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<document id="8FC8A31C2946C4F680E619DF683F1E0B" ID-DOI="10.11646/zootaxa.3718.3.2" ID-GBIF-Dataset="5a826d17-bd9a-4646-9603-f9a45a23ac5c" ID-ISSN="1175-5326" ID-Zenodo-Dep="247941" ID-ZooBank="003A16B5-34C5-48F8-A41C-8B784F3B06CB" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460686059778" checkinUser="plazi" docAuthor="Darby, Michael" docDate="2013" docId="187B8785405F09173BE6FF1AFC1AF1E1" docLanguage="en" docName="zt03718p266.pdf" docOrigin="Zootaxa 3718 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dacrysoma Grebennikov, sp. n." docType="treatment" docVersion="7" lastPageNumber="249" masterDocId="E442FFFD4055091D3B71FF8DFF8CF608" masterDocTitle="Studies of Madagascan Ptiliidae (Coleoptera) 1: The Tribe Discheramocephalini including eighteen new species" masterLastPageNumber="266" masterPageNumber="239" pageNumber="249" updateTime="1698351236098" updateUser="plazi">
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<mods:title id="5F4368AA769A0A7D9449EF37C220C3FF">Studies of Madagascan Ptiliidae (Coleoptera) 1: The Tribe Discheramocephalini including eighteen new species</mods:title>
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<mods:namePart id="42B21E4FAEBF48DB8D43B0531A33737F">Darby, Michael</mods:namePart>
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<mods:date id="68998DD4793360D1FD2B4BFF66411016">2013</mods:date>
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<treatment id="187B8785405F09173BE6FF1AFC1AF1E1" ID-DOI="http://doi.org/10.5281/zenodo.6154593" ID-GBIF-Taxon="119579906" ID-Zenodo-Dep="6154593" LSID="urn:lsid:plazi:treatment:187B8785405F09173BE6FF1AFC1AF1E1" httpUri="http://treatment.plazi.org/id/187B8785405F09173BE6FF1AFC1AF1E1" lastPageNumber="249" pageId="10" pageNumber="249">
<subSubSection id="D8C86518405F09173BE6FF1AFC1AF1E1" pageId="10" pageNumber="249" type="nomenclature">
<paragraph id="906D3693405F09173BE6FF1AFE47F6BA" blockId="10.[151,459,151,178]" box="[151,459,151,178]" pageId="10" pageNumber="249">
<heading id="CB2581FF405F09173BE6FF1AFE47F6BA" bold="true" box="[151,459,151,178]" fontSize="11" level="1" pageId="10" pageNumber="249" reason="1">
<taxonomicName id="57D24D10405F09173BE6FF1AFE47F6BA" ID-CoL="8X78W" authority="Grebennikov" authorityName="Grebennikov" box="[151,459,151,178]" class="Insecta" family="Ptiliidae" genus="Dacrysoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="A2A6EA81405F09173BE6FF1AFE47F6BA" bold="true" box="[151,459,151,178]" pageId="10" pageNumber="249">
<emphasis id="A2A6EA81405F09173BE6FF1AFEACF6B9" bold="true" box="[151,288,151,177]" italics="true" pageId="10" pageNumber="249">Dacrysoma</emphasis>
Grebennikov
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="906D3693405F09173BE6FF55FE5EF6E6" blockId="10.[151,466,216,238]" box="[151,466,216,238]" pageId="10" pageNumber="249">Grebennikov (2009: 128–130).</paragraph>
<paragraph id="906D3693405F09173BE6FE99FBFBF77B" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">
The generic description of Grebennikov is derived from 37 morphological characters used to test the validity of
<taxonomicName id="57D24D10405F09173BE6FEBAFE09F758" box="[151,389,311,336]" class="Insecta" family="Ptiliidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="249" phylum="Arthropoda" rank="tribe" tribe="Discheramocephalini">Discheramocephalini</taxonomicName>
as a monophyletic tribe and focuses, as a result, mainly on absent features rather than those which are present. The following revises some details of his description and adds others.
</paragraph>
<paragraph id="906D3693405F09173BB6FEF0FB20F4FE" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">
Similar to
<taxonomicName id="57D24D10405F09173A4CFEF0FE21F79D" box="[317,429,381,405]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="A2A6EA81405F09173A4CFEF0FE21F79D" box="[317,429,381,405]" italics="true" pageId="10" pageNumber="249">Cissidium</emphasis>
</taxonomicName>
in possessing a dumb-bell shaped antennomere 11 (although the dumb-bells of
<taxonomicName id="57D24D10405F09173E58FEF3FA10F79D" box="[1321,1436,381,405]" class="Insecta" family="Ptiliidae" genus="Dacrysoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="249" phylum="Arthropoda" rank="species" species="fabrum" status="sp. nov.">
<emphasis id="A2A6EA81405F09173E58FEF3FA10F79D" box="[1321,1436,381,405]" italics="true" pageId="10" pageNumber="249">D. fabrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B99557FA405F09173BE6FE2DFF5BF7B1" box="[151,215,416,441]" pageId="10" pageNumber="249" rank="species">
<emphasis id="A2A6EA81405F09173BE6FE2DFF5BF7B1" bold="true" box="[151,215,416,441]" pageId="10" pageNumber="249">sp. n.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="57D24D10405F09173A7CFE2CFE3FF7B1" box="[269,435,417,441]" class="Insecta" family="Ptiliidae" genus="Dacrysoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="249" phylum="Arthropoda" rank="species" species="denticulatm" status="sp. nov.">
<emphasis id="A2A6EA81405F09173A7CFE2CFE3FF7B1" box="[269,435,417,441]" italics="true" pageId="10" pageNumber="249">D. denticulatm</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B99557FA405F09173ACAFE2DFE77F7B1" box="[443,507,416,441]" pageId="10" pageNumber="249" rank="species">
<emphasis id="A2A6EA81405F09173ACAFE2DFE77F7B1" bold="true" box="[443,507,416,441]" pageId="10" pageNumber="249">sp. n.</emphasis>
</taxonomicNameLabel>
are represented by only a slight median narrowing of the segment) but differing in the presence of a clearly visible horizontal perforation of the mesoventral keel and of two further perforations towards the mesoventral anterior corners, absence of serrations on the hind coxal margin of the meso/meteventral suture and a single, clearly defined, indentation of the posterior margin of the pronotum opposite the scutellum. The ventral surface of the back of the head has a narrow band of +/-100 microscopic punctures and the dorsal surface a similar band of +/- 30 punctures. Similar punctures are present on the pre-mentum and mentum. Arms of the metendosternite very long and thin reaching almost to the anterior corners of the metaventrum.
<collectingCountry id="E8C57603405F09173E68FDF9FADDF485" box="[1305,1361,628,653]" name="India" pageId="10" pageNumber="249">Hind</collectingCountry>
coxae separated by approximately one quarter of the metaventral width, coxal plates small. Spermathcae ranging from globular to tubular. Sexual dimorphism is present in the genus in the form of reduced eyes and an increased number of hairs on antennomeres 10–11 of some males, and micropterism/ apterism in some females.
</paragraph>
<paragraph id="906D3693405F09173BB6FC8DFDCDF568" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">
There are two described species
<taxonomicName id="57D24D10405F0917394BFC8CFC07F511" authority="Grebennikov" authorityName="Grebennikov" box="[570,907,768,793]" class="Insecta" family="Ptiliidae" genus="Dacrysoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="249" phylum="Arthropoda" rank="species" species="usambarense">
<emphasis id="A2A6EA81405F0917394BFC8CFD7DF511" box="[570,753,769,793]" italics="true" pageId="10" pageNumber="249">D. usambarense</emphasis>
Grebennikov
</taxonomicName>
from
<collectingCountry id="E8C57603405F091738A3FC8DFBBBF511" box="[978,1079,768,793]" name="Tanzania" pageId="10" pageNumber="249">Tanzania</collectingCountry>
and
<taxonomicName id="57D24D10405F09173F01FC8CFAD1F511" authority="Grebennikov" authorityName="Grebennikov" box="[1136,1373,768,793]" class="Insecta" family="Ptiliidae" genus="Dacrysoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="249" phylum="Arthropoda" rank="species" species="felis">
<emphasis id="A2A6EA81405F09173F01FC8CFB4FF511" box="[1136,1219,769,793]" italics="true" pageId="10" pageNumber="249">D. felis</emphasis>
Grebennikov
</taxonomicName>
from
<collectingCountry id="E8C57603405F09173BE6FCA9FE92F535" box="[151,286,804,829]" name="Madagascar" pageId="10" pageNumber="249">Madagascar</collectingCountry>
(Antsiranana, Manongarivo Reserve). The new species described here appear to establish
<collectingCountry id="E8C57603405F09173E65FCA9FA17F535" box="[1300,1435,804,829]" name="Madagascar" pageId="10" pageNumber="249">Madagascar</collectingCountry>
as the centre for
<taxonomicName id="57D24D10405F09173A20FCC5FE5EF557" box="[337,466,840,863]" class="Insecta" family="Ptiliidae" genus="Dacrysoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="A2A6EA81405F09173A20FCC5FE5EF557" box="[337,466,840,863]" italics="true" pageId="10" pageNumber="249">Dacrysoma</emphasis>
</taxonomicName>
diversity.
</paragraph>
<paragraph id="906D3693405F09173BB6FCE7FB9EF218" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">
<emphasis id="A2A6EA81405F09173BB6FCE7FE8AF58B" bold="true" box="[199,262,874,899]" pageId="10" pageNumber="249">Note.</emphasis>
the form of the antennae, genitalia and pronota provide the best means of distinguishing species but the last require very careful orientation so that they are viewed from directly above (some of the figs. 56–61 are not viewed in this way). In normal view, when the pronotum slopes down at the front, the lateral margins can easily appear to be angled and/or to have straight edges particularly in the posterior half. Similarly, the tip of the aedeagus is best viewed from several angles to be sure of correctly determining its shape.
</paragraph>
<paragraph id="906D3693405F09173BB6FB97FEABF25E" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">Unless stated otherwise all species share the following features in addition to those mentioned in the generic descriptions:</paragraph>
<paragraph id="906D3693405F09173BB6FBEDFBB3F295" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">General: appearing polished and shining between the foveae, with no, or little, visible evidence of microsculpture or cuticular structure except around the mesoventral keel and collar.</paragraph>
<paragraph id="906D3693405F09173BB6FB2AFBC0F2EA" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">
Vestiture: long, yellow/golden, recumbent and overlapping particularly on the pronotum. None of the species possesses the mixture of short and long hairs evident in the African
<emphasis id="A2A6EA81405F091738FEFB46FBC0F2EA" box="[911,1100,1226,1250]" italics="true" pageId="10" pageNumber="249">
<taxonomicName id="57D24D10405F091738FEFB46FBCBF2EA" box="[911,1095,1226,1250]" class="Insecta" family="Ptiliidae" genus="Dacrysoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="249" phylum="Arthropoda" rank="species" species="usambarense">D. usambarense</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="906D3693405F09173BB6FB60FDD2F345" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">Head: vertex with a shallow depression running across the forehead immediately above the eyes and small groups of +/- 10 hairs towards the lateral margins inside and below the eyes, occasionally one or more placed inside the eye margin. Labrum with +/- 6 hairs.</paragraph>
<paragraph id="906D3693405F09173BB6FAD5FB77F378" blockId="10.[151,1437,276,2025]" box="[199,1275,1367,1392]" pageId="10" pageNumber="249">Eyes: larger in males than females in all species where both sexes have been available for study.</paragraph>
<paragraph id="906D3693405F09173BB6FAF6FE09F3D1" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">
Pronotum: cordate, with fine to coarse shallow foveae with separation sparse to close (
<figureCitation id="08E92A16405F09173FFEFAF7FA9EF39B" box="[1167,1298,1402,1427]" captionStart="FIGURES 56 – 61" captionStartId="14.[151,264,1953,1976]" captionTargetBox="[251,1335,201,1926]" captionTargetId="figure@14.[246,1341,193,1932]" captionTargetPageId="14" captionText="FIGURES 56 – 61. Dacrysoma spp. pronota. 56, D. ravelosoni (x 515); 57, D. nigerrimum (x 530); 58, D. ranomafanense (x 555); 59, D. fabrum (x 555); 60, D. varium (x 525); 61, D. rahanitriniainae (x 635)." httpUri="https://zenodo.org/record/247947/files/figure.png" pageId="10" pageNumber="249">Figs 56–61</figureCitation>
), pubescent, lateral margins with narrow raised ridges sometimes extending a short distance onto the posterior margin before the central emargination.
</paragraph>
<paragraph id="906D3693405F09173BB6FA69FCA1F3F5" blockId="10.[151,1437,276,2025]" box="[199,813,1508,1533]" pageId="10" pageNumber="249">Scutellum: pubescent, without any distinctive features.</paragraph>
<paragraph id="906D3693405F09173BB6F98AFB5CF04B" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">Elytra: elliptical/oblong elliptical, finely punctured, shallowly foveate with separation fine/moderate, and pubescent, the pubescence a little shorter than on the pronotum; without any distinctive features.</paragraph>
<paragraph id="906D3693405F09173BB6F9C0FE19F081" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">Meso/metaventral suture: lateral arms not, or only partially visible except in slide preparations when they appear as an apodeme.</paragraph>
<paragraph id="906D3693405F09173BB6F918FC5FF0A5" blockId="10.[151,1437,276,2025]" box="[199,979,1684,1709]" pageId="10" pageNumber="249">
Mesoventrum: glabrous, shoulders short and sharply keeled (
<figureCitation id="08E92A16405F09173803F919FC4AF0A5" box="[882,966,1684,1709]" captionStart="FIGURES 35 – 40" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[228,1303,210,1917]" captionTargetId="figure@12.[210,1318,195,1934]" captionTargetPageId="12" captionText="FIGURES 35 – 40. Dacrysoma spp .. 35, D. felis; 36, D. denticulatum, mesoventral process showing perforation (x 850); 37, D. denticulatum, mesoventral process showing perforation (x 1020); 38, D. varium aedeagus (x 1260); 39, D. ravelosoni aedeagus (x 1180); 40, D. fusum aedeagus (x 1440)." httpUri="https://zenodo.org/record/247945/files/figure.png" pageId="10" pageNumber="249">Fig. 35</figureCitation>
).
</paragraph>
<paragraph id="906D3693405F09173BB6F93AFBABF0D8" blockId="10.[151,1437,276,2025]" box="[199,1063,1719,1744]" pageId="10" pageNumber="249">Mesoventral keel: dorsal surface with +/- 12 backwardly directed hairs/setae.</paragraph>
<paragraph id="906D3693405F09173BB6F956FE9FF11E" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">Metaventrum: pubescence denser towards the margins, vertex often appearing glabrous because hairs have rubbed off.</paragraph>
<paragraph id="906D3693405F09173BB6F8ACFC9AF131" blockId="10.[151,1437,276,2025]" box="[199,790,1824,1849]" pageId="10" pageNumber="249">Legs: long and thin, without any distinctive features.</paragraph>
<paragraph id="906D3693405F09173BB6F8C9FD4AF155" blockId="10.[151,1437,276,2025]" box="[199,710,1860,1885]" pageId="10" pageNumber="249">
Wings: when present of the usual Ptiliid
<typeStatus id="4F698831405F091739FFF8C8FD4EF155" box="[654,706,1861,1885]" pageId="10" pageNumber="249">type</typeStatus>
.
</paragraph>
<paragraph id="906D3693405F09173BB6F8EAFC1AF1E1" blockId="10.[151,1437,276,2025]" pageId="10" pageNumber="249">
Abdomen: All segments punctured and pubescent. Tergite IX longer than other dorsal segments, Tergite X (pygidium) with no distinctive features in either sex. Ventrite VIII in both sexes with groups of hairs at lateral margins appearing as two pale patches.
<collectingCountry id="E8C57603405F09173914F820FD13F1CE" box="[613,671,1965,1990]" name="Malta" pageId="10" pageNumber="249">Male</collectingCountry>
ventrite IX very small and difficult to detect, rounded and with a short, forwardly pointed projection; without any distinctive features.
</paragraph>
</subSubSection>
</treatment>
</document>
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