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<document id="45BD3A42968CE876B4789FDC4CF2D385" ID-DOI="10.5281/zenodo.185598" ID-GBIF-Dataset="b319448e-6bd0-4017-b484-de6a6dd87601" ID-ISSN="1175-5326" ID-Zenodo-Dep="185598" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459994774502" checkinUser="plazi" docAuthor="Fresneda, Javier, Bourdeau, Charles &amp; Faille, Arnaud" docDate="2009" docId="0D1987CFB874FFB1FF654DBDFD0C1B44" docLanguage="en" docName="zt01993p016.pdf" docOrigin="Zootaxa 1993" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Baronniesia delioti Fresneda, Bourdeau &amp; Faille, sp. n." docType="treatment" docVersion="9" lastPageNumber="13" masterDocId="F120FFB7B873FFBDFFF24E28FFC41F3C" masterDocTitle="Baronniesia delioti gen. n. sp. n., a new subterranean Leptodirini from the French Pyrenees (Coleoptera: Leiodidae: Cholevinae)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="8" updateTime="1698232777016" updateUser="plazi">
<mods:mods id="E3A1121ED1DB8BF6BB6E0D23FB26DD0F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7E133B1D04248F203F9F6BBA31B89364">
<mods:title id="DF06B6B6DE1B18D3C7254B8118CCC8A0">Baronniesia delioti gen. n. sp. n., a new subterranean Leptodirini from the French Pyrenees (Coleoptera: Leiodidae: Cholevinae)</mods:title>
</mods:titleInfo>
<mods:name id="4667DD97E2849621E0F62FDAA7837711" type="personal">
<mods:role id="A53700AAF7F3E12211EBD1849563D6A5">
<mods:roleTerm id="721D8735E8D48807F1EE3FC675B9693E">Author</mods:roleTerm>
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<mods:namePart id="EA1C91A4370403529B808FA44AD598F8">Fresneda, Javier</mods:namePart>
</mods:name>
<mods:name id="8932C8C7634CE45CDD202C0E0E18219D" type="personal">
<mods:role id="6806699BAC77477B513C863A8125A4D2">
<mods:roleTerm id="F504CDF654E41F7A9EB5D9A31BC676AF">Author</mods:roleTerm>
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<mods:namePart id="3075AA98B0C44240A923F347113AE6D6">Bourdeau, Charles</mods:namePart>
</mods:name>
<mods:name id="2C6FFB8A9A6E78BF655E70403715083E" type="personal">
<mods:role id="12C51FC26D6B369B5C00B3B60EDD4037">
<mods:roleTerm id="5C39F57A542E0824A0DAE8CF52FFB372">Author</mods:roleTerm>
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<mods:namePart id="35A3E0B287A037832B9CC1262EEBFE00">Faille, Arnaud</mods:namePart>
</mods:name>
<mods:typeOfResource id="48D585C01AE0A43E0B1C44D4B94C8194">text</mods:typeOfResource>
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<mods:titleInfo id="667D1CEDC52F819D78FCB7E0AA77026E">
<mods:title id="B0FCC42C22D08C0FF7087B6241C2810E">Zootaxa</mods:title>
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<mods:part id="6497E585C2ADAC8F7F7F4BFEF6404350">
<mods:date id="3C4191A3CA2F971988669A438A4A2F59">2009</mods:date>
<mods:detail id="A97BA31C512CBE4C3AB4CA3121F7D3BB" type="volume">
<mods:number id="54691D3327E9B00B9295BD7C628E89B0">1993</mods:number>
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<mods:start id="33CC41A7EB42750A9641E9105AE0EEC7">1</mods:start>
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<mods:classification id="084B43B18B4613EF17F8FF41921B11EE">journal article</mods:classification>
<mods:identifier id="1C0610A91CEBD979BAC8D20894B5A1FE" type="DOI">10.5281/zenodo.185598</mods:identifier>
<mods:identifier id="1C97F718E422472FA6F281FB0D3D4552" type="GBIF-Dataset">b319448e-6bd0-4017-b484-de6a6dd87601</mods:identifier>
<mods:identifier id="477F037A311F1B4FBA735B12C397EA8E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1C4AC8251D739AA857E6B8BB733D266D" type="Zenodo-Dep">185598</mods:identifier>
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<treatment id="0D1987CFB874FFB1FF654DBDFD0C1B44" ID-DOI="http://doi.org/10.5281/zenodo.6215571" ID-GBIF-Taxon="119380242" ID-Zenodo-Dep="6215571" LSID="urn:lsid:plazi:treatment:0D1987CFB874FFB1FF654DBDFD0C1B44" httpUri="http://treatment.plazi.org/id/0D1987CFB874FFB1FF654DBDFD0C1B44" lastPageId="12" lastPageNumber="13" pageId="7" pageNumber="8">
<subSubSection id="CDAA6552B874FFBAFF654DBDFE491CEA" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="850F36D9B874FFBAFF654DBDFCFC1C93" blockId="7.[151,824,917,982]" box="[151,824,917,943]" pageId="7" pageNumber="8">
<heading id="DE4781B5B874FFBAFF654DBDFCFC1C93" bold="true" box="[151,824,917,943]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B7C4EACBB874FFBAFF654DBDFCFC1C93" bold="true" box="[151,824,917,943]" pageId="7" pageNumber="8">
<taxonomicName id="42B04D5AB874FFBAFF654DBDFD2B1C93" ID-CoL="68DT6" authority="Fresneda, Bourdeau &amp; Faille" authorityName="Fresneda, Bourdeau &amp; Faille" box="[151,751,917,943]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="delioti" status="sp. nov.">
<emphasis id="B7C4EACBB874FFBAFF654DBDFEB91C93" bold="true" box="[151,381,917,943]" italics="true" pageId="7" pageNumber="8">Baronniesia delioti</emphasis>
Fresneda, Bourdeau &amp; Faille
</taxonomicName>
<taxonomicNameLabel id="ACF757B0B874FFBAFD044DBDFCFC1C93" box="[758,824,917,943]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="850F36D9B874FFBAFF654D94FE491CEA" blockId="7.[151,824,917,982]" box="[151,397,956,982]" pageId="7" pageNumber="8">
(
<figureCitation id="1D8B2A5CB874FFBAFF6D4D94FED41CEA" box="[159,272,956,982]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 4" captionStart-2="FIGURES 5 – 6" captionStartId-0="2.[151,269,1590,1614]" captionStartId-1="3.[151,269,1501,1525]" captionStartId-2="4.[151,276,1409,1433]" captionTargetBox-0="[168,1413,341,1573]" captionTargetBox-1="[375,1213,477,1469]" captionTargetBox-2="[441,1145,181,1397]" captionTargetId-0="figure@2.[159,1427,330,1586]" captionTargetId-1="figure@3.[365,1221,465,1477]" captionTargetId-2="figure@4.[441,1146,181,1398]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 – 2. Habitus of Baronniesia delioti gen. n. sp. n. 1 ― male; 2 ― female." captionText-1="FIGURES 3 – 4. Aedeagus of Baronniesia delioti gen. n. sp. n. 3 — dorsal view; 4 — lateral view." captionText-2="FIGURES 5 – 6. Inner sac of aedeagus of Leptodirini. 5 — Baronniesia delioti gen. n. sp. n. and 6 — Antrocharis querilhaci dispar Abeille de Perrin. BDAM, dorsal sacs of apex of median area; BRA, reinforcement bands; FDM, dorsal feather-like structures; FVM, ventral feather-like structures; NL, connection nodule; FAPY, apical fibers of PY; IVP, inverted “ V ” feather-like structures; PY, “ piece en Y ”." httpUri-0="https://zenodo.org/record/185599/files/figure.png" httpUri-1="https://zenodo.org/record/185600/files/figure.png" httpUri-2="https://zenodo.org/record/185601/files/figure.png" pageId="7" pageNumber="8">Figs. 1–5</figureCitation>
,
<figureCitation id="1D8B2A5CB874FFBAFEEE4D94FEE91CEA" box="[284,301,956,982]" captionStart="FIGURES 7 – 14" captionStartId="5.[151,279,1785,1809]" captionTargetBox="[164,1428,301,1753]" captionTargetId="figure@5.[151,1436,286,1761]" captionTargetPageId="5" captionText="FIGURES 7 – 14. Apices of lateral styles of aedeagus of Leptodirini. 7 — Baronniesia delioti gen. n. sp. n.; 8 — Paratroglophyes jeanneli Coiffait; 9 — Troglophyes aubryi aubryi Coiffait; 10 — Troglophyes gavoyi Abeille de Perrin; 11 — Parvospeonomus delarouzeei Fairmaire; 12 — Phacomorphus (Phacomorphoides) sioberi sioberi (Coiffait); 13 — Speonomus curvipes subcurvipes Abeille de Perrin; 14 — Paraspeonomus vandeli Coiffait." httpUri="https://zenodo.org/record/185602/files/figure.png" pageId="7" pageNumber="8">7</figureCitation>
,
<figureCitation id="1D8B2A5CB874FFBAFEC84D94FE401CEA" box="[314,388,956,982]" captionStart="FIGURES 15 – 16" captionStartId="8.[151,275,1121,1145]" captionTargetBox="[151,1435,194,1097]" captionTargetId="figure@8.[151,1436,194,1097]" captionTargetPageId="8" captionText="FIGURES 15 – 16. Baronniesia delioti gen. n. sp. n. 15 — urite VIII of female; 16 — female genital segment with spermathecal complex." httpUri="https://zenodo.org/record/185603/files/figure.png" pageId="7" pageNumber="8">15–16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CDAA6552B874FFBAFF654A20FE2C1AF0" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="850F36D9B874FFBAFF654A20FB161B1F" blockId="7.[151,1438,1032,1484]" box="[151,1234,1032,1059]" pageId="7" pageNumber="8">
<emphasis id="B7C4EACBB874FFBAFF654A20FEF81B1E" bold="true" box="[151,316,1032,1058]" pageId="7" pageNumber="8">
<typeStatus id="5A0B887BB874FFBAFF654A20FF101B1E" box="[151,212,1032,1058]" pageId="7" pageNumber="8">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="FDA77649B874FFBAFEB64A21FE531B1F" box="[324,407,1033,1059]" name="France" pageId="7" pageNumber="8">France</collectingCountry>
, Hautes-Pyrénées, Esparros, subterranean river of Artigaléou-Arodets.
</paragraph>
<paragraph id="850F36D9B874FFBAFF344A07FC6C1BAB" blockId="7.[151,1438,1032,1484]" pageId="7" pageNumber="8">
<emphasis id="B7C4EACBB874FFBAFF344A07FDAE1B75" bold="true" box="[198,618,1071,1097]" pageId="7" pageNumber="8">
<typeStatus id="5A0B887BB874FFBAFF344A07FEC71B75" box="[198,259,1071,1097]" pageId="7" pageNumber="8">Type</typeStatus>
series.
<typeStatus id="5A0B887BB874FFBAFE904A07FE1C1B75" box="[354,472,1071,1097]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="FDA77649B874FFBAFE144A07FDA11B75" box="[486,613,1071,1097]" name="France" pageId="7" pageNumber="8">FRANCE</collectingCountry>
:
</emphasis>
ɗ, Esparros (massif of Baronnies), subterranean river of Artigaléou- Arodets,
<date id="F10E1019B874FFBAFEF04A7EFEB91B4C" box="[258,381,1110,1136]" pageId="7" pageNumber="8" value="2007-05-10">10.V.2007</date>
, Bourdeau &amp; Faille leg. (Muséum National d’Histoire Naturelle, Paris).
<emphasis id="B7C4EACBB874FFBAFB3B4A7EFA941B4C" bold="true" box="[1225,1360,1110,1136]" pageId="7" pageNumber="8">
<typeStatus id="5A0B887BB874FFBAFB3B4A7EFA8E1B4C" box="[1225,1354,1110,1136]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
:
</emphasis>
33ɗɗ and 61ΨΨ, same locality, data and collectors (authors’ collections).
</paragraph>
<paragraph id="850F36D9B874FFBAFF344A8CFCAC1A64" blockId="7.[151,1438,1032,1484]" pageId="7" pageNumber="8">
<typeStatus id="5A0B887BB874FFBAFF344A8CFEF71B82" box="[198,307,1188,1214]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
description.
<emphasis id="B7C4EACBB874FFBAFE394A8CFCB81B81" box="[459,892,1188,1213]" italics="true" pageId="7" pageNumber="8">
Habitus and size.
<taxonomicName id="42B04D5AB874FFBAFD6E4A8CFCB81B81" box="[668,892,1188,1213]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="delioti" status="sp. nov.">Baronniesia delioti</taxonomicName>
</emphasis>
<taxonomicNameLabel id="ACF757B0B874FFBAFC714A8CFC061B82" box="[899,962,1188,1214]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
is one of the largest species of Pyrenean Leptodirini. Length from anterior edge of pronotum to apex of elytra: 4.54 mm (see Variability). Anophthalmous and depigmented; outline oval, stocky, with pronotum wider than elytra (
<figureCitation id="1D8B2A5CB874FFBAFB164AD9FAE81A37" box="[1252,1324,1265,1291]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,269,1590,1614]" captionTargetBox="[168,1413,341,1573]" captionTargetId="figure@2.[159,1427,330,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Baronniesia delioti gen. n. sp. n. 1 ― male; 2 ― female." httpUri="https://zenodo.org/record/185599/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
). All the surface covered by yellowish pubescence, thin and soft; elytral pubescence longer. Punctation of head thin; mandibles very robust with a subapical tooth at internal edge.
</paragraph>
<paragraph id="850F36D9B874FFBAFF344B4DFE2C1AF0" blockId="7.[151,1438,1032,1484]" pageId="7" pageNumber="8">
<emphasis id="B7C4EACBB874FFBAFF344B4DFEF91A42" box="[198,317,1381,1406]" italics="true" pageId="7" pageNumber="8">Antennae.</emphasis>
Length 3.50 mm; antennal articles gradually dilated from fourth to ninth, tenth and eleventh articles not dilated; ventral side of the ninth article concave, dorsal side strongly convex. For measurements of antennal articles see Table 1.
</paragraph>
</subSubSection>
<paragraph id="850F36D9B874FFBAFF654BD7FA9E19C8" pageId="7" pageNumber="8">
<table id="F7B0C479B8740042FF654BD7FAAD19C8" box="[151,1385,1535,1780]" gridcols="9" gridrows="6" pageId="7" pageNumber="8">
<tr id="3B80349BB8740042FF654BD7FAAD192B" box="[151,1385,1535,1559]" gridrow="0" pageId="7" pageNumber="8">
<th id="78515DE7B8740042FF654BD7FF2B192B" box="[151,239,1535,1559]" gridcol="0" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="B7C4EACBB874FFBAFF654BD7FF2B192B" bold="true" box="[151,239,1535,1559]" pageId="7" pageNumber="8">TABLE</emphasis>
</th>
<th id="78515DE7B8740042FF044BD7FECC192B" box="[246,264,1535,1559]" gridcol="1" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="B7C4EACBB874FFBAFF044BD7FECC192B" bold="true" box="[246,264,1535,1559]" pageId="7" pageNumber="8">1.</emphasis>
</th>
<th id="78515DE7B8740042FEE34BD7FE68192B" box="[273,428,1535,1559]" gridcol="2" gridrow="0" pageId="7" pageNumber="8">Measurements</th>
<th id="78515DE7B8740042FE414BD7FD5E192B" box="[435,666,1535,1559]" gridcol="3" gridrow="0" pageId="7" pageNumber="8">of antennal articles of</th>
<th id="78515DE7B8740042FD534BD7FCA4192B" box="[673,864,1535,1559]" gridcol="4" gridrow="0" pageId="7" pageNumber="8">the holotype male</th>
<th id="78515DE7B8740042FC954BD7FC49192B" box="[871,909,1535,1559]" gridcol="5" gridrow="0" pageId="7" pageNumber="8">and</th>
<th id="78515DE7B8740042FC664BD7FB53192B" box="[916,1175,1535,1559]" gridcol="6" gridrow="0" pageId="7" pageNumber="8">a single female paratype</th>
<th id="78515DE7B8740042FB6C4BD7FB38192B" box="[1182,1276,1535,1559]" gridcol="7" gridrow="0" pageId="7" pageNumber="8">(40 units</th>
<th id="78515DE7B8740042FAF14BD7FAAD192B" box="[1283,1385,1535,1559]" gridcol="8" gridrow="0" pageId="7" pageNumber="8">= 1 mm).</th>
</tr>
<tr id="3B80349BB8740042FF65481AFAAD1975" box="[151,1385,1586,1609]" gridrow="1" pageId="7" pageNumber="8" rowspan-1="1" rowspan-2="1">
<th id="78515DE7B8740042FF65481AFF2B1975" box="[151,239,1586,1609]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">Article</th>
<td id="78515DE7B8740042FE41481AFD5E1975" box="[435,666,1586,1609]" gridcol="3" gridrow="1" pageId="7" pageNumber="8">I II III</td>
<td id="78515DE7B8740042FD53481AFCA41975" box="[673,864,1586,1609]" gridcol="4" gridrow="1" pageId="7" pageNumber="8">IV V</td>
<td id="78515DE7B8740042FC95481AFC491975" box="[871,909,1586,1609]" gridcol="5" gridrow="1" pageId="7" pageNumber="8">VI</td>
<td id="78515DE7B8740042FC66481AFB531975" box="[916,1175,1586,1609]" gridcol="6" gridrow="1" pageId="7" pageNumber="8">VII VIII</td>
<td id="78515DE7B8740042FB6C481AFB381975" box="[1182,1276,1586,1609]" gridcol="7" gridrow="1" pageId="7" pageNumber="8">IX</td>
<td id="78515DE7B8740042FAF1481AFAAD1975" box="[1283,1385,1586,1609]" gridcol="8" gridrow="1" pageId="7" pageNumber="8">X XI</td>
</tr>
<tr id="3B80349BB8740042FF654875FAAD1948" box="[151,1385,1629,1652]" gridrow="2" pageId="7" pageNumber="8" rowspan-1="1">
<th id="78515DE7B8740042FF654875FF2B1948" box="[151,239,1629,1652]" gridcol="0" gridrow="2" pageId="7" pageNumber="8">Male</th>
<td id="78515DE7B8740042FEE34875FE681948" box="[273,428,1629,1652]" gridcol="2" gridrow="2" pageId="7" pageNumber="8">Length</td>
<td id="78515DE7B8740042FE414875FD5E1948" box="[435,666,1629,1652]" gridcol="3" gridrow="2" pageId="7" pageNumber="8">17 10 8</td>
<td id="78515DE7B8740042FD534875FCA41948" box="[673,864,1629,1652]" gridcol="4" gridrow="2" pageId="7" pageNumber="8">10 12</td>
<td id="78515DE7B8740042FC954875FC491948" box="[871,909,1629,1652]" gridcol="5" gridrow="2" pageId="7" pageNumber="8">11</td>
<td id="78515DE7B8740042FC664875FB531948" box="[916,1175,1629,1652]" gridcol="6" gridrow="2" pageId="7" pageNumber="8">13 12</td>
<td id="78515DE7B8740042FB6C4875FB381948" box="[1182,1276,1629,1652]" gridcol="7" gridrow="2" pageId="7" pageNumber="8">15</td>
<td id="78515DE7B8740042FAF14875FAAD1948" box="[1283,1385,1629,1652]" gridcol="8" gridrow="2" pageId="7" pageNumber="8">7 14</td>
</tr>
<tr id="3B80349BB8740042FF6548A0FAAD19A3" box="[151,1385,1672,1695]" gridrow="3" pageId="7" pageNumber="8" rowspan-0="1" rowspan-1="1">
<td id="78515DE7B8740042FEE348A0FE6819A3" box="[273,428,1672,1695]" gridcol="2" gridrow="3" pageId="7" pageNumber="8">Width</td>
<td id="78515DE7B8740042FE4148A0FD5E19A3" box="[435,666,1672,1695]" gridcol="3" gridrow="3" pageId="7" pageNumber="8">6 4 4</td>
<td id="78515DE7B8740042FD5348A0FCA419A3" box="[673,864,1672,1695]" gridcol="4" gridrow="3" pageId="7" pageNumber="8">6 8</td>
<td id="78515DE7B8740042FC9548A0FC4919A3" box="[871,909,1672,1695]" gridcol="5" gridrow="3" pageId="7" pageNumber="8">10</td>
<td id="78515DE7B8740042FC6648A0FB5319A3" box="[916,1175,1672,1695]" gridcol="6" gridrow="3" pageId="7" pageNumber="8">14 14</td>
<td id="78515DE7B8740042FB6C48A0FB3819A3" box="[1182,1276,1672,1695]" gridcol="7" gridrow="3" pageId="7" pageNumber="8">13</td>
<td id="78515DE7B8740042FAF148A0FAAD19A3" box="[1283,1385,1672,1695]" gridcol="8" gridrow="3" pageId="7" pageNumber="8">6 5</td>
</tr>
<tr id="3B80349BB8740042FF65489AFAAD19F5" box="[151,1385,1714,1737]" gridrow="4" pageId="7" pageNumber="8" rowspan-1="1">
<th id="78515DE7B8740042FF65489AFF2B19F5" box="[151,239,1714,1737]" gridcol="0" gridrow="4" pageId="7" pageNumber="8">Female</th>
<td id="78515DE7B8740042FEE3489AFE6819F5" box="[273,428,1714,1737]" gridcol="2" gridrow="4" pageId="7" pageNumber="8">Length</td>
<td id="78515DE7B8740042FE41489AFD5E19F5" box="[435,666,1714,1737]" gridcol="3" gridrow="4" pageId="7" pageNumber="8">13 10 8</td>
<td id="78515DE7B8740042FD53489AFCA419F5" box="[673,864,1714,1737]" gridcol="4" gridrow="4" pageId="7" pageNumber="8">9 10</td>
<td id="78515DE7B8740042FC95489AFC4919F5" box="[871,909,1714,1737]" gridcol="5" gridrow="4" pageId="7" pageNumber="8">10</td>
<td id="78515DE7B8740042FC66489AFB5319F5" box="[916,1175,1714,1737]" gridcol="6" gridrow="4" pageId="7" pageNumber="8">11 8</td>
<td id="78515DE7B8740042FB6C489AFB3819F5" box="[1182,1276,1714,1737]" gridcol="7" gridrow="4" pageId="7" pageNumber="8">11</td>
<td id="78515DE7B8740042FAF1489AFAAD19F5" box="[1283,1385,1714,1737]" gridcol="8" gridrow="4" pageId="7" pageNumber="8">7 12</td>
</tr>
<tr id="3B80349BB8740042FF6548F5FAAD19C8" box="[151,1385,1757,1780]" gridrow="5" pageId="7" pageNumber="8" rowspan-0="1" rowspan-1="1">
<td id="78515DE7B8740042FEE348F5FE6819C8" box="[273,428,1757,1780]" gridcol="2" gridrow="5" pageId="7" pageNumber="8">Width</td>
<td id="78515DE7B8740042FE4148F5FD5E19C8" box="[435,666,1757,1780]" gridcol="3" gridrow="5" pageId="7" pageNumber="8">4 3 3</td>
<td id="78515DE7B8740042FD5348F5FCA419C8" box="[673,864,1757,1780]" gridcol="4" gridrow="5" pageId="7" pageNumber="8">3 4</td>
<td id="78515DE7B8740042FC9548F5FC4919C8" box="[871,909,1757,1780]" gridcol="5" gridrow="5" pageId="7" pageNumber="8">4</td>
<td id="78515DE7B8740042FC6648F5FB5319C8" box="[916,1175,1757,1780]" gridcol="6" gridrow="5" pageId="7" pageNumber="8">5 3</td>
<td id="78515DE7B8740042FB6C48F5FB3819C8" box="[1182,1276,1757,1780]" gridcol="7" gridrow="5" pageId="7" pageNumber="8">5</td>
<td id="78515DE7B8740042FAF148F5FAAD19C8" box="[1283,1385,1757,1780]" gridcol="8" gridrow="5" pageId="7" pageNumber="8">5 5</td>
</tr>
</table>
</paragraph>
<subSubSection id="CDAA6552B874FFB5FF344917FC541921" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="850F36D9B874FFBAFF344917FE531843" blockId="7.[151,1436,1854,2035]" pageId="7" pageNumber="8">
<emphasis id="B7C4EACBB874FFBAFF344917FE861864" box="[198,322,1855,1880]" italics="true" pageId="7" pageNumber="8">Pronotum.</emphasis>
Strongly transverse (length 1.25 mm, width 2.30 mm), sides very arcuate, maximum width before posterior edge.
</paragraph>
<paragraph id="850F36D9B874FFBAFF3449A4FDD218CF" blockId="7.[151,1436,1854,2035]" pageId="7" pageNumber="8">
<emphasis id="B7C4EACBB874FFBAFF3449A4FED31899" box="[198,279,1932,1957]" italics="true" pageId="7" pageNumber="8">Elytra.</emphasis>
One and half time longer than wide, strongly convex, with dehiscent apices; each elytron regularly curved at apex. Punctation strong and rough, in basal area arranged in transverse striae; in apical half with punctures distributed irregularly.
</paragraph>
<caption id="D1CF6651B87BFFB5FF654A49FE491BA7" httpUri="https://zenodo.org/record/185603/files/figure.png" pageId="8" pageNumber="9" targetBox="[151,1435,194,1097]" targetPageId="8">
<paragraph id="850F36D9B87BFFB5FF654A49FE491BA7" blockId="8.[151,1436,1121,1179]" pageId="8" pageNumber="9">
<emphasis id="B7C4EACBB87BFFB5FF654A49FEAC1B45" bold="true" box="[151,360,1121,1145]" pageId="8" pageNumber="9">FIGURES 15–16.</emphasis>
<taxonomicName id="42B04D5AB87BFFB5FE834A4AFD891B45" box="[369,589,1122,1145]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="delioti" status="gen. nov.">
<emphasis id="B7C4EACBB87BFFB5FE834A4AFD891B45" box="[369,589,1122,1145]" italics="true" pageId="8" pageNumber="9">Baronniesia delioti</emphasis>
</taxonomicName>
<emphasis id="B7C4EACBB87BFFB5FDA64A49FD2B1B45" bold="true" box="[596,751,1121,1145]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="ACF757B0B87BFFB5FDA64A49FD621B45" box="[596,678,1121,1145]" pageId="8" pageNumber="9" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
15—urite VIII of female; 16—female genital segment with spermathecal complex.
</paragraph>
</caption>
<paragraph id="850F36D9B87BFFB5FF344AE6FB331A32" blockId="8.[151,1437,1229,2029]" pageId="8" pageNumber="9">
<emphasis id="B7C4EACBB87BFFB5FF344AE6FEC11BDB" box="[198,261,1230,1255]" italics="true" pageId="8" pageNumber="9">Legs.</emphasis>
Proportionally long; mesotibiae arched, metatibiae straight. Male protarsomeres 1–3 strongly dilated, the first wider than apex of protibia; onychium as long as protarsomeres 1–4 combined.
</paragraph>
<paragraph id="850F36D9B87BFFB5FF344B33FB201A09" blockId="8.[151,1437,1229,2029]" box="[198,1252,1307,1333]" pageId="8" pageNumber="9">Mesoventral carina, aedeagus and male genital segment as in the description of the genus.</paragraph>
<paragraph id="850F36D9B87BFFB5FF344B69FECA1AF3" blockId="8.[151,1437,1229,2029]" pageId="8" pageNumber="9">
<emphasis id="B7C4EACBB87BFFB5FF344B69FE601A67" bold="true" box="[198,420,1345,1371]" pageId="8" pageNumber="9">Female diagnosis.</emphasis>
Habitus as in
<figureCitation id="1D8B2A5CB87BFFB5FDBC4B6AFD511A60" box="[590,661,1346,1372]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,269,1590,1614]" captionTargetBox="[168,1413,341,1573]" captionTargetId="figure@2.[159,1427,330,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Baronniesia delioti gen. n. sp. n. 1 ― male; 2 ― female." httpUri="https://zenodo.org/record/185599/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
; body smaller than in male (see Variability), pronotum as wide as elytra; outline perfectly oval. Antennae filiform, articles not dilated. For measurements of antennal articles see Table 1. Protarsi tetramerous, non dilated. Urosternite VIII and spermathecal complex as in the description of the genus.
</paragraph>
<paragraph id="850F36D9B87BFFB5FF344BF4FC541921" blockId="8.[151,1437,1229,2029]" pageId="8" pageNumber="9">
<emphasis id="B7C4EACBB87BFFB5FF344BF4FE961ACA" bold="true" box="[198,338,1500,1526]" pageId="8" pageNumber="9">Variability.</emphasis>
Body length (measured from anterior edge of pronotum to apex of elytra): male: 4.02–4.54 mm, average 4.30 mm; female 4.01–4.40 mm; average 4.16 mm.
</paragraph>
</subSubSection>
<subSubSection id="CDAA6552B87BFFB5FF344801FEF11956" pageId="8" pageNumber="9" type="etymology">
<paragraph id="850F36D9B87BFFB5FF344801FEF11956" blockId="8.[151,1437,1229,2029]" pageId="8" pageNumber="9">
<emphasis id="B7C4EACBB87BFFB5FF344801FE97197F" bold="true" box="[198,339,1577,1603]" pageId="8" pageNumber="9">Etymology.</emphasis>
This new species is dedicated to our entomologist colleague and friend Philippe Déliot, who died recently.
</paragraph>
</subSubSection>
<subSubSection id="CDAA6552B87BFFB7FF34485EFE9E1A06" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="distribution">
<paragraph id="850F36D9B87BFFB5FF34485EFB4518D1" blockId="8.[151,1437,1229,2029]" pageId="8" pageNumber="9">
<emphasis id="B7C4EACBB87BFFB5FF34485EFE3A19AC" bold="true" box="[198,510,1654,1680]" pageId="8" pageNumber="9">Distribution and ecology.</emphasis>
The new species is only known from a single cave located in the central part of the Pyrenean chain, south of Esparros. The total length of the cave of Artigaléou is more than one kilometer, with two levels of chalk-stoned fossil galleries before reaching the subterranean river. Specimens of
<taxonomicName id="42B04D5AB87BFFB5FAC248ECFA5819E1" box="[1328,1436,1732,1757]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="delioti">
<emphasis id="B7C4EACBB87BFFB5FAC248ECFA5819E1" box="[1328,1436,1732,1757]" italics="true" pageId="8" pageNumber="9">B. delioti</emphasis>
</taxonomicName>
were found only in traps placed near the river. Apparently, the species does not occur in the deeper parts of the cave, and it seems likely that the specimens found were carried from deep soil by water together with
<taxonomicName id="42B04D5AB87BFFB5FF654910FD8D186E" authority="Coiffait, 1950" authorityName="Coiffait" authorityYear="1950" box="[151,585,1848,1874]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bastideus">
<emphasis id="B7C4EACBB87BFFB5FF654910FE5C186D" box="[151,408,1848,1873]" italics="true" pageId="8" pageNumber="9">Speonomus bastideus</emphasis>
<bibRefCitation id="E1214B28B87BFFB5FE524910FD8D186E" author="Coiffait" box="[416,585,1848,1874]" pageId="8" pageNumber="14" refString="Coiffait, H. (1950 a) Un Speonomus nouveau de la vallee d'Aure (Hautes-Pyrenees). Notes Biospeologiques, 5, 63 - 64." type="journal article" year="1950" yearSuffix="a">Coiffait, 1950</bibRefCitation>
</taxonomicName>
a and the species of Trechini mentioned below. Hypogean fauna of the Baronnies has never been studied in detail, and there are scarce faunistic data from only three other caves (
<bibRefCitation id="E1214B28B87BFFB5FF6D49ADFE8618A3" author="Jeannel" box="[159,322,1925,1951]" pageId="8" pageNumber="15" refString="Jeannel, R. (1928) Monographie des Trechinae. Morphologie comparee et distribution d'un groupe de Coleopteres. Troisieme Livraison: les Trechini cavernicoles. L'Abeille, 35, 1 - 808." type="journal article" year="1928">Jeannel 1928</bibRefCitation>
,
<bibRefCitation id="E1214B28B87BFFB5FEBD49ADFDC718A3" author="Coiffait" box="[335,515,1925,1951]" pageId="8" pageNumber="14" refString="Coiffait, H. (1950 a) Un Speonomus nouveau de la vallee d'Aure (Hautes-Pyrenees). Notes Biospeologiques, 5, 63 - 64." type="journal article" year="1950" yearSuffix="a">Coiffait 1950a</bibRefCitation>
). To better characterize the ecology of
<taxonomicName id="42B04D5AB87BFFB5FC2E49AEFBAA18A3" box="[988,1134,1926,1951]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87BFFB5FC2E49AEFBAA18A3" box="[988,1134,1926,1951]" italics="true" pageId="8" pageNumber="9">Baronniesia</emphasis>
</taxonomicName>
it would be necessary to investigate other localities where it could live, probably as an MSS or endogean species together with
<taxonomicName id="42B04D5AB87BFFB5FF6549FBFE7718D0" box="[151,435,2003,2028]" class="Insecta" family="Carabidae" genus="Geotrechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="discontignyi">
<emphasis id="B7C4EACBB87BFFB5FF6549FBFE7718D0" box="[151,435,2003,2028]" italics="true" pageId="8" pageNumber="9">Geotrechus discontignyi</emphasis>
</taxonomicName>
, a typical endogeous ground beetle also found near the traps.
</paragraph>
<caption id="D1CF6651B87AFFB4FF65494BFEE718DE" httpUri="https://zenodo.org/record/185604/files/figure.png" pageId="9" pageNumber="10" targetBox="[233,1355,186,1872]" targetPageId="9">
<paragraph id="850F36D9B87AFFB4FF65494BFEE718DE" blockId="9.[151,1436,1891,2018]" pageId="9" pageNumber="10">
<emphasis id="B7C4EACBB87AFFB4FF65494BFEE31847" bold="true" box="[151,295,1891,1915]" pageId="9" pageNumber="10">FIGURE 17.</emphasis>
Phylogram of the strict consensus of the 330 trees obtained after successive reweighting of the characters (in the tree, the character weights were reset to unity, so the scale of the branch lengths is uniform across the tree). Taxa with identical character states were lumped in a single terminal taxon except the genera of the
<taxonomicName id="42B04D5AB87AFFB4FB504980FAE61883" box="[1186,1314,1960,1983]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87AFFB4FB504980FAE61883" box="[1186,1314,1960,1983]" italics="true" pageId="9" pageNumber="10">Antrocharis</emphasis>
</taxonomicName>
group (see Appendix 2).
</paragraph>
</caption>
<caption id="D1CF6651B879FFB7FF654A17FD751B45" httpUri="https://zenodo.org/record/185605/files/figure.png" pageId="10" pageNumber="11" targetBox="[152,1434,194,1062]" targetPageId="10">
<paragraph id="850F36D9B879FFB7FF654A17FD751B45" blockId="10.[151,1436,1087,1145]" pageId="10" pageNumber="11">
<emphasis id="B7C4EACBB879FFB7FF654A17FEED1B6B" bold="true" box="[151,297,1087,1111]" pageId="10" pageNumber="11">FIGURE 18.</emphasis>
Distribution of troglobitic Trechini in Pyrenees.
<taxonomicName id="42B04D5AB879FFB7FCB64A68FB171B6A" authority="Bonvouloir" authorityName="Bonvouloir" box="[836,1235,1087,1111]" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subGenus" subGenus="Aphaenops">
<emphasis id="B7C4EACBB879FFB7FCB64A68FC791B6B" box="[836,957,1088,1111]" italics="true" pageId="10" pageNumber="11">Aphaenops</emphasis>
(
<emphasis id="B7C4EACBB879FFB7FC3F4A68FB821B6B" box="[973,1094,1088,1111]" italics="true" pageId="10" pageNumber="11">Aphaenops</emphasis>
) Bonvouloir
</taxonomicName>
(continuous line);
<taxonomicName id="42B04D5AB879FFB7FF654A4AFDEC1B45" authority="Coiffait" authorityName="Coiffait" box="[151,552,1122,1145]" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subGenus" subGenus="Cerbaphaenops">
<emphasis id="B7C4EACBB879FFB7FF654A4AFEC91B45" box="[151,269,1122,1145]" italics="true" pageId="10" pageNumber="11">Aphaenops</emphasis>
(
<emphasis id="B7C4EACBB879FFB7FEEF4A4AFE021B45" box="[285,454,1122,1145]" italics="true" pageId="10" pageNumber="11">Cerbaphaenops</emphasis>
) Coiffait
</taxonomicName>
(doted line).
</paragraph>
</caption>
<paragraph id="850F36D9B879FFB7FF344A83FE9E1A06" blockId="10.[151,1437,1195,2034]" pageId="10" pageNumber="11">
<emphasis id="B7C4EACBB879FFB7FF344A83FE251BF9" bold="true" box="[198,481,1195,1221]" pageId="10" pageNumber="11">Biogeographical notes.</emphasis>
In the area where
<taxonomicName id="42B04D5AB879FFB7FD4B4A84FCE01BF9" box="[697,804,1196,1221]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="delioti">
<emphasis id="B7C4EACBB879FFB7FD4B4A84FCE01BF9" box="[697,804,1196,1221]" italics="true" pageId="10" pageNumber="11">B. delioti</emphasis>
</taxonomicName>
was discovered, the only other known Leptodirini are the species of the
<emphasis id="B7C4EACBB879FFB7FE824AFAFDCE1BD7" box="[368,522,1234,1259]" italics="true" pageId="10" pageNumber="11">speluncarum</emphasis>
group of the genus
<taxonomicName id="42B04D5AB879FFB7FD084AFAFC451BD7" box="[762,897,1234,1259]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB879FFB7FD084AFAFC451BD7" box="[762,897,1234,1259]" italics="true" pageId="10" pageNumber="11">Speonomus</emphasis>
</taxonomicName>
, distributed between the Lez valley (Ariège) and the western Pyrenees, where they have also colonized the southern range in Navarra until the Echo valley (Huesca,
<collectingCountry id="FDA77649B879FFB7FEF74B08FE891A06" box="[261,333,1312,1338]" name="Spain" pageId="10" pageNumber="11">Spain</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="CDAA6552B879FFB7FF344B6EFF2A1980" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="850F36D9B879FFB7FF344B6EFDD91AC7" blockId="10.[151,1437,1195,2034]" pageId="10" pageNumber="11">
<taxonomicName id="42B04D5AB879FFB7FF344B6EFE6F1A63" ID-CoL="68DT6" box="[198,427,1350,1375]" class="Insecta" family="Leiodidae" genus="Baronniesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="delioti">
<emphasis id="B7C4EACBB879FFB7FF344B6EFE6F1A63" box="[198,427,1350,1375]" italics="true" pageId="10" pageNumber="11">Baronniesia delioti</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="42B04D5AB879FFB7FD674B6EFC511A63" ID-CoL="6Z96W" box="[661,917,1350,1375]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bastideus">
<emphasis id="B7C4EACBB879FFB7FD674B6EFC511A63" box="[661,917,1350,1375]" italics="true" pageId="10" pageNumber="11">Speonomus bastideus</emphasis>
</taxonomicName>
(
<emphasis id="B7C4EACBB879FFB7FC584B6EFB851A63" box="[938,1089,1350,1375]" italics="true" pageId="10" pageNumber="11">speluncarum</emphasis>
group) and three species of Trechini:
<taxonomicName id="42B04D5AB879FFB7FEFE4B45FD191ABB" ID-CoL="8LGNF" authority="Gaudin, 1945" authorityName="Gaudin" authorityYear="1945" box="[268,733,1389,1415]" class="Insecta" family="Carabidae" genus="Hydraphaenops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B7C4EACBB879FFB7FEFE4B45FDEA1ABA" box="[268,558,1389,1414]" italics="true" pageId="10" pageNumber="11">Hydraphaenops elegans</emphasis>
<bibRefCitation id="E1214B28B879FFB7FDC54B45FD191ABB" author="Gaudin" box="[567,733,1389,1415]" pageId="10" pageNumber="14" refString="Gaudin, L. (1945) Troglobies nouveaux des Pyrenees francaises (Col., Carabidae). Miscellanea Entomologica, 42 (1), 19 - 24." type="journal article" year="1945">Gaudin, 1945</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="42B04D5AB879FFB7FD194B45FC6C1ABA" box="[747,936,1389,1414]" class="Insecta" family="Carabidae" genus="Cerbaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB879FFB7FD194B45FC6C1ABA" box="[747,936,1389,1414]" italics="true" pageId="10" pageNumber="11">Cerbaphaenops</emphasis>
</taxonomicName>
aff.
<emphasis id="B7C4EACBB879FFB7FC134B45FBF01ABA" box="[993,1076,1389,1414]" italics="true" pageId="10" pageNumber="11">aeacus</emphasis>
<bibRefCitation id="E1214B28B879FFB7FBCE4B45FB1C1ABB" author="Saulcy" box="[1084,1240,1389,1415]" pageId="10" pageNumber="15" refString="Saulcy, F. de. (1864) Faune francaise et europeenne. Descriptions et remarques. Annales de la Societe Entomologique de France, 4 (4), 253 - 260." type="journal article" year="1864">Saulcy, 1864</bibRefCitation>
and
<taxonomicName id="42B04D5AB879FFB7FAE14B45FDA81A91" ID-CoL="8M87W" authority="Cabidoche, 1967" authorityName="Cabidoche" authorityYear="1967" class="Insecta" family="Carabidae" genus="Geotrechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subSpecies" species="discontignyi" subSpecies="canteti">
<emphasis id="B7C4EACBB879FFB7FAE14B45FE491A91" italics="true" pageId="10" pageNumber="11">Geotrechus discontignyi canteti</emphasis>
<bibRefCitation id="E1214B28B879FFB7FE674BBBFDA81A91" author="Cabidoche" box="[405,620,1427,1453]" pageId="10" pageNumber="14" refString="Cabidoche, M. (1967) Coleopteres troglobies et endoges des Pyrenees Occidentales (Col. Carab.). Annales de Speleologie, 22 (3), 647 - 658." type="journal article" year="1967">Cabidoche, 1967</bibRefCitation>
</taxonomicName>
. The subterranean river of Artigaléou-Arodets is the third known locality of
<emphasis id="B7C4EACBB879FFB7FEE64B92FDD31AEF" box="[276,535,1466,1491]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="42B04D5AB879FFB7FEE64B92FDD71AEF" ID-CoL="6Z96W" box="[276,531,1466,1491]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bastideus">Speonomus bastideus</taxonomicName>
,
</emphasis>
which was so far only known from the caves of Labastide and Diable Rouge, both in Banios, Hautes-Pyrénées.
</paragraph>
<paragraph id="850F36D9B879FFB7FF344820FF2A1980" blockId="10.[151,1437,1195,2034]" pageId="10" pageNumber="11">
It may be expected that the two main unrelated groups of cave
<taxonomicName id="42B04D5AB879FFB7FC244820FA58191E" ID-CoL="C2L" authority="Leptodirini and Trechini" authorityName="Leptodirini and Trechini" box="[982,1436,1544,1570]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="order">Coleoptera (Leptodirini and Trechini)</taxonomicName>
living in the same geographical area show some similarities in their patterns of distribution and colonization of the subterranean environment. Both live in the same underground ecosystem, characterized by strict and constant features, and share the area affected by the same climatic conditions and a particular paleogeographic history.
</paragraph>
</subSubSection>
<subSubSection id="CDAA6552B879FFB1FF3448E1FD0C1B44" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
<paragraph id="850F36D9B879FFB1FF3448E1FBA21EA6" blockId="10.[151,1437,1195,2034]" lastBlockId="12.[151,1437,152,1144]" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
The pattern of distribution of the Pyrenean Trechini is presented in
<figureCitation id="1D8B2A5CB879FFB7FC1848E1FB8619DF" box="[1002,1090,1737,1763]" captionStart="FIGURE 18" captionStartId="10.[151,256,1087,1111]" captionTargetBox="[152,1434,194,1062]" captionTargetId="figure@10.[152,1435,194,1063]" captionTargetPageId="10" captionText="FIGURE 18. Distribution of troglobitic Trechini in Pyrenees. Aphaenops (Aphaenops) Bonvouloir (continuous line); Aphaenops (Cerbaphaenops) Coiffait (doted line)." httpUri="https://zenodo.org/record/185605/files/figure.png" pageId="10" pageNumber="11">Fig. 18</figureCitation>
. The species of the subgenus
<taxonomicName id="42B04D5AB879FFB7FF6548D8FE391836" authority="Bonvouloir, 1862" authorityName="Bonvouloir" authorityYear="1862" box="[151,509,1776,1802]" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB879FFB7FF6548D8FEDA1835" box="[151,286,1776,1801]" italics="true" pageId="10" pageNumber="11">Aphaenops</emphasis>
<bibRefCitation id="E1214B28B879FFB7FED448D8FE391836" author="Bonvouloir" box="[294,509,1776,1802]" pageId="10" pageNumber="14" refString="Bonvouloir, H. de (1862) Description d'un genre nouveau et de deux especes nouvelles de coleopteres de France. Annales de la Societe entomologique de France, (4) 1 [1861], 567 - 571." type="journal article" year="1862">Bonvouloir, 1862</bibRefCitation>
</taxonomicName>
have radiated in the western part of the Pyrenean chain, and their diversity increases from Lourdes to the west, the majority of them (including the endogean
<taxonomicName id="42B04D5AB879FFB7FB9E493EFF12186B" authority="Cabidoche, 1965" authorityName="Cabidoche" authorityYear="1965" class="Insecta" family="Carabidae" genus="Geaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB879FFB7FB9E493EFACB1813" box="[1132,1295,1814,1839]" italics="true" pageId="10" pageNumber="11">Geaphaenops</emphasis>
<bibRefCitation id="E1214B28B879FFB7FAE4493EFF12186B" author="Cabidoche" pageId="10" pageNumber="14" refString="Cabidoche, M. (1965) Sur les Aphaenops du groupe rhadamanthus (Col. Carab.). Annales de Speleologie, 20 (4), 523 - 528." type="journal article" year="1965">Cabidoche, 1965</bibRefCitation>
</taxonomicName>
) diversified between Lourdes and Saint-Jean-Pied-de-Port (
<bibRefCitation id="E1214B28B879FFB7FC4D4915FB97186B" author="Faille" box="[959,1107,1853,1879]" pageId="10" pageNumber="14" refString="Faille, A. (2006) Endemisme et adaptation a la vie cavernicole chez les Trechinae Pyreneens (Coleoptera: Carabidae). Approches moleculaire et morphometrique. These inedite. Museum National d'Histoire Naturelle, Paris. 319 pp." type="book" year="2006">Faille 2006</bibRefCitation>
). The eastern-most French species of the subgenus
<taxonomicName id="42B04D5AB879FFB7FE3A494CFD951841" box="[456,593,1892,1917]" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB879FFB7FE3A494CFD951841" box="[456,593,1892,1917]" italics="true" pageId="10" pageNumber="11">Aphaenops</emphasis>
</taxonomicName>
is
<taxonomicName id="42B04D5AB879FFB7FD8A494CFBDE1841" authority="Bonvouloir, 1862" authorityName="Bonvouloir" authorityYear="1862" box="[632,1050,1891,1917]" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="leschenaulti">
<emphasis id="B7C4EACBB879FFB7FD8A494CFCF71841" box="[632,819,1892,1917]" italics="true" pageId="10" pageNumber="11">A. leschenaulti</emphasis>
<bibRefCitation id="E1214B28B879FFB7FCCE494BFBDE1841" author="Bonvouloir" box="[828,1050,1891,1917]" pageId="10" pageNumber="14" refString="Bonvouloir, H. de (1862) Description d'un genre nouveau et de deux especes nouvelles de coleopteres de France. Annales de la Societe entomologique de France, (4) 1 [1861], 567 - 571." type="journal article" year="1862">Bonvouloir, 1862</bibRefCitation>
</taxonomicName>
, known from the caves around Bagnères-de-Bigorre. On the contrary, the species of the subgenus
<taxonomicName id="42B04D5AB879FFB7FC2C49A2FA9E1898" authority="Coiffait, 1962" authorityName="Coiffait" authorityYear="1962" box="[990,1370,1930,1956]" class="Insecta" family="Carabidae" genus="Cerbaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB879FFB7FC2C49A2FB65189F" box="[990,1185,1930,1955]" italics="true" pageId="10" pageNumber="11">Cerbaphaenops</emphasis>
Coiffait, 1962
</taxonomicName>
have radiated in the eastern part of the Pyrenees between Ariège valley and Bigorre area, and their diversity decreases from east to west (
<bibRefCitation id="E1214B28B879FFB7FE1549F0FDB618CE" author="Faille" box="[487,626,2008,2034]" pageId="10" pageNumber="14" refString="Faille, A. (2006) Endemisme et adaptation a la vie cavernicole chez les Trechinae Pyreneens (Coleoptera: Carabidae). Approches moleculaire et morphometrique. These inedite. Museum National d'Histoire Naturelle, Paris. 319 pp." type="book" year="2006">Faille 2006</bibRefCitation>
). The surrounding area of Bagnères-de-Bigorre is the contact area between the two main radiations of subterranean cave Trechini,
<taxonomicName id="42B04D5AB87FFFB1FC694EB0FBDB1F8D" box="[923,1055,152,177]" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FC694EB0FBDB1F8D" box="[923,1055,152,177]" italics="true" pageId="12" pageNumber="13">Aphaenops</emphasis>
</taxonomicName>
in the narrow sense in the west and
<taxonomicName id="42B04D5AB87FFFB1FF3E4E97FE4A1FE4" box="[204,398,191,216]" class="Insecta" family="Carabidae" genus="Cerbaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FF3E4E97FE4A1FE4" box="[204,398,191,216]" italics="true" pageId="12" pageNumber="13">Cerbaphaenops</emphasis>
</taxonomicName>
in the central-eastern Pyrenees. In this area, the two lineages are represented by
<taxonomicName id="42B04D5AB87FFFB1FA704E97FE2F1FC2" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="crypticola" subGenus="Cerbaphaenops">
<emphasis id="B7C4EACBB87FFFB1FA704E97FA501FE4" box="[1410,1428,191,216]" italics="true" pageId="12" pageNumber="13">A</emphasis>
. (
<emphasis id="B7C4EACBB87FFFB1FF534ECDFE2F1FC2" box="[161,491,229,254]" italics="true" pageId="12" pageNumber="13">Cerbaphaenops) crypticola</emphasis>
</taxonomicName>
ssp.
<emphasis id="B7C4EACBB87FFFB1FDD84ECDFDB91FC2" box="[554,637,229,254]" italics="true" pageId="12" pageNumber="13">aeacus</emphasis>
<bibRefCitation id="E1214B28B87FFFB1FD744ECDFCE51FC3" author="Saulcy" box="[646,801,229,255]" pageId="12" pageNumber="15" refString="Saulcy, F. de. (1864) Faune francaise et europeenne. Descriptions et remarques. Annales de la Societe Entomologique de France, 4 (4), 253 - 260." type="journal article" year="1864">Saulcy, 1864</bibRefCitation>
and
<taxonomicName id="42B04D5AB87FFFB1FCAF4ECDFA9E1FC3" authority="Saulcy, 1864" authorityName="Saulcy" authorityYear="1864" box="[861,1370,229,255]" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="leschenaulti" subGenus="Aphaenops">
<emphasis id="B7C4EACBB87FFFB1FCAF4ECDFCAB1FC2" box="[861,879,229,254]" italics="true" pageId="12" pageNumber="13">A</emphasis>
. (
<emphasis id="B7C4EACBB87FFFB1FC784ECDFB771FC2" box="[906,1203,229,254]" italics="true" pageId="12" pageNumber="13">Aphaenops) leschenaulti</emphasis>
<bibRefCitation id="E1214B28B87FFFB1FB494ECDFA9E1FC3" author="Saulcy" box="[1211,1370,229,255]" pageId="12" pageNumber="15" refString="Saulcy, F. de. (1864) Faune francaise et europeenne. Descriptions et remarques. Annales de la Societe Entomologique de France, 4 (4), 253 - 260." type="journal article" year="1864">Saulcy, 1864</bibRefCitation>
</taxonomicName>
, both sympatric in the cavities around Bagnères-de-Bigorre, in particular the Castelmouly and Tuco caves (
<bibRefCitation id="E1214B28B87FFFB1FAB04F24FF121E70" author="Jeannel" pageId="12" pageNumber="15" refString="Jeannel, R. (1928) Monographie des Trechinae. Morphologie comparee et distribution d'un groupe de Coleopteres. Troisieme Livraison: les Trechini cavernicoles. L'Abeille, 35, 1 - 808." type="journal article" year="1928">Jeannel 1928</bibRefCitation>
). While the species of the subgenus
<taxonomicName id="42B04D5AB87FFFB1FD764F1AFCC31E77" box="[644,775,306,331]" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FD764F1AFCC31E77" box="[644,775,306,331]" italics="true" pageId="12" pageNumber="13">Aphaenops</emphasis>
</taxonomicName>
on the northern slope of the range do not reach the area east of Bagnères-de-Bigorre, on the southern slope some species of this subgenus are distributed further east, till the Ribagorçana valley, where
<taxonomicName id="42B04D5AB87FFFB1FDDA4FA8FC2C1EA6" authority="Escola &amp; Cancio, 1983" authorityName="Escola &amp; Cancio" authorityYear="1983" box="[552,1000,384,410]" class="Insecta" family="Carabidae" genus="Aphaenops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="catalonicus">
<emphasis id="B7C4EACBB87FFFB1FDDA4FA8FD141EA5" box="[552,720,384,409]" italics="true" pageId="12" pageNumber="13">A. catalonicus</emphasis>
<bibRefCitation id="E1214B28B87FFFB1FD254FA8FC2C1EA6" author="Escola" box="[727,1000,384,410]" pageId="12" pageNumber="14" refString="Escola, O. &amp; Cancio, E. (1983) Un nou Aphaenops de la Ribagorca (Coleoptera Trechinae). Speleon, 26 - 27, 75 - 77." type="book chapter" year="1983">Escolà &amp; Canció, 1983</bibRefCitation>
</taxonomicName>
is present.
</paragraph>
<caption id="D1CF6651B878FFB6FF654909FAC418D0" httpUri="https://zenodo.org/record/185606/files/figure.png" pageId="11" pageNumber="12" targetBox="[405,1215,199,1805]" targetPageId="11">
<paragraph id="850F36D9B878FFB6FF654909FAC418D0" blockId="11.[151,1437,1825,2028]" pageId="11" pageNumber="12">
<emphasis id="B7C4EACBB878FFB6FF654909FEE31805" bold="true" box="[151,295,1825,1849]" pageId="11" pageNumber="12">FIGURE 19.</emphasis>
Map of the Pyrenees with distribution areas of taxa with different models of lateral styles of the aedeagus in Leptodirini. The
<taxonomicName id="42B04D5AB878FFB6FE88496DFDC51860" box="[378,513,1861,1884]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB878FFB6FE88496DFDC51860" box="[378,513,1861,1884]" italics="true" pageId="11" pageNumber="12">Antrocharis</emphasis>
</taxonomicName>
type (continuous line):
<taxonomicName id="42B04D5AB878FFB6FCE4496DFB0C1860" authority="Abeille" authorityName="Abeille" box="[790,1224,1861,1884]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="querilhaci" subSpecies="dispar">
<emphasis id="B7C4EACBB878FFB6FCE4496DFBA81860" box="[790,1132,1861,1884]" italics="true" pageId="11" pageNumber="12">Antrocharis querilhaci dispar</emphasis>
Abeille
</taxonomicName>
de Perrin (a). The
<taxonomicName id="42B04D5AB878FFB6FF654941FED918BC" box="[151,285,1897,1920]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB878FFB6FF654941FED918BC" box="[151,285,1897,1920]" italics="true" pageId="11" pageNumber="12">Troglophyes</emphasis>
</taxonomicName>
type (dots and dashes):
<taxonomicName id="42B04D5AB878FFB6FDDA4941FBEF18BC" authority="Dupre" authorityName="Dupre" box="[552,1067,1897,1920]" class="Insecta" family="Leiodidae" genus="Phacomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fratyi" subGenus="Phacomorphus">
<emphasis id="B7C4EACBB878FFB6FDDA4941FD0818BC" box="[552,716,1897,1920]" italics="true" pageId="11" pageNumber="12">Phacomorphus</emphasis>
(
<emphasis id="B7C4EACBB878FFB6FD2F4941FC4518BC" box="[733,897,1897,1920]" italics="true" pageId="11" pageNumber="12">Phacomorphus</emphasis>
)
<emphasis id="B7C4EACBB878FFB6FC604941FC0A18BC" box="[914,974,1897,1920]" italics="true" pageId="11" pageNumber="12">fratyi</emphasis>
(Dupré)
</taxonomicName>
(b),
<taxonomicName id="42B04D5AB878FFB6FB904941FEA11898" authority="Escola &amp; Fresneda" authorityName="Escola &amp; Fresneda" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ere" subGenus="Speonomus">
<emphasis id="B7C4EACBB878FFB6FB904941FB1A18BC" box="[1122,1246,1897,1920]" italics="true" pageId="11" pageNumber="12">Speonomus</emphasis>
(
<emphasis id="B7C4EACBB878FFB6FB1C4941FAAD18BC" box="[1262,1385,1897,1920]" italics="true" pageId="11" pageNumber="12">Speonomus</emphasis>
)
<emphasis id="B7C4EACBB878FFB6FA884941FA5818BC" box="[1402,1436,1897,1920]" italics="true" pageId="11" pageNumber="12">ere</emphasis>
Escolà &amp; Fresneda
</taxonomicName>
(c),
<taxonomicName id="42B04D5AB878FFB6FE6B49A5FD701898" authority="Jeannel" authorityName="Jeannel" box="[409,692,1933,1956]" class="Insecta" family="Leiodidae" genus="Perriniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="faurai">
<emphasis id="B7C4EACBB878FFB6FE6B49A5FD9F1898" box="[409,603,1933,1956]" italics="true" pageId="11" pageNumber="12">Perriniella faurai</emphasis>
Jeannel
</taxonomicName>
(d),
<taxonomicName id="42B04D5AB878FFB6FD1849A5FB421898" authority="Auroux &amp; Belles" authorityName="Auroux &amp; Belles" box="[746,1158,1933,1956]" class="Insecta" family="Leiodidae" genus="Bellesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="espanyoli">
<emphasis id="B7C4EACBB878FFB6FD1849A5FC751898" box="[746,945,1933,1956]" italics="true" pageId="11" pageNumber="12">Bellesia espanyoli</emphasis>
(Auroux &amp; Bellés)
</taxonomicName>
(e) and
<taxonomicName id="42B04D5AB878FFB6FB1349A5FEA318F4" authority="Espanol" authorityName="Espanol" class="Insecta" family="Leiodidae" genus="Parvospeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="urgellesi">
<emphasis id="B7C4EACBB878FFB6FB1349A5FF3218F4" italics="true" pageId="11" pageNumber="12">Parvospeonomus urgellesi</emphasis>
(Español)
</taxonomicName>
(f). The
<taxonomicName id="42B04D5AB878FFB6FE384999FD8A18F4" box="[458,590,1969,1992]" class="Insecta" family="Leiodidae" genus="Trocharanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB878FFB6FE384999FD8A18F4" box="[458,590,1969,1992]" italics="true" pageId="11" pageNumber="12">Trocharanis</emphasis>
</taxonomicName>
type (doted line):
<taxonomicName id="42B04D5AB878FFB6FCE94999FADE18F4" box="[795,1306,1969,1992]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="curvipes" subGenus="Speonomus" subSpecies="subcurvipes">
<emphasis id="B7C4EACBB878FFB6FCE94999FC5318F4" box="[795,919,1969,1992]" italics="true" pageId="11" pageNumber="12">Speonomus</emphasis>
(
<emphasis id="B7C4EACBB878FFB6FC554999FBE718F4" box="[935,1059,1969,1992]" italics="true" pageId="11" pageNumber="12">Speonomus</emphasis>
)
<emphasis id="B7C4EACBB878FFB6FBC64999FADE18F4" box="[1076,1306,1969,1992]" italics="true" pageId="11" pageNumber="12">curvipes subcurvipes</emphasis>
</taxonomicName>
(Abeille de Perrin) (g),
<taxonomicName id="42B04D5AB878FFB6FEE649FDFDEC18D0" authority="Jeannel" authorityName="Jeannel" box="[276,552,2005,2028]" class="Insecta" family="Leiodidae" genus="Salgadoia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brieti">
<emphasis id="B7C4EACBB878FFB6FEE649FDFE0518D0" box="[276,449,2005,2028]" italics="true" pageId="11" pageNumber="12">Salgadoia brieti</emphasis>
(Jeannel)
</taxonomicName>
(h) and
<taxonomicName id="42B04D5AB878FFB6FD7049FDFB1D18D0" authority="Reitter" authorityName="Reitter" box="[642,1241,2005,2028]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="ferreri" subGenus="Troglocharinus" subSpecies="ferreri">
<emphasis id="B7C4EACBB878FFB6FD7049FDFCE118D0" box="[642,805,2005,2028]" italics="true" pageId="11" pageNumber="12">Troglocharinus</emphasis>
(
<emphasis id="B7C4EACBB878FFB6FCC749FDFC1D18D0" box="[821,985,2005,2028]" italics="true" pageId="11" pageNumber="12">Troglocharinus</emphasis>
)
<emphasis id="B7C4EACBB878FFB6FC1A49FDFBB318D0" box="[1000,1143,2005,2028]" italics="true" pageId="11" pageNumber="12">ferreri ferreri</emphasis>
(Reitter)
</taxonomicName>
(i).
</paragraph>
</caption>
<paragraph id="850F36D9B87FFFB1FF344F8EFC361C20" blockId="12.[151,1437,152,1144]" pageId="12" pageNumber="13">
The pattern of diversification of the Pyrenean Leptodirini is quite similar, although with some differences (
<figureCitation id="1D8B2A5CB87FFFB1FF524FE5FF3D1EDB" box="[160,249,461,487]" captionStart="FIGURE 19" captionStartId="11.[151,255,1825,1849]" captionTargetBox="[405,1215,199,1805]" captionTargetId="figure@11.[395,1226,192,1810]" captionTargetPageId="11" captionText="FIGURE 19. Map of the Pyrenees with distribution areas of taxa with different models of lateral styles of the aedeagus in Leptodirini. The Antrocharis type (continuous line): Antrocharis querilhaci dispar Abeille de Perrin (a). The Troglophyes type (dots and dashes): Phacomorphus (Phacomorphus) fratyi (Dupré) (b), Speonomus (Speonomus) ere Escolà &amp; Fresneda (c), Perriniella faurai Jeannel (d), Bellesia espanyoli (Auroux &amp; Bellés) (e) and Parvospeonomus urgellesi (Español) (f). The Trocharanis type (doted line): Speonomus (Speonomus) curvipes subcurvipes (Abeille de Perrin) (g), Salgadoia brieti (Jeannel) (h) and Troglocharinus (Troglocharinus) ferreri ferreri (Reitter) (i)." httpUri="https://zenodo.org/record/185606/files/figure.png" pageId="12" pageNumber="13">Fig. 19</figureCitation>
). The species of
<taxonomicName id="42B04D5AB87FFFB1FE314FE5FD8D1EDA" box="[451,585,461,486]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FE314FE5FD8D1EDA" box="[451,585,461,486]" italics="true" pageId="12" pageNumber="13">Speonomus</emphasis>
</taxonomicName>
of the
<emphasis id="B7C4EACBB87FFFB1FD6D4FE5FCFC1EDA" box="[671,824,461,486]" italics="true" pageId="12" pageNumber="13">speluncarum</emphasis>
group are distributed from the western Pyrenees to the Aure valley (Sarrancolin, Lortet) and the Labastide area (Artigaléou, Labastide cave). Members of the
<emphasis id="B7C4EACBB87FFFB1FF654C32FEF71D0F" box="[151,307,538,563]" italics="true" pageId="12" pageNumber="13">speluncarum</emphasis>
group are found again in Ariège, with
<taxonomicName id="42B04D5AB87FFFB1FCE04C32FB961D08" authority="Gers, 1989" authorityName="Gers" authorityYear="1989" box="[786,1106,538,564]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="orgibetensis">
<emphasis id="B7C4EACBB87FFFB1FCE04C32FC001D0F" box="[786,964,538,563]" italics="true" pageId="12" pageNumber="13">S. orgibetensis</emphasis>
<bibRefCitation id="E1214B28B87FFFB1FC3F4C32FB961D08" author="Gers" box="[973,1106,538,564]" pageId="12" pageNumber="14" refString="Gers, C. (1989) Speonomus orgibetensis, n. sp., coleoptere Bathysciinae troglobie du milieu souterrain superficiel. Annales de la Societe entomologique de France (N. S.), 25 (1), 105 - 116." type="journal article" year="1989">Gers, 1989</bibRefCitation>
</taxonomicName>
(
<emphasis id="B7C4EACBB87FFFB1FB974C32FAC51D0F" box="[1125,1281,538,563]" italics="true" pageId="12" pageNumber="13">speluncarum</emphasis>
group
<emphasis id="B7C4EACBB87FFFB1FAAB4C32FA581D0F" box="[1369,1436,538,563]" italics="true" pageId="12" pageNumber="13">sensu</emphasis>
<bibRefCitation id="E1214B28B87FFFB1FF654C69FEEC1D67" author="Gers" box="[151,296,577,603]" pageId="12" pageNumber="14" refString="Gers, C. (1989) Speonomus orgibetensis, n. sp., coleoptere Bathysciinae troglobie du milieu souterrain superficiel. Annales de la Societe entomologique de France (N. S.), 25 (1), 105 - 116." type="journal article" year="1989">Gers, 1989</bibRefCitation>
:
<figureCitation id="1D8B2A5CB87FFFB1FEC44C69FEBA1D67" box="[310,382,577,603]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,269,1590,1614]" captionTargetBox="[168,1413,341,1573]" captionTargetId="figure@2.[159,1427,330,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Baronniesia delioti gen. n. sp. n. 1 ― male; 2 ― female." httpUri="https://zenodo.org/record/185599/files/figure.png" pageId="12" pageNumber="13">Fig. 1</figureCitation>
D), the two species of the
<taxonomicName id="42B04D5AB87FFFB1FD2A4C69FCF41D66" box="[728,816,577,602]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ehlersi">
<emphasis id="B7C4EACBB87FFFB1FD2A4C69FCF41D66" box="[728,816,577,602]" italics="true" pageId="12" pageNumber="13">ehlersi</emphasis>
</taxonomicName>
group,
<taxonomicName id="42B04D5AB87FFFB1FC664C69FBB01D67" authority="Abeille" authorityName="Abeille" box="[916,1140,577,603]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ehlersi">
<emphasis id="B7C4EACBB87FFFB1FC664C69FBCA1D66" box="[916,1038,577,602]" italics="true" pageId="12" pageNumber="13">S. ehlersi</emphasis>
Abeille
</taxonomicName>
de Perrin, 1872 and
<taxonomicName id="42B04D5AB87FFFB1FA774C69FDF81DBE" authority="Gers &amp; Dupuis, 1988" authorityName="Gers &amp; Dupuis" authorityYear="1988" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="opisthonoxus">
<emphasis id="B7C4EACBB87FFFB1FA774C69FEF01DBD" italics="true" pageId="12" pageNumber="13">S. opisthonoxus</emphasis>
<bibRefCitation id="E1214B28B87FFFB1FECE4C40FDF81DBE" author="Gers" box="[316,572,616,642]" pageId="12" pageNumber="14" refString="Gers, C. &amp; Dupuis, M. (1988) Une nouvelle espece remarquable du genre Speonomus (Coleoptera, Bathysciinae)." type="book" year="1988">Gers &amp; Dupuis, 1988</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="E1214B28B87FFFB1FDBF4C40FC721DBE" author="Gers" box="[589,950,616,642]" pageId="12" pageNumber="14" refString="Gers, C. &amp; Dupuis, M. (1988) Une nouvelle espece remarquable du genre Speonomus (Coleoptera, Bathysciinae)." type="book" year="1988">
Gers &amp; Dupuis 1988:
<figureCitation id="1D8B2A5CB87FFFB1FCAB4C40FC5B1DBE" box="[857,927,616,642]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,269,1501,1525]" captionTargetBox="[375,1213,477,1469]" captionTargetId="figure@3.[365,1221,465,1477]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Aedeagus of Baronniesia delioti gen. n. sp. n. 3 — dorsal view; 4 — lateral view." httpUri="https://zenodo.org/record/185600/files/figure.png" pageId="12" pageNumber="13">Fig. 3</figureCitation>
C
</bibRefCitation>
), and the subgenus
<taxonomicName id="42B04D5AB87FFFB1FB514C40FA9A1DBD" box="[1187,1374,616,641]" class="Insecta" family="Leiodidae" genus="Machaeroscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FB514C40FA9A1DBD" box="[1187,1374,616,641]" italics="true" pageId="12" pageNumber="13">Machaeroscelis</emphasis>
</taxonomicName>
. The distribution area of the
<emphasis id="B7C4EACBB87FFFB1FE5A4CA7FDFB1D94" box="[424,575,655,680]" italics="true" pageId="12" pageNumber="13">speluncarum</emphasis>
group and related Leptodirini are thus extended to the east. The presence of other genera apparently related to the
<emphasis id="B7C4EACBB87FFFB1FD864C9DFCCF1DF2" box="[628,779,693,718]" italics="true" pageId="12" pageNumber="13">speluncarum</emphasis>
group (
<taxonomicName id="42B04D5AB87FFFB1FC9B4C9DFC3F1DF2" box="[873,1019,693,718]" class="Insecta" family="Leiodidae" genus="Ceretophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FC9B4C9DFC3F1DF2" box="[873,1019,693,718]" italics="true" pageId="12" pageNumber="13">Ceretophyes</emphasis>
</taxonomicName>
,
<taxonomicName id="42B04D5AB87FFFB1FBFB4C9DFB171DF2" box="[1033,1235,693,718]" class="Insecta" family="Leiodidae" genus="Parvospeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FBFB4C9DFB171DF2" box="[1033,1235,693,718]" italics="true" pageId="12" pageNumber="13">Parvospeonomus</emphasis>
</taxonomicName>
,
<taxonomicName id="42B04D5AB87FFFB1FB104C9DFAA61DF2" box="[1250,1378,693,718]" class="Insecta" family="Leiodidae" genus="Perriniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FB104C9DFAA61DF2" box="[1250,1378,693,718]" italics="true" pageId="12" pageNumber="13">Perriniella</emphasis>
</taxonomicName>
, and
<taxonomicName id="42B04D5AB87FFFB1FF654CF4FEF51DC9" box="[151,305,732,757]" class="Insecta" family="Leiodidae" genus="Troglophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FF654CF4FEF51DC9" box="[151,305,732,757]" italics="true" pageId="12" pageNumber="13">Troglophyes</emphasis>
</taxonomicName>
) in the eastern part of the Pyrenees is very remarkable. A revision of the north Pyrenean Leptodirini would be necessary to elucidate this biogeographical enigma.
</paragraph>
<paragraph id="850F36D9B87FFFB1FF344D01FD0C1B44" blockId="12.[151,1437,152,1144]" pageId="12" pageNumber="13">
When Leptodirini are compared to troglobitic Trechini, the main difference in their distribution is that in Leptodirini the contact area of the two groups, the western one (
<taxonomicName id="42B04D5AB87FFFB1FC454D78FB861C55" box="[951,1090,848,873]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FC454D78FB861C55" box="[951,1090,848,873]" italics="true" pageId="12" pageNumber="13">Speonomus</emphasis>
</taxonomicName>
of
<emphasis id="B7C4EACBB87FFFB1FB9F4D78FACF1C55" box="[1133,1291,848,873]" italics="true" pageId="12" pageNumber="13">speluncarum</emphasis>
and
<taxonomicName id="42B04D5AB87FFFB1FABA4D78FA581C55" box="[1352,1436,848,873]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ehlersi">
<emphasis id="B7C4EACBB87FFFB1FABA4D78FA581C55" box="[1352,1436,848,873]" italics="true" pageId="12" pageNumber="13">ehlersi</emphasis>
</taxonomicName>
groups,
<taxonomicName id="42B04D5AB87FFFB1FF094D5FFE781CAC" box="[251,444,887,912]" class="Insecta" family="Leiodidae" genus="Machaeroscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FF094D5FFE781CAC" box="[251,444,887,912]" italics="true" pageId="12" pageNumber="13">Machaeroscelis</emphasis>
</taxonomicName>
,
<taxonomicName id="42B04D5AB87FFFB1FE3F4D5FFDA11CAC" box="[461,613,887,912]" class="Insecta" family="Leiodidae" genus="Bathysciella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FE3F4D5FFDA11CAC" box="[461,613,887,912]" italics="true" pageId="12" pageNumber="13">Bathysciella</emphasis>
</taxonomicName>
and
<taxonomicName id="42B04D5AB87FFFB1FD514D5FFC981CAC" box="[675,860,887,912]" class="Insecta" family="Leiodidae" genus="Phacomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FD514D5FFC981CAC" box="[675,860,887,912]" italics="true" pageId="12" pageNumber="13">Phacomorphus</emphasis>
</taxonomicName>
) and the central-eastern one (
<emphasis id="B7C4EACBB87FFFB1FB224D5FFEED1C8A" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="42B04D5AB87FFFB1FB224D5FFA5C1CAC" box="[1232,1432,887,912]" class="Insecta" family="Leiodidae" genus="Paraspeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Paraspeonomus</taxonomicName>
,
<taxonomicName id="42B04D5AB87FFFB1FF654DB5FEED1C8A" box="[151,297,925,950]" class="Insecta" family="Leiodidae" genus="Trocharanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Trocharanis</taxonomicName>
</emphasis>
and
<taxonomicName id="42B04D5AB87FFFB1FE964DB5FE281C8A" box="[356,492,925,950]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FE964DB5FE281C8A" box="[356,492,925,950]" italics="true" pageId="12" pageNumber="13">Speonomus</emphasis>
</taxonomicName>
of the
<emphasis id="B7C4EACBB87FFFB1FDB64DB5FD6B1C8A" box="[580,687,925,950]" italics="true" pageId="12" pageNumber="13">pyreneus</emphasis>
group) is extended to the east until Ariège. The area between Ariège and Bigorre seems to be an overlapping area of distribution for the Leptodirini of the
<taxonomicName id="42B04D5AB87FFFB1FB264DECFAA61CE1" box="[1236,1378,964,989]" class="Insecta" family="Leiodidae" genus="Trocharanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FB264DECFAA61CE1" box="[1236,1378,964,989]" italics="true" pageId="12" pageNumber="13">Trocharanis</emphasis>
</taxonomicName>
<typeStatus id="5A0B887BB87FFFB1FA9B4DECFA581CE2" box="[1385,1436,964,990]" pageId="12" pageNumber="13">type</typeStatus>
and those of the
<taxonomicName id="42B04D5AB87FFFB1FEAB4DC2FE1B1B3F" box="[345,479,1002,1027]" class="Insecta" family="Leiodidae" genus="Speonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FEAB4DC2FE1B1B3F" box="[345,479,1002,1027]" italics="true" pageId="12" pageNumber="13">Speonomus</emphasis>
</taxonomicName>
of the
<emphasis id="B7C4EACBB87FFFB1FDC04DC2FD0D1B3F" box="[562,713,1002,1027]" italics="true" pageId="12" pageNumber="13">speluncarum</emphasis>
group. The lineage of the
<taxonomicName id="42B04D5AB87FFFB1FBF24DC2FB4F1B3F" box="[1024,1163,1002,1027]" class="Insecta" family="Leiodidae" genus="Antrocharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B7C4EACBB87FFFB1FBF24DC2FB4F1B3F" box="[1024,1163,1002,1027]" italics="true" pageId="12" pageNumber="13">Antrocharis</emphasis>
</taxonomicName>
group of genera seems to have originated in this contact area, but we do not know whether from the western or the eastern group. The coincidence of the same contact area for the eastern and western Pyrenean groups of Leptodirini and Trechini suggests the same geologic and climatic causes.
</paragraph>
</subSubSection>
</treatment>
</document>
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