367 lines
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367 lines
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<document id="B8F9D812F1AE011057C9C0A0F2B533C5" ID-DOI="10.11646/zootaxa.5497.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="13618054" ID-ZooBank="8F0CBBEF-DA94-4A6D-B7A0-3250F31D3A87" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725025523655" checkinUser="plazi" docAuthor="Jałoszyński, Paweł & Szawaryn, Karol" docDate="2024" docId="DD22F04CCE42A519F08D41A2FB17FE8F" docLanguage="en" docName="zootaxa.5497.1.7.pdf" docOrigin="Zootaxa 5497 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Loeblitoides latus Jaloszynski & Szawaryn 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="129" masterDocId="211B8834CE41A51EF01A4135FFDCFFEA" masterDocTitle="Characters of a new species of the Cretaceous genus † Loeblitoides affirm affinities to the extant ‘ Syndicus group’ of Stenichnini (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="132" masterPageNumber="123" pageNumber="126" updateTime="1725026637147" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="414A0E87AAFEF25859D34BD15DB0BC79">Characters of a new species of the Cretaceous genus † Loeblitoides affirm affinities to the extant ‘ Syndicus group’ of Stenichnini (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:name id="D89A132839515E61FAE8165C0980F08C" type="personal">
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<mods:roleTerm id="2994BBE6B7C8AE32EA3CAEEDA63F9835">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5ACB3F40E31D2B4C9DEA1A369F51FFC5">Jałoszyński, Paweł</mods:namePart>
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<mods:affiliation id="12BCA87418390099A0A503565AB5EB5D">Museum of Natural History, University of Wrocław, Sienkiewicza 21, 50 - 335 Wrocław, Poland.</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="0EE084866763B58D8FBFAFED3912BF83">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0ECA9152287B1BC6948BB36F3443494E">Szawaryn, Karol</mods:namePart>
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<mods:affiliation id="56782E2A133A90EE8D7FFCB34BBFEEB7">Museum and Institute of Zoology, Polish Academy of Sciences, Twarda 51 / 55, 00 - 818 Warszawa, Poland.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="A27B2A18F00D2786404FB01C62A2E9E5">text</mods:typeOfResource>
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<mods:title id="72B089C30B8B36B24A0E40CDCA2B0975">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F9BCF52F1409565749C42A2415A2C279">
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<mods:date id="A221D4E49BAE16E9AAFAAEFFE4D1A642">2024</mods:date>
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<mods:number id="5412849FF01EE9590E24E3274AE5EBB0">2024-08-22</mods:number>
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<mods:number id="7AEF70D9872743F12ED7FAC18686D8FB">5497</mods:number>
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<mods:number id="0C409747EE220A39DC9F50EF06C9A8B7">1</mods:number>
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<mods:url id="55B9929BE4C7226EA4DE1E97BD9EA332">http://dx.doi.org/10.11646/zootaxa.5497.1.7</mods:url>
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<mods:classification id="5EB5B92589F479C49EB6BF4C99A310C7">journal article</mods:classification>
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<mods:identifier id="FD8EA7F3FD932A7D21BDB109BA3A94A6" type="DOI">10.11646/zootaxa.5497.1.7</mods:identifier>
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<mods:identifier id="FBC8EE2573CA39FD4E208D40547F98BD" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="DD22F04CCE42A519F08D41A2FB17FE8F" LSID="urn:lsid:plazi:treatment:DD22F04CCE42A519F08D41A2FB17FE8F" httpUri="http://treatment.plazi.org/id/DD22F04CCE42A519F08D41A2FB17FE8F" lastPageId="7" lastPageNumber="129" pageId="3" pageNumber="126">
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<subSubSection id="1D9112D1CE42A51DF08D41A2FCC5FF5B" box="[151,793,150,178]" pageId="3" pageNumber="126" type="nomenclature">
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<paragraph id="5534415ACE42A51DF08D41A2FCC5FF5B" blockId="3.[151,793,150,213]" box="[151,793,150,178]" pageId="3" pageNumber="126">
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<heading id="0E7CF636CE42A51DF08D41A2FCC5FF5B" bold="true" box="[151,793,150,178]" fontSize="11" level="1" pageId="3" pageNumber="126" reason="1">
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<emphasis id="67FF9D48CE42A51DF08D41A2FCC5FF5B" bold="true" box="[151,793,150,178]" pageId="3" pageNumber="126">
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†
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<taxonomicName id="928B3AD9CE42A51DF0BC41A2FD68FF5B" authority="Jaloszynski & Szawaryn" authorityName="Jaloszynski & Szawaryn" authorityYear="2024" box="[166,692,150,178]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
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<emphasis id="67FF9D48CE42A51DF0BC41A2FEA7FF58" bold="true" box="[166,379,151,178]" italics="true" pageId="3" pageNumber="126">Loeblitoides latus</emphasis>
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Jałoszyński & Szawaryn
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</taxonomicName>
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<taxonomicNameLabel id="7CCC2033CE42A51DF2A641A2FCC5FF5B" box="[700,793,151,177]" pageId="3" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1D9112D1CE42A51DF08D418EFEC5FF3F" box="[151,281,187,213]" pageId="3" pageNumber="126" type="description">
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<paragraph id="5534415ACE42A51DF08D418EFEC5FF3F" blockId="3.[151,793,150,213]" box="[151,281,187,213]" pageId="3" pageNumber="126">
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(
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<figureCitation id="CDB05DDFCE42A51DF085418EFECCFF3F" box="[159,272,187,213]" captionStart-0="FIGURES 1–7" captionStart-1="FIGURES 8–11" captionStart-2="FIGURES 12–13" captionStart-3="FIGURES 14–19" captionStartId-0="3.[151,264,1928,1952]" captionStartId-1="4.[151,264,1903,1927]" captionStartId-2="5.[151,264,1279,1303]" captionStartId-3="6.[151,264,1628,1652]" captionTargetBox-0="[151,1435,453,1905]" captionTargetBox-1="[158,1430,275,1880]" captionTargetBox-2="[151,1435,181,1255]" captionTargetBox-3="[151,1436,777,1605]" captionTargetId-0="figure-100@3.[151,1435,453,1905]" captionTargetId-1="figure-54@4.[158,1430,273,1880]" captionTargetId-2="figure-17@5.[151,1435,181,1255]" captionTargetId-3="figure-226@6.[151,1436,777,1605]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 1–7. †Loeblitoides latus sp. nov., holotype; light micrographs. Lateral habitus (1), pronotum and elytral base in dorsal view (2), head in dorsal view (3), anterior body part in ventral view (4), distal region of maxillary palp (5), mesotarsus (6), and mesotarsomere 5 (7)." captionText-1="FIGURES 8–11. †Loeblitoides latus sp. nov., holotype (8–10), and Syndicus echinatus Jałoszyński (11). Sketch of habitus in lateral view (8); μCT reconstructions of habitus in dorsal (9) and ventral (10) views; left elytral articulating lobe in dorsal view, scanning electron micrography (11). Abbreviations: a3‒11, antennomere 3‒11; bef, basal elytral fovea; ce, compound eye; fcg, frontoclypeal groove; pd, pedicel; pn, pronotum; sc, scape; scs, scutellar shield." captionText-2="FIGURES 12–13. †Loeblitoides latus sp. nov., holotype; μCT reconstructions of habitus in right (12) and left (13) lateral views." captionText-3="FIGURES 14–19. †Loeblitoides latus sp. nov., holotype, μCT reconstructions. Body in anterior (14), anteroventral (15), posterolateral (16), anterodorsal (17), anteroventral (18), and posteroventral (19) views. Abbreviations: ce, compound eye; cl, clypeus; cx1, procoxa; cx2, mesocoxa; fcg, frontoclypeal groove; fr, frons; hr, hypostomal ridge; tm, temple; v3, metaventrite; vt, vertex." figureDoi-0="http://doi.org/10.5281/zenodo.13618056" figureDoi-1="http://doi.org/10.5281/zenodo.13618058" figureDoi-2="http://doi.org/10.5281/zenodo.13618060" figureDoi-3="http://doi.org/10.5281/zenodo.13618062" httpUri-0="https://zenodo.org/record/13618056/files/figure.png" httpUri-1="https://zenodo.org/record/13618058/files/figure.png" httpUri-2="https://zenodo.org/record/13618060/files/figure.png" httpUri-3="https://zenodo.org/record/13618062/files/figure.png" pageId="3" pageNumber="126">Figs 1–19</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="1D9112D1CE42A51DF08D4036FF2FFE8F" pageId="3" pageNumber="126" type="materials_examined">
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<paragraph id="5534415ACE42A51DF08D4036FF2FFE8F" blockId="3.[151,1437,259,430]" pageId="3" pageNumber="126">
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<emphasis id="67FF9D48CE42A51DF08D4036FE99FEF7" bold="true" box="[151,325,259,285]" pageId="3" pageNumber="126">Type material.</emphasis>
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<materialsCitation id="E5E34B07CE42A51DF1564036FF2FFE8F" collectionCode="GPIH" collectorName="Carsten Grohn" country="Myanmar" location="Myanmar" pageId="3" pageNumber="126" specimenCode="CCHH 11930" specimenCount="2" typeStatus="holotype">
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<emphasis id="67FF9D48CE42A51DF1564036FE1EFEF7" bold="true" box="[332,450,259,285]" pageId="3" pageNumber="126">
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<typeStatus id="8A30FFF8CE42A51DF1564036FE61FEF7" box="[332,445,259,285]" pageId="3" pageNumber="126" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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(
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<emphasis id="67FF9D48CE42A51DF1C84036FDA0FEF7" bold="true" box="[466,636,259,285]" pageId="3" pageNumber="126">
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<collectingCountry id="2D9C01CACE42A51DF1C84036FDACFEF7" box="[466,624,259,285]" name="Myanmar" pageId="3" pageNumber="126">MYANMAR</collectingCountry>
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):
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</emphasis>
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sex unknown, label: “
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<location id="50541781CE42A51DF36E4036FC38FEF7" LSID="urn:lsid:plazi:treatment:DD22F04CCE42A519F08D41A2FB17FE8F:50541781CE42A51DF36E4036FC38FEF7" box="[884,996,259,285]" country="Myanmar" name="Myanmar" pageId="3" pageNumber="126">Myanmar</location>
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amber /
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<collectionCode id="339AD99FCE42A51DF45C4036FB5BFEF7" box="[1094,1159,259,285]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="3" pageNumber="126">GPIH</collectionCode>
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no. 5076 / (coll.
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<collectorName id="F87E248CCE42A51DF55D4036FF3DFEAB" pageId="3" pageNumber="126">Carsten Gröhn</collectorName>
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,
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<specimenCode id="052DE921CE42A51DF0F44012FE57FEAB" box="[238,395,295,321]" collectionCode="CCHH" pageId="3" pageNumber="126">CCHH 11930</specimenCode>
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) /
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<taxonomicName id="928B3AD9CE42A51DF1B04012FDABFEAB" authorityName="Jaloszynski" authorityYear="2019" box="[426,631,295,321]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="genus">
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<emphasis id="67FF9D48CE42A51DF1B04012FDABFEAB" bold="true" box="[426,631,295,321]" italics="true" pageId="3" pageNumber="126">LOEBLITOIDES</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="928B3AD9CE42A51DF2954012FBF7FEAB" authority="Jaloszynski & Szawaryn, 2024" authorityName="Jaloszynski & Szawaryn" authorityYear="2024" box="[655,1067,295,322]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="latus">
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<emphasis id="67FF9D48CE42A51DF2954012FD1BFEA8" bold="true" box="[655,711,295,322]" italics="true" pageId="3" pageNumber="126">latus</emphasis>
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Jałoszyński & Szawaryn, 2024
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</taxonomicName>
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/
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<emphasis id="67FF9D48CE42A51DF4594012FB2FFEAB" bold="true" box="[1091,1267,295,321]" pageId="3" pageNumber="126">
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<typeStatus id="8A30FFF8CE42A51DF4594012FB2FFEAB" box="[1091,1267,295,321]" pageId="3" pageNumber="126" type="holotype">HOLOTYPUS</typeStatus>
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</emphasis>
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” [red, printed] (
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<collectionCode id="339AD99FCE42A51DF085407EFF3BFE8F" box="[159,231,331,357]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="3" pageNumber="126">GPIH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1D9112D1CE42A51DF0DD405AFAB7FE47" pageId="3" pageNumber="126" type="diagnosis">
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<paragraph id="5534415ACE42A51DF0DD405AFAB7FE47" blockId="3.[151,1437,259,430]" pageId="3" pageNumber="126">
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<emphasis id="67FF9D48CE42A51DF0DD405AFE9EFE63" bold="true" box="[199,322,367,393]" pageId="3" pageNumber="126">Diagnosis.</emphasis>
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Body stout, with pronotal length / width clearly below 1.5 and the elytral length / width below 1.7; antennomere 3 about 3 times as long as broad in lateral view; antennomeres 8‒10 each about as long as broad.
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</paragraph>
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</subSubSection>
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<caption id="01F411D2CE42A51DF08D46BDFE1EF803" ID-DOI="http://doi.org/10.5281/zenodo.13618056" ID-Zenodo-Dep="13618056" httpUri="https://zenodo.org/record/13618056/files/figure.png" pageId="3" pageNumber="126" startId="3.[151,264,1928,1952]" targetBox="[151,1435,453,1905]" targetPageId="3" targetType="figure">
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<paragraph id="5534415ACE42A51DF08D46BDFE1EF803" blockId="3.[151,1437,1928,2025]" pageId="3" pageNumber="126">
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<emphasis id="67FF9D48CE42A51DF08D46BDFEE1F84A" bold="true" box="[151,317,1928,1952]" pageId="3" pageNumber="126">FIGURES 1–7.</emphasis>
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†
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<taxonomicName id="928B3AD9CE42A51DF14846BDFDD5F84A" authorityName="Jałoszyński & Szawaryn" authorityYear="2024" box="[338,521,1928,1952]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
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<emphasis id="67FF9D48CE42A51DF14846BDFDD5F84A" box="[338,521,1928,1952]" italics="true" pageId="3" pageNumber="126">Loeblitoides latus</emphasis>
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</taxonomicName>
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<emphasis id="67FF9D48CE42A51DF20846BCFDB9F84B" bold="true" box="[530,613,1929,1953]" pageId="3" pageNumber="126">
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<taxonomicNameLabel id="7CCC2033CE42A51DF20846BCFDB9F84B" box="[530,613,1929,1953]" pageId="3" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype; light micrographs. Lateral habitus (
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<emphasis id="67FF9D48CE42A51DF45746BDFB86F84A" bold="true" box="[1101,1114,1928,1952]" pageId="3" pageNumber="126">1</emphasis>
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), pronotum and elytral base in dorsal view (
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<emphasis id="67FF9D48CE42A51DF1074699FEF6F82E" bold="true" box="[285,298,1964,1988]" pageId="3" pageNumber="126">2</emphasis>
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), head in dorsal view (
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<emphasis id="67FF9D48CE42A51DF20D4699FDF8F82E" bold="true" box="[535,548,1964,1988]" pageId="3" pageNumber="126">3</emphasis>
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), anterior body part in ventral view (
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<emphasis id="67FF9D48CE42A51DF3BB4699FC72F82E" bold="true" box="[929,942,1964,1988]" pageId="3" pageNumber="126">4</emphasis>
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), distal region of maxillary palp (
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<emphasis id="67FF9D48CE42A51DF5134698FACAF82F" bold="true" box="[1289,1302,1965,1989]" pageId="3" pageNumber="126">5</emphasis>
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), mesotarsus (
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<emphasis id="67FF9D48CE42A51DF0BA46E5FF71F802" bold="true" box="[160,173,2000,2024]" pageId="3" pageNumber="126">6</emphasis>
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), and mesotarsomere 5 (
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<emphasis id="67FF9D48CE42A51DF1BD46E4FE68F803" bold="true" box="[423,436,2001,2025]" pageId="3" pageNumber="126">7</emphasis>
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).
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</paragraph>
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</caption>
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<subSubSection id="1D9112D1CE45A518F0DD41A3FEADFD8B" lastPageId="6" lastPageNumber="129" pageId="4" pageNumber="127" type="description">
|
|||
|
<paragraph id="5534415ACE45A51AF0DD41A3FEC0FF13" blockId="4.[151,1436,150,249]" pageId="4" pageNumber="127">
|
|||
|
<emphasis id="67FF9D48CE45A51AF0DD41A3FE85FF5A" bold="true" box="[199,345,150,176]" pageId="4" pageNumber="127">Description.</emphasis>
|
|||
|
Body (
|
|||
|
<figureCitation id="CDB05DDFCE45A51AF1AA41A2FE25FF5B" box="[432,505,151,177]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1928,1952]" captionTargetBox="[151,1435,453,1905]" captionTargetId="figure-100@3.[151,1435,453,1905]" captionTargetPageId="3" captionText="FIGURES 1–7. †Loeblitoides latus sp. nov., holotype; light micrographs. Lateral habitus (1), pronotum and elytral base in dorsal view (2), head in dorsal view (3), anterior body part in ventral view (4), distal region of maxillary palp (5), mesotarsus (6), and mesotarsomere 5 (7)." figureDoi="http://doi.org/10.5281/zenodo.13618056" httpUri="https://zenodo.org/record/13618056/files/figure.png" pageId="4" pageNumber="127">Figs 1</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="CDB05DDFCE45A51AF21041A2FD99FF5B" box="[522,581,151,177]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[158,1430,275,1880]" captionTargetId="figure-54@4.[158,1430,273,1880]" captionTargetPageId="4" captionText="FIGURES 8–11. †Loeblitoides latus sp. nov., holotype (8–10), and Syndicus echinatus Jałoszyński (11). Sketch of habitus in lateral view (8); μCT reconstructions of habitus in dorsal (9) and ventral (10) views; left elytral articulating lobe in dorsal view, scanning electron micrography (11). Abbreviations: a3‒11, antennomere 3‒11; bef, basal elytral fovea; ce, compound eye; fcg, frontoclypeal groove; pd, pedicel; pn, pronotum; sc, scape; scs, scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.13618058" httpUri="https://zenodo.org/record/13618058/files/figure.png" pageId="4" pageNumber="127">8–10</figureCitation>
|
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|
,
|
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|
<figureCitation id="CDB05DDFCE45A51AF24941A2FD47FF5B" box="[595,667,151,177]" captionStart="FIGURES 12–13" captionStartId="5.[151,264,1279,1303]" captionTargetBox="[151,1435,181,1255]" captionTargetId="figure-17@5.[151,1435,181,1255]" captionTargetPageId="5" captionText="FIGURES 12–13. †Loeblitoides latus sp. nov., holotype; μCT reconstructions of habitus in right (12) and left (13) lateral views." figureDoi="http://doi.org/10.5281/zenodo.13618060" httpUri="https://zenodo.org/record/13618060/files/figure.png" pageId="4" pageNumber="127">12–13</figureCitation>
|
|||
|
; Supplementary animation 1) moderately slender, strongly convex, dark brown, length
|
|||
|
<quantity id="9273ECBFCE45A51AF169418EFE05FF3F" box="[371,473,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" pageId="4" pageNumber="127" unit="mm" value="2.33">2.33 mm</quantity>
|
|||
|
, constriction between head and pronotum deep, between pronotum and elytra shallow but distinct.
|
|||
|
</paragraph>
|
|||
|
<caption id="01F411D2CE45A51AF08D465AFC62F819" ID-DOI="http://doi.org/10.5281/zenodo.13618058" ID-Zenodo-Dep="13618058" httpUri="https://zenodo.org/record/13618058/files/figure.png" pageId="4" pageNumber="127" startId="4.[151,264,1903,1927]" targetBox="[158,1430,275,1880]" targetPageId="4" targetType="figure">
|
|||
|
<paragraph id="5534415ACE45A51AF08D465AFC62F819" blockId="4.[151,1437,1903,2036]" pageId="4" pageNumber="127">
|
|||
|
<emphasis id="67FF9D48CE45A51AF08D465AFE9BF86D" bold="true" box="[151,327,1903,1927]" pageId="4" pageNumber="127">FIGURES 8–11.</emphasis>
|
|||
|
†
|
|||
|
<taxonomicName id="928B3AD9CE45A51AF141465AFDCCF86D" authorityName="Jałoszyński & Szawaryn" authorityYear="2024" box="[347,528,1903,1927]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
|
|||
|
<emphasis id="67FF9D48CE45A51AF141465AFDCCF86D" box="[347,528,1903,1927]" italics="true" pageId="4" pageNumber="127">Loeblitoides latus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<emphasis id="67FF9D48CE45A51AF20D465AFDB5F86D" bold="true" box="[535,617,1903,1927]" pageId="4" pageNumber="127">
|
|||
|
<taxonomicNameLabel id="7CCC2033CE45A51AF20D465AFDB5F86D" box="[535,617,1903,1927]" pageId="4" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
</emphasis>
|
|||
|
, holotype (
|
|||
|
<emphasis id="67FF9D48CE45A51AF2C5465AFCCEF86D" bold="true" box="[735,786,1903,1927]" pageId="4" pageNumber="127">8–10</emphasis>
|
|||
|
), and
|
|||
|
<taxonomicName id="928B3AD9CE45A51AF34E465AFB4EF86D" authority="Jaloszynski" authorityName="Jaloszynski" authorityYear="2004" box="[852,1170,1903,1927]" class="Insecta" family="Staphylinidae" genus="Syndicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="species" species="echinatus">
|
|||
|
<emphasis id="67FF9D48CE45A51AF34E465AFBCAF86D" box="[852,1046,1903,1927]" italics="true" pageId="4" pageNumber="127">Syndicus echinatus</emphasis>
|
|||
|
Jałoszyński
|
|||
|
</taxonomicName>
|
|||
|
(
|
|||
|
<emphasis id="67FF9D48CE45A51AF4B8465AFB66F86D" bold="true" box="[1186,1210,1903,1927]" pageId="4" pageNumber="127">11</emphasis>
|
|||
|
). Sketch of habitus in lateral view (
|
|||
|
<emphasis id="67FF9D48CE45A51AF10646A6FEF5F841" bold="true" box="[284,297,1939,1963]" pageId="4" pageNumber="127">8</emphasis>
|
|||
|
); μCT reconstructions of habitus in dorsal (
|
|||
|
<emphasis id="67FF9D48CE45A51AF2FE46A6FD2DF841" bold="true" box="[740,753,1939,1963]" pageId="4" pageNumber="127">9</emphasis>
|
|||
|
) and ventral (
|
|||
|
<emphasis id="67FF9D48CE45A51AF36546A6FC44F841" bold="true" box="[895,920,1939,1963]" pageId="4" pageNumber="127">10</emphasis>
|
|||
|
) views; left elytral articulating lobe in dorsal view, scanning electron micrography (
|
|||
|
<emphasis id="67FF9D48CE45A51AF1FA4682FE24F825" bold="true" box="[480,504,1975,1999]" pageId="4" pageNumber="127">11</emphasis>
|
|||
|
). Abbreviations: a3‒11, antennomere 3‒11; bef, basal elytral fovea; ce, compound eye; fcg, frontoclypeal groove; pd, pedicel; pn, pronotum; sc, scape; scs, scutellar shield.
|
|||
|
</paragraph>
|
|||
|
</caption>
|
|||
|
<caption id="01F411D2CE44A51BF08D45CAFF05FAD1" ID-DOI="http://doi.org/10.5281/zenodo.13618060" ID-Zenodo-Dep="13618060" httpUri="https://zenodo.org/record/13618060/files/figure.png" pageId="5" pageNumber="128" startId="5.[151,264,1279,1303]" targetBox="[151,1435,181,1255]" targetPageId="5" targetType="figure">
|
|||
|
<paragraph id="5534415ACE44A51BF08D45CAFF05FAD1" blockId="5.[151,1436,1279,1339]" pageId="5" pageNumber="128">
|
|||
|
<emphasis id="67FF9D48CE44A51BF08D45CAFE8BFAFD" bold="true" box="[151,343,1279,1303]" pageId="5" pageNumber="128">FIGURES 12–13.</emphasis>
|
|||
|
†
|
|||
|
<taxonomicName id="928B3AD9CE44A51BF17745CAFDF8FAFD" authorityName="Jałoszyński & Szawaryn" authorityYear="2024" box="[365,548,1279,1303]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="128" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
|
|||
|
<emphasis id="67FF9D48CE44A51BF17745CAFDF8FAFD" box="[365,548,1279,1303]" italics="true" pageId="5" pageNumber="128">Loeblitoides latus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<emphasis id="67FF9D48CE44A51BF23445CAFD5EFAFD" bold="true" box="[558,642,1279,1303]" pageId="5" pageNumber="128">
|
|||
|
<taxonomicNameLabel id="7CCC2033CE44A51BF23445CAFD5EFAFD" box="[558,642,1279,1303]" pageId="5" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
</emphasis>
|
|||
|
, holotype; μCT reconstructions of habitus in right (
|
|||
|
<emphasis id="67FF9D48CE44A51BF4BF45CAFB62FAFD" bold="true" box="[1189,1214,1279,1303]" pageId="5" pageNumber="128">12</emphasis>
|
|||
|
) and left (
|
|||
|
<emphasis id="67FF9D48CE44A51BF52845CAFA97FAFD" bold="true" box="[1330,1355,1279,1303]" pageId="5" pageNumber="128">13</emphasis>
|
|||
|
) lateral views.
|
|||
|
</paragraph>
|
|||
|
</caption>
|
|||
|
<paragraph id="5534415ACE44A51BF0DD44B8FB66F94E" blockId="5.[151,1437,1421,2023]" pageId="5" pageNumber="128">
|
|||
|
Head (
|
|||
|
<figureCitation id="CDB05DDFCE44A51BF10B44B8FE84FA42" box="[273,344,1421,1448]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1928,1952]" captionTargetBox="[151,1435,453,1905]" captionTargetId="figure-100@3.[151,1435,453,1905]" captionTargetPageId="3" captionText="FIGURES 1–7. †Loeblitoides latus sp. nov., holotype; light micrographs. Lateral habitus (1), pronotum and elytral base in dorsal view (2), head in dorsal view (3), anterior body part in ventral view (4), distal region of maxillary palp (5), mesotarsus (6), and mesotarsomere 5 (7)." figureDoi="http://doi.org/10.5281/zenodo.13618056" httpUri="https://zenodo.org/record/13618056/files/figure.png" pageId="5" pageNumber="128">Figs 3</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="CDB05DDFCE44A51BF17E44BBFE77FA42" box="[356,427,1422,1448]" captionStart="FIGURES 14–19" captionStartId="6.[151,264,1628,1652]" captionTargetBox="[151,1436,777,1605]" captionTargetId="figure-226@6.[151,1436,777,1605]" captionTargetPageId="6" captionText="FIGURES 14–19. †Loeblitoides latus sp. nov., holotype, μCT reconstructions. Body in anterior (14), anteroventral (15), posterolateral (16), anterodorsal (17), anteroventral (18), and posteroventral (19) views. Abbreviations: ce, compound eye; cl, clypeus; cx1, procoxa; cx2, mesocoxa; fcg, frontoclypeal groove; fr, frons; hr, hypostomal ridge; tm, temple; v3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.13618062" httpUri="https://zenodo.org/record/13618062/files/figure.png" pageId="5" pageNumber="128">14–19</figureCitation>
|
|||
|
) flattened; length
|
|||
|
<quantity id="9273ECBFCE44A51BF26D44B8FD01FA42" box="[631,733,1421,1448]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="128" unit="mm" value="0.5">0.50 mm</quantity>
|
|||
|
, width
|
|||
|
<quantity id="9273ECBFCE44A51BF33444B8FC48FA42" box="[814,916,1421,1448]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="128" unit="mm" value="0.35">0.35 mm</quantity>
|
|||
|
; vertex strongly transverse and weakly, evenly convex; tempora weakly rounded, strongly convergent posteriorly, about as long as eye (best visible in
|
|||
|
<figureCitation id="CDB05DDFCE44A51BF5204484FA51FA26" box="[1338,1421,1457,1484]" captionStart="FIGURES 12–13" captionStartId="5.[151,264,1279,1303]" captionTargetBox="[151,1435,181,1255]" captionTargetId="figure-17@5.[151,1435,181,1255]" captionTargetPageId="5" captionText="FIGURES 12–13. †Loeblitoides latus sp. nov., holotype; μCT reconstructions of habitus in right (12) and left (13) lateral views." figureDoi="http://doi.org/10.5281/zenodo.13618060" httpUri="https://zenodo.org/record/13618060/files/figure.png" pageId="5" pageNumber="128">Fig. 13</figureCitation>
|
|||
|
); frons posteriorly confluent with vertex, transverse, its anterior margin with broad and short anteromedian expansion over frontoclypeal groove; supra-antennal tubercles weakly elevated; clypeus subtrapezoidal, distinctly transverse, weakly convex, weakly narrowing anterad, on entire width demarcated posteriorly by deep and sharply marked frontoclypeal groove; compound eyes large, strongly convex, moderately finely faceted. Punctures on vertex and frons poorly visible, setae on dorsal surface of head sparse and short, suberect. Maxillary palps (
|
|||
|
<figureCitation id="CDB05DDFCE44A51BF4A34750FB26F96A" box="[1209,1274,1637,1664]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1928,1952]" captionTargetBox="[151,1435,453,1905]" captionTargetId="figure-100@3.[151,1435,453,1905]" captionTargetPageId="3" captionText="FIGURES 1–7. †Loeblitoides latus sp. nov., holotype; light micrographs. Lateral habitus (1), pronotum and elytral base in dorsal view (2), head in dorsal view (3), anterior body part in ventral view (4), distal region of maxillary palp (5), mesotarsus (6), and mesotarsomere 5 (7)." figureDoi="http://doi.org/10.5281/zenodo.13618056" httpUri="https://zenodo.org/record/13618056/files/figure.png" pageId="5" pageNumber="128">Fig. 5</figureCitation>
|
|||
|
) with strongly elongate, slender palpomere 3 and with strongly elongate subconical and pointed palpomere 4.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="5534415ACE44A51BF0DD479BFD7CF8B2" blockId="5.[151,1437,1421,2023]" pageId="5" pageNumber="128">
|
|||
|
Antennae (
|
|||
|
<figureCitation id="CDB05DDFCE44A51BF1584798FE74F922" box="[322,424,1709,1736]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1928,1952]" captionTargetBox="[151,1435,453,1905]" captionTargetId="figure-100@3.[151,1435,453,1905]" captionTargetPageId="3" captionText="FIGURES 1–7. †Loeblitoides latus sp. nov., holotype; light micrographs. Lateral habitus (1), pronotum and elytral base in dorsal view (2), head in dorsal view (3), anterior body part in ventral view (4), distal region of maxillary palp (5), mesotarsus (6), and mesotarsomere 5 (7)." figureDoi="http://doi.org/10.5281/zenodo.13618056" httpUri="https://zenodo.org/record/13618056/files/figure.png" pageId="5" pageNumber="128">Figs 2–4</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="CDB05DDFCE44A51BF1AE479BFE1EF922" box="[436,450,1710,1736]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[158,1430,275,1880]" captionTargetId="figure-54@4.[158,1430,273,1880]" captionTargetPageId="4" captionText="FIGURES 8–11. †Loeblitoides latus sp. nov., holotype (8–10), and Syndicus echinatus Jałoszyński (11). Sketch of habitus in lateral view (8); μCT reconstructions of habitus in dorsal (9) and ventral (10) views; left elytral articulating lobe in dorsal view, scanning electron micrography (11). Abbreviations: a3‒11, antennomere 3‒11; bef, basal elytral fovea; ce, compound eye; fcg, frontoclypeal groove; pd, pedicel; pn, pronotum; sc, scape; scs, scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.13618058" httpUri="https://zenodo.org/record/13618058/files/figure.png" pageId="5" pageNumber="128">8</figureCitation>
|
|||
|
) slender, weakly and gradually broadening from antennomere 1 to 10, length
|
|||
|
<quantity id="9273ECBFCE44A51BF52B479BFA45F922" box="[1329,1433,1710,1736]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="5" pageNumber="128" unit="mm" value="1.23">1.23 mm</quantity>
|
|||
|
. Antennomeres 3–5 each with conspicuous mesal spine inserted on distinct projection or papilla, also antennomere 6 with similar seta; antennomeres 1–6 each distinctly elongate, with antennomere 3 longest, about 3 times as long as broad in lateral view (not counting mesal tubercle), antennomere 7 indistinctly elongate, 8–10 each about as long as broad, 11 minute, tightly sitting on apex of 10.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="5534415ACE44A518F0DD4657FF20FF13" blockId="5.[151,1437,1421,2023]" lastBlockId="6.[151,1437,151,753]" lastPageId="6" lastPageNumber="129" pageId="5" pageNumber="128">
|
|||
|
Pronotum (
|
|||
|
<figureCitation id="CDB05DDFCE44A51BF15D4654FE4EF896" box="[327,402,1889,1916]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1928,1952]" captionTargetBox="[151,1435,453,1905]" captionTargetId="figure-100@3.[151,1435,453,1905]" captionTargetPageId="3" captionText="FIGURES 1–7. †Loeblitoides latus sp. nov., holotype; light micrographs. Lateral habitus (1), pronotum and elytral base in dorsal view (2), head in dorsal view (3), anterior body part in ventral view (4), distal region of maxillary palp (5), mesotarsus (6), and mesotarsomere 5 (7)." figureDoi="http://doi.org/10.5281/zenodo.13618056" httpUri="https://zenodo.org/record/13618056/files/figure.png" pageId="5" pageNumber="128">Figs 2</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="CDB05DDFCE44A51BF1BA4657FE73F896" box="[416,431,1890,1916]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[158,1430,275,1880]" captionTargetId="figure-54@4.[158,1430,273,1880]" captionTargetPageId="4" captionText="FIGURES 8–11. †Loeblitoides latus sp. nov., holotype (8–10), and Syndicus echinatus Jałoszyński (11). Sketch of habitus in lateral view (8); μCT reconstructions of habitus in dorsal (9) and ventral (10) views; left elytral articulating lobe in dorsal view, scanning electron micrography (11). Abbreviations: a3‒11, antennomere 3‒11; bef, basal elytral fovea; ce, compound eye; fcg, frontoclypeal groove; pd, pedicel; pn, pronotum; sc, scape; scs, scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.13618058" httpUri="https://zenodo.org/record/13618058/files/figure.png" pageId="5" pageNumber="128">9</figureCitation>
|
|||
|
) distinctly but not strongly elongate, broadest near anterior 1/4, length
|
|||
|
<quantity id="9273ECBFCE44A51BF4FC4654FA93F896" box="[1254,1359,1889,1916]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="5" pageNumber="128" unit="mm" value="0.58">0.58 mm</quantity>
|
|||
|
, width
|
|||
|
<quantity id="9273ECBFCE44A51BF08D46B3FF21F84A" box="[151,253,1926,1952]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="128" unit="mm" value="0.4">0.40 mm</quantity>
|
|||
|
, pronotal index (length/width) 1.44. Anterior margin and sides in anterior fourth strongly, evenly rounded, lateral margins broadly and very shallowly concave behind middle; posterior corners well-marked, nearly rightangled, blunt; posterior margin nearly straight. Basal pronotal cavity poorly visible, presumably filled with dirt, about as long as half-length of pronotum and as wide as about 1/3 of pronotal width. Lateral impressions on pronotal disc strongly elongate and sharply demarcated. Punctures on pronotal disc indiscernible, setae short and sparse, suberect.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="5534415ACE47A518F0DD4036FF0BFE47" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="129">
|
|||
|
Elytra (
|
|||
|
<figureCitation id="CDB05DDFCE47A518F1004036FEA2FEF7" box="[282,382,259,285]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1928,1952]" captionTargetBox="[151,1435,453,1905]" captionTargetId="figure-100@3.[151,1435,453,1905]" captionTargetPageId="3" captionText="FIGURES 1–7. †Loeblitoides latus sp. nov., holotype; light micrographs. Lateral habitus (1), pronotum and elytral base in dorsal view (2), head in dorsal view (3), anterior body part in ventral view (4), distal region of maxillary palp (5), mesotarsus (6), and mesotarsomere 5 (7)." figureDoi="http://doi.org/10.5281/zenodo.13618056" httpUri="https://zenodo.org/record/13618056/files/figure.png" pageId="6" pageNumber="129">Figs 1, 2</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="CDB05DDFCE47A518F1934036FE18FEF7" box="[393,452,259,285]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[158,1430,275,1880]" captionTargetId="figure-54@4.[158,1430,273,1880]" captionTargetPageId="4" captionText="FIGURES 8–11. †Loeblitoides latus sp. nov., holotype (8–10), and Syndicus echinatus Jałoszyński (11). Sketch of habitus in lateral view (8); μCT reconstructions of habitus in dorsal (9) and ventral (10) views; left elytral articulating lobe in dorsal view, scanning electron micrography (11). Abbreviations: a3‒11, antennomere 3‒11; bef, basal elytral fovea; ce, compound eye; fcg, frontoclypeal groove; pd, pedicel; pn, pronotum; sc, scape; scs, scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.13618058" httpUri="https://zenodo.org/record/13618058/files/figure.png" pageId="6" pageNumber="129">8–10</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="CDB05DDFCE47A518F1D54036FE30FEF7" box="[463,492,259,285]" captionStart="FIGURES 12–13" captionStartId="5.[151,264,1279,1303]" captionTargetBox="[151,1435,181,1255]" captionTargetId="figure-17@5.[151,1435,181,1255]" captionTargetPageId="5" captionText="FIGURES 12–13. †Loeblitoides latus sp. nov., holotype; μCT reconstructions of habitus in right (12) and left (13) lateral views." figureDoi="http://doi.org/10.5281/zenodo.13618060" httpUri="https://zenodo.org/record/13618060/files/figure.png" pageId="6" pageNumber="129">12</figureCitation>
|
|||
|
) together oval, broadest slightly in front of middle; length
|
|||
|
<quantity id="9273ECBFCE47A518F4614037FB3DFEF7" box="[1147,1249,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="6" pageNumber="129" unit="mm" value="1.25">1.25 mm</quantity>
|
|||
|
, width
|
|||
|
<quantity id="9273ECBFCE47A518F5284037FA44FEF7" box="[1330,1432,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="129" unit="mm" value="0.75">0.75 mm</quantity>
|
|||
|
, elytral index (length/width) 1.67; elytral apices rounded together; humeral calli poorly developed, basal elytral impressions shallow, each elytron with two deep and asetose basal foveae connected by inversely U-shaped deep groove convex anteriorly (
|
|||
|
<figureCitation id="CDB05DDFCE47A518F1D6405AFDC9FE63" box="[460,533,367,393]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[158,1430,275,1880]" captionTargetId="figure-54@4.[158,1430,273,1880]" captionTargetPageId="4" captionText="FIGURES 8–11. †Loeblitoides latus sp. nov., holotype (8–10), and Syndicus echinatus Jałoszyński (11). Sketch of habitus in lateral view (8); μCT reconstructions of habitus in dorsal (9) and ventral (10) views; left elytral articulating lobe in dorsal view, scanning electron micrography (11). Abbreviations: a3‒11, antennomere 3‒11; bef, basal elytral fovea; ce, compound eye; fcg, frontoclypeal groove; pd, pedicel; pn, pronotum; sc, scape; scs, scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.13618058" httpUri="https://zenodo.org/record/13618058/files/figure.png" pageId="6" pageNumber="129">Fig. 9</figureCitation>
|
|||
|
). Punctures fine and inconspicuous; elytra covered with long, dense suberect setae.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="5534415ACE47A518F0DD4082FCD6FE1F" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="129">
|
|||
|
Mesoscutellar shield (
|
|||
|
<figureCitation id="CDB05DDFCE47A518F1DA4082FDD9FE3B" box="[448,517,439,465]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[158,1430,275,1880]" captionTargetId="figure-54@4.[158,1430,273,1880]" captionTargetPageId="4" captionText="FIGURES 8–11. †Loeblitoides latus sp. nov., holotype (8–10), and Syndicus echinatus Jałoszyński (11). Sketch of habitus in lateral view (8); μCT reconstructions of habitus in dorsal (9) and ventral (10) views; left elytral articulating lobe in dorsal view, scanning electron micrography (11). Abbreviations: a3‒11, antennomere 3‒11; bef, basal elytral fovea; ce, compound eye; fcg, frontoclypeal groove; pd, pedicel; pn, pronotum; sc, scape; scs, scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.13618058" httpUri="https://zenodo.org/record/13618058/files/figure.png" pageId="6" pageNumber="129">Fig. 9</figureCitation>
|
|||
|
) elongate subtriangular, only its very tip projecting posteriorly beyond step-wise border between elytral articulating lobe and elytral disc.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="5534415ACE47A518F0DD40CAFEADFD8B" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="129">
|
|||
|
Legs slender and lacking peculiar characters except for protibiae (
|
|||
|
<figureCitation id="CDB05DDFCE47A518F3A140CAFBCCFDF3" box="[955,1040,511,537]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1903,1927]" captionTargetBox="[158,1430,275,1880]" captionTargetId="figure-54@4.[158,1430,273,1880]" captionTargetPageId="4" captionText="FIGURES 8–11. †Loeblitoides latus sp. nov., holotype (8–10), and Syndicus echinatus Jałoszyński (11). Sketch of habitus in lateral view (8); μCT reconstructions of habitus in dorsal (9) and ventral (10) views; left elytral articulating lobe in dorsal view, scanning electron micrography (11). Abbreviations: a3‒11, antennomere 3‒11; bef, basal elytral fovea; ce, compound eye; fcg, frontoclypeal groove; pd, pedicel; pn, pronotum; sc, scape; scs, scutellar shield." figureDoi="http://doi.org/10.5281/zenodo.13618058" httpUri="https://zenodo.org/record/13618058/files/figure.png" pageId="6" pageNumber="129">Fig. 10</figureCitation>
|
|||
|
; best visible in right tibia) slightly curved inwards (i.e., mesally) in distal regions; all tarsi with tarsomeres distally reducing in length (
|
|||
|
<figureCitation id="CDB05DDFCE47A518F4F04316FAF1FDD7" box="[1258,1325,547,573]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1928,1952]" captionTargetBox="[151,1435,453,1905]" captionTargetId="figure-100@3.[151,1435,453,1905]" captionTargetPageId="3" captionText="FIGURES 1–7. †Loeblitoides latus sp. nov., holotype; light micrographs. Lateral habitus (1), pronotum and elytral base in dorsal view (2), head in dorsal view (3), anterior body part in ventral view (4), distal region of maxillary palp (5), mesotarsus (6), and mesotarsomere 5 (7)." figureDoi="http://doi.org/10.5281/zenodo.13618056" httpUri="https://zenodo.org/record/13618056/files/figure.png" pageId="6" pageNumber="129">Fig. 6</figureCitation>
|
|||
|
) and with long claws (
|
|||
|
<figureCitation id="CDB05DDFCE47A518F1054372FEBFFD8B" box="[287,355,583,609]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1928,1952]" captionTargetBox="[151,1435,453,1905]" captionTargetId="figure-100@3.[151,1435,453,1905]" captionTargetPageId="3" captionText="FIGURES 1–7. †Loeblitoides latus sp. nov., holotype; light micrographs. Lateral habitus (1), pronotum and elytral base in dorsal view (2), head in dorsal view (3), anterior body part in ventral view (4), distal region of maxillary palp (5), mesotarsus (6), and mesotarsomere 5 (7)." figureDoi="http://doi.org/10.5281/zenodo.13618056" httpUri="https://zenodo.org/record/13618056/files/figure.png" pageId="6" pageNumber="129">Fig. 7</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="1D9112D1CE47A518F0DD435EFE06FD43" pageId="6" pageNumber="129" type="etymology">
|
|||
|
<paragraph id="5534415ACE47A518F0DD435EFE06FD43" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="129">
|
|||
|
<emphasis id="67FF9D48CE47A518F0DD435EFE92FD6F" bold="true" box="[199,334,619,645]" pageId="6" pageNumber="129">Etymology.</emphasis>
|
|||
|
The Latin adjective
|
|||
|
<taxonomicName id="928B3AD9CE47A518F236435EFDBEFD6F" authorityName="Jałoszyński & Szawaryn" authorityYear="2024" box="[556,610,619,645]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="species" species="latus">
|
|||
|
<emphasis id="67FF9D48CE47A518F236435EFDBEFD6F" box="[556,610,619,645]" italics="true" pageId="6" pageNumber="129">latus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
, broad, refers to the much stouter body of this species than in the previously known †
|
|||
|
<taxonomicName id="928B3AD9CE47A518F0E243BAFE0FFD42" box="[248,467,655,681]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="species" species="separatus">
|
|||
|
<emphasis id="67FF9D48CE47A518F0E243BAFE0FFD42" box="[248,467,655,681]" italics="true" pageId="6" pageNumber="129">Loeblites separatus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="1D9112D1CE47A518F0DD4387FE30FD1B" pageId="6" pageNumber="129" type="distribution">
|
|||
|
<paragraph id="5534415ACE47A518F0DD4387FE30FD1B" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="129">
|
|||
|
<emphasis id="67FF9D48CE47A518F0DD4387FEBDFD26" bold="true" box="[199,353,690,716]" pageId="6" pageNumber="129">Distribution.</emphasis>
|
|||
|
Hukawng Valley near Tanai village,
|
|||
|
<collectingRegion id="974F8FB8CE47A518F32F4386FC0DFD27" box="[821,977,691,717]" country="Myanmar" name="Kachin" pageId="6" pageNumber="129">Kachin State</collectingRegion>
|
|||
|
, northern
|
|||
|
<collectingCountry id="2D9C01CACE47A518F4494386FB18FD27" box="[1107,1220,691,717]" name="Myanmar" pageId="6" pageNumber="129">Myanmar</collectingCountry>
|
|||
|
; late Albian-early Cenomanian, mid-Cretaceous.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<caption id="01F411D2CE47A518F08D4769FF22F90A" ID-DOI="http://doi.org/10.5281/zenodo.13618062" ID-Zenodo-Dep="13618062" httpUri="https://zenodo.org/record/13618062/files/figure.png" pageId="6" pageNumber="129" startId="6.[151,264,1628,1652]" targetBox="[151,1436,777,1605]" targetPageId="6" targetType="figure">
|
|||
|
<paragraph id="5534415ACE47A518F08D4769FF22F90A" blockId="6.[151,1437,1628,1760]" pageId="6" pageNumber="129">
|
|||
|
<emphasis id="67FF9D48CE47A518F08D4769FE84F99E" bold="true" box="[151,344,1628,1652]" pageId="6" pageNumber="129">FIGURES 14–19.</emphasis>
|
|||
|
†
|
|||
|
<taxonomicName id="928B3AD9CE47A518F1754769FDF4F99E" authorityName="Jałoszyński & Szawaryn" authorityYear="2024" box="[367,552,1628,1652]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
|
|||
|
<emphasis id="67FF9D48CE47A518F1754769FDF4F99E" box="[367,552,1628,1652]" italics="true" pageId="6" pageNumber="129">Loeblitoides latus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<emphasis id="67FF9D48CE47A518F2294769FD54F99E" bold="true" box="[563,648,1628,1652]" pageId="6" pageNumber="129">
|
|||
|
<taxonomicNameLabel id="7CCC2033CE47A518F2294769FD54F99E" box="[563,648,1628,1652]" pageId="6" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
</emphasis>
|
|||
|
, holotype, μCT reconstructions. Body in anterior (
|
|||
|
<emphasis id="67FF9D48CE47A518F4B34769FB1EF99E" bold="true" box="[1193,1218,1628,1652]" pageId="6" pageNumber="129">14</emphasis>
|
|||
|
), anteroventral (
|
|||
|
<emphasis id="67FF9D48CE47A518F56E4769FA51F99E" bold="true" box="[1396,1421,1628,1652]" pageId="6" pageNumber="129">15</emphasis>
|
|||
|
), posterolateral (
|
|||
|
<emphasis id="67FF9D48CE47A518F12B47B5FE96F972" bold="true" box="[305,330,1664,1688]" pageId="6" pageNumber="129">16</emphasis>
|
|||
|
), anterodorsal (
|
|||
|
<emphasis id="67FF9D48CE47A518F1F747B5FDDAF972" bold="true" box="[493,518,1664,1688]" pageId="6" pageNumber="129">17</emphasis>
|
|||
|
), anteroventral (
|
|||
|
<emphasis id="67FF9D48CE47A518F2A847B5FD17F972" bold="true" box="[690,715,1664,1688]" pageId="6" pageNumber="129">18</emphasis>
|
|||
|
), and posteroventral (
|
|||
|
<emphasis id="67FF9D48CE47A518F3B447B5FC1BF972" bold="true" box="[942,967,1664,1688]" pageId="6" pageNumber="129">19</emphasis>
|
|||
|
) views. Abbreviations: ce, compound eye; cl, clypeus; cx1, procoxa; cx2, mesocoxa; fcg, frontoclypeal groove; fr, frons; hr, hypostomal ridge; tm, temple; v3, metaventrite; vt, vertex.
|
|||
|
</paragraph>
|
|||
|
</caption>
|
|||
|
<subSubSection id="1D9112D1CE47A519F0DD4606FB17FE8F" lastPageId="7" lastPageNumber="130" pageId="6" pageNumber="129" type="discussion">
|
|||
|
<paragraph id="5534415ACE47A519F0DD4606FB17FE8F" blockId="6.[151,1437,1842,2013]" lastBlockId="7.[151,1437,151,357]" lastPageId="7" lastPageNumber="130" pageId="6" pageNumber="129">
|
|||
|
<emphasis id="67FF9D48CE47A518F0DD4606FEE7F8A7" bold="true" box="[199,315,1843,1869]" pageId="6" pageNumber="129">Remarks.</emphasis>
|
|||
|
†
|
|||
|
<taxonomicName id="928B3AD9CE47A518F14E4607FDFCF8A6" authorityName="Jałoszyński & Szawaryn" authorityYear="2024" box="[340,544,1842,1868]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="species" species="latus">
|
|||
|
<emphasis id="67FF9D48CE47A518F14E4607FDFCF8A6" box="[340,544,1842,1868]" italics="true" pageId="6" pageNumber="129">Loeblitoides latus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
can be easily distinguished from †
|
|||
|
<taxonomicName id="928B3AD9CE47A518F3A54606FB91F8A6" box="[959,1101,1843,1868]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="species" species="separatus">
|
|||
|
<emphasis id="67FF9D48CE47A518F3A54606FB91F8A6" box="[959,1101,1843,1868]" italics="true" pageId="6" pageNumber="129">L. separatus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
by its stouter pronotum and elytra, and different proportions of antennomeres. In †
|
|||
|
<taxonomicName id="928B3AD9CE47A518F31B4662FC8AF89A" authorityName="Jałoszyński & Szawaryn" authorityYear="2024" box="[769,854,1878,1904]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="species" species="latus">
|
|||
|
<emphasis id="67FF9D48CE47A518F31B4662FC8AF89A" box="[769,854,1878,1904]" italics="true" pageId="6" pageNumber="129">L. latus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
, the pronotal ratio length/width is merely 1.44 (vs. 2.00 in †
|
|||
|
<taxonomicName id="928B3AD9CE47A518F0E6464EFE5BF87E" box="[252,391,1915,1940]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="species" species="separatus">
|
|||
|
<emphasis id="67FF9D48CE47A518F0E6464EFE5BF87E" box="[252,391,1915,1940]" italics="true" pageId="6" pageNumber="129">L. separatus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
), and the elytral ratio equals 1.67 (vs.
|
|||
|
<quantity id="9273ECBFCE47A518F327464EFC50F87E" box="[829,908,1914,1941]" metricMagnitude="-2" metricUnit="m" metricValue="5.2832" pageId="6" pageNumber="129" unit="in" value="2.08">2.08 in</quantity>
|
|||
|
†
|
|||
|
<taxonomicName id="928B3AD9CE47A518F3BB464EFBF1F87E" box="[929,1069,1915,1940]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="species" species="separatus">
|
|||
|
<emphasis id="67FF9D48CE47A518F3BB464EFBF1F87E" box="[929,1069,1915,1940]" italics="true" pageId="6" pageNumber="129">L. separatus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
). The antennomere
|
|||
|
<quantity id="9273ECBFCE47A518F50A464EFAE0F87E" box="[1296,1340,1914,1941]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="129" unit="in" value="3.0">3 in</quantity>
|
|||
|
the new species is clearly more elongate, while antennomeres 7‒10 are much stouter, each almost as long as broad (each clearly elongate in †
|
|||
|
<taxonomicName id="928B3AD9CE47A518F16046F6FDD8F836" box="[378,516,1987,2012]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="species" species="separatus">
|
|||
|
<emphasis id="67FF9D48CE47A518F16046F6FDD8F836" box="[378,516,1987,2012]" italics="true" pageId="6" pageNumber="129">L. separatus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
). The differences in proportions of the pronotum and elytra cannot be attributed to a different sex of the same species, as such profound differences are not known to characterize sexes in any known extant
|
|||
|
<taxonomicName id="928B3AD9CE46A519F0F9418EFEBFFF3F" box="[227,355,187,213]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="tribe" tribe="Stenichnini">Stenichnini</taxonomicName>
|
|||
|
species. Also, the shapes of antennomeres in otherwise unmodified antennae are too different to represent sexual dimorphism (which in extant
|
|||
|
<taxonomicName id="928B3AD9CE46A519F28541EAFCC3FF13" box="[671,799,223,249]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="tribe" tribe="Stenichnini">Stenichnini</taxonomicName>
|
|||
|
is either not expressed in antennal features or evident by some conspicuously modified antennomeres in males). †
|
|||
|
<taxonomicName id="928B3AD9CE46A519F3154036FC0AFEF7" authorityName="Jałoszyński & Szawaryn" authorityYear="2024" box="[783,982,259,285]" class="Insecta" family="Staphylinidae" genus="Loeblitoides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="species" species="latus">
|
|||
|
<emphasis id="67FF9D48CE46A519F3154036FC0AFEF7" box="[783,982,259,285]" italics="true" pageId="7" pageNumber="130">Loeblitoides latus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
has also a distinctly broader neck region than that in the slenderer †
|
|||
|
<taxonomicName id="928B3AD9CE46A519F1D24012FD88FEAA" box="[456,596,295,320]" class="Insecta" family="Staphylinidae" genus="Loeblites" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="species" species="separatus">
|
|||
|
<emphasis id="67FF9D48CE46A519F1D24012FD88FEAA" box="[456,596,295,320]" italics="true" pageId="7" pageNumber="130">L. separatus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
. Still, the neck is slightly narrower than half width of the head (including eyes), and therefore the generic diagnosis does not need to be modified to include this variation.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|