<documentid="325F920E7FEE1B8AE82CEA5E4D318FA2"ID-DOI="10.5852/ejt.2017.359"ID-GBIF-Dataset="87920495-c1bf-4bdb-b153-50e709c1d6c2"ID-ISSN="2118-9773"ID-Zenodo-Dep="3855694"IM.bibliography_approvedBy="juliana"IM.illustrations_approvedBy="juliana"IM.materialsCitations_approvedBy="juliana"IM.metadata_approvedBy="juliana"IM.tables_approvedBy="juliana"IM.taxonomicNames_approvedBy="juliana"IM.treatmentCitations_approvedBy="juliana"IM.treatments_approvedBy="juliana"checkinTime="1590500868344"checkinUser="carolina"docAuthor="d’Acoz, Cédric d’Udekem & Verheye, Marie L."docDate="2017"docId="4A5A879BFFA36857FE58FB1ECA26FE3C"docLanguage="en"docName="ejt-2017-359.pdf.imf"docOrigin="European Journal of Taxonomy 359"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Epimeria (Drakepimeria) havermansiana d’Acoz & Verheye 2017, subgen. et sp. nov."docType="treatment"docVersion="16"lastPageNumber="40"masterDocId="B663FFE3FF86687FFFF7FFE8CF17FF89"masterDocTitle="Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea)"masterLastPageNumber="553"masterPageNumber="1"pageNumber="38"updateTime="1736878097187"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-3.0"zenodo-license-figures="CC-BY-3.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="C439E269A0BC97537F05B2CA0191932C">Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea)</mods:title>
<mods:affiliationid="A340E537403B5971A91DA6CA43307BA5">Royal Belgian Institute of Natural Sciences, Service Heritage, Rue Vautier 29, B- 1000 Brussels, Belgium.</mods:affiliation>
<mods:affiliationid="663BAAAB061E6E374B0479C5B1B687BB">Royal Belgian Institute of Natural Sciences, Operational direction Taxonomy and Phylogeny, Rue Vautier 29, B- 1000 Brussels, Belgium.</mods:affiliation>
<taxonomicNameid="05F34D0EFFA3685AFE58FB1ECCB3FA99"ID-CoL="6G662"authority="Verheye, 2017"authorityName="d’Acoz & Verheye"authorityYear="2017"box="[431,932,1269,1296]"class="Malacostraca"family="Epimeriidae"genus="Epimeria"higherTaxonomySource="GBIF"kingdom="Animalia"order="Amphipoda"pageId="37"pageNumber="38"phylum="Arthropoda"rank="species"species="havermansiana"status="subgen. et sp. nov."subGenus="Drakepimeria">
<taxonomicNameLabelid="EBB457E4FFA3685AFC5BFB1DCB92FA99"box="[940,1157,1269,1296]"pageId="37"pageNumber="38"rank="species"typeStatus="paratype">subgen. et sp. nov.</taxonomicNameLabel>
<bibRefCitationid="A6624B7CFFA3685AFD28FA6ACCC4FA15"author="Verheye"box="[735,979,1410,1437]"etAl="et al."firstAuthor="Verheye"pageId="37"pageNumber="38"pagination="925 - 945"refId="ref146163"refString="Verheye M., Backeljau T. & d'Udekem d'Acoz C. 2016 a. Looking beneath the tip of the iceberg: diversification of the genus Epimeria on the Antarctic shelf (Crustacea, Amphipoda). In: Gutt J., David B. & Isla E. (eds) High environmental variability and steep biological gradients in the waters off the northern Antarctic Peninsula. Polar Biology 39 (5): 925 - 945, online supplementary material https: // doi. org / 10.1007 / s 00300 - 016 - 1910 - 5"type="journal article"typeStatus="paratype"year="2016"yearSuffix="a">
The species is dedicated to Charlotte Havermans (formerly RBINS, currently Alfred-Wegener-Institute Helmholtz-Zentrum für Polar- und Meeresforschung), who collected the
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, INV. 132700);
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<paragraphid="C24C368DFFA06859FF4AF972CD74F95E"blockId="38.[189,1399,1689,1751]"pageId="38"pageNumber="39">ROSTRUM. Long, reaching about tip of article 1 of peduncle of antenna 1 (teeth excluded), moderately curved, sharp-tipped in lateral view.</paragraph>
<paragraphid="C24C368DFFA06859FF4AF8E8CDE8F893"blockId="38.[189,767,1791,1818]"box="[189,767,1791,1818]"pageId="38"pageNumber="39">EYE. Very large, broadly elliptic (nearly circular).</paragraph>
<paragraphid="C24C368DFFA06858FF4AF8AACDD5FE19"blockId="38.[189,1399,1857,2027]"lastBlockId="39.[189,1399,267,400]"lastPageId="39"lastPageNumber="40"pageId="38"pageNumber="39">PEREION–PLEOSOME TOOTH PATTERN. Pereionite 1 with indistinct posterior bump, with pair of low noncarinate dorsolateral swellings; pereionite 2 slightly narrower than pereionite 1, with small but distinct blunt mid-dorsal tooth and pair of low dorsolateral swellings; pereionite 3 with medium-sized blunttipped mid-dorsal tooth and pair of well-developed dorsolateral blunt teeth; pereionite 4 to pereionite 7 with large, broad, regularly curved, acute-tipped mid-dorsal tooth (dorsal length of teeth slightly and gradually increasing posteriorly) and pair of well developed conical dorsolateral teeth (pleonites 1–2 without second pair of dorsolateral teeth); pleonite 3 with dorsal sharp carina with weak median notch and produced posteriorly into an a broad acute and sharp triangular tooth, and pair of fairly large dorsolateral non-carinate acute-tipped teeth.</paragraph>
<paragraphid="C24C368DFFA16858FF4AFE5FCDE4FE58"blockId="39.[189,755,439,465]"box="[189,755,439,465]"pageId="39"pageNumber="40">COXAE 1–3. Strongly carinate and distally sharp.</paragraph>
<paragraphid="C24C368DFFA16858FF4AFE11CDF9FDF4"blockId="39.[189,1398,505,638]"pageId="39"pageNumber="40">COXA 4. Anterodorsal + anteroventral borders forming a curve projecting forward (anterodorsal and anteroventral borders becoming straight respectively only at their upper and lower extremities); ventral tooth very sharp and very long; lateral carina without tooth, without distinct angularity, carina very distant from margin of coxa at its deepest point.</paragraph>
<paragraphid="C24C368DFFA16858FF4AFD4DCC0DFD6A"blockId="39.[189,1399,677,739]"pageId="39"pageNumber="40">COXA 5. With sharp and narrowly triangular carinate, lateral tooth pointing backwards (its lateral border is nearly parallel to body axis or weakly divergent).</paragraph>
<paragraphid="C24C368DFFA16858FF4AFC87CE13FC24"blockId="39.[189,1398,879,941]"pageId="39"pageNumber="40">COXA 7. With ventral and posterior border straight, converging to form a sharp acute, nearly squared angle.</paragraph>
<paragraphid="C24C368DFFA16858FF4AFC3DCB64FC67"blockId="39.[189,1139,980,1007]"box="[189,1139,980,1007]"pageId="39"pageNumber="40">EPIMERAL PLATES 1–3. Posteroventral angle produced into a very long sharp tooth.</paragraph>
<paragraphid="C24C368DFFA16858FF4AFBFFCCA1FBDA"blockId="39.[189,1398,1046,1107]"pageId="39"pageNumber="40">UROSOME TOOTH PATTERN. Urosomite 1 with strong triangular process pointing upwards; urosomite 2 with pair of small sharp posterior dorsolateral teeth pointing upwards.</paragraph>
<paragraphid="C24C368DFFA16858FF4AFB94CC93FB1C"blockId="39.[189,900,1147,1174]"box="[189,900,1147,1174]"pageId="39"pageNumber="40">TELSON. Cleft on 0.25; tips of lobes sharp, broadly V-shaped.</paragraph>
<paragraphid="C24C368DFFA16858FF4AFB55CFE8FAEC"blockId="39.[189,1398,1213,1381]"pageId="39"pageNumber="40">PEDUNCLE OF ANTENNA 1. Article 1 with long lateral and medial teeth reaching beyond mid and nearly mid of article 2 (teeth excluded) and long ventral tooth nearly reaching tip of article 2 (teeth excluded); article 2 with very long lateral and medial teeth overreaching tip of article 3, with ventral tooth slightly overreaching tip of article 3 (tooth excluded); article 3 with long ventral tooth, about as long as article itself.</paragraph>
<paragraphid="C24C368DFFA16858FF4AFA64CE0CFA43"blockId="39.[189,1399,1420,1482]"pageId="39"pageNumber="40">GNATHOPODS 1–2. Carpus and propodus of normal slenderness; propodus not narrowing distally, palm distinct.</paragraph>
<paragraphid="C24C368DFFA16858FF4AFA1ACEBFF9FF"blockId="39.[189,1398,1521,1654]"pageId="39"pageNumber="40">PEREIOPODS 5–7. Merus, carpus and propodus fairly slender; basis of pereiopods 5–6 of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth strong; basis of pereiopod 7 very broad with posterodistal tooth angulate, followed more proximally by small concavity, directed posteriorly.</paragraph>
<taxonomicNameid="05F34D0EFFA16858FF4AF846CEF3F84E"authority="Verheye, 2017"authorityName="d’Acoz & Verheye"authorityYear="2017"box="[189,484,1965,1991]"class="Malacostraca"family="Epimeriidae"genus="Epimeria"higherTaxonomySource="GBIF"kingdom="Animalia"order="Amphipoda"pageId="39"pageNumber="40"phylum="Arthropoda"rank="species"species="havermansiana"status="sp. nov.">
<taxonomicNameid="05F34D0EFFA16858FC25F846CB43F84E"authority="Verheye, 2017"authorityName="d’Acoz & Verheye"authorityYear="2017"box="[978,1108,1965,1991]"class="Malacostraca"family="Epimeriidae"genus="Epimeria"higherTaxonomySource="GBIF"kingdom="Animalia"order="Amphipoda"pageId="39"pageNumber="40"phylum="Arthropoda"rank="species"species="pandora"status="sp. nov.">
<bibRefCitationid="A6624B7CFFA16858FB88F839CA65F862"author="Verheye"box="[1151,1394,2001,2027]"etAl="et al."firstAuthor="Verheye"pageId="39"pageNumber="40"pagination="925 - 945"refId="ref146163"refString="Verheye M., Backeljau T. & d'Udekem d'Acoz C. 2016 a. Looking beneath the tip of the iceberg: diversification of the genus Epimeria on the Antarctic shelf (Crustacea, Amphipoda). In: Gutt J., David B. & Isla E. (eds) High environmental variability and steep biological gradients in the waters off the northern Antarctic Peninsula. Polar Biology 39 (5): 925 - 945, online supplementary material https: // doi. org / 10.1007 / s 00300 - 016 - 1910 - 5"type="journal article"typeStatus="paratype"year="2016"yearSuffix="a">
<figureCitationid="5AC82A08FFAE6857FE9FFEE3CED8FEAC"box="[360,463,267,293]"captionStart="Fig"captionStartId="551.[189,232,1824,1850]"captionTargetBox="[343,1219,266,1752]"captionTargetId="graphics@551.[343,1120,270,1752]"captionTargetPageId="551"captionText="Fig. 342. Phylogenetic tree obtained by Bayesian analysis of the concatenated COI and 28S sequences, updated from Verheye et al. (2016). Bayesian posterior probabilities and bootstrap values (from the Maximum Likelihood analysis) are indicated above the nodes of interest. Bootstrap values inferior to 50 are not indicated. Besides the species names are indicated the MOTUs codes used in Verheye et al. (2016). The Genbank accession number of the outgroup species: Gnathiphimedia sexdentata (Schellenberg, 1926) are KU870835 (COI) and KU759609 (28S)."figureDoi="http://doi.org/10.5281/zenodo.3856377"httpUri="https://zenodo.org/record/3856377/files/figure.png"pageId="40"pageNumber="41"typeStatus="paratype">Fig. 342</figureCitation>
<taxonomicNameid="05F34D0EFFAE6857FD51FEE4CC6BFEAC"authority="Verheye, 2017"authorityName="d’Acoz & Verheye"authorityYear="2017"box="[678,892,267,293]"class="Malacostraca"family="Epimeriidae"genus="Epimeria"higherTaxonomySource="GBIF"kingdom="Animalia"order="Amphipoda"pageId="40"pageNumber="41"phylum="Arthropoda"rank="species"species="havermansiana"status="sp. nov.">
may have a trace of posterior middorsal bump, which is absent in
<taxonomicNameid="05F34D0EFFAE6857FDB1FED8CDDBFEC0"authority="Verheye, 2017"authorityName="d’Acoz & Verheye"authorityYear="2017"box="[582,716,303,329]"class="Malacostraca"family="Epimeriidae"genus="Epimeria"higherTaxonomySource="GBIF"kingdom="Animalia"order="Amphipoda"pageId="40"pageNumber="41"phylum="Arthropoda"rank="species"species="pandora"status="sp. nov.">
<taxonomicNameid="05F34D0EFFAE6857FBFFFED8CBF7FEC0"authority="Verheye, 2017"authorityName="d’Acoz & Verheye"authorityYear="2017"box="[1032,1248,303,329]"class="Malacostraca"family="Epimeriidae"genus="Epimeria"higherTaxonomySource="GBIF"kingdom="Animalia"order="Amphipoda"pageId="40"pageNumber="41"phylum="Arthropoda"rank="species"species="havermansiana"status="sp. nov.">
<taxonomicNameid="05F34D0EFFAE6857FE2EFEBCCD4AFEE4"authority="Verheye, 2017"authorityName="d’Acoz & Verheye"authorityYear="2017"box="[473,605,339,365]"class="Malacostraca"family="Epimeriidae"genus="Epimeria"higherTaxonomySource="GBIF"kingdom="Animalia"order="Amphipoda"pageId="40"pageNumber="41"phylum="Arthropoda"rank="species"species="pandora"status="sp. nov.">
<taxonomicNameid="05F34D0EFFAE6857FBFEFEBCCBC8FEE4"authority="Verheye, 2017"authorityName="d’Acoz & Verheye"authorityYear="2017"box="[1033,1247,339,365]"class="Malacostraca"family="Epimeriidae"genus="Epimeria"higherTaxonomySource="GBIF"kingdom="Animalia"order="Amphipoda"pageId="40"pageNumber="41"phylum="Arthropoda"rank="species"species="havermansiana"status="sp. nov.">
, the dorsolateral teeth of the pereion pleosome and urosomite 2 are larger than in
<taxonomicNameid="05F34D0EFFAE6857FBCDFE90CBABFE18"authority="Verheye, 2017"authorityName="d’Acoz & Verheye"authorityYear="2017"box="[1082,1212,375,401]"class="Malacostraca"family="Epimeriidae"genus="Epimeria"higherTaxonomySource="GBIF"kingdom="Animalia"order="Amphipoda"pageId="40"pageNumber="41"phylum="Arthropoda"rank="species"species="pandora"status="sp. nov.">
<taxonomicNameid="05F34D0EFFAE6857FDB3FE73CDDFFE3C"authority="Verheye, 2017"authorityName="d’Acoz & Verheye"authorityYear="2017"box="[580,712,411,437]"class="Malacostraca"family="Epimeriidae"genus="Epimeria"higherTaxonomySource="GBIF"kingdom="Animalia"order="Amphipoda"pageId="40"pageNumber="41"phylum="Arthropoda"rank="species"species="pandora"status="sp. nov.">