treatments-xml/data/72/32/87/72328796E90AFF9FFF78932D7D910339.xml

<document id="755634280E4799D7003B01F924887232" ID-DOI="10.5281/zenodo.199703" ID-GBIF-Dataset="a6568cca-bec7-4f60-8832-ebd6e001b4b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="199703" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460344064441" checkinUser="plazi" docAuthor="Benine, Ricardo C., Lopes, Guilherme A. M. &amp; Ron, Ernesto" docDate="2010" docId="72328796E90AFF9FFF78932D7D910339" docLanguage="en" docName="zt02715p067.pdf" docOrigin="Zootaxa 2715" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ctenobrycon oliverai Benine, Lopes &amp; Ron, 2010, new species" docType="treatment" docVersion="8" lastPageNumber="66" masterDocId="8E0BFFEEE90BFF98FFEF96217C040761" masterDocTitle="A new species of Ctenobrycon Eigenmann, 1908 (Characiformes: Characidae) from the río Orinoco basin, Venezuela" masterLastPageNumber="67" masterPageNumber="59" pageNumber="60" updateTime="1698261789361" updateUser="plazi">
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<mods:title id="1C06E88C6C7729D6C83C70E4CF5CBEA1">A new species of Ctenobrycon Eigenmann, 1908 (Characiformes: Characidae) from the río Orinoco basin, Venezuela</mods:title>
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<mods:namePart id="991FB600779A7461AF868A5D4F17B638">Benine, Ricardo C.</mods:namePart>
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<mods:namePart id="629E6CD3C2115F2BD0B7744D8723642B">Lopes, Guilherme A. M.</mods:namePart>
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<mods:namePart id="C9CEFDB05A223866EEAF90D14957B599">Ron, Ernesto</mods:namePart>
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<mods:date id="00337F1B1F8607891A34096066EED796">2010</mods:date>
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<treatment id="72328796E90AFF9FFF78932D7D910339" ID-DOI="http://doi.org/10.5281/zenodo.5668677" ID-GBIF-Taxon="119406948" ID-Zenodo-Dep="5668677" LSID="urn:lsid:plazi:treatment:72328796E90AFF9FFF78932D7D910339" httpUri="http://treatment.plazi.org/id/72328796E90AFF9FFF78932D7D910339" lastPageId="7" lastPageNumber="66" pageId="1" pageNumber="60">
<subSubSection id="B281650BE90AFF99FF78932D7D640227" pageId="1" pageNumber="60" type="nomenclature">
<paragraph id="FA243680E90AFF99FF78932D7E360244" blockId="1.[151,562,1291,1350]" box="[151,562,1291,1318]" pageId="1" pageNumber="60">
<heading id="A16C81ECE90AFF99FF78932D7E360244" bold="true" box="[151,562,1291,1318]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
<emphasis id="C8EFEA92E90AFF99FF78932D7E360244" bold="true" box="[151,562,1291,1318]" pageId="1" pageNumber="60">
<taxonomicName id="3D9B4D03E90AFF99FF78932D7D900247" ID-CoL="3256L" box="[151,404,1292,1318]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="oliverai" status="sp. nov.">
<emphasis id="C8EFEA92E90AFF99FF78932D7D900247" bold="true" box="[151,404,1292,1318]" italics="true" pageId="1" pageNumber="60">Ctenobrycon oliverai</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D3DC57E9E90AFF99FE4C932A7E360244" box="[419,562,1291,1317]" pageId="1" pageNumber="60" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FA243680E90AFF99FF78930F7D640227" blockId="1.[151,562,1291,1350]" box="[151,352,1326,1350]" pageId="1" pageNumber="60">
<tableCitation id="B719033BE90AFF99FF78930F7CED0227" box="[151,233,1326,1350]" captionStart="TABLE 1" captionStartId="2.[151,239,152,176]" captionText="TABLE 1. Morphometric data for Ctenobrycon oliverai, new species." pageId="1" pageNumber="60">Table 1</tableCitation>
, 
<figureCitation id="62A02A05E90AFF99FF17930F7D640227" box="[248,352,1326,1350]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,255,976,1000]" captionStartId-1="4.[151,255,1426,1450]" captionTargetBox-0="[151,1436,323,952]" captionTargetBox-1="[312,1275,381,1398]" captionTargetId-0="figure@3.[151,1436,314,952]" captionTargetId-1="figure@4.[307,1280,375,1405]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Ctenobrycon oliverai, new species, holotype, MZUSP 50130, 55.4 mm SL, río Apure, West of Ciudad de Apure, State of Apure, Venezuela." captionText-1="FIGURE 2. Ctenobrycon oliverai, paratype, LBP 3061, 50.7 mm SL, premaxillary, maxillary and dentary; left side, lateral view (maxillary and dentary) or frontal view (premaxillary). Scale bar = 1 mm." httpUri-0="https://zenodo.org/record/199704/files/figure.png" httpUri-1="https://zenodo.org/record/199705/files/figure.png" pageId="1" pageNumber="60">Figs. 1–2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B281650BE90AFF99FF7893567F09017F" pageId="1" pageNumber="60" type="materials_examined">
<paragraph id="FA243680E90AFF99FF7893567DF102D2" blockId="1.[151,1437,1398,2034]" pageId="1" pageNumber="60">
<emphasis id="C8EFEA92E90AFF99FF7893567D0F02EE" bold="true" box="[151,267,1399,1423]" pageId="1" pageNumber="60">
<typeStatus id="25208822E90AFF99FF7893567D0202EE" box="[151,262,1399,1423]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="9C8AAE45E90AFF99FEF993577D7702EF" box="[278,371,1398,1422]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="60">MZUSP</collectionCode>
50130, 
<quantity id="3D639B65E90AFF99FE3993577E3A02EF" box="[470,574,1398,1422]" metricMagnitude="-2" metricUnit="m" metricValue="5.4399999999999995" pageId="1" pageNumber="60" unit="mm" value="54.4">54.4 mm</quantity>
SL, río Apure, West of Ciudad de Apure, State of Apure, 
<collectingCountry id="828C7610E90AFF99FB149357797602EF" box="[1275,1394,1398,1422]" name="Venezuela" pageId="1" pageNumber="60">Venezuela</collectingCountry>
. 
<date id="8E251040E90AFF99FA6F93577D3402D3" pageId="1" pageNumber="60" value="1982-01-28">28 January 1982</date>
. 
<collectingCountry id="828C7610E90AFF99FED493BB7D5602D3" box="[315,338,1434,1458]" name="Iceland" pageId="1" pageNumber="60">O</collectingCountry>
. Costillo 
<emphasis id="C8EFEA92E90AFF99FE5293BD7DF102D2" box="[445,501,1435,1459]" italics="true" pageId="1" pageNumber="60">et al.</emphasis>
</paragraph>
<paragraph id="FA243680E90AFF99FF28939E7F09017F" blockId="1.[151,1437,1398,2034]" pageId="1" pageNumber="60">
<emphasis id="C8EFEA92E90AFF99FF28939E7D4202B6" bold="true" box="[199,326,1471,1495]" pageId="1" pageNumber="60">
<typeStatus id="25208822E90AFF99FF28939E7D4502B6" box="[199,321,1471,1495]" pageId="1" pageNumber="60" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="9C8AAE45E90AFF99FEA2939F7DAE02B7" box="[333,426,1470,1494]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="60">MZUSP</collectionCode>
27979, 15, 
<quantity id="3D639B65E90AFF99FDDF939F7EE602B7" box="[560,738,1470,1494]" metricMagnitude="-2" metricUnit="m" metricValue="4.725" metricValueMax="5.58" metricValueMin="3.87" pageId="1" pageNumber="60" unit="mm" value="47.25" valueMax="55.8" valueMin="38.7">38.7 – 55.8 mm</quantity>
SL, collected with the 
<typeStatus id="25208822E90AFF99FC08939F784902B7" box="[999,1101,1470,1494]" pageId="1" pageNumber="60" type="holotype">holotype</typeStatus>
. LBP 3061, 11, 43.8 – 54.6, 1 C&amp;S, 
<quantity id="3D639B65E90AFF99FF3093C37D41029B" box="[223,325,1506,1530]" metricMagnitude="-2" metricUnit="m" metricValue="5.029999999999999" pageId="1" pageNumber="60" unit="mm" value="50.3">50.3 mm</quantity>
SL, Rio Orinoco, Ciudad de Caicara del Orinoco, 
<geoCoordinate id="9FAF5047E90AFF99FC7F93C3783D029B" box="[912,1081,1506,1530]" direction="north" orientation="latitude" pageId="1" pageNumber="60" precision="1" value="7.636555">07°38’11.6” N</geoCoordinate>
, 66°19’04.2”. State of Bolívar, 
<collectingCountry id="828C7610E90AFF99FF7890277D0A017F" box="[151,270,1542,1566]" name="Venezuela" pageId="1" pageNumber="60">Venezuela</collectingCountry>
. 0 
<date id="8E251040E90AFF99FEC890277DD3017F" box="[295,471,1542,1566]" pageId="1" pageNumber="60" value="2005-10-03">3 October 2005</date>
. A. Granado &amp; C. Oliveira.
</paragraph>
</subSubSection>
<subSubSection id="B281650BE90AFF99FF28900B7E1C0197" pageId="1" pageNumber="60" type="diagnosis">
<paragraph id="FA243680E90AFF99FF28900B7E1C0197" blockId="1.[151,1437,1398,2034]" pageId="1" pageNumber="60">
<emphasis id="C8EFEA92E90AFF99FF28900B7D450122" bold="true" box="[199,321,1578,1603]" pageId="1" pageNumber="60">Diagnosis.</emphasis>
<taxonomicName id="3D9B4D03E90AFF99FEA2900A7E440122" box="[333,576,1579,1603]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90AFF99FEA2900A7E440122" box="[333,576,1579,1603]" italics="true" pageId="1" pageNumber="60">Ctenobrycon oliverai</emphasis>
</taxonomicName>
is distinguished from all congeners and 
<taxonomicName id="3D9B4D03E90AFF99FBC7900A79420122" box="[1064,1350,1579,1603]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90AFF99FBC7900A79420122" box="[1064,1350,1579,1603]" italics="true" pageId="1" pageNumber="60">Psellogrammus kennedyi</emphasis>
</taxonomicName>
by the number of scale rows between dorsal-fin origin and lateral line (14 – 15 
<emphasis id="C8EFEA92E90AFF99FC3A90717FF00106" box="[981,1012,1616,1639]" italics="true" pageId="1" pageNumber="60">vs.</emphasis>
<quantity id="3D639B65E90AFF99FC13906F78740107" box="[1020,1136,1614,1638]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.302" metricValueMin="2.794" pageId="1" pageNumber="60" unit="in" value="12.0" valueMax="13.0" valueMin="11.0">11 – 13 in</quantity>
<taxonomicName id="3D9B4D03E90AFF99FB96906E78F10107" box="[1145,1269,1614,1638]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90AFF99FB96906E78F10107" box="[1145,1269,1614,1638]" italics="true" pageId="1" pageNumber="60">C. spilurus</emphasis>
</taxonomicName>
; 
<quantity id="3D639B65E90AFF99FAEA906F797E0107" box="[1285,1402,1614,1638]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="1" pageNumber="60" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">11 – 12 in</quantity>
<taxonomicName id="3D9B4D03E90AFF99FA6D906E7CD301EA" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E90AFF99FA6D906E7CD301EA" italics="true" pageId="1" pageNumber="60">C. alleni</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90AFF99FEFC90557D9101EA" box="[275,405,1651,1675]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90AFF99FEFC90557D9101EA" box="[275,405,1651,1675]" italics="true" pageId="1" pageNumber="60">P. kennedyi</emphasis>
</taxonomicName>
). 
<taxonomicName id="3D9B4D03E90AFF99FE5F90527EA601EA" box="[432,674,1651,1675]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90AFF99FE5F90527EA601EA" box="[432,674,1651,1675]" italics="true" pageId="1" pageNumber="60">Ctenobrycon oliverai</emphasis>
</taxonomicName>
is further distinguished from 
<taxonomicName id="3D9B4D03E90AFF99FC109052786601EA" box="[1023,1122,1651,1675]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E90AFF99FC109052786601EA" box="[1023,1122,1651,1675]" italics="true" pageId="1" pageNumber="60">C. alleni</emphasis>
</taxonomicName>
by the number of humeral blotches (one 
<emphasis id="C8EFEA92E90AFF99FED690B97D5401CE" box="[313,336,1688,1711]" italics="true" pageId="1" pageNumber="60">vs</emphasis>
. two, respectively). The following characters may be usefull in distinguishing 
<taxonomicName id="3D9B4D03E90AFF99FB0E90B6795F01CF" box="[1249,1371,1686,1710]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90AFF99FB0E90B6795F01CF" box="[1249,1371,1686,1710]" italics="true" pageId="1" pageNumber="60">C. oliverai</emphasis>
</taxonomicName>
from congeners: greatest body depth (48.0–58.3% of SL 
<emphasis id="C8EFEA92E90AFF99FD07909D7EFB01B2" box="[744,767,1724,1747]" italics="true" pageId="1" pageNumber="60">vs</emphasis>
. 41.3–53.0 in 
<taxonomicName id="3D9B4D03E90AFF99FC47909A782101B2" box="[936,1061,1723,1747]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90AFF99FC47909A782101B2" box="[936,1061,1723,1747]" italics="true" pageId="1" pageNumber="60">C. spilurus</emphasis>
</taxonomicName>
; 41.1–51.0% of SL in 
<taxonomicName id="3D9B4D03E90AFF99FADE909A799001B2" box="[1329,1428,1723,1747]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E90AFF99FADE909A799001B2" box="[1329,1428,1723,1747]" italics="true" pageId="1" pageNumber="60">C. alleni</emphasis>
</taxonomicName>
; 42.4–50.5% of SL in 
<taxonomicName id="3D9B4D03E90AFF99FE6790C17E030197" box="[392,519,1758,1783]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90AFF99FE6790C17E030197" box="[392,519,1758,1783]" italics="true" pageId="1" pageNumber="60">P. kennedyi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="B281650BE90AFF9BFF2891237DC001B5" lastPageId="3" lastPageNumber="62" pageId="1" pageNumber="60" type="description">
<paragraph id="FA243680E90AFF99FF28912379330093" blockId="1.[151,1437,1398,2034]" pageId="1" pageNumber="60">
<emphasis id="C8EFEA92E90AFF99FF2891237D5D007A" bold="true" box="[199,345,1794,1819]" pageId="1" pageNumber="60">Description.</emphasis>
Morphometric data for 
<taxonomicName id="3D9B4D03E90AFF99FD8591227F5E007A" box="[618,858,1795,1819]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90AFF99FD8591227F5E007A" box="[618,858,1795,1819]" italics="true" pageId="1" pageNumber="60">Ctenobrycon oliverai</emphasis>
</taxonomicName>
are summarized in 
<tableCitation id="B719033BE90AFF99FBD291237896007B" box="[1085,1170,1794,1818]" captionStart="TABLE 1" captionStartId="2.[151,239,152,176]" captionText="TABLE 1. Morphometric data for Ctenobrycon oliverai, new species." pageId="1" pageNumber="60">Table 1</tableCitation>
. Deep bodied. Greatest body depth at dorsal-fin origin. Dorsal profile of head concave. Dorsal profile of body strongly convex from tip of supraoccipital spine to dorsal-fin origin, dorsal-fin base posteroventrally slanted, straight or slightly convex from posterior terminus of dorsal-fin base to end of adipose fin, and concave along caudal peduncle. Ventral body profile convex from tip of lower jaw to anal-fin origin, anal-fin base posterodorsally slanted, concave along caudal peduncle. Prepelvic region transversally flattened, more so proximal to pelvic-fin insertion. Postpelvic region transversally flattened proximal to pelvic-fin insertion becoming somewhat obtuse toward anal-fin origin.
</paragraph>
<caption id="AEE46608E909FF9AFF7896B97F8607CE" box="[151,898,152,176]" pageId="2" pageNumber="61">
<paragraph id="FA243680E909FF9AFF7896B97F8607CE" blockId="2.[151,1233,152,312]" box="[151,898,152,176]" pageId="2" pageNumber="61">
<emphasis id="C8EFEA92E909FF9AFF7896B97D0C07D1" bold="true" box="[151,264,152,176]" pageId="2" pageNumber="61">TABLE 1.</emphasis>
Morphometric data for 
<emphasis id="C8EFEA92E909FF9AFDE596B87EF007D1" box="[522,756,153,176]" italics="true" pageId="2" pageNumber="61">
<taxonomicName id="3D9B4D03E909FF9AFDE596B87EF407D1" box="[522,752,153,176]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="oliverai" status="sp. nov.">Ctenobrycon oliverai</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="D3DC57E9E909FF9AFD1496B97F7A07CE" box="[763,894,152,175]" pageId="2" pageNumber="61" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="FA243680E909FF9AFD9696ED78080782" blockId="2.[151,1233,152,312]" box="[633,1036,204,227]" pageId="2" pageNumber="61">
<typeStatus id="25208822E909FF9AFD9696ED7ED20782" box="[633,726,204,227]" pageId="2" pageNumber="61" type="holotype">holotype</typeStatus>
N 
<typeStatus id="25208822E909FF9AFC4996ED78080782" box="[934,1036,204,227]" pageId="2" pageNumber="61" type="paratype">paratypes</typeStatus>
</paragraph>
<paragraph id="FA243680E909FF9AFC4996D77DB703D9" blockId="2.[151,1233,152,312]" lastBlockId="2.[159,1222,1185,1379]" pageId="2" pageNumber="61">Limits mean Standard length (mm) 55.4 28 39.4–59.9 50.1 Percentage of head length</paragraph>
<paragraph id="FA243680E909FF9AFF70975778C20302" pageId="2" pageNumber="61">
<table id="889BC420E9090067FF70975778C20302" box="[159,1222,374,1123]" gridcols="4" gridrows="18" pageId="2" pageNumber="61">
<tr id="44AB34C2E9090067FF70975778C206EC" box="[159,1222,374,397]" gridrow="0" pageId="2" pageNumber="61" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="077A5DBEE9090067FF7097577EAC06EC" box="[159,680,374,397]" gridcol="0" gridrow="0" pageId="2" pageNumber="61">Percentage of standard length</th>
</tr>
<tr id="44AB34C2E9090067FF70978078C206D9" box="[159,1222,417,440]" gridrow="1" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7097807EAC06D9" box="[159,680,417,440]" gridcol="0" gridrow="1" pageId="2" pageNumber="61">Greatest depth 52.2</th>
<td id="077A5DBEE9090067FCE097807F2E06D9" box="[783,810,417,440]" gridcol="1" gridrow="1" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC499780781606D9" box="[934,1042,417,440]" gridcol="2" gridrow="1" pageId="2" pageNumber="61">48.0–58.3</td>
<td id="077A5DBEE9090067FB78978078C206D9" box="[1175,1222,417,440]" gridcol="3" gridrow="1" pageId="2" pageNumber="61">53.0</td>
</tr>
<tr id="44AB34C2E9090067FF7097ED78C20682" box="[159,1222,460,483]" gridrow="2" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7097ED7EAC0682" box="[159,680,460,483]" gridcol="0" gridrow="2" pageId="2" pageNumber="61">Snout to dorsal-fin origin 54.0</th>
<td id="077A5DBEE9090067FCE097ED7F2E0682" box="[783,810,460,483]" gridcol="1" gridrow="2" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC4997ED78160682" box="[934,1042,460,483]" gridcol="2" gridrow="2" pageId="2" pageNumber="61">52.2–57.1</td>
<td id="077A5DBEE9090067FB7897ED78C20682" box="[1175,1222,460,483]" gridcol="3" gridrow="2" pageId="2" pageNumber="61">54.1</td>
</tr>
<tr id="44AB34C2E9090067FF7097D778C2056C" box="[159,1222,502,525]" gridrow="3" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7097D77EAC056C" box="[159,680,502,525]" gridcol="0" gridrow="3" pageId="2" pageNumber="61">Snout to pectoral-fin origin 26.3</th>
<td id="077A5DBEE9090067FCE097D77F2E056C" box="[783,810,502,525]" gridcol="1" gridrow="3" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC4997D77816056C" box="[934,1042,502,525]" gridcol="2" gridrow="3" pageId="2" pageNumber="61">25.3–27.9</td>
<td id="077A5DBEE9090067FB7897D778C2056C" box="[1175,1222,502,525]" gridcol="3" gridrow="3" pageId="2" pageNumber="61">26.6</td>
</tr>
<tr id="44AB34C2E9090067FF70940078C20559" box="[159,1222,545,568]" gridrow="4" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7094007EAC0559" box="[159,680,545,568]" gridcol="0" gridrow="4" pageId="2" pageNumber="61">Snout to pelvic-fin origin 42.1</th>
<td id="077A5DBEE9090067FCE094007F2E0559" box="[783,810,545,568]" gridcol="1" gridrow="4" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC49940078160559" box="[934,1042,545,568]" gridcol="2" gridrow="4" pageId="2" pageNumber="61">40.9–47.0</td>
<td id="077A5DBEE9090067FB78940078C20559" box="[1175,1222,545,568]" gridcol="3" gridrow="4" pageId="2" pageNumber="61">43.4</td>
</tr>
<tr id="44AB34C2E9090067FF70946D78C20502" box="[159,1222,588,611]" gridrow="5" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF70946D7EAC0502" box="[159,680,588,611]" gridcol="0" gridrow="5" pageId="2" pageNumber="61">Snout to anal-fin origin 59.3</th>
<td id="077A5DBEE9090067FCE0946D7F2E0502" box="[783,810,588,611]" gridcol="1" gridrow="5" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC49946D78160502" box="[934,1042,588,611]" gridcol="2" gridrow="5" pageId="2" pageNumber="61">55.4–64.1</td>
<td id="077A5DBEE9090067FB78946D78C20502" box="[1175,1222,588,611]" gridcol="3" gridrow="5" pageId="2" pageNumber="61">58.7</td>
</tr>
<tr id="44AB34C2E9090067FF70945778C205EC" box="[159,1222,630,653]" gridrow="6" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7094577EAC05EC" box="[159,680,630,653]" gridcol="0" gridrow="6" pageId="2" pageNumber="61">Caudal peduncle depth 10.0</th>
<td id="077A5DBEE9090067FCE094577F2E05EC" box="[783,810,630,653]" gridcol="1" gridrow="6" pageId="2" pageNumber="61">24</td>
<td id="077A5DBEE9090067FC499457781605EC" box="[934,1042,630,653]" gridcol="2" gridrow="6" pageId="2" pageNumber="61">9.2–11.7</td>
<td id="077A5DBEE9090067FB78945778C205EC" box="[1175,1222,630,653]" gridcol="3" gridrow="6" pageId="2" pageNumber="61">10.1</td>
</tr>
<tr id="44AB34C2E9090067FF70948078C205D9" box="[159,1222,673,696]" gridrow="7" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7094807EAC05D9" box="[159,680,673,696]" gridcol="0" gridrow="7" pageId="2" pageNumber="61">Caudal peduncle length 2.3</th>
<td id="077A5DBEE9090067FCE094807F2E05D9" box="[783,810,673,696]" gridcol="1" gridrow="7" pageId="2" pageNumber="61">23</td>
<td id="077A5DBEE9090067FC499480781605D9" box="[934,1042,673,696]" gridcol="2" gridrow="7" pageId="2" pageNumber="61">1.6–3.0</td>
<td id="077A5DBEE9090067FB78948078C205D9" box="[1175,1222,673,696]" gridcol="3" gridrow="7" pageId="2" pageNumber="61">2.2</td>
</tr>
<tr id="44AB34C2E9090067FF7094ED78C20582" box="[159,1222,716,739]" gridrow="8" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7094ED7EAC0582" box="[159,680,716,739]" gridcol="0" gridrow="8" pageId="2" pageNumber="61">Pectoral-fin length 23.1</th>
<td id="077A5DBEE9090067FCE094ED7F2E0582" box="[783,810,716,739]" gridcol="1" gridrow="8" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC4994ED78160582" box="[934,1042,716,739]" gridcol="2" gridrow="8" pageId="2" pageNumber="61">22.2–24.6</td>
<td id="077A5DBEE9090067FB7894ED78C20582" box="[1175,1222,716,739]" gridcol="3" gridrow="8" pageId="2" pageNumber="61">23.4</td>
</tr>
<tr id="44AB34C2E9090067FF7094D778C2046C" box="[159,1222,758,781]" gridrow="9" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7094D77EAC046C" box="[159,680,758,781]" gridcol="0" gridrow="9" pageId="2" pageNumber="61">Pelvic-fin length 17.1</th>
<td id="077A5DBEE9090067FCE094D77F2E046C" box="[783,810,758,781]" gridcol="1" gridrow="9" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC4994D77816046C" box="[934,1042,758,781]" gridcol="2" gridrow="9" pageId="2" pageNumber="61">13.4–18.5</td>
<td id="077A5DBEE9090067FB7894D778C2046C" box="[1175,1222,758,781]" gridcol="3" gridrow="9" pageId="2" pageNumber="61">16.9</td>
</tr>
<tr id="44AB34C2E9090067FF70950078C20459" box="[159,1222,801,824]" gridrow="10" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7095007EAC0459" box="[159,680,801,824]" gridcol="0" gridrow="10" pageId="2" pageNumber="61">Dorsal-fin length 35.1</th>
<td id="077A5DBEE9090067FCE095007F2E0459" box="[783,810,801,824]" gridcol="1" gridrow="10" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC49950078160459" box="[934,1042,801,824]" gridcol="2" gridrow="10" pageId="2" pageNumber="61">31.9–36.5</td>
<td id="077A5DBEE9090067FB78950078C20459" box="[1175,1222,801,824]" gridcol="3" gridrow="10" pageId="2" pageNumber="61">34.2</td>
</tr>
<tr id="44AB34C2E9090067FF70956D78C20402" box="[159,1222,844,867]" gridrow="11" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF70956D7EAC0402" box="[159,680,844,867]" gridcol="0" gridrow="11" pageId="2" pageNumber="61">Dorsal-fin base 14.2</th>
<td id="077A5DBEE9090067FCE0956D7F2E0402" box="[783,810,844,867]" gridcol="1" gridrow="11" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC49956D78160402" box="[934,1042,844,867]" gridcol="2" gridrow="11" pageId="2" pageNumber="61">12.5–14.7</td>
<td id="077A5DBEE9090067FB78956D78C20402" box="[1175,1222,844,867]" gridcol="3" gridrow="11" pageId="2" pageNumber="61">13.3</td>
</tr>
<tr id="44AB34C2E9090067FF70955778C204EC" box="[159,1222,886,909]" gridrow="12" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7095577EAC04EC" box="[159,680,886,909]" gridcol="0" gridrow="12" pageId="2" pageNumber="61">Anal-fin length 13.0</th>
<td id="077A5DBEE9090067FCE095577F2E04EC" box="[783,810,886,909]" gridcol="1" gridrow="12" pageId="2" pageNumber="61">24</td>
<td id="077A5DBEE9090067FC499557781604EC" box="[934,1042,886,909]" gridcol="2" gridrow="12" pageId="2" pageNumber="61">10.9–16.1</td>
<td id="077A5DBEE9090067FB78955778C204EC" box="[1175,1222,886,909]" gridcol="3" gridrow="12" pageId="2" pageNumber="61">13.4</td>
</tr>
<tr id="44AB34C2E9090067FF70958078C204D9" box="[159,1222,929,952]" gridrow="13" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7095807EAC04D9" box="[159,680,929,952]" gridcol="0" gridrow="13" pageId="2" pageNumber="61">Anal-fin base 49.6</th>
<td id="077A5DBEE9090067FCE095807F2E04D9" box="[783,810,929,952]" gridcol="1" gridrow="13" pageId="2" pageNumber="61">27</td>
<td id="077A5DBEE9090067FC499580781604D9" box="[934,1042,929,952]" gridcol="2" gridrow="13" pageId="2" pageNumber="61">45.9–52.0</td>
<td id="077A5DBEE9090067FB78958078C204D9" box="[1175,1222,929,952]" gridcol="3" gridrow="13" pageId="2" pageNumber="61">48.8</td>
</tr>
<tr id="44AB34C2E9090067FF7095ED78C20482" box="[159,1222,972,995]" gridrow="14" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7095ED7EAC0482" box="[159,680,972,995]" gridcol="0" gridrow="14" pageId="2" pageNumber="61">Eye to dorsal-fin origin 43.9</th>
<td id="077A5DBEE9090067FCE095ED7F2E0482" box="[783,810,972,995]" gridcol="1" gridrow="14" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC4995ED78160482" box="[934,1042,972,995]" gridcol="2" gridrow="14" pageId="2" pageNumber="61">40.9–45.8</td>
<td id="077A5DBEE9090067FB7895ED78C20482" box="[1175,1222,972,995]" gridcol="3" gridrow="14" pageId="2" pageNumber="61">42.9</td>
</tr>
<tr id="44AB34C2E9090067FF7095D778C2036C" box="[159,1222,1014,1037]" gridrow="15" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7095D77EAC036C" box="[159,680,1014,1037]" gridcol="0" gridrow="15" pageId="2" pageNumber="61">Dorsal-fin origin to caudal–fin origin 53.3</th>
<td id="077A5DBEE9090067FCE095D77F2E036C" box="[783,810,1014,1037]" gridcol="1" gridrow="15" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC4995D77816036C" box="[934,1042,1014,1037]" gridcol="2" gridrow="15" pageId="2" pageNumber="61">52.8–57.5</td>
<td id="077A5DBEE9090067FB7895D778C2036C" box="[1175,1222,1014,1037]" gridcol="3" gridrow="15" pageId="2" pageNumber="61">54.6</td>
</tr>
<tr id="44AB34C2E9090067FF70920078C20359" box="[159,1222,1057,1080]" gridrow="16" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF7092007EAC0359" box="[159,680,1057,1080]" gridcol="0" gridrow="16" pageId="2" pageNumber="61">Head length 24.2</th>
<td id="077A5DBEE9090067FCE092007F2E0359" box="[783,810,1057,1080]" gridcol="1" gridrow="16" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC49920078160359" box="[934,1042,1057,1080]" gridcol="2" gridrow="16" pageId="2" pageNumber="61">23.2–25.7</td>
<td id="077A5DBEE9090067FB78920078C20359" box="[1175,1222,1057,1080]" gridcol="3" gridrow="16" pageId="2" pageNumber="61">24.7</td>
</tr>
<tr id="44AB34C2E9090067FF70926D78C20302" box="[159,1222,1100,1123]" gridrow="17" pageId="2" pageNumber="61">
<th id="077A5DBEE9090067FF70926D7EAC0302" box="[159,680,1100,1123]" gridcol="0" gridrow="17" pageId="2" pageNumber="61">Head depth 19.9</th>
<td id="077A5DBEE9090067FCE0926D7F2E0302" box="[783,810,1100,1123]" gridcol="1" gridrow="17" pageId="2" pageNumber="61">28</td>
<td id="077A5DBEE9090067FC49926D78160302" box="[934,1042,1100,1123]" gridcol="2" gridrow="17" pageId="2" pageNumber="61">19.0–22.0</td>
<td id="077A5DBEE9090067FB78926D78C20302" box="[1175,1222,1100,1123]" gridcol="3" gridrow="17" pageId="2" pageNumber="61">20.1</td>
</tr>
</table>
</paragraph>
<paragraph id="FA243680E909FF9AFF4A92ED7EAC0117" blockId="2.[159,1222,1185,1379]" lastBlockId="2.[151,1436,1486,2015]" pageId="2" pageNumber="61">
Snout length 25.2 28 21.5–26.9 24.5 Upper jaw length 35.6 28 34.6–39.8 36.3 Horizontal orbital diameter 38.6 28 37.7–43.1 40.6 Least interorbital width 41.6 28 37.4–42.8 40.2 Mouth terminal. Maxillary not surpassing vertical through anterior margin of orbit. Premaxillary teeth in to rows; outer row with 3 (2), 4* (25), 5 (1) tricuspid teeth, midcentral cusps longer than others; inner tooth row with 5* (27), 6 (1) teeth with 3 to 6 cusps, midcentral cusps longer than others. Maxillary with 1* (27) or 2 (1) pentacuspidate teeth. Dentary with 5 teeth with 3 to 5 cusps usually midcentral cusps longer than others, followed by 1 to 3 small teeth, with 1 to 3 cusps (
<figureCitation id="62A02A05E909FF9AFDBA907F7E9F0117" box="[597,667,1630,1654]" captionStart="FIGURE 2" captionStartId="4.[151,255,1426,1450]" captionTargetBox="[312,1275,381,1398]" captionTargetId="figure@4.[307,1280,375,1405]" captionTargetPageId="4" captionText="FIGURE 2. Ctenobrycon oliverai, paratype, LBP 3061, 50.7 mm SL, premaxillary, maxillary and dentary; left side, lateral view (maxillary and dentary) or frontal view (premaxillary). Scale bar = 1 mm." httpUri="https://zenodo.org/record/199705/files/figure.png" pageId="2" pageNumber="61">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="FA243680E909FF9AFF2890A37E6D01DF" blockId="2.[151,1436,1486,2015]" pageId="2" pageNumber="61">Nostrils closer to anterior orbital margins than to each other. Supraoccipital process elongate, its tip surpasses the vertical through origin of pectoral fin.</paragraph>
<paragraph id="FA243680E909FF9AFF2890EB7E5E002F" blockId="2.[151,1436,1486,2015]" pageId="2" pageNumber="61">Dorsal-fin rays ii,9. Pectoral-fin rays i,11 (3), i,12* (17), i,13 (8). Tip of pectoral fin exceed anterior half of length of adpressed pelvic fin. Adipose fin well developed. Pelvic-fin rays i,7, when adpressed, its tip extends up to first branched ray of anal fin. Anal-fin rays iv, 39 (2), 40 (1), 41 (1), 42 (2), 43 (8), 44 (7), 45 (3*), 46 (1). Principal caudal-fin rays i,17,i. Caudal fin forked.</paragraph>
<paragraph id="FA243680E909FF9AFF26917B7F4500BE" blockId="2.[151,1436,1486,2015]" pageId="2" pageNumber="61">Spinoid scales. Lateral line complete, 51 (1), 52 (2*), 53 (3), 54 (2), 55 (2), 56 (5), 58 (1). Scale rows between dorsal-fin origin and lateral line 14 (17*), 15 (7), scale rows between lateral line and pelvic-fin origin 11 (11*), 12 (13). Circumpeduncular scale rows 18 (1*), 19(5), 20 (2), 21 (1). Scale sheath along anal-fin base in a single series, extending posteriorly between 33–42* branched anal-fin ray.</paragraph>
<paragraph id="FA243680E908FF9BFF2896B67D4407B5" blockId="3.[151,1436,151,284]" pageId="3" pageNumber="62">First gill arch with 14* (12), 15 (3), 16 (1) on upper limb and 8* (11), 9 (5) on lower limb. Total vertebrae 32, supraneurals 4.</paragraph>
<paragraph id="FA243680E908FF9BFF2896FE7E57067D" blockId="3.[151,1436,151,284]" pageId="3" pageNumber="62">
<emphasis id="C8EFEA92E908FF9BFF2896FE7DBC0799" bold="true" box="[199,440,223,248]" pageId="3" pageNumber="62">Sexual dimorphism.</emphasis>
No secondary sexually dimorphic feature, such as bony hooks on anal and pelvic fins, were observed in 
<taxonomicName id="3D9B4D03E908FF9BFEB297257E48067D" box="[349,588,260,284]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E908FF9BFEB297257E48067D" box="[349,588,260,284]" italics="true" pageId="3" pageNumber="62">Ctenobrycon oliverai</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="AEE46608E908FF9BFF7895F17E000366" httpUri="https://zenodo.org/record/199704/files/figure.png" pageId="3" pageNumber="62" targetBox="[151,1436,323,952]" targetPageId="3">
<paragraph id="FA243680E908FF9BFF7895F17E000366" blockId="3.[151,1436,976,1031]" pageId="3" pageNumber="62">
<emphasis id="C8EFEA92E908FF9BFF7895F17D1E0489" bold="true" box="[151,282,976,1000]" pageId="3" pageNumber="62">FIGURE 1.</emphasis>
<taxonomicName id="3D9B4D03E908FF9BFECC95F07E030489" box="[291,519,977,1000]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="oliverai" status="sp. nov.">
<emphasis id="C8EFEA92E908FF9BFECC95F07E030489" box="[291,519,977,1000]" italics="true" pageId="3" pageNumber="62">Ctenobrycon oliverai</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D3DC57E9E908FF9BFDF995F17E9F0486" box="[534,667,976,999]" pageId="3" pageNumber="62" rank="species">new species</taxonomicNameLabel>
, holotype, MZUSP 50130, 55.4 mm SL, río Apure, West of Ciudad de Apure, State of Apure, Venezuela.
</paragraph>
</caption>
<paragraph id="FA243680E908FF9BFF2692157D7303BC" blockId="3.[151,1437,1076,2037]" pageId="3" pageNumber="62">
<emphasis id="C8EFEA92E908FF9BFF2692157D96032C" bold="true" box="[201,402,1076,1101]" pageId="3" pageNumber="62">Color in alcohol.</emphasis>
Overall coloration yellowish. Mid-dorsal line darker. Scattered small dark chromatophores on dorsal surface of head from upper lip to tip of supraoccipital spine. Infraorbitals, preopercle, and opercle retaining guanine. Lower lip well delimited by dark chromatophores. Small dark chormatophores delineating inferior margin of eyes. Ventral portion of head with very few dark chromatophores, more concentrated on branchiostegal rays.
</paragraph>
<paragraph id="FA243680E908FF9BFF2692C97F1202F1" blockId="3.[151,1437,1076,2037]" pageId="3" pageNumber="62">Wedge-shaped humeral mark extending horizontally from fourth to sixth scale posterior to opercle and vertically from the first to seventh scale series above lateral line. A two scales-deep silver midlateral stripe extending on portion of body beginning at vertical through the first branched dorsal-fin ray, with stripe narrowing posteriorly up to a vertical oval-shaped dark spot on terminus of caudal peduncle. Limits of the erector/depressor muscles of the anal fin outlined by dark chromatophores.</paragraph>
<paragraph id="FA243680E908FF9BFF2693BD7DC001B5" blockId="3.[151,1437,1076,2037]" pageId="3" pageNumber="62">Dorsal fin with scattered small dark chromatophores uniformly distributed in its interradial membrane; second unbranched dorsal-fin ray dark pigmented with chromatophores distributed along its whole extension; first branched dorsal-fin ray with dark pigments restricted to its distal half; further dorsal-fin rays hyaline. Anal fin with few dark chromatophores uniformly distributed in its interradial membrane in most specimens; anal-fin hyaline or with very few sparsely distributed dark chromatophores in few specimens. Caudal fin with small chromatophores sparsely distributed in its interradial membrane. Adipose fin hyaline or with small chromatophores, more concentrated in its distal half. Pectoral fins with small chromatophores homogeneously distributed along the unbranched ray; small chromatophores in the distal half of the first six branched rays; interradial membrane hyaline. Pelvic fin hyaline.</paragraph>
</subSubSection>
<subSubSection id="B281650BE908FF9BFF2890C179040199" box="[199,1280,1760,1785]" pageId="3" pageNumber="62" type="distribution">
<paragraph id="FA243680E908FF9BFF2890C179040199" blockId="3.[151,1437,1076,2037]" box="[199,1280,1760,1785]" pageId="3" pageNumber="62">
<emphasis id="C8EFEA92E908FF9BFF2890C17D640198" bold="true" box="[199,352,1760,1785]" pageId="3" pageNumber="62">Distribution.</emphasis>
Known from the río Orinoco and río Apure, río Orinoco basin, 
<collectingCountry id="828C7610E908FF9BFBC490C1789A0199" box="[1067,1182,1760,1784]" name="Venezuela" pageId="3" pageNumber="62">Venezuela</collectingCountry>
(
<figureCitation id="62A02A05E908FF9BFB4190C178F60199" box="[1198,1266,1760,1784]" captionStart="FIGURE 3" captionStartId="4.[151,255,1973,1997]" captionTargetBox="[451,1132,1538,1945]" captionTargetId="figure@4.[448,1138,1533,1952]" captionTargetPageId="4" captionText="FIGURE 3. Map of northern South America indicating the known geographic distribution of Ctenobrycon oliverai n. sp.. Map by M. J. Weitzman." httpUri="https://zenodo.org/record/199706/files/figure.png" pageId="3" pageNumber="62">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B281650BE908FF9CFF2891247F5F0621" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62" type="etymology">
<paragraph id="FA243680E908FF9BFF2891247F180021" blockId="3.[151,1437,1076,2037]" pageId="3" pageNumber="62">
<emphasis id="C8EFEA92E908FF9BFF2891247D4A007C" bold="true" box="[199,334,1797,1821]" pageId="3" pageNumber="62">Etymology.</emphasis>
The specific name is in honor of Claudio de Oliveira, the collector of the new species and a great contributor to our knowledge of Neotropical Ichthyology.
</paragraph>
<paragraph id="FA243680E908FF9CFF28916D7F5F0621" blockId="3.[151,1437,1076,2037]" lastBlockId="4.[151,1436,151,320]" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
<emphasis id="C8EFEA92E908FF9BFF28916D7DD50004" bold="true" box="[199,465,1868,1893]" pageId="3" pageNumber="62">Multivariate analysis.</emphasis>
The results of the size-free canonical variate analysis revealed that 
<taxonomicName id="3D9B4D03E908FF9BFAE4916C7CEA00E8" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E908FF9BFAE4916C7CEA00E8" italics="true" pageId="3" pageNumber="62">Ctenobrycon oliverai</emphasis>
</taxonomicName>
can be discriminated from 
<taxonomicName id="3D9B4D03E908FF9BFDD891507E9800E8" box="[567,668,1905,1929]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E908FF9BFDD891507E9800E8" box="[567,668,1905,1929]" italics="true" pageId="3" pageNumber="62">C. alleni</emphasis>
</taxonomicName>
, 
<emphasis id="C8EFEA92E908FF9BFD4191507FD900E8" box="[686,989,1905,1929]" italics="true" pageId="3" pageNumber="62">
<taxonomicName id="3D9B4D03E908FF9BFD4191507F2900E8" box="[686,813,1905,1929]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="spilurus">C. spilurus</taxonomicName>
– 
<taxonomicName id="3D9B4D03E908FF9BFCD491507FD900E8" box="[827,989,1905,1929]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="hauxwellianus">hauxwellianus</taxonomicName>
</emphasis>
and from 
<taxonomicName id="3D9B4D03E908FF9BFBB1915378E500E8" box="[1118,1249,1905,1929]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E908FF9BFBB1915378E500E8" box="[1118,1249,1905,1929]" italics="true" pageId="3" pageNumber="62">P. kennedyi</emphasis>
</taxonomicName>
in the first and second canonical variate axis that explains 68.23% and 23.22% of the total variance of the data, respectively (
<figureCitation id="62A02A05E908FF9BFA8191B57CAC00B1" captionStart="FIGURE 4" captionStartId="6.[151,255,1197,1221]" captionTargetBox="[163,1375,422,1146]" captionTargetId="figure@6.[151,1436,338,1176]" captionTargetPageId="6" captionText="FIGURE 4. Scatter diagram of scores of specimens on first and second axis of Size-free Canonical Variate Analysis of Ctenobrycon oliverai (red crosses), C. alleni (green Diamonds), C. spilurus-hauxwellianus (blue squares) and Psellogrammus kennedyi (violet circles)." httpUri="https://zenodo.org/record/199707/files/figure.png" pageId="3" pageNumber="62">Fig. 4</figureCitation>
; 
<tableCitation id="B719033BE908FF9BFF5991997D0B00B1" box="[182,271,1976,2000]" captionStart="TABLE 2" captionStartId="5.[151,239,151,175]" captionTargetBox="[151,1436,234,796]" captionTargetPageId="5" captionText="TABLE 2. Loadings of variables in the first and second size-free Canonical Varietes of Ctenobrycon oliverai, C. alleni, C. spilurus, C. hauxwellianus and Psellogrammus kennedyi." httpUri="http://table.plazi.org/id/AEE46608E90EFF9DFF7896B67F1C07AE" pageId="3" pageNumber="62" tableUuid="AEE46608E90EFF9DFF7896B67F1C07AE">Table 2</tableCitation>
). 
<taxonomicName id="3D9B4D03E908FF9BFEC991987E1C00B0" box="[294,536,1977,2001]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E908FF9BFEC991987E1C00B0" box="[294,536,1977,2001]" italics="true" pageId="3" pageNumber="62">Ctenobrycon oliverai</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E908FF9BFDBB91987EB300B0" box="[596,695,1977,2001]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E908FF9BFDBB91987EB300B0" box="[596,695,1977,2001]" italics="true" pageId="3" pageNumber="62">C. alleni</emphasis>
</taxonomicName>
have greater anal fin-length, least interorbital width, and eye to origin of dorsal-fin distance (higher positive values of CV1, p&lt;0.05) than that found in 
<emphasis id="C8EFEA92E908FF9BFB9A91FC79980094" box="[1141,1436,2013,2037]" italics="true" pageId="3" pageNumber="62">C. spilurus-hauxwellianus</emphasis>
and 
<taxonomicName id="3D9B4D03E90FFF9CFF2796B97DE507D1" box="[200,481,152,176]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90FFF9CFF2796B97DE507D1" box="[200,481,152,176]" italics="true" pageId="4" pageNumber="63">Psellogrammus kennedyi</emphasis>
</taxonomicName>
. These last species in turn present higher snout to dorsal-fin origin distance than 
<taxonomicName id="3D9B4D03E90FFF9CFA6D96B97CEA07B5" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90FFF9CFA6D96B97CEA07B5" italics="true" pageId="4" pageNumber="63">C. oliverai</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90FFF9CFEC9969C7D7B07B5" box="[294,383,188,212]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="aleni">
<emphasis id="C8EFEA92E90FFF9CFEC9969C7D7B07B5" box="[294,383,188,212]" italics="true" pageId="4" pageNumber="63">C. aleni</emphasis>
</taxonomicName>
(higher negative value of CV1, p&lt;0.05). In addition, 
<taxonomicName id="3D9B4D03E90FFF9CFC32969C785107B5" box="[989,1109,188,212]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90FFF9CFC32969C785107B5" box="[989,1109,188,212]" italics="true" pageId="4" pageNumber="63">C. oliverai</emphasis>
</taxonomicName>
could be discriminated from the other three species on the second canonical axis, based on higher body depth (higher positive values of CV2, p&lt;0.05) whereas, 
<emphasis id="C8EFEA92E90FFF9CFE8697247E97067D" box="[361,659,260,284]" italics="true" pageId="4" pageNumber="63">C. spilurus-hauxwellianus</emphasis>
, 
<taxonomicName id="3D9B4D03E90FFF9CFD4A97247F0F067D" box="[677,779,260,284]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E90FFF9CFD4A97247F0F067D" box="[677,779,260,284]" italics="true" pageId="4" pageNumber="63">C. alleni</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90FFF9CFCA597247FCA067D" box="[842,974,260,284]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90FFF9CFCA597247FCA067D" box="[842,974,260,284]" italics="true" pageId="4" pageNumber="63">P. kennedyi</emphasis>
</taxonomicName>
presented greater head depth and head length than 
<taxonomicName id="3D9B4D03E90FFF9CFEF497097D960621" box="[283,402,296,320]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90FFF9CFEF497097D960621" box="[283,402,296,320]" italics="true" pageId="4" pageNumber="63">C. oliverai</emphasis>
</taxonomicName>
(higher negative value of CV2, p&lt;0.05).
</paragraph>
</subSubSection>
<caption id="AEE46608E90FFF9CFF7893B3782902AB" httpUri="https://zenodo.org/record/199705/files/figure.png" pageId="4" pageNumber="63" targetBox="[312,1275,381,1398]" targetPageId="4">
<paragraph id="FA243680E90FFF9CFF7893B3782902AB" blockId="4.[151,1436,1426,1482]" pageId="4" pageNumber="63">
<emphasis id="C8EFEA92E90FFF9CFF7893B37D1F02CB" bold="true" box="[151,283,1426,1450]" pageId="4" pageNumber="63">FIGURE 2.</emphasis>
<taxonomicName id="3D9B4D03E90FFF9CFECA93B27E0F02CB" box="[293,523,1427,1450]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90FFF9CFECA93B27E0F02CB" box="[293,523,1427,1450]" italics="true" pageId="4" pageNumber="63">Ctenobrycon oliverai</emphasis>
</taxonomicName>
, paratype, LBP 3061, 50.7 mm SL, premaxillary, maxillary and dentary; left side, lateral view (maxillary and dentary) or frontal view (premaxillary). Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="AEE46608E90FFF9CFF7891947DCF0091" httpUri="https://zenodo.org/record/199706/files/figure.png" pageId="4" pageNumber="63" targetBox="[451,1132,1538,1945]" targetPageId="4">
<paragraph id="FA243680E90FFF9CFF7891947DCF0091" blockId="4.[151,1436,1973,2033]" pageId="4" pageNumber="63">
<emphasis id="C8EFEA92E90FFF9CFF7891947D1F00AC" bold="true" box="[151,283,1973,1997]" pageId="4" pageNumber="63">FIGURE 3.</emphasis>
Map of northern South America indicating the known geographic distribution of 
<taxonomicName id="3D9B4D03E90FFF9CFB759197797B00AC" box="[1178,1407,1974,1997]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="oliverai" status="sp. nov.">
<emphasis id="C8EFEA92E90FFF9CFB759197797B00AC" box="[1178,1407,1974,1997]" italics="true" pageId="4" pageNumber="63">Ctenobrycon oliverai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3DC57E9E90FFF9CFA6791977CB10090" pageId="4" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
. Map by M. J. Weitzman.
</paragraph>
</caption>
<caption id="AEE46608E90EFF9DFF7896B67F1C07AE" ID-Table-UUID="AEE46608E90EFF9DFF7896B67F1C07AE" httpUri="http://table.plazi.org/id/AEE46608E90EFF9DFF7896B67F1C07AE" pageId="5" pageNumber="64" targetBox="[151,1436,234,796]" targetIsTable="true" targetPageId="5">
<paragraph id="FA243680E90EFF9DFF7896B67F1C07AE" blockId="5.[151,1436,151,207]" pageId="5" pageNumber="64">
<emphasis id="C8EFEA92E90EFF9DFF7896B67D0D07CE" bold="true" box="[151,265,151,175]" pageId="5" pageNumber="64">TABLE 2.</emphasis>
Loadings of variables in the first and second size-free Canonical Varietes of 
<taxonomicName id="3D9B4D03E90EFF9DFBA896B9792F07CE" box="[1095,1323,152,175]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90EFF9DFBA896B9792F07CE" box="[1095,1323,152,175]" italics="true" pageId="5" pageNumber="64">Ctenobrycon oliverai</emphasis>
</taxonomicName>
, 
<taxonomicName id="3D9B4D03E90EFF9DFAD696B9799107CE" box="[1337,1429,152,175]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E90EFF9DFAD696B9799107CE" box="[1337,1429,152,175]" italics="true" pageId="5" pageNumber="64">C. alleni</emphasis>
</taxonomicName>
, 
<taxonomicName id="3D9B4D03E90EFF9DFF7896997D0807AE" box="[151,268,184,207]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90EFF9DFF7896997D0807AE" box="[151,268,184,207]" italics="true" pageId="5" pageNumber="64">C. spilurus</emphasis>
</taxonomicName>
, 
<taxonomicName id="3D9B4D03E90EFF9DFEF696997DD607AE" box="[281,466,184,207]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="hauxwellianus">
<emphasis id="C8EFEA92E90EFF9DFEF696997DD607AE" box="[281,466,184,207]" italics="true" pageId="5" pageNumber="64">C. hauxwellianus</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90EFF9DFDE996997F1507AE" box="[518,785,184,207]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90EFF9DFDE996997F1507AE" box="[518,785,184,207]" italics="true" pageId="5" pageNumber="64">Psellogrammus kennedyi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="FA243680E90EFF9DFD2896CB7945046E" pageId="5" pageNumber="64">
<table id="889BC420E90E0067FF7896CB7998047D" box="[151,1436,234,796]" gridcols="5" gridrows="8" pageId="5" pageNumber="64">
<tr id="44AB34C2E90E0067FF7896CB79980660" box="[151,1436,234,257]" gridrow="0" pageId="5" pageNumber="64" rowspan-0="1">
<th id="077A5DBEE90E0067FD2896CB7F1D0660" box="[711,793,234,257]" gridcol="1" gridrow="0" pageId="5" pageNumber="64">CV1</th>
<th id="077A5DBEE90E0067FC9296CB7FD00660" box="[893,980,234,257]" gridcol="2" gridrow="0" pageId="5" pageNumber="64">Prob.</th>
<th id="077A5DBEE90E0067FBDB96CB78820660" box="[1076,1158,234,257]" gridcol="3" gridrow="0" pageId="5" pageNumber="64">CV2</th>
<th id="077A5DBEE90E0067FB0496CB79980660" box="[1259,1436,234,257]" gridcol="4" gridrow="0" pageId="5" pageNumber="64">Prob.</th>
</tr>
<tr id="44AB34C2E90E0067FF78973479980638" box="[151,1436,277,345]" gridrow="1" pageId="5" pageNumber="64">
<th id="077A5DBEE90E0067FF7897347E2D0638" box="[151,553,277,345]" gridcol="0" gridrow="1" pageId="5" pageNumber="64">Body depth Snout to dorsal-fin origin</th>
<td id="077A5DBEE90E0067FD2897347F1D0638" box="[711,793,277,345]" gridcol="1" gridrow="1" pageId="5" pageNumber="64">0.3625 -0.0934</td>
<td id="077A5DBEE90E0067FC9297347FD00638" box="[893,980,277,345]" gridcol="2" gridrow="1" pageId="5" pageNumber="64">0.6293ns 0.0115*</td>
<td id="077A5DBEE90E0067FBDB973478820638" box="[1076,1158,277,345]" gridcol="3" gridrow="1" pageId="5" pageNumber="64">0.3242 0.0718</td>
<td id="077A5DBEE90E0067FB04973479980638" box="[1259,1436,277,345]" gridcol="4" gridrow="1" pageId="5" pageNumber="64">0.0435* 0.0181*</td>
</tr>
<tr id="44AB34C2E90E0067FF78974C799806D5" box="[151,1436,365,436]" gridrow="2" pageId="5" pageNumber="64">
<th id="077A5DBEE90E0067FF78974C7E2D06D5" box="[151,553,365,436]" gridcol="0" gridrow="2" pageId="5" pageNumber="64">Snout to anal-fin origin Pelvic-fin length</th>
<td id="077A5DBEE90E0067FD28974C7F1D06D5" box="[711,793,365,436]" gridcol="1" gridrow="2" pageId="5" pageNumber="64">0.1383 0.1751</td>
<td id="077A5DBEE90E0067FC92974C7FD006D5" box="[893,980,365,436]" gridcol="2" gridrow="2" pageId="5" pageNumber="64">0.5631ns 0.0749ns</td>
<td id="077A5DBEE90E0067FBDB974C788206D5" box="[1076,1158,365,436]" gridcol="3" gridrow="2" pageId="5" pageNumber="64">-0.5182 0.1891</td>
<td id="077A5DBEE90E0067FB04974C799806D5" box="[1259,1436,365,436]" gridcol="4" gridrow="2" pageId="5" pageNumber="64">0.0000* 0.8191ns</td>
</tr>
<tr id="44AB34C2E90E0067FF7897E979980680" box="[151,1436,456,481]" gridrow="3" pageId="5" pageNumber="64">
<th id="077A5DBEE90E0067FF7897E97E2D0680" box="[151,553,456,481]" gridcol="0" gridrow="3" pageId="5" pageNumber="64">Anal-fin length</th>
<td id="077A5DBEE90E0067FD2897E97F1D0680" box="[711,793,456,481]" gridcol="1" gridrow="3" pageId="5" pageNumber="64">0.7619</td>
<td id="077A5DBEE90E0067FC9297E97FD00680" box="[893,980,456,481]" gridcol="2" gridrow="3" pageId="5" pageNumber="64">0.0000*</td>
<td id="077A5DBEE90E0067FBDB97E978820680" box="[1076,1158,456,481]" gridcol="3" gridrow="3" pageId="5" pageNumber="64">-0.0966</td>
<td id="077A5DBEE90E0067FB0497E979980680" box="[1259,1436,456,481]" gridcol="4" gridrow="3" pageId="5" pageNumber="64">0.0620ns</td>
</tr>
<tr id="44AB34C2E90E0067FF7897D479980558" box="[151,1436,501,569]" gridrow="4" pageId="5" pageNumber="64">
<th id="077A5DBEE90E0067FF7897D47E2D0558" box="[151,553,501,569]" gridcol="0" gridrow="4" pageId="5" pageNumber="64">Anal-fin base Eye to dorsal-fin origin</th>
<td id="077A5DBEE90E0067FD2897D47F1D0558" box="[711,793,501,569]" gridcol="1" gridrow="4" pageId="5" pageNumber="64">0.1178 0.1283</td>
<td id="077A5DBEE90E0067FC9297D47FD00558" box="[893,980,501,569]" gridcol="2" gridrow="4" pageId="5" pageNumber="64">0.8807ns 0.0135*</td>
<td id="077A5DBEE90E0067FBDB97D478820558" box="[1076,1158,501,569]" gridcol="3" gridrow="4" pageId="5" pageNumber="64">0.5496 -0.2729</td>
<td id="077A5DBEE90E0067FB0497D479980558" box="[1259,1436,501,569]" gridcol="4" gridrow="4" pageId="5" pageNumber="64">0.0978ns 0.0001*</td>
</tr>
<tr id="44AB34C2E90E0067FF78946C799805F5" box="[151,1436,589,660]" gridrow="5" pageId="5" pageNumber="64">
<th id="077A5DBEE90E0067FF78946C7E2D05F5" box="[151,553,589,660]" gridcol="0" gridrow="5" pageId="5" pageNumber="64">Dorsal-fin origin to caudal-fin origin Head depth</th>
<td id="077A5DBEE90E0067FD28946C7F1D05F5" box="[711,793,589,660]" gridcol="1" gridrow="5" pageId="5" pageNumber="64">0.1385 0.2928</td>
<td id="077A5DBEE90E0067FC92946C7FD005F5" box="[893,980,589,660]" gridcol="2" gridrow="5" pageId="5" pageNumber="64">0.3194ns 0.4602ns</td>
<td id="077A5DBEE90E0067FBDB946C788205F5" box="[1076,1158,589,660]" gridcol="3" gridrow="5" pageId="5" pageNumber="64">0.0673 -0.3413</td>
<td id="077A5DBEE90E0067FB04946C799805F5" box="[1259,1436,589,660]" gridcol="4" gridrow="5" pageId="5" pageNumber="64">0.1675ns 0.0000*</td>
</tr>
<tr id="44AB34C2E90E0067FF789489799805A0" box="[151,1436,680,705]" gridrow="6" pageId="5" pageNumber="64">
<th id="077A5DBEE90E0067FF7894897E2D05A0" box="[151,553,680,705]" gridcol="0" gridrow="6" pageId="5" pageNumber="64">Head length</th>
<td id="077A5DBEE90E0067FD2894897F1D05A0" box="[711,793,680,705]" gridcol="1" gridrow="6" pageId="5" pageNumber="64">0.1448</td>
<td id="077A5DBEE90E0067FC9294897FD005A0" box="[893,980,680,705]" gridcol="2" gridrow="6" pageId="5" pageNumber="64">0.9687ns</td>
<td id="077A5DBEE90E0067FBDB9489788205A0" box="[1076,1158,680,705]" gridcol="3" gridrow="6" pageId="5" pageNumber="64">-0.0230</td>
<td id="077A5DBEE90E0067FB049489799805A0" box="[1259,1436,680,705]" gridcol="4" gridrow="6" pageId="5" pageNumber="64">0.0026*</td>
</tr>
<tr id="44AB34C2E90E0067FF7894F47998047D" box="[151,1436,725,796]" gridrow="7" pageId="5" pageNumber="64">
<th id="077A5DBEE90E0067FF7894F47E2D047D" box="[151,553,725,796]" gridcol="0" gridrow="7" pageId="5" pageNumber="64">Least interobital width Upper jaw length</th>
<td id="077A5DBEE90E0067FD2894F47F1D047D" box="[711,793,725,796]" gridcol="1" gridrow="7" pageId="5" pageNumber="64">0.2652 0.0523</td>
<td id="077A5DBEE90E0067FC9294F47FD0047D" box="[893,980,725,796]" gridcol="2" gridrow="7" pageId="5" pageNumber="64">0.0043* 0.2473ns</td>
<td id="077A5DBEE90E0067FBDB94F47882047D" box="[1076,1158,725,796]" gridcol="3" gridrow="7" pageId="5" pageNumber="64">0.2115 0.1822</td>
<td id="077A5DBEE90E0067FB0494F47998047D" box="[1259,1436,725,796]" gridcol="4" gridrow="7" pageId="5" pageNumber="64">0.3815ns 0.1082ns</td>
</tr>
</table>
</paragraph>
<subSubSection id="B281650BE90EFF9FFF2695A4780E055D" lastPageId="7" lastPageNumber="66" pageId="5" pageNumber="64" type="discussion">
<paragraph id="FA243680E90EFF9DFF2695A47E1B0160" blockId="5.[151,1437,901,2005]" pageId="5" pageNumber="64">
<emphasis id="C8EFEA92E90EFF9DFF2695A47D4A04FF" bold="true" box="[201,334,901,926]" pageId="5" pageNumber="64">Discussion.</emphasis>
<bibRefCitation id="9E0A4B71E90EFF9DFEB695A47E9F04FC" author="Eigenmann" box="[345,667,901,925]" pageId="5" pageNumber="64" refString="Eigenmann, C. H. (1927) The American Characidae. Memoirs of the Museum of the Comparative Zoology, Harward University, 43, 311 - 428." type="journal article" year="1927">Eigenmann (1927:330–336)</bibRefCitation>
, in the first revisionary study after the original description of the genus, recognized 
<taxonomicName id="3D9B4D03E90EFF9DFE9C95887F0B04A0" authority="Cope" authorityName="Cope" box="[371,783,937,961]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="hauxwellianus">
<emphasis id="C8EFEA92E90EFF9DFE9C95887EB604A0" box="[371,690,937,961]" italics="true" pageId="5" pageNumber="64">Ctenobrycon hauxwellianus</emphasis>
(Cope)
</taxonomicName>
, from Amazon basin; 
<taxonomicName id="3D9B4D03E90EFF9DFBF1958B795104A0" authority="Valenciennes" authorityName="Valenciennes" box="[1054,1365,937,961]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90EFF9DFBF1958B789A04A0" box="[1054,1182,937,961]" italics="true" pageId="5" pageNumber="64">C. spilurus</emphasis>
(Valenciennes)
</taxonomicName>
, from 
<collectingCountry id="828C7610E90EFF9DFF7895EC7D0E0484" box="[151,266,973,997]" name="Venezuela" pageId="5" pageNumber="64">Venezuela</collectingCountry>
and 
<collectingCountry id="828C7610E90EFF9DFEAB95EC7DB70484" box="[324,435,973,997]" name="Suriname" pageId="5" pageNumber="64">Suriname</collectingCountry>
; and 
<taxonomicName id="3D9B4D03E90EFF9DFE1E95EF7F5E0484" authority="Steindachner" authorityName="Steindachner" box="[497,858,973,997]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="multiradiatus">
<emphasis id="C8EFEA92E90EFF9DFE1E95EF7EAE0484" box="[497,682,973,997]" italics="true" pageId="5" pageNumber="64">C. multiradiatus</emphasis>
(Steindachner)
</taxonomicName>
, from Amazon basin and possibly 
<collectingCountry id="828C7610E90EFF9DFB0695EF79550487" box="[1257,1361,974,998]" name="Paraguay" pageId="5" pageNumber="64">Paraguay</collectingCountry>
basin. This author diagnosed 
<taxonomicName id="3D9B4D03E90EFF9DFE7695D37E590368" box="[409,605,1009,1033]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="hauxwellianus">
<emphasis id="C8EFEA92E90EFF9DFE7695D37E590368" box="[409,605,1009,1033]" italics="true" pageId="5" pageNumber="64">C. hauxwellianus</emphasis>
</taxonomicName>
from 
<taxonomicName id="3D9B4D03E90EFF9DFD4C95D37F1A0368" box="[675,798,1009,1033]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90EFF9DFD4C95D37F1A0368" box="[675,798,1009,1033]" italics="true" pageId="5" pageNumber="64">C. spilurus</emphasis>
</taxonomicName>
uniquely by the average of the body depth (2.0 times in SL versus 2.5 times in SL, respectively). Eigenmann, however, was not sure about the validity of 
<taxonomicName id="3D9B4D03E90EFF9DFB3292377991034C" box="[1245,1429,1045,1069]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="multiradiatus">
<emphasis id="C8EFEA92E90EFF9DFB3292377991034C" box="[1245,1429,1045,1069]" italics="true" pageId="5" pageNumber="64">C. multiradiatus</emphasis>
</taxonomicName>
, since he considered that the diagnostic characters used by Steindachner could be purely individual variations and also pointed out that the forms from 
<collectingCountry id="828C7610E90EFF9DFDA7927F7EB40317" box="[584,688,1118,1142]" name="Paraguay" pageId="5" pageNumber="64">Paraguay</collectingCountry>
were most likely 
<taxonomicName id="3D9B4D03E90EFF9DFC6A927C79500317" authority="Eigenmann &amp; MacAtee" authorityName="Eigenmann &amp; MacAtee" box="[901,1364,1117,1142]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E90EFF9DFC6A927C78320314" box="[901,1078,1117,1141]" italics="true" pageId="5" pageNumber="64">Astyanax alleni</emphasis>
Eigenmann &amp; MacAtee
</taxonomicName>
, or 
<taxonomicName id="3D9B4D03E90EFF9DFA6B927C7D8203F8" authority="Eigenmann" authorityName="Eigenmann" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="pelegrini">
<emphasis id="C8EFEA92E90EFF9DFA6B927C7CF803F8" italics="true" pageId="5" pageNumber="64">A. pelegrini</emphasis>
Eigenmann
</taxonomicName>
or even 
<taxonomicName id="3D9B4D03E90EFF9DFE1E92A0783303F8" authority="Eigenmann. Gery (1977)" authorityName="Eigenmann. Gery" authorityYear="1977" box="[497,1079,1153,1177]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90EFF9DFE1E92A07F0803F8" box="[497,780,1153,1177]" italics="true" pageId="5" pageNumber="64">Psellogrammus kennedyi</emphasis>
Eigenmann. 
<bibRefCitation id="9E0A4B71E90EFF9DFC4792A0783303F8" author="Gery" box="[936,1079,1153,1177]" pageId="5" pageNumber="64" refString="Gery, J. (1977) Characoids of the World. THF publications, Neptune City, NJ, 672 pp." type="book" year="1977">Géry (1977)</bibRefCitation>
</taxonomicName>
, reallocated 
<taxonomicName id="3D9B4D03E90EFF9DFB2492A0797803F8" box="[1227,1404,1153,1177]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E90EFF9DFB2492A0797803F8" box="[1227,1404,1153,1177]" italics="true" pageId="5" pageNumber="64">Astyanax alleni</emphasis>
</taxonomicName>
in 
<taxonomicName id="3D9B4D03E90EFF9DFF7892847D2C03DC" box="[151,296,1189,1213]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="C8EFEA92E90EFF9DFF7892847D2C03DC" box="[151,296,1189,1213]" italics="true" pageId="5" pageNumber="64">Ctenobrycon</emphasis>
</taxonomicName>
but, due to the great overlap in the characters, named the subspecies 
<taxonomicName id="3D9B4D03E90EFF9DFBDA9287791603DC" box="[1077,1298,1189,1213]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="subSpecies" species="spilurus" subSpecies="spilurus">
<emphasis id="C8EFEA92E90EFF9DFBDA9287791603DC" box="[1077,1298,1189,1213]" italics="true" pageId="5" pageNumber="64">C. spilurus spilurus</emphasis>
</taxonomicName>
, 
<taxonomicName id="3D9B4D03E90EFF9DFACF92877D3D0380" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="subSpecies" species="spilurus" subSpecies="hauxwellianus">
<emphasis id="C8EFEA92E90EFF9DFACF92877D3D0380" italics="true" pageId="5" pageNumber="64">C. spilurus hauxwellianus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="3D9B4D03E90EFF9DFE9792EB7E380380" box="[376,572,1225,1249]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="subSpecies" species="spilurus" subSpecies="alleni">
<emphasis id="C8EFEA92E90EFF9DFE9792EB7E380380" box="[376,572,1225,1249]" italics="true" pageId="5" pageNumber="64">C. spilurus alleni</emphasis>
</taxonomicName>
. Besides, this author also cited 
<taxonomicName id="3D9B4D03E90EFF9DFC4C92EB785F0380" box="[931,1115,1225,1249]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="multiradiatus">
<emphasis id="C8EFEA92E90EFF9DFC4C92EB785F0380" box="[931,1115,1225,1249]" italics="true" pageId="5" pageNumber="64">C. multiradiatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90EFF9DFB7B92EB79320380" box="[1172,1334,1226,1249]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="correntinus">
<emphasis id="C8EFEA92E90EFF9DFB7B92EB79320380" box="[1172,1334,1226,1249]" italics="true" pageId="5" pageNumber="64">C. correntinus</emphasis>
</taxonomicName>
as valid, but argued that these were probably synonyms of 
<taxonomicName id="3D9B4D03E90EFF9DFCE592CF7FD20264" box="[778,982,1261,1285]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="hauxwellianus">
<emphasis id="C8EFEA92E90EFF9DFCE592CF7FD20264" box="[778,982,1261,1285]" italics="true" pageId="5" pageNumber="64">C. hauxwellianus</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90EFF9DFBF292CC78FC0264" box="[1053,1272,1261,1285]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="pelegrini">
<emphasis id="C8EFEA92E90EFF9DFBF292CC78FC0264" box="[1053,1272,1261,1285]" italics="true" pageId="5" pageNumber="64">Astyanax pelegrini</emphasis>
</taxonomicName>
, respectively. Notwithstanding the 
<typeStatus id="25208822E90EFF9DFE6393337DEA024B" box="[396,494,1298,1322]" pageId="5" pageNumber="64" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="3D9B4D03E90EFF9DFDF093337E9B0248" box="[543,671,1297,1321]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90EFF9DFDF093337E9B0248" box="[543,671,1297,1321]" italics="true" pageId="5" pageNumber="64">C. spilurus</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90EFF9DFD0F93337FAC0248" box="[736,936,1297,1321]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="hauxwellianus">
<emphasis id="C8EFEA92E90EFF9DFD0F93337FAC0248" box="[736,936,1297,1321]" italics="true" pageId="5" pageNumber="64">C. hauxwellianus</emphasis>
</taxonomicName>
are inadequately preserved, which further hinders the process of delineating these species, we herein recognized 
<taxonomicName id="3D9B4D03E90EFF9DFC5593177918022C" authority="Cope, 1870" authorityName="Cope" authorityYear="1870" box="[954,1308,1333,1357]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="hauxwellianus">
<emphasis id="C8EFEA92E90EFF9DFC559317787A022C" box="[954,1150,1333,1357]" italics="true" pageId="5" pageNumber="64">C. hauxwellianus</emphasis>
(
<bibRefCitation id="9E0A4B71E90EFF9DFB6193147917022C" author="Cope" box="[1166,1299,1333,1357]" pageId="5" pageNumber="64" refString="Cope, E. D. (1870) Contribution to the ichthyology of the Maranon. Proceedings of the American Philosophical Society, 11, 559 - 570." type="journal article" year="1870">Cope, 1870</bibRefCitation>
)
</taxonomicName>
as a junior synonym of 
<taxonomicName id="3D9B4D03E90EFF9DFECB937B7E5D0210" authority="Vallenciennes" authorityName="Vallenciennes" box="[292,601,1369,1393]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90EFF9DFECB937B7DA40210" box="[292,416,1369,1393]" italics="true" pageId="5" pageNumber="64">C. spilurus</emphasis>
(Vallenciennes)
</taxonomicName>
, based on the examination of the comparative lots of 
<taxonomicName id="3D9B4D03E90EFF9DFB579378794D0210" box="[1208,1353,1369,1393]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="C8EFEA92E90EFF9DFB579378794D0210" box="[1208,1353,1369,1393]" italics="true" pageId="5" pageNumber="64">Ctenobrycon</emphasis>
</taxonomicName>
, which did reveal a substantial overlap for all morphometric and meristic characters, not supporting the diagnostic characters presented by 
<bibRefCitation id="9E0A4B71E90EFF9DFE4193807E8602D8" author="Eigenmann" box="[430,642,1441,1465]" pageId="5" pageNumber="64" refString="Eigenmann, C. H. (1927) The American Characidae. Memoirs of the Museum of the Comparative Zoology, Harward University, 43, 311 - 428." type="journal article" year="1927">Eigenmann (1927)</bibRefCitation>
for 
<taxonomicName id="3D9B4D03E90EFF9DFD5793837F3202D8" box="[696,822,1441,1465]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90EFF9DFD5793837F3202D8" box="[696,822,1441,1465]" italics="true" pageId="5" pageNumber="64">C. spilurus</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90EFF9DFC9C9383783D02D8" box="[883,1081,1441,1465]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="hauxwellianus">
<emphasis id="C8EFEA92E90EFF9DFC9C9383783D02D8" box="[883,1081,1441,1465]" italics="true" pageId="5" pageNumber="64">C. hauxwellianus</emphasis>
</taxonomicName>
. 
<taxonomicName id="3D9B4D03E90EFF9DFBA59380792002D8" box="[1098,1316,1441,1465]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E90EFF9DFBA59380792002D8" box="[1098,1316,1441,1465]" italics="true" pageId="5" pageNumber="64">Ctenobrycon alleni</emphasis>
</taxonomicName>
, however, could be easily distinguished from congeners by presenting two humeral blotches versus a single one, a condition already described by Britski, 2007.
</paragraph>
<paragraph id="FA243680E90EFF9DFF26902C7D050040" blockId="5.[151,1437,901,2005]" pageId="5" pageNumber="64">
Taking into account this great overlap in characters, a given count that falls totally out of the known range of 
<taxonomicName id="3D9B4D03E90EFF9DFF7890107D2C0128" box="[151,296,1585,1609]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="C8EFEA92E90EFF9DFF7890107D2C0128" box="[151,296,1585,1609]" italics="true" pageId="5" pageNumber="64">Ctenobrycon</emphasis>
</taxonomicName>
is strong evidence that it is related to a different species. From the 235 examined comparative specimens, all of them presented up to 13 scale rows above the lateral line (
<emphasis id="C8EFEA92E90EFF9DFC0E90777FFB010C" box="[993,1023,1622,1645]" italics="true" pageId="5" pageNumber="64">vs.</emphasis>
<quantity id="3D639B65E90EFF9DFBE990747874010C" box="[1030,1136,1621,1645]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="3.81" metricValueMin="3.556" pageId="5" pageNumber="64" unit="in" value="14.5" valueMax="15.0" valueMin="14.0">14 –15 in</quantity>
<taxonomicName id="3D9B4D03E90EFF9DFB99907778EA010C" box="[1142,1262,1621,1645]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90EFF9DFB99907778EA010C" box="[1142,1262,1621,1645]" italics="true" pageId="5" pageNumber="64">C. oliverai</emphasis>
</taxonomicName>
). Moreover, no comparative material from río Orinoco drainage has more than 12 scale rows above lateral line, which enhance our decision of considering this species as new rather than a population which would only widen the variation range observed for 
<taxonomicName id="3D9B4D03E90EFF9DFEDF90E07E2101B8" box="[304,549,1729,1753]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90EFF9DFEDF90E07E2101B8" box="[304,549,1729,1753]" italics="true" pageId="5" pageNumber="64">Ctenobrycon spilurus</emphasis>
</taxonomicName>
. The results of the multivariate analysis corroborate our hypothesis showing statistically significant evidences which discriminate 
<taxonomicName id="3D9B4D03E90EFF9DFD1590C47FEF019C" box="[762,1003,1765,1789]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90EFF9DFD1590C47FEF019C" box="[762,1003,1765,1789]" italics="true" pageId="5" pageNumber="64">Ctenobrycon oliverai</emphasis>
</taxonomicName>
from 
<taxonomicName id="3D9B4D03E90EFF9DFBDB90C778B6019C" box="[1076,1202,1765,1789]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90EFF9DFBDB90C778B6019C" box="[1076,1202,1765,1789]" italics="true" pageId="5" pageNumber="64">C. spilurus</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90EFF9DFB0190C47CFF0040" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90EFF9DFB0190C47CFF0040" italics="true" pageId="5" pageNumber="64">Psellogrammus kennedyi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="FA243680E90EFF9EFF26910C7E8A067D" blockId="5.[151,1437,901,2005]" lastBlockId="6.[151,1436,151,284]" lastPageId="6" lastPageNumber="65" pageId="5" pageNumber="64">
<bibRefCitation id="9E0A4B71E90EFF9DFF26910C7DA50024" box="[201,417,1837,1861]" pageId="5" pageNumber="64" refString="Lima, F. C. T., Malabarba, L. R., Buckup, P. A., Silva, J. F. P., Vari, R. P., Harold, A., Benine, R., Oyakawa, O. T., Pavanelli, C. S., Menezes, N. A., Lucena, C. A. S., Reis, R. E., Langeani, F., Casatti, L., Bertaco, V. A., Moreira, C. R. &amp; Lucinda, P. H. F. (2003) Genera Incertae Sedis in Characidae. In: Reis, R. E., Kullander, S. O. &amp; Ferraris, Jr., C. J. (Ed.) Check List of the Freshwater fishes of South and Central America. Edipucrs, Porto Alegre, 106 - 169." type="book chapter">
Lima 
<emphasis id="C8EFEA92E90EFF9DFEE0910F7D480024" box="[271,332,1837,1861]" italics="true" pageId="5" pageNumber="64">et al.</emphasis>
(2003)
</bibRefCitation>
listed 
<taxonomicName id="3D9B4D03E90EFF9DFE1B910F7FB30024" authority="Holmberg" authorityName="Holmberg" box="[500,951,1837,1861]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="correntinus">
<emphasis id="C8EFEA92E90EFF9DFE1B910F7F330024" box="[500,823,1838,1861]" italics="true" pageId="5" pageNumber="64">Tetragonopterus correntinus</emphasis>
Holmberg
</taxonomicName>
, 
<taxonomicName id="3D9B4D03E90EFF9DFC29910F78D20024" authority="Pelegrin" authorityName="Pelegrin" box="[966,1238,1837,1861]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="gibbicervix">
<emphasis id="C8EFEA92E90EFF9DFC29910F78670024" box="[966,1123,1837,1861]" italics="true" pageId="5" pageNumber="64">T. gibbicervix</emphasis>
, Pelegrin
</taxonomicName>
, 
<taxonomicName id="3D9B4D03E90EFF9DFB0A910F7D2D0008" authority="Steindachner" authorityName="Steindachner" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="multiradiatus">
<emphasis id="C8EFEA92E90EFF9DFB0A910F79980024" box="[1253,1436,1837,1861]" italics="true" pageId="5" pageNumber="64">T. multiradiatus</emphasis>
Steindachner
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90EFF9DFE8E91707EB90008" authority="Eigenmann" authorityName="Eigenmann" box="[353,701,1873,1897]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="pelegrini">
<emphasis id="C8EFEA92E90EFF9DFE8E91707E310008" box="[353,565,1873,1897]" italics="true" pageId="5" pageNumber="64">Astyanax pelegrini</emphasis>
Eigenmann
</taxonomicName>
as 
<emphasis id="C8EFEA92E90EFF9DFD0B91737FC70008" box="[740,963,1873,1897]" italics="true" pageId="5" pageNumber="64">species inquirendae</emphasis>
in 
<taxonomicName id="3D9B4D03E90EFF9DFC069170787E0008" box="[1001,1146,1873,1897]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="C8EFEA92E90EFF9DFC069170787E0008" box="[1001,1146,1873,1897]" italics="true" pageId="5" pageNumber="64">Ctenobrycon</emphasis>
</taxonomicName>
. Nonetheless, 
<bibRefCitation id="9E0A4B71E90EFF9DFACE91707D0C00EC" pageId="5" pageNumber="64" refString="Mirande, J. M., Azpelicueta, M. M. &amp; Aguilera, G. (2006) Redescription of Astyanax correntinus (Holmberg, 1891) (Teleostei: Characiformes: Caracidae), more than one hundred years after original description. Zoologische Abhandlungen (Dresden), 55, 9 - 15." type="journal article">
Mirande 
<emphasis id="C8EFEA92E90EFF9DFA6791737CB000EC" italics="true" pageId="5" pageNumber="64">et al.</emphasis>
(2006)
</bibRefCitation>
considered both 
<taxonomicName id="3D9B4D03E90EFF9DFE2191577F0B00EC" box="[462,783,1910,1933]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="correntinus">
<emphasis id="C8EFEA92E90EFF9DFE2191577F0B00EC" box="[462,783,1910,1933]" italics="true" pageId="5" pageNumber="64">Tetragonopterus correntinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90EFF9DFCA59154782300EC" box="[842,1063,1909,1933]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="pellegrini">
<emphasis id="C8EFEA92E90EFF9DFCA59154782300EC" box="[842,1063,1909,1933]" italics="true" pageId="5" pageNumber="64">Astyanax pellegrini</emphasis>
</taxonomicName>
, species of 
<taxonomicName id="3D9B4D03E90EFF9DFB5E9154791C00EC" box="[1201,1304,1909,1933]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="C8EFEA92E90EFF9DFB5E9154791C00EC" box="[1201,1304,1909,1933]" italics="true" pageId="5" pageNumber="64">Astyanax</emphasis>
</taxonomicName>
rather than 
<taxonomicName id="3D9B4D03E90EFF9DFF7891B87D2C00D0" box="[151,296,1945,1969]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="C8EFEA92E90EFF9DFF7891B87D2C00D0" box="[151,296,1945,1969]" italics="true" pageId="5" pageNumber="64">Ctenobrycon</emphasis>
</taxonomicName>
, since these do not display the spinoid scales, which promptly separate these taxa from our new species (that presents spinoid scales). According to Mirande’s (2010) examinations, the scales of the belly of 
<taxonomicName id="3D9B4D03E90DFF9EFF7896B97DB507D1" box="[151,433,152,176]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90DFF9EFF7896B97DB507D1" box="[151,433,152,176]" italics="true" pageId="6" pageNumber="65">Psellogrammus kennedyi</emphasis>
</taxonomicName>
have simple flattened serrations restricted to the margin of the scales similar to that of crenate scales in the classification of Roberts (1993). However, our examinations indicated that the form of the scales of the belly in both 
<taxonomicName id="3D9B4D03E90DFF9EFE5196C17E4B0799" box="[446,591,224,248]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="C8EFEA92E90DFF9EFE5196C17E4B0799" box="[446,591,224,248]" italics="true" pageId="6" pageNumber="65">Ctenobrycon</emphasis>
</taxonomicName>
and 
<taxonomicName id="3D9B4D03E90DFF9EFD6A96C17F370799" box="[645,819,224,248]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="C8EFEA92E90DFF9EFD6A96C17F370799" box="[645,819,224,248]" italics="true" pageId="6" pageNumber="65">Psellogrammus</emphasis>
</taxonomicName>
species are, in fact, spinoid, with acute projections not restricted to the margin of the scales (
<figureCitation id="62A02A05E90DFF9EFDD397257E84067D" box="[572,640,260,284]" captionStart="FIGURE 5" captionStartId="6.[151,255,1831,1855]" captionTargetBox="[505,1087,1342,1803]" captionTargetId="figure@6.[489,1098,1336,1810]" captionTargetPageId="6" captionText="FIGURE 5. Spinoid scale from the prepelvic area of Ctenobrycon spilurus, LIRP 4999, 35.7 mm SL." httpUri="https://zenodo.org/record/199708/files/figure.png" pageId="6" pageNumber="65">Fig. 5</figureCitation>
).
</paragraph>
<caption id="AEE46608E90DFF9EFF78928C7E4F0265" httpUri="https://zenodo.org/record/199707/files/figure.png" pageId="6" pageNumber="65" targetBox="[163,1375,422,1146]" targetPageId="6">
<paragraph id="FA243680E90DFF9EFF78928C7E4F0265" blockId="6.[151,1436,1197,1285]" pageId="6" pageNumber="65">
<emphasis id="C8EFEA92E90DFF9EFF78928C7D1E03A4" bold="true" box="[151,282,1197,1221]" pageId="6" pageNumber="65">FIGURE 4.</emphasis>
Scatter diagram of scores of specimens on first and second axis of Size-free Canonical Variate Analysis of 
<taxonomicName id="3D9B4D03E90DFF9EFF7892EF7D800384" box="[151,388,1230,1253]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="oliverai">
<emphasis id="C8EFEA92E90DFF9EFF7892EF7D800384" box="[151,388,1230,1253]" italics="true" pageId="6" pageNumber="65">Ctenobrycon oliverai</emphasis>
</taxonomicName>
(red crosses), 
<taxonomicName id="3D9B4D03E90DFF9EFDD392EF7EA50384" box="[572,673,1230,1253]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E90DFF9EFDD392EF7EA50384" box="[572,673,1230,1253]" italics="true" pageId="6" pageNumber="65">C. alleni</emphasis>
</taxonomicName>
(green Diamonds), 
<emphasis id="C8EFEA92E90DFF9EFC7B92EF78B10384" box="[916,1205,1230,1253]" italics="true" pageId="6" pageNumber="65">C. spilurus-hauxwellianus</emphasis>
(blue squares) and 
<taxonomicName id="3D9B4D03E90DFF9EFF7892CF7DA60264" box="[151,418,1262,1285]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90DFF9EFF7892CF7DA60264" box="[151,418,1262,1285]" italics="true" pageId="6" pageNumber="65">Psellogrammus kennedyi</emphasis>
</taxonomicName>
(violet circles).
</paragraph>
</caption>
<caption id="AEE46608E90DFF9EFF78910678D7005F" box="[151,1235,1831,1855]" httpUri="https://zenodo.org/record/199708/files/figure.png" pageId="6" pageNumber="65" targetBox="[505,1087,1342,1803]" targetPageId="6">
<paragraph id="FA243680E90DFF9EFF78910678D7005F" blockId="6.[151,1235,1831,1855]" box="[151,1235,1831,1855]" pageId="6" pageNumber="65">
<emphasis id="C8EFEA92E90DFF9EFF7891067D1E005E" bold="true" box="[151,282,1831,1855]" pageId="6" pageNumber="65">FIGURE 5.</emphasis>
Spinoid scale from the prepelvic area of 
<taxonomicName id="3D9B4D03E90DFF9EFD3E91067FB3005F" box="[721,951,1831,1854]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90DFF9EFD3E91067FB3005F" box="[721,951,1831,1854]" italics="true" pageId="6" pageNumber="65">Ctenobrycon spilurus</emphasis>
</taxonomicName>
, LIRP 4999, 35.7 mm SL.
</paragraph>
</caption>
<paragraph id="FA243680E90DFF9EFF26914A7DA200AA" blockId="6.[151,1436,1899,1995]" pageId="6" pageNumber="65">
Examination of images of three 
<typeStatus id="25208822E90DFF9EFDA2914D7EAB00E5" box="[589,687,1900,1924]" pageId="6" pageNumber="65" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="3D9B4D03E90DFF9EFD0F914D782700E2" box="[736,1059,1899,1923]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="gibbicervix">
<emphasis id="C8EFEA92E90DFF9EFD0F914D782700E2" box="[736,1059,1899,1923]" italics="true" pageId="6" pageNumber="65">Tetragonopterus gibbicervix</emphasis>
</taxonomicName>
, made it clear that this species presents 12 scale rows above lateral line, which distinguishes it from our new species (that presents 14–15 scale rows above lateral line).
</paragraph>
<paragraph id="FA243680E90CFF9FFF2696B6780E055D" blockId="7.[151,1437,151,1113]" pageId="7" pageNumber="66">
<bibRefCitation id="9E0A4B71E90CFF9FFF2696B67D5B07CE" author="Gery" box="[201,351,151,175]" pageId="7" pageNumber="66" refString="Gery, J. (2006) Note on the orthography of Astyanax pellegrini Eigenmann 1907. Zoologische Abhandlungen (Dresden), 56, 47." type="journal article" year="2006">Géry (2006)</bibRefCitation>
affirmed that 
<taxonomicName id="3D9B4D03E90CFF9FFDCB96B8782807CE" authority="Steindachner" authorityName="Steindachner" box="[548,1068,151,176]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="multiradiatus">
<emphasis id="C8EFEA92E90CFF9FFDCB96B87F8207D1" box="[548,902,152,176]" italics="true" pageId="7" pageNumber="66">Tetragonopterus multiradiatus</emphasis>
Steindachner
</taxonomicName>
is a junior synonym of 
<taxonomicName id="3D9B4D03E90CFF9FFA6D96B97D3D07B5" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="hauxwellianus">
<emphasis id="C8EFEA92E90CFF9FFA6D96B97D3D07B5" italics="true" pageId="7" pageNumber="66">C. hauxwellianus</emphasis>
</taxonomicName>
, but did not expose the reasons for considering so, nor de he mention the paper where this new combination was proposed, if it exists at all. Considering that the 
<typeStatus id="25208822E90CFF9FFC6296C17FEB0799" box="[909,1007,224,248]" pageId="7" pageNumber="66" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="3D9B4D03E90CFF9FFBF796C079740799" box="[1048,1392,224,248]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="multiradiatus">
<emphasis id="C8EFEA92E90CFF9FFBF796C079740799" box="[1048,1392,224,248]" italics="true" pageId="7" pageNumber="66">Tetragonopterus multiradiatus</emphasis>
</taxonomicName>
are possibly lost (see 
<bibRefCitation id="9E0A4B71E90CFF9FFE8397257E39067D" author="Lima" box="[364,573,260,284]" pageId="7" pageNumber="66" refString="Lima, F. C. T., Malabarba, L. R., Buckup, P. A., Silva, J. F. P., Vari, R. P., Harold, A., Benine, R., Oyakawa, O. T., Pavanelli, C. S., Menezes, N. A., Lucena, C. A. S., Reis, R. E., Langeani, F., Casatti, L., Bertaco, V. A., Moreira, C. R. &amp; Lucinda, P. H. F. (2003) Genera Incertae Sedis in Characidae. In: Reis, R. E., Kullander, S. O. &amp; Ferraris, Jr., C. J. (Ed.) Check List of the Freshwater fishes of South and Central America. Edipucrs, Porto Alegre, 106 - 169." type="book chapter" year="2003">
Lima 
<emphasis id="C8EFEA92E90CFF9FFE5D97247DEB067D" box="[434,495,260,284]" italics="true" pageId="7" pageNumber="66">et al.</emphasis>
, 2003
</bibRefCitation>
), not much information can be retrieved but from Steindachner’s original description and putative topotypes. The eight examined specimens from the 
<typeStatus id="25208822E90CFF9FFBD69709786D0621" box="[1081,1129,296,320]" pageId="7" pageNumber="66">type</typeStatus>
locality (MZUSP 27765, município de Tefé, AM, 
<collectingCountry id="828C7610E90CFF9FFE45976D7DF40605" box="[426,496,332,356]" name="Brazil" pageId="7" pageNumber="66">Brazil</collectingCountry>
) presented one tooth in the maxillary bone, 45 to 49 scales in the lateral line, and 11 to 12 scale rows above lateral line and 9 to 10 scale rows below lateral line, perfectly fitting with 
<taxonomicName id="3D9B4D03E90CFF9FFB359751799806E9" box="[1242,1436,368,392]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="hauxwellianus">
<emphasis id="C8EFEA92E90CFF9FFB359751799806E9" box="[1242,1436,368,392]" italics="true" pageId="7" pageNumber="66">C. hauxwellianus</emphasis>
</taxonomicName>
(= 
<taxonomicName id="3D9B4D03E90CFF9FFF5997B47D3506CD" box="[182,305,404,428]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90CFF9FFF5997B47D3506CD" box="[182,305,404,428]" italics="true" pageId="7" pageNumber="66">C. spilurus</emphasis>
</taxonomicName>
), according to the key presented by Eigenmann. In fact, none of the herein examined morphotypes of 
<taxonomicName id="3D9B4D03E90CFF9FFF7897997D2C06B1" box="[151,296,440,464]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="66" phylum="Chordata" rank="genus">
<emphasis id="C8EFEA92E90CFF9FFF7897997D2C06B1" box="[151,296,440,464]" italics="true" pageId="7" pageNumber="66">Ctenobrycon</emphasis>
</taxonomicName>
presented toothless maxillaries and 41–42 lateral line scales as described for 
<taxonomicName id="3D9B4D03E90CFF9FFB519798797106B1" box="[1214,1397,440,464]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="multiradiatus">
<emphasis id="C8EFEA92E90CFF9FFB519798797106B1" box="[1214,1397,440,464]" italics="true" pageId="7" pageNumber="66">T. multiradiatus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="9E0A4B71E90CFF9FFF7897FD7D7E0695" author="Steindachner" box="[151,378,476,500]" pageId="7" pageNumber="66" refString="Steindachner, F. (1876) Ichthyologische Beitrage (V). Sitzungsberichte Akademie der Wissenschaften inWien, 74, 49 - 240, pls. 1 - 15." type="journal article" year="1876">Steindachner (1876)</bibRefCitation>
and, thus, there are not enough elements to assure that 
<taxonomicName id="3D9B4D03E90CFF9FFC0C97FC789E0695" box="[995,1178,476,500]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="multiradiatus">
<emphasis id="C8EFEA92E90CFF9FFC0C97FC789E0695" box="[995,1178,476,500]" italics="true" pageId="7" pageNumber="66">C. multiradiatus</emphasis>
</taxonomicName>
is a junior synonym of 
<taxonomicName id="3D9B4D03E90CFF9FFF7894217D170579" box="[151,275,512,536]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90CFF9FFF7894217D170579" box="[151,275,512,536]" italics="true" pageId="7" pageNumber="66">C. spilurus</emphasis>
</taxonomicName>
or even if it is a species of 
<taxonomicName id="3D9B4D03E90CFF9FFDBF94217EE50579" box="[592,737,512,536]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="66" phylum="Chordata" rank="genus">
<emphasis id="C8EFEA92E90CFF9FFDBF94217EE50579" box="[592,737,512,536]" italics="true" pageId="7" pageNumber="66">Ctenobrycon</emphasis>
</taxonomicName>
, if we consider that no mention was made on the presence of spinoid scales in 
<taxonomicName id="3D9B4D03E90CFF9FFEB894047E0B055D" box="[343,527,548,572]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="multiradiatus">
<emphasis id="C8EFEA92E90CFF9FFEB894047E0B055D" box="[343,527,548,572]" italics="true" pageId="7" pageNumber="66">C. multiradiatus</emphasis>
</taxonomicName>
, be it in its original description or elsewhere.
</paragraph>
</subSubSection>
<subSubSection id="B281650BE90CFF9FFF2894667D910339" pageId="7" pageNumber="66" type="materials_examined">
<paragraph id="FA243680E90CFF9FFF2894667D910339" blockId="7.[151,1437,151,1113]" pageId="7" pageNumber="66">
<emphasis id="C8EFEA92E90CFF9FFF2894667DDC0501" bold="true" box="[199,472,583,608]" pageId="7" pageNumber="66">Comparative material.</emphasis>
<taxonomicName id="3D9B4D03E90CFF9FFE3094697EBC0501" box="[479,696,584,608]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="alleni">
<emphasis id="C8EFEA92E90CFF9FFE3094697EBC0501" box="[479,696,584,608]" italics="true" pageId="7" pageNumber="66">Ctenobrycon alleni</emphasis>
</taxonomicName>
: 
<collectionCode id="9C8AAE45E90CFF9FFD2894697F00053E" box="[711,772,584,607]" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="7" pageNumber="66">LIRP</collectionCode>
3786 (n=1), 
<collectingCountry id="828C7610E90CFF9FFC7794667FDA053E" box="[920,990,583,607]" name="Brazil" pageId="7" pageNumber="66">Brasil</collectingCountry>
, Mato Grosso do Sul, Porto Manga, rio 
<collectingCountry id="828C7610E90CFF9FFF78944D7CF905E5" box="[151,253,620,644]" name="Paraguay" pageId="7" pageNumber="66">Paraguai</collectingCountry>
; 
<collectionCode id="9C8AAE45E90CFF9FFEE7944D7D6105E5" box="[264,357,620,644]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="66">MZUSP</collectionCode>
54023 (34) (1 C&amp;S), 
<collectingCountry id="828C7610E90CFF9FFD8F944D7EC105E5" box="[608,709,620,644]" name="Paraguay" pageId="7" pageNumber="66">Paraguai</collectingCountry>
, Concepcion, Puerto Itacua, rio 
<collectingCountry id="828C7610E90CFF9FFBDF944D789205E5" box="[1072,1174,620,644]" name="Paraguay" pageId="7" pageNumber="66">Paraguai</collectingCountry>
; 
<taxonomicName id="3D9B4D03E90CFF9FFB4D944D799005E5" box="[1186,1428,620,644]" class="Actinopterygii" family="Characidae" genus="Ctenobrycon" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90CFF9FFB4D944D799005E5" box="[1186,1428,620,644]" italics="true" pageId="7" pageNumber="66">Ctenobrycon spilurus</emphasis>
</taxonomicName>
: 
<collectingCountry id="828C7610E90CFF9FFF7894AE7CE405C6" box="[151,224,655,679]" name="Brazil" pageId="7" pageNumber="66">Brazil</collectingCountry>
: 
<collectionCode id="9C8AAE45E90CFF9FFF0294AE7D4E05C6" box="[237,330,655,679]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="66">MZUSP</collectionCode>
5156 (28), Roraima, Surumu, rio Surumu; 
<collectionCode id="9C8AAE45E90CFF9FFCAA94AE7FA605C6" box="[837,930,655,679]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="66">MZUSP</collectionCode>
5601 (22), Pará, Oriximiná, rio Trombetas, Lago Parú; Amazonas: 
<collectionCode id="9C8AAE45E90CFF9FFE4794947DE105AD" box="[424,485,693,716]" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="7" pageNumber="66">LIRP</collectionCode>
4999 (21) (3 C&amp;S), Janauari, Lago Terra Preta; 
<collectionCode id="9C8AAE45E90CFF9FFBCC9494786405AD" box="[1059,1120,693,716]" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="7" pageNumber="66">LIRP</collectionCode>
4965 (10), Janauaca, Lago Castanho; 
<collectionCode id="9C8AAE45E90CFF9FFEFD94F97D4A058E" box="[274,334,728,751]" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="7" pageNumber="66">LIRP</collectionCode>
4985 (2), Camaleão, 
<collectingCountry id="828C7610E90CFF9FFDBF94F67E78058E" box="[592,636,727,751]" name="Iceland" pageId="7" pageNumber="66">Ilha</collectingCountry>
de Marchantaria, rio Amazonas; 
<collectionCode id="9C8AAE45E90CFF9FFBEA94F67866058E" box="[1029,1122,727,751]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="66">MZUSP</collectionCode>
27765 (8), Tefé, baixo rio Japurá; 
<collectionCode id="9C8AAE45E90CFF9FFF1F94DD7D490475" box="[240,333,764,788]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="66">MZUSP</collectionCode>
54495 (8), 
<collectingCountry id="828C7610E90CFF9FFE3B94DD7E310475" box="[468,565,764,788]" name="Ecuador" pageId="7" pageNumber="66">Equador</collectingCountry>
, Napo, rio Yasuní; 
<collectingCountry id="828C7610E90CFF9FFCFE94DD7F5C0475" box="[785,856,764,788]" name="Brazil" pageId="7" pageNumber="66">Brazil</collectingCountry>
, Acre, Cruzeiro do Sul, rio Moa: LBP 4047 (4), (1 C&amp;S) LBP 4151 (2), rio Japiim: LBP 4095 (20); 
<collectingCountry id="828C7610E90CFF9FFD0195007F200459" box="[750,804,801,824]" name="Peru" pageId="7" pageNumber="66">Peru</collectingCountry>
, Ucayali, Província de Coronel Portilho, rio Ucayali: 
<collectionCode id="9C8AAE45E90CFF9FFF7895657CF0043D" box="[151,244,836,860]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="66">MZUSP</collectionCode>
25996 (1); 
<collectionCode id="9C8AAE45E90CFF9FFE6795657DE1043D" box="[392,485,836,860]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="66">MZUSP</collectionCode>
26242 (8); 
<collectingCountry id="828C7610E90CFF9FFD9595657EEC043D" box="[634,744,836,860]" name="Suriname" pageId="7" pageNumber="66">Suriname</collectingCountry>
, Paramaribo: 
<collectionCode id="9C8AAE45E90CFF9FFC6195657FD1043D" LSID="urn:lsid:biocol.org:col:34784" box="[910,981,836,860]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="7" pageNumber="66">ANSP</collectionCode>
137053 (3); 
<collectingCountry id="828C7610E90CFF9FFB8C956578E5043D" box="[1123,1249,836,860]" name="Venezuela" pageId="7" pageNumber="66">Venezuela</collectingCountry>
, Guárico: 
<collectionCode id="9C8AAE45E90CFF9FFA8E95657998043D" box="[1377,1436,836,860]" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="7" pageNumber="66">MCP</collectionCode>
15138 (5), Camaguán, río Portuguesa; 
<collectionCode id="9C8AAE45E90CFF9FFD8795467EC1041E" box="[616,709,871,895]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="66">MZUSP</collectionCode>
74698 (3), San Fernando, río Guárico; Bolívar, Caicara Del Orinoco: LBP 2232 (11), (2 C&amp;S), río Orinoco; LBP 2222 (2), Laguna de Castilleros; 
<taxonomicName id="3D9B4D03E90CFF9FFB9595AD799004C5" box="[1146,1428,908,932]" class="Actinopterygii" family="Characidae" genus="Psellogrammus" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="kennedyi">
<emphasis id="C8EFEA92E90CFF9FFB9595AD799004C5" box="[1146,1428,908,932]" italics="true" pageId="7" pageNumber="66">Psellogrammus kennedyi</emphasis>
</taxonomicName>
: LBP 3220 (24) (3 C&amp;S), 
<collectingCountry id="828C7610E90CFF9FFE56958E7DFB04A6" box="[441,511,943,967]" name="Brazil" pageId="7" pageNumber="66">Brasil</collectingCountry>
, Mato Grosso, Nobres, rio Cuiabazinho, Lagoa Marginal; 
<taxonomicName id="3D9B4D03E90CFF9FFB7B9590799004A9" box="[1172,1428,944,968]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="alleni">
<emphasis id="C8EFEA92E90CFF9FFB7B9590799004A9" box="[1172,1428,944,968]" italics="true" pageId="7" pageNumber="66">Tetragonopterus alleni</emphasis>
</taxonomicName>
: 
<collectionCode id="9C8AAE45E90CFF9FFF7895F47CEC048D" LSID="urn:lsid:biocol.org:col:34795" box="[151,232,981,1004]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="66">FMNH</collectionCode>
52634, 
<typeStatus id="25208822E90CFF9FFEA095F47DB6048C" box="[335,434,981,1005]" pageId="7" pageNumber="66" type="paratype">paratype</typeStatus>
, 
<collectingCountry id="828C7610E90CFF9FFE2B95F57E0E048D" box="[452,522,980,1004]" name="Brazil" pageId="7" pageNumber="66">Brasil</collectingCountry>
, Mato Grosso, Corumbá. 
<taxonomicName id="3D9B4D03E90CFF9FFCAB95F47883048D" box="[836,1159,980,1004]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="gibbicervix">
<emphasis id="C8EFEA92E90CFF9FFCAB95F47883048D" box="[836,1159,980,1004]" italics="true" pageId="7" pageNumber="66">Tetragonopterus gibbicervix</emphasis>
</taxonomicName>
: 
<collectionCode id="9C8AAE45E90CFF9FFB7395F578E2048D" box="[1180,1254,980,1004]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="66">NMW</collectionCode>
57516, 
<typeStatus id="25208822E90CFF9FFAD295F47993048C" box="[1341,1431,981,1005]" pageId="7" pageNumber="66" type="syntype">syntype</typeStatus>
, 
<collectingCountry id="828C7610E90CFF9FFF7895D67CDA036E" box="[151,222,1015,1039]" name="Brazil" pageId="7" pageNumber="66">Brazil</collectingCountry>
, Amazonas, Teffé (photo), 
<collectionCode id="9C8AAE45E90CFF9FFDE295D87E660371" LSID="urn:lsid:biocol.org:col:34988" box="[525,610,1017,1040]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="66">MNHN</collectionCode>
1909-182, 
<typeStatus id="25208822E90CFF9FFD0D95D87F470370" box="[738,835,1017,1041]" pageId="7" pageNumber="66" type="paratype">paratype</typeStatus>
(photo); 
<collectionCode id="9C8AAE45E90CFF9FFC4595D878040371" LSID="urn:lsid:biocol.org:col:34988" box="[938,1024,1017,1040]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="66">MNHN</collectionCode>
1909-320/321, 2 
<typeStatus id="25208822E90CFF9FFB2A95D879340370" box="[1221,1328,1017,1041]" pageId="7" pageNumber="66" type="paratype">paratypes</typeStatus>
(photos); 
<taxonomicName id="3D9B4D03E90CFF9FFF78923C7DFF0355" box="[151,507,1052,1076]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="hauxwellianus">
<emphasis id="C8EFEA92E90CFF9FFF78923C7DFF0355" box="[151,507,1052,1076]" italics="true" pageId="7" pageNumber="66">Tetragonopterus hauxwellianus</emphasis>
</taxonomicName>
: 
<collectionCode id="9C8AAE45E90CFF9FFDE2923D7E510355" LSID="urn:lsid:biocol.org:col:34784" box="[525,597,1052,1076]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="7" pageNumber="66">ANSP</collectionCode>
8138-8142, 5 
<typeStatus id="25208822E90CFF9FFCEC923C7F750354" box="[771,881,1053,1077]" pageId="7" pageNumber="66" type="paratype">paratypes</typeStatus>
, Pebas, 
<collectingCountry id="828C7610E90CFF9FFC3C923C780E0355" box="[979,1034,1053,1076]" name="Peru" pageId="7" pageNumber="66">Peru</collectingCountry>
. 
<taxonomicName id="3D9B4D03E90CFF9FFBF8923C79300355" box="[1047,1332,1052,1076]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="spilurus">
<emphasis id="C8EFEA92E90CFF9FFBF8923C79300355" box="[1047,1332,1052,1076]" italics="true" pageId="7" pageNumber="66">Tetragonopterus spilurus</emphasis>
</taxonomicName>
: 
<collectionCode id="9C8AAE45E90CFF9FFAA9923C799F0355" LSID="urn:lsid:biocol.org:col:34988" box="[1350,1435,1053,1076]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="66">MNHN</collectionCode>
5341, 
<typeStatus id="25208822E90CFF9FFF3292607D300338" box="[221,308,1089,1113]" pageId="7" pageNumber="66" type="syntype">syntype</typeStatus>
(photo).
</paragraph>
</subSubSection>
</treatment>
</document>