumber="660">P. soriana</emphasis>
</taxonomicName>
in which the second neural is hexagonal possess the short sides anteriorly positioned. The presence of a rectangular second neural, recognized in other specimens of 
<taxonomicName id="DCB64D69FFFEDC5EFE81376ED780473E" box="[282,400,1879,1900]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFFEDC5EFE81376ED780473E" bold="true" box="[282,400,1879,1900]" italics="true" pageId="12" pageNumber="660">P. soriana</emphasis>
</taxonomicName>
, is shared with 
<taxonomicName id="DCB64D69FFFEDC5EFDC4376ED5BB408E" box="[607,939,198,1901]" class="Reptilia" family="Testudinidae" genus="Fontainechelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="cassouleti">
<emphasis id="29C2EAF8FFFEDC5EFDC4376ED5BB408E" bold="true" box="[607,939,198,1901]" italics="true" pageId="12" pageNumber="660">Fontainechelon cassouleti</emphasis>
</taxonomicName>
. The third neural 
<taxonomicName id="DCB64D69FFFEDC5EFB3230FED30F408E" box="[1193,1311,199,220]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFFEDC5EFB3230FED30F408E" bold="true" box="[1193,1311,199,220]" italics="true" pageId="12" pageNumber="660">P. soriana</emphasis>
</taxonomicName>
is always hexagonal, their short sides being anteriorly positioned. However, the short sides are posteriorly positioned in the specimens of 
<taxonomicName id="DCB64D69FFFEDC5EFB32311AD3254165" box="[1193,1333,290,312]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFB32311AD3254165" bold="true" box="[1193,1333,290,312]" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
in which the third neural is hexagonal. The presence of a rectangular fourth neural, recognized in other specimens of 
<taxonomicName id="DCB64D69FFFEDC5EFC2F3147D25141C1" box="[948,1089,382,403]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFC2F3147D25141C1" bold="true" box="[948,1089,382,403]" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
, is shared with 
<taxonomicName id="DCB64D69FFFEDC5EFAB33147D38D41C1" box="[1320,1437,382,403]" class="Reptilia" family="Testudinidae" genus="Hadrianus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="corsoni">
<emphasis id="29C2EAF8FFFEDC5EFAB33147D38D41C1" bold="true" box="[1320,1437,382,403]" italics="true" pageId="12" pageNumber="660">H. corsoni</emphasis>
</taxonomicName>
, 
<taxonomicName id="DCB64D69FFFEDC5EFCA631A5D5DB41E0" box="[829,971,412,434]" class="Testudines" family="Testudinidae" genus="Chersine" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="maurini">
<emphasis id="29C2EAF8FFFEDC5EFCA631A5D5DB41E0" bold="true" box="[829,971,412,434]" italics="true" pageId="12" pageNumber="660">
<collectingCountry id="63A1767AFFFEDC5EFCA631A5D57041E0" box="[829,864,412,434]" name="Switzerland" pageId="12" pageNumber="660">Ch</collectingCountry>
. maurini
</emphasis>
</taxonomicName>
, ‘ 
<emphasis id="29C2EAF8FFFEDC5EFC7E31A4D24E41E0" bold="true" box="[997,1118,413,434]" italics="true" pageId="12" pageNumber="660">Erginemys</emphasis>
’ 
<emphasis id="29C2EAF8FFFEDC5EFBEA31A5D2D441E0" bold="true" box="[1137,1220,412,434]" italics="true" pageId="12" pageNumber="660">bruneti</emphasis>
and the members of 
<emphasis id="29C2EAF8FFFEDC5EFCF93182D5E64183" bold="true" box="[866,1014,443,465]" italics="true" pageId="12" pageNumber="660">Titanochelon</emphasis>
, being longer than wide in 
<taxonomicName id="DCB64D69FFFEDC5EFAFF3182D58141BD" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFAFF3182D58141BD" bold="true" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
, 
<taxonomicName id="DCB64D69FFFEDC5EFC3931E3D20841BD" box="[930,1048,474,495]" class="Reptilia" family="Testudinidae" genus="Hadrianus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="corsoni">
<emphasis id="29C2EAF8FFFEDC5EFC3931E3D20841BD" bold="true" box="[930,1048,474,495]" italics="true" pageId="12" pageNumber="660">H. corsoni</emphasis>
</taxonomicName>
and 
<taxonomicName id="DCB64D69FFFEDC5EFBCE31E0D2F341BD" box="[1109,1251,473,495]" class="Testudines" family="Testudinidae" genus="Chersine" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="maurini">
<emphasis id="29C2EAF8FFFEDC5EFBCE31E0D2F341BD" bold="true" box="[1109,1251,473,495]" italics="true" pageId="12" pageNumber="660">
<collectingCountry id="63A1767AFFFEDC5EFBCE31E0D26841BD" box="[1109,1144,473,495]" name="Switzerland" pageId="12" pageNumber="660">Ch</collectingCountry>
. maurini
</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1B0936EAFFFEDC5EFCCD31C1D2C94351" blockId="12.[829,1444,198,954]" pageId="12" pageNumber="660">
The gular protrusion of 
<taxonomicName id="DCB64D69FFFEDC5EFB0631C0D339425F" box="[1181,1321,504,526]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFB0631C0D339425F" bold="true" box="[1181,1321,504,526]" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
is much longer than that of 
<taxonomicName id="DCB64D69FFFEDC5EFBB6322ED2B4427E" box="[1069,1188,535,556]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFFEDC5EFBB6322ED2B4427E" bold="true" box="[1069,1188,535,556]" italics="true" pageId="12" pageNumber="660">P. soriana</emphasis>
</taxonomicName>
. For this reason, the length of the epiplastral symphysis is much longer in the first of these two taxa, being even longer than the entoplastron in some specimens. The presence of a very short dorsal epiplastral lip in 
<taxonomicName id="DCB64D69FFFEDC5EFA9F32A8D36A42F4" box="[1284,1402,657,678]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFFEDC5EFA9F32A8D36A42F4" bold="true" box="[1284,1402,657,678]" italics="true" pageId="12" pageNumber="660">P. soriana</emphasis>
</taxonomicName>
, in the sagittal plane, contrasts with the long lip observed in 
<taxonomicName id="DCB64D69FFFEDC5EFC4132F7D27642B1" box="[986,1126,718,739]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFC4132F7D27642B1" bold="true" box="[986,1126,718,739]" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
, longer than half of the length of the epiplastral symphysis.
</paragraph>
<paragraph id="1B0936EAFFFEDC5EFCCD3332D27F43EB" blockId="12.[829,1444,198,954]" pageId="12" pageNumber="660">
The medial region of the pectoral scutes of 
<taxonomicName id="DCB64D69FFFEDC5EFAFF3335D581436D" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFAFF3335D581436D" bold="true" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
is long. In this sense, it can be as long as the entoplastron, or even longer than this plate in some specimens. This condition is shared with 
<taxonomicName id="DCB64D69FFFEDC5EFACC335ED59743C9" class="Reptilia" family="Testudinidae" genus="Fontainechelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="cassouleti">
<emphasis id="29C2EAF8FFFEDC5EFACC335ED59743C9" bold="true" italics="true" pageId="12" pageNumber="660">F. cassouleti</emphasis>
</taxonomicName>
. The pectoral scutes of 
<taxonomicName id="DCB64D69FFFEDC5EFB3933BFD30943C8" box="[1186,1305,901,923]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="soriana">
<emphasis id="29C2EAF8FFFEDC5EFB3933BFD30943C8" bold="true" box="[1186,1305,901,923]" italics="true" pageId="12" pageNumber="660">P. soriana</emphasis>
</taxonomicName>
are shorter than those of 
<taxonomicName id="DCB64D69FFFEDC5EFC47339DD27843EB" box="[988,1128,932,953]" class="Reptilia" family="Testudinidae" genus="Pelorochelon" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="660" phylum="Chordata" rank="species" species="eocaenica">
<emphasis id="29C2EAF8FFFEDC5EFC47339DD27843EB" bold="true" box="[988,1128,932,953]" italics="true" pageId="12" pageNumber="660">P. eocaenica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>