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179 lines
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<document id="5B43FEB39585455DEC0C0E1D772C84E5" ID="10.11646/zootaxa.4162.1.7" ID-DOI="10.11646/zootaxa.4162.1.7" ID-GBIF-Dataset="9f3ae59e-4c5b-4342-99b1-19f650f3aeae" ID-ISSN="1175-5326" ID-Zenodo-Dep="254692" ID-ZooBank="9E192699-C74F-44EB-BEE2-C7298A88DBA5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1473440432386" checkinUser="plazi" docAuthor="Dellacasa, Marco & Dellacasa, Giovanni" docDate="2016" docId="FF798790C2760F7BFF31FC45F54D1605" docLanguage="en" docName="zootaxa.4162.1.7.pdf" docOrigin="Zootaxa 4162 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phaeaphodius himalorectus Ahrens & Stebnicka 1997" docType="treatment" docVersion="14" lastPageNumber="149" masterDocId="0340FFE8C2700F7DFFA6FFC9F1611161" masterDocTitle="Systematic revision of the genus Phaeaphodius Reitter, 1892 (Coleoptera: Scarabaeidae: Aphodiinae)" masterLastPageNumber="163" masterPageNumber="143" pageNumber="149" updateTime="1698384649814" updateUser="plazi">
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<mods:title id="586FB338657BDC02FE671D2669562280">Systematic revision of the genus Phaeaphodius Reitter, 1892 (Coleoptera: Scarabaeidae: Aphodiinae)</mods:title>
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<mods:namePart id="1B2283B2A9B889EE80FF27E02D02E8D2">Dellacasa, Marco</mods:namePart>
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<mods:namePart id="8F405557DD3946CFF10570C575593C84">Dellacasa, Giovanni</mods:namePart>
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<mods:typeOfResource id="CA9B36558CFC933FB322F3695F3CCB5C">text</mods:typeOfResource>
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<mods:title id="8238235879F5573CCEFF48DE8759B991">Zootaxa</mods:title>
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<mods:part id="DBBE9E7E96279524FF26D70D55CAE379">
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<mods:date id="F056119CE26F555B4B96F5A3EB56A97A">2016</mods:date>
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<mods:detail id="5204E27B9BE82D5A2393FC9C4D8654EB" type="volume">
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<mods:number id="B54F1443EBB5EDB354752EA2143A650C">4162</mods:number>
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<mods:number id="3692F83B1A3ED0EF551CAF3F6D82261B">1</mods:number>
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<mods:identifier id="3811523E242129AF8ABA8425404899FF" type="DOI">10.11646/zootaxa.4162.1.7</mods:identifier>
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<treatment id="FF798790C2760F7BFF31FC45F54D1605" ID-DOI="http://doi.org/10.5281/zenodo.6070479" ID-GBIF-Taxon="124483505" ID-Zenodo-Dep="6070479" LSID="urn:lsid:plazi:treatment:FF798790C2760F7BFF31FC45F54D1605" httpUri="http://treatment.plazi.org/id/FF798790C2760F7BFF31FC45F54D1605" lastPageNumber="149" pageId="6" pageNumber="149">
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<subSubSection id="3FCA650DC2760F7BFF31FC45F04C12A6" pageId="6" pageNumber="149" type="nomenclature">
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<paragraph id="776F3686C2760F7BFF31FC45F22812C7" blockId="6.[151,841,908,967]" box="[151,841,908,934]" pageId="6" pageNumber="149">
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<heading id="2C2781EAC2760F7BFF31FC45F22812C7" bold="true" box="[151,841,908,934]" fontSize="11" level="1" pageId="6" pageNumber="149" reason="1">
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<taxonomicName id="B0D04D05C2760F7BFF31FC45F22812C7" authority="Ahrens & Stebnicka, 1997" authorityName="Ahrens & Stebnicka" authorityYear="1997" box="[151,841,908,934]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="149" phylum="Arthropoda" rank="species" species="himalorectus">
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<emphasis id="45A4EA94C2760F7BFF31FC45F22812C7" bold="true" box="[151,841,908,934]" pageId="6" pageNumber="149">
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<emphasis id="45A4EA94C2760F7BFF31FC45F08812C7" bold="true" box="[151,489,908,934]" italics="true" pageId="6" pageNumber="149">Phaeaphodius himalorectus</emphasis>
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(
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<bibRefCitation id="13414B77C2760F7BFE5EFC45F22112C7" author="Ahrens" box="[504,832,908,934]" pageId="6" pageNumber="149" refString="Ahrens, D. & Stebnicka, Z. (1997) On the Aphodiinae of the Nepal-Himalayas. Stuttgarter Beitrage zur Naturkunde, Serie A, 552, 1 - 17." type="journal article" year="1997">Ahrens & Stebnicka, 1997</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="776F3686C2760F7BFF31FC67F04C12A6" blockId="6.[151,841,908,967]" box="[151,301,942,967]" pageId="6" pageNumber="149">
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(
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<figureCitation id="EFEB2A03C2760F7BFF39FC67F04712A6" box="[159,294,942,967]" captionStart="FIGURES 13 – 23" captionStartId="7.[151,264,1885,1908]" captionTargetBox="[202,1385,206,1855]" captionTargetId="figure@7.[185,1402,193,1865]" captionTargetPageId="7" captionText="FIGURES 13 – 23. Phaeaphodius himalorectus (Ahrens & Stebnicka, 1997) (Nepal, Himalaya, Myagdi District, 2100 – 2300 m, paratype). 13, epipharynx; 14 – 15. habitus (scale bar and morphological details); 16 – 17, aedeagus (dorsal and lateral views). Phaeaphodius jouravliowi (Reitter, 1907) (Ural). 18, epipharynx; 19 – 21, habitus (morphological details, scale bar, and color variations); 22 – 23, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254694/files/figure.png" pageId="6" pageNumber="149">Figs. 13–17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="3FCA650DC2760F7BFF31FC3FF276154A" pageId="6" pageNumber="149" type="reference_group">
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<paragraph id="776F3686C2760F7BFF31FC3FF5A6156D" blockId="6.[151,1223,1014,1067]" box="[151,1223,1014,1036]" pageId="6" pageNumber="149">
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<treatmentCitationGroup id="57C011A8C2760F7BFF31FC3FF5A6156D" box="[151,1223,1014,1036]" pageId="6" pageNumber="149">
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<taxonomicName id="B0D04D05C2760F7BFF31FC3FF205156D" authority="Ahrens & Stebnicka, 1997: 12" authorityName="Ahrens & Stebnicka" authorityPageNumber="12" authorityYear="1997" box="[151,868,1014,1036]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="149" phylum="Arthropoda" rank="species" species="himalorectus" subGenus="Phaeaphodius">
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<emphasis id="45A4EA94C2760F7BFF31FC3FF349156D" box="[151,552,1014,1036]" italics="true" pageId="6" pageNumber="149">Aphodius (Phaeaphodius) himalorectus</emphasis>
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<treatmentCitation id="F6711097C2760F7BFD88FC3FF205156D" author="Ahrens" box="[558,868,1014,1036]" page="12" pageId="6" pageNumber="149" year="1997">
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<bibRefCitation id="13414B77C2760F7BFD88FC3FF25F156D" author="Ahrens" box="[558,830,1014,1036]" pageId="6" pageNumber="149" refString="Ahrens, D. & Stebnicka, Z. (1997) On the Aphodiinae of the Nepal-Himalayas. Stuttgarter Beitrage zur Naturkunde, Serie A, 552, 1 - 17." type="journal article" year="1997">Ahrens & Stebnicka, 1997</bibRefCitation>
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: 12
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="F6711097C2760F7BFCC8FC3FF5A2156D" author="Dellacasa" box="[878,1219,1014,1036]" page="133" pageId="6" pageNumber="149" year="2006">
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<bibRefCitation id="13414B77C2760F7BFCC8FC3FF5F1156D" author="Dellacasa" box="[878,1168,1014,1036]" pageId="6" pageNumber="149" refString="Dellacasa, M. & Dellacasa, G. (2006) Scarabaeidae: Aphodiinae: Aphodiini (Aphodiina). In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Apollo Books, Stenstrup, pp. 105 - 142." type="book" year="2006">Dellacasa & Dellacasa 2006</bibRefCitation>
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: 133
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="776F3686C2760F7BFF31FBDCF276154A" blockId="6.[151,1223,1014,1067]" box="[151,791,1045,1067]" pageId="6" pageNumber="149">
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<treatmentCitationGroup id="57C011A8C2760F7BFF31FBDCF276154A" box="[151,791,1045,1067]" pageId="6" pageNumber="149">
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<taxonomicName id="B0D04D05C2760F7BFF31FBDCF272154A" authority="Dellacasa & Dellacasa 2016: 144" authorityName="Dellacasa & Dellacasa" authorityPageNumber="144" authorityYear="2016" box="[151,787,1045,1067]" class="Insecta" family="Aphodiidae" genus="Phaeaphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="149" phylum="Arthropoda" rank="species" species="himalorectus">
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<emphasis id="45A4EA94C2760F7BFF31FBDCF0D0154A" box="[151,433,1045,1067]" italics="true" pageId="6" pageNumber="149">Phaeaphodius himalorectus</emphasis>
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:
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<treatmentCitation id="F6711097C2760F7BFE18FBDCF272154A" ID-DOI="http://doi.org/10.5281/zenodo.6070479" ID-Zenodo-Dep="6070479" author="Dellacasa" box="[446,787,1045,1067]" httpUri="http://treatment.plazi.org/id/FF798790C2760F7BFF31FC45F54D1605" page="144" pageId="6" pageNumber="149" year="2016">
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<bibRefCitation id="13414B77C2760F7BFE18FBDCF380154A" author="Dellacasa" box="[446,737,1045,1067]" pageId="6" pageNumber="149" refString="Dellacasa, M. & Dellacasa, G. (2016) Scarabaeidae: Aphodiinae: Aphodiini (Aphodiina). In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea and Byrrhoidea. Revised and updated edition. Brill, Leiden; Boston, pp. 98 - 155." type="journal article" year="2016">Dellacasa & Dellacasa 2016</bibRefCitation>
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: 144
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="3FCA650DC2760F7BFF31FB9EF5E615F4" pageId="6" pageNumber="149" type="materials_examined">
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<paragraph id="776F3686C2760F7BFF31FB9EF5561511" blockId="6.[151,1437,1111,1893]" box="[151,1079,1111,1136]" pageId="6" pageNumber="149">
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<materialsCitation id="C7B83CDBC2760F7BFF31FB9EF5561511" ID-GBIF-Occurrence="1306553621" box="[151,1079,1111,1136]" country="Nepal" elevation="2800" location="Nepal-Himalaya Annapurna Mountains" pageId="6" pageNumber="149" specimenCount="1" typeStatus="holotype">
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<emphasis id="45A4EA94C2760F7BFF31FB9EF04C1511" bold="true" box="[151,301,1111,1136]" pageId="6" pageNumber="149">
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<typeStatus id="A86B8824C2760F7BFF31FB9EF1B31511" box="[151,210,1111,1136]" pageId="6" pageNumber="149">Type</typeStatus>
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locality
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</emphasis>
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. Ghorepani,
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<quantity id="B0289B63C2760F7BFE66FB91F376150E" box="[448,535,1112,1136]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="149" unit="m" value="2800.0">
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<elevation id="FCFDD1B5C2760F7BFE66FB91F376150E" box="[448,535,1112,1136]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="149" unit="m" value="2800.0">2800 m</elevation>
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</quantity>
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,
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<location id="720F605DC2760F7BFD85FB9EF2801511" LSID="urn:lsid:plazi:treatment:FF798790C2760F7BFF31FC45F54D1605:720F605DC2760F7BFD85FB9EF2801511" box="[547,993,1111,1136]" country="Nepal" name="Nepal-Himalaya Annapurna Mountains" pageId="6" pageNumber="149">Nepal-Himalaya Annapurna Mountains</location>
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,
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<collectingCountry id="0FC77616C2760F7BFC4AFB9EF5521511" box="[1004,1075,1111,1136]" name="Nepal" pageId="6" pageNumber="149">Nepal</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="776F3686C2760F7BFF61FBB5F5E615F4" blockId="6.[151,1437,1111,1893]" box="[199,1159,1148,1173]" pageId="6" pageNumber="149">
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<emphasis id="45A4EA94C2760F7BFF61FBB5F0E015F4" bold="true" box="[199,385,1148,1173]" pageId="6" pageNumber="149">Type repository</emphasis>
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. Staatliches Museum für Naturkunde, Stuttgart (paratype examined).
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</paragraph>
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</subSubSection>
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<subSubSection id="3FCA650DC2760F7BFF61FB56F54617B4" pageId="6" pageNumber="149" type="description">
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<paragraph id="776F3686C2760F7BFF61FB56F54617B4" blockId="6.[151,1437,1111,1893]" pageId="6" pageNumber="149">
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<emphasis id="45A4EA94C2760F7BFF61FB56F00C15D9" bold="true" box="[199,365,1183,1208]" pageId="6" pageNumber="149">Redescription</emphasis>
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. Length 5.5–6.0 mm; oval elongate, moderately convex, shiny, almost glabrous. Head and pronotum piceous; elytra dark reddish brown, sometimes slightly paler at suture and apex; legs and antennal club reddish brown. Head with epistome coarsely, densely, somewhat irregularly punctate; punctation denser and confluent toward clypeal margin, anteriorly with arcuate transverse carina more-or-less distinctly raised; clypeus feebly sinuate at middle, round on sides, thinly bordered, edge feebly upturned, glabrous; genae obtusely angulate, densely setose, distinctly protruding from the eyes; frontal suture nearly obsolete, only with vestiges of tubercles; front coarsely, not densely, evenly punctate. Pronotum transverse, moderately convex, dually punctate; large punctures five to six times larger than small punctures, irregularly closer on sides; small punctures almost evenly scattered throughout, more superficial on disc; sides feebly arcuate, thinly bordered, edge sparsely setose; hind angles subtruncate; basal margin bisinuate, not bordered, laterally crenulate by few medium-size punctures. Scutellum triangular with curved sides, superficially minutely punctate on basal third. Elytra moderately convex, oblong oval, deeply striate; striae coarsely punctate, crenulate; interstriae flat on disc, very convex and with short straight pale setae on preapical declivity. Metatibiae superior spur slightly shorter than first metatarsomere; latter as long as following 3 metatarsomeres combined. Male: pronotum more transverse, less coarsely and less densely punctate; protibiae spur S-shaped, apically hooked; aedeagus
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<figureCitation id="EFEB2A03C2760F7BFCFBF95EF28617D1" box="[861,999,1687,1712]" captionStart="FIGURES 13 – 23" captionStartId="7.[151,264,1885,1908]" captionTargetBox="[202,1385,206,1855]" captionTargetId="figure@7.[185,1402,193,1865]" captionTargetPageId="7" captionText="FIGURES 13 – 23. Phaeaphodius himalorectus (Ahrens & Stebnicka, 1997) (Nepal, Himalaya, Myagdi District, 2100 – 2300 m, paratype). 13, epipharynx; 14 – 15. habitus (scale bar and morphological details); 16 – 17, aedeagus (dorsal and lateral views). Phaeaphodius jouravliowi (Reitter, 1907) (Ural). 18, epipharynx; 19 – 21, habitus (morphological details, scale bar, and color variations); 22 – 23, aedeagus (dorsal and lateral views)." httpUri="https://zenodo.org/record/254694/files/figure.png" pageId="6" pageNumber="149">Figs. 16–17</figureCitation>
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. Female: pronotum narrower apically, more coarsely and more densely punctate; protibiae spur thin and almost straight.
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</paragraph>
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</subSubSection>
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<subSubSection id="3FCA650DC2760F7BFF61F916F2231799" box="[199,834,1759,1784]" pageId="6" pageNumber="149" type="distribution">
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<paragraph id="776F3686C2760F7BFF61F916F2231799" blockId="6.[151,1437,1111,1893]" box="[199,834,1759,1784]" pageId="6" pageNumber="149">
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<emphasis id="45A4EA94C2760F7BFF61F916F0381799" bold="true" box="[199,345,1759,1784]" pageId="6" pageNumber="149">Distribution</emphasis>
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.
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<collectingCountry id="0FC77616C2760F7BFEC1F916F0CB1799" box="[359,426,1759,1784]" name="Nepal" pageId="6" pageNumber="149">Nepal</collectingCountry>
|
|||
|
|
(Himalaya: Annapurna Mountains).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="3FCA650DC2760F7BFF61F8CDF54D1605" pageId="6" pageNumber="149" type="materials_examined">
|
|||
|
|
<paragraph id="776F3686C2760F7BFF61F8CDF36B1621" blockId="6.[151,1437,1111,1893]" pageId="6" pageNumber="149">
|
|||
|
|
<materialsCitation id="C7B83CDBC2760F7BFF61F8CDF36B1621" ID-GBIF-Occurrence="1306553625" collectingDate="1994-03-20" collectionCode="DCG" collectorName="D. Ahrens" country="Nepal" county="Phalante" elevation="2200" location="Material" municipality="Myagdi District" pageId="6" pageNumber="149" specimenCount="11" typeStatus="paratype">
|
|||
|
|
<emphasis id="45A4EA94C2760F7BFF61F8CDF0C8167C" bold="true" box="[199,425,1796,1821]" pageId="6" pageNumber="149">
|
|||
|
|
<location id="720F605DC2760F7BFF61F8CDF04E167C" LSID="urn:lsid:plazi:treatment:FF798790C2760F7BFF31FC45F54D1605:720F605DC2760F7BFF61F8CDF04E167C" box="[199,303,1796,1821]" country="Nepal" county="Phalante" municipality="Myagdi District" name="Material" pageId="6" pageNumber="149">Material</location>
|
|||
|
|
examined
|
|||
|
|
</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="45A4EA94C2760F7BFE1EF8CDF371167C" bold="true" box="[440,528,1796,1821]" pageId="6" pageNumber="149">
|
|||
|
|
<collectingCountry id="0FC77616C2760F7BFE1EF8CDF369167A" box="[440,520,1796,1821]" name="Nepal" pageId="6" pageNumber="149">NEPAL</collectingCountry>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
<location id="720F605DC2760F7BFDBEF8CDF3E9167C" LSID="urn:lsid:plazi:treatment:FF798790C2760F7BFF31FC45F54D1605:720F605DC2760F7BFDBEF8CDF3E9167C" box="[536,648,1796,1821]" country="Nepal" county="Phalante" municipality="Myagdi District" name="Himalaya" pageId="6" pageNumber="149">Himalaya</location>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="970BACFCC2760F7BFD35F8CDF228167C" box="[659,841,1796,1821]" pageId="6" pageNumber="149">Myagdi District</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<collectingCounty id="9E0E4E0AC2760F7BFCF2F8CDF2D6167C" box="[852,951,1796,1821]" pageId="6" pageNumber="149">Phalante</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<quantity id="B0289B63C2760F7BFC65F8CDF501167D" box="[963,1120,1796,1820]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="6" pageNumber="149" unit="m" value="2200.0" valueMax="2300.0" valueMin="2100.0">
|
|||
|
|
<elevation id="FCFDD1B5C2760F7BFC65F8CDF501167D" box="[963,1120,1796,1820]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="6" pageNumber="149" unit="m" value="2200.0" valueMax="2300.0" valueMin="2100.0">2100–2300 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="036E1046C2760F7BFBCAF8CDF58D167D" box="[1132,1260,1796,1820]" pageId="6" pageNumber="149" value="1994-03-20">
|
|||
|
|
<collectingDate id="132AE9AEC2760F7BFBCAF8CDF58D167D" box="[1132,1260,1796,1820]" pageId="6" pageNumber="149" value="1994-03-20">20.III.1994</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="DA255350C2760F7BFB5EF8CCF40D167C" box="[1272,1388,1796,1821]" pageId="6" pageNumber="149">D. Ahrens</collectorName>
|
|||
|
|
leg. (
|
|||
|
|
<specimenCount id="61D6FD0FC2760F7BFF06F8E1F05B1621" box="[160,314,1831,1856]" pageId="6" pageNumber="149" type="generic">11 specimens</specimenCount>
|
|||
|
|
,
|
|||
|
|
<typeStatus id="A86B8824C2760F7BFEE3F8E1F0D21621" box="[325,435,1832,1856]" pageId="6" pageNumber="149" type="paratype">paratypes</typeStatus>
|
|||
|
|
,
|
|||
|
|
<collectionCode id="11C1AE43C2760F7BFE18F8E1F09F1621" box="[446,510,1832,1856]" pageId="6" pageNumber="149">DCG</collectionCode>
|
|||
|
|
).
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="776F3686C2760F7BFF61F885F54D1605" blockId="6.[151,1437,1111,1893]" box="[199,1068,1868,1893]" pageId="6" pageNumber="149">
|
|||
|
|
<emphasis id="45A4EA94C2760F7BFF61F885F0221604" bold="true" box="[199,323,1868,1893]" pageId="6" pageNumber="149">Bionomics</emphasis>
|
|||
|
|
. All specimens examined were collected in spring, above
|
|||
|
|
<quantity id="B0289B63C2760F7BFC76F885F5491605" box="[976,1064,1868,1892]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="149" unit="m" value="2000.0">2000 m</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|