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<mods:title id="85263A9EC18FB55EA5E310CED39C412E">Redescription of two species of Naineris (Annelida, Orbiniidae) with multiple dorsal organs and description of a new species from the NE Pacific</mods:title>
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<mods:affiliation id="93B7711F8D4B17EF1C4E32B3CBE3CDDE">Graduate program in Oceanic Coastal Systems (PGSISCO), Federal University of Paraná, Pontal do Paraná, Paraná, Brazil &amp; Departamento de Invertebrados, Campus de Ensino e Pesquisa do Museu Nacional, Universidade Federal do Rio de Janeiro, 20941 - 160, São Cristóvão, Rio de Janeiro, Rio de Janeiro, Brazil</mods:affiliation>
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<taxonomicName id="0EBCDDA4FF824F48FF12E5B0834A85B1" ID-CoL="45KRC" authority="Hartman, 1951" authorityName="Hartman" authorityYear="1951" box="[151,557,151,178]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="2" pageNumber="397" phylum="Annelida" rank="species" species="bicornis">
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<emphasis id="FBC87A35FF824F48FF12E5B0800E85B2" bold="true" box="[151,361,151,178]" italics="true" pageId="2" pageNumber="397">Naineris bicornis</emphasis>
<bibRefCitation id="AD2DDBD6FF824F48FEF5E5B0834A85B1" author="Hartman, O." box="[368,557,151,177]" pageId="2" pageNumber="397" pagination="7 - 124" refId="ref8904" refString="Hartman, O. (1951) The littoral marine annelids of the Gulf of Mexico. Publications of the Institute of Marine Science, University of Texas, 2, 7 - 124, 27 pls." type="journal article" year="1951">Hartman, 1951</bibRefCitation>
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<paragraph id="C903A627FF824F48FF12E59C806C85D5" blockId="2.[151,557,151,213]" box="[151,267,187,213]" pageId="2" pageNumber="397">
(
<figureCitation id="5187BAA2FF824F48FF1AE59C806585D5" box="[159,258,187,213]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1498,1522]" captionStartId-1="4.[151,250,1532,1556]" captionStartId-2="5.[151,250,1297,1321]" captionTargetBox-0="[151,1435,333,1472]" captionTargetBox-1="[152,1426,181,1507]" captionTargetBox-2="[152,1436,274,1274]" captionTargetId-0="figure-61@2.[151,1436,332,1472]" captionTargetId-1="figure-21@4.[151,1436,181,1508]" captionTargetId-2="figure-78@5.[151,1436,273,1274]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." captionText-1="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." captionText-2="FIGURE 3. Naineris bicornis (USNM 1284137).A, anterior end; B, abdominal segments, dorsal view, multiple dorsal organs; C, thoracic neurochaetae; D, abdominal neurochaetae. a—acicula, cc—crenulate capillaries, do—dorsal organs, pe—peristomium, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 400 μm; B: 100 μm.; C–D: 20 μm)." figureDoi-0="http://doi.org/10.5281/zenodo.13234905" figureDoi-1="http://doi.org/10.5281/zenodo.13234907" figureDoi-2="http://doi.org/10.5281/zenodo.13234909" httpUri-0="https://zenodo.org/record/13234905/files/figure.png" httpUri-1="https://zenodo.org/record/13234907/files/figure.png" httpUri-2="https://zenodo.org/record/13234909/files/figure.png" pageId="2" pageNumber="397">Figs 1–3</figureCitation>
)
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<treatmentCitation id="481D8036FF824F48FF12E42783558418" author="Hartman, O." box="[151,562,255,280]" page="74" pageId="2" pageNumber="397" year="1951">
<taxonomicName id="0EBCDDA4FF824F48FF12E42783558418" ID-CoL="45KRC" authority="Hartman, 1951: 72 - 74" authorityName="Hartman" authorityPageNumber="72 - 74" authorityYear="1951" box="[151,562,255,280]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="2" pageNumber="397" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF824F48FF12E42780208418" box="[151,327,256,280]" italics="true" pageId="2" pageNumber="397">Naineris bicornis</emphasis>
<bibRefCitation id="AD2DDBD6FF824F48FEC9E42783558418" author="Hartman, O." box="[332,562,255,280]" pageId="2" pageNumber="397" pagination="7 - 124" refId="ref8904" refString="Hartman, O. (1951) The littoral marine annelids of the Gulf of Mexico. Publications of the Institute of Marine Science, University of Texas, 2, 7 - 124, 27 pls." type="journal article" year="1951">Hartman, 1951: 72–74</bibRefCitation>
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, pl. 19, figs 1–6; 1957: 304–305, pl. 40, figs 1–6; 1959: 366.— 
<treatmentCitation id="481D8036FF824F48FB22E5D881B88436" author="Perkins, T. H. &amp; Savage, T." page="43" pageId="2" pageNumber="397" year="1975">
<bibRefCitation id="AD2DDBD6FF824F48FB22E5D881B88436" author="Perkins, T. H. &amp; Savage, T." pageId="2" pageNumber="397" pagination="1 - 62" refId="ref9399" refString="Perkins, T. H. &amp; Savage, T. (1975) A bibliography and checklist of polychaetous annelids of Florida, the Gulf of Mexico, and the Caribbean region. Florida Marine Research Publications, 14, 1 - 62." type="journal article" year="1975">Perkins &amp; Savage 1975: 43</bibRefCitation>
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.— 
<treatmentCitation id="481D8036FF824F48FF78E43980D78436" author="Taylor, J. L." box="[253,432,286,310]" page="1" pageId="2" pageNumber="397" year="1984">
<bibRefCitation id="AD2DDBD6FF824F48FF78E43980D78436" author="Taylor, J. L." box="[253,432,286,310]" pageId="2" pageNumber="397" pagination="1 - 38" refId="ref9638" refString="Taylor, J. L. (1984) Chapter 1 Orbiniidae. In: Uebelacker, J. M. &amp; Johnson, P. G. (Eds.), Taxonomic guide to the polychaete of the Northern Gulf of Mexico. Vol. I. Barry A. Vittor &amp; Associates, Inc., Inc. Mobile, Alabama, pp. 1 - 38." type="book chapter" year="1984">Taylor 1984: 1–9</bibRefCitation>
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, figs 1–5, 1–6a–f.— 
<treatmentCitation id="481D8036FF824F48FDFCE439821D8436" author="Salazar-Vallejo, S. I." box="[633,890,286,310]" page="26" pageId="2" pageNumber="397" year="1996">
<bibRefCitation id="AD2DDBD6FF824F48FDFCE439821D8436" author="Salazar-Vallejo, S. I." box="[633,890,286,310]" pageId="2" pageNumber="397" pagination="11 - 50" refId="ref9504" refString="Salazar-Vallejo, S. I. (1996) Lista de especies y bibliografia de poliquetos (Polychaeta) del Gran Caribe. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Serie Zoologia, 67, 11 - 50." type="journal article" year="1996">Salazar-Vallejo 1996: 26</bibRefCitation>
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.— 
<treatmentCitation id="481D8036FF824F48FC12E43985238436" author="Perkins, T. H." box="[919,1092,286,310]" page="88" pageId="2" pageNumber="397" year="1998">
<bibRefCitation id="AD2DDBD6FF824F48FC12E43985238436" author="Perkins, T. H." box="[919,1092,286,310]" pageId="2" pageNumber="397" pagination="79 - 122" refId="ref9326" refString="Perkins, T. H. (1998) Checklist of shallow-water marine polychaetous Annelida of Florida. In: Camp, D. K., Lyons, W. G. &amp; Perkins, T. H. (Eds.), Checklists of selected shallow - water marine invertebrates of Florida. Florida Marine Research Institute, Technical Report, TR- 3, pp. 79 - 122." type="journal article" year="1998">Perkins 1998: 88</bibRefCitation>
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.—Felder &amp; Camp 2009: 764.
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</paragraph>
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<emphasis id="FBC87A35FF824F48FF12E0FD807780F2" bold="true" box="[151,272,1498,1522]" pageId="2" pageNumber="397">FIGURE 1</emphasis>
. 
<taxonomicName id="0EBCDDA4FF824F48FE9AE0FC80B480F2" authorityName="Hartman" authorityYear="1951" box="[287,467,1498,1522]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="2" pageNumber="397" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF824F48FE9AE0FC80B480F2" box="[287,467,1498,1522]" italics="true" pageId="2" pageNumber="397">Naineris bicornis</emphasis>
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. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm).
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<emphasis id="FBC87A35FF824F48FF12E3F380E083EE" bold="true" box="[151,391,1748,1774]" pageId="2" pageNumber="397">Material examined.</emphasis>
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(
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674).
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— 
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off 
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, 
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2421, coll. 
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, 
<collectorName id="6449C3F1FF824F48FCBAE23A82DC8236" box="[831,955,1820,1847]" pageId="2" pageNumber="397">H. Kritzler</collectorName>
, 
<collectorName id="6449C3F1FF824F48FC42E23A85268236" box="[967,1089,1820,1847]" pageId="2" pageNumber="397">D. Garlick</collectorName>
, 
<date id="BD0280E7FF824F48FBC8E23A85DD8237" box="[1101,1210,1820,1847]" pageId="2" pageNumber="397" value="1975-09">
<collectingDate id="AD46790FFF824F48FBC8E23A85DD8237" box="[1101,1210,1820,1847]" pageId="2" pageNumber="397" value="1975-09">Sep 1975</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF824F48FB43E23B84408236" box="[1222,1319,1820,1846]" degrees="29.62" direction="north" orientation="latitude" pageId="2" pageNumber="397" precision="555" value="29.62">29.62°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF824F48FAB6E23A84FF8237" box="[1331,1432,1821,1847]" degrees="84.28" direction="west" orientation="longitude" pageId="2" pageNumber="397" precision="555" value="-84.28">84.28°W</geoCoordinate>
, 
<quantity id="0E440BC2FF824F48FF12E26681B1825B" box="[151,214,1857,1883]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="397" unit="m" value="19.0">
<elevation id="42914114FF824F48FF12E26681B1825B" box="[151,214,1857,1883]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="397" unit="m" value="19.0">19 m</elevation>
</quantity>
, box core (1, 
<specimenCode id="991A0E5CFF824F48FE05E2668324825B" box="[384,579,1856,1883]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="397" type="Museum">USNM 1284135</specimenCode>
);
</materialsCitation>
<materialsCitation id="79D4AC7AFF824F48FDDEE266821C827F" collectingDate="1975-09" collectionCode="MAFLA" collectorName="Sta. &amp; D. Savelle &amp; H. Kritzler &amp; D. Garlick" country="United States of America" county="Franklin County" elevation="19" latitude="29.91" location="Gulf of Mexico" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="2" pageNumber="397" specimenCode="USNM 1284134" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF824F48FDDEE26683DC825B" box="[603,699,1857,1883]" pageId="2" pageNumber="397">MAFLA</collectionCode>
<collectorName id="6449C3F1FF824F48FD43E2668396825B" box="[710,753,1857,1883]" pageId="2" pageNumber="397">Sta.</collectorName>
2421, coll. 
<collectorName id="6449C3F1FF824F48FC07E2668299825A" box="[898,1022,1856,1883]" pageId="2" pageNumber="397">D. Savelle</collectorName>
, 
<collectorName id="6449C3F1FF824F48FB88E26685EA825A" box="[1037,1165,1856,1883]" pageId="2" pageNumber="397">H. Kritzler</collectorName>
, 
<collectorName id="6449C3F1FF824F48FB19E266847E825A" box="[1180,1305,1856,1883]" pageId="2" pageNumber="397">D. Garlick</collectorName>
, 
<date id="BD0280E7FF824F48FAADE26684FF825B" box="[1320,1432,1856,1883]" pageId="2" pageNumber="397" value="1975-09">
<collectingDate id="AD46790FFF824F48FAADE26684FF825B" box="[1320,1432,1856,1883]" pageId="2" pageNumber="397" value="1975-09">Sep 1975</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF824F48FF12E2438056827E" box="[151,305,1892,1918]" degrees="29.616667" direction="north" orientation="latitude" pageId="2" pageNumber="397" precision="1" value="29.616667">29.616667°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF824F48FEBEE24280B0827F" box="[315,471,1893,1919]" degrees="84.283333" direction="west" orientation="longitude" pageId="2" pageNumber="397" precision="1" value="-84.28333">84.283333°W</geoCoordinate>
, 
<quantity id="0E440BC2FF824F48FE67E242837B827F" box="[482,540,1893,1919]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="397" unit="m" value="19.0">
<elevation id="42914114FF824F48FE67E242837B827F" box="[482,540,1893,1919]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="397" unit="m" value="19.0">19 m</elevation>
</quantity>
, box core (1, 
<specimenCode id="991A0E5CFF824F48FD34E2428208827F" box="[689,879,1893,1919]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="397" type="Museum">USNM 1284134</specimenCode>
);
</materialsCitation>
<materialsCitation id="79D4AC7AFF824F48FC04E24282CE82A3" collectingDate="1950-02-04" collectionCode="MAFLA" collectorName="Sta. &amp; D. Savelle &amp; H. Kritzler &amp; D. Garlick" country="United States of America" county="Franklin County" elevation="19" latitude="29.91" location="Gulf of Mexico" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="2" pageNumber="397" specimenCode="USNM 1284133" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF824F48FC04E2428281827F" box="[897,998,1893,1919]" pageId="2" pageNumber="397">MAFLA</collectionCode>
, 
<collectorName id="6449C3F1FF824F48FC75E242857B827F" box="[1008,1052,1893,1919]" pageId="2" pageNumber="397">Sta.</collectorName>
2420, coll. 
<collectorName id="6449C3F1FF824F48FB18E2428473827E" box="[1181,1300,1892,1919]" pageId="2" pageNumber="397">D. Savelle</collectorName>
, 
<collectorName id="6449C3F1FF824F48FA9BE24284FF827E" box="[1310,1432,1892,1919]" pageId="2" pageNumber="397">H. Kritzler</collectorName>
, 
<collectorName id="6449C3F1FF824F48FF12E2AE807682A2" box="[151,273,1928,1955]" pageId="2" pageNumber="397">D. Garlick</collectorName>
, 1975, 
<geoCoordinate id="AC88C0E0FF824F48FEE7E2AF808782A2" box="[354,480,1928,1954]" degrees="29.6994" direction="north" orientation="latitude" pageId="2" pageNumber="397" precision="5" value="29.6994">29.6994°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF824F48FE6EE2AF830B82A2" box="[491,620,1928,1954]" degrees="84.1836" direction="west" orientation="longitude" pageId="2" pageNumber="397" precision="5" value="-84.1836">84.1836°W</geoCoordinate>
, 
<quantity id="0E440BC2FF824F48FDFDE2AE83D682A3" box="[632,689,1929,1955]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="397" unit="m" value="19.0">
<elevation id="42914114FF824F48FDFDE2AE83D682A3" box="[632,689,1929,1955]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="397" unit="m" value="19.0">19 m</elevation>
</quantity>
(1, 
<specimenCode id="991A0E5CFF824F48FD5BE2AE82FA82A3" box="[734,925,1929,1955]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="397" type="Museum">USNM 1284133</specimenCode>
);
</materialsCitation>
<materialsCitation id="79D4AC7AFF824F48FC34E2AE828982C7" collectingDate="1975-06" collectionCode="MAFLA" collectorName="Sta. &amp; Savelle &amp; H. Kritzler &amp; D. Garlick" country="United States of America" county="Franklin County" elevation="14" latitude="29.91" location="Gulf of Mexico" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="2" pageNumber="397" specimenCode="USNM 1284132" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF824F48FC34E2AE857182A3" box="[945,1046,1929,1955]" pageId="2" pageNumber="397">MAFLA</collectionCode>
, 
<collectorName id="6449C3F1FF824F48FBA4E2AE852B82A3" box="[1057,1100,1929,1955]" pageId="2" pageNumber="397">Sta.</collectorName>
2420, D. 
<collectorName id="6449C3F1FF824F48FB39E2AF847682A2" box="[1212,1297,1928,1954]" pageId="2" pageNumber="397">Savelle</collectorName>
, 
<collectorName id="6449C3F1FF824F48FA98E2AE84FF82A2" box="[1309,1432,1928,1955]" pageId="2" pageNumber="397">H. Kritzler</collectorName>
, 
<collectorName id="6449C3F1FF824F48FF12E28A807582C6" box="[151,274,1964,1991]" pageId="2" pageNumber="397">D. Garlick</collectorName>
, 
<date id="BD0280E7FF824F48FE9AE28A80ED82C7" box="[287,394,1964,1991]" pageId="2" pageNumber="397" value="1975-06">
<collectingDate id="AD46790FFF824F48FE9AE28A80ED82C7" box="[287,394,1964,1991]" pageId="2" pageNumber="397" value="1975-06">Jun 1975</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF824F48FE12E28B837382C6" box="[407,532,1964,1990]" degrees="29.6994" direction="north" orientation="latitude" pageId="2" pageNumber="397" precision="5" value="29.6994">29.6994°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF824F48FDA4E28B83C582C6" box="[545,674,1964,1990]" degrees="84.1836" direction="west" orientation="longitude" pageId="2" pageNumber="397" precision="5" value="-84.1836">84.1836°W</geoCoordinate>
, 
<quantity id="0E440BC2FF824F48FD2AE28A838B82C7" box="[687,748,1965,1991]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="397" unit="m" value="14.0">
<elevation id="42914114FF824F48FD2AE28A838B82C7" box="[687,748,1965,1991]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="397" unit="m" value="14.0">14 m</elevation>
</quantity>
, (1, 
<specimenCode id="991A0E5CFF824F48FCA4E28A828582C7" box="[801,994,1965,1991]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="397" type="Museum">USNM 1284132</specimenCode>
);
</materialsCitation>
<materialsCitation id="79D4AC7AFF824F48FC72E28A855382EB" collectingDate="1950-02-04" collectionCode="MAFLA" collectorName="Sta. &amp; D. Savelle &amp; H. Kritzler &amp; D. Garlick" country="United States of America" county="Franklin County" elevation="14" latitude="29.91" location="Gulf of Mexico" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="2" pageNumber="397" specimenCode="USNM 1284131" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF824F48FC72E28A853B82C7" box="[1015,1116,1965,1991]" pageId="2" pageNumber="397">MAFLA</collectionCode>
, 
<collectorName id="6449C3F1FF824F48FBECE28A85F382C7" box="[1129,1172,1965,1991]" pageId="2" pageNumber="397">Sta.</collectorName>
2420, coll. 
<collectorName id="6449C3F1FF824F48FA9AE28A84FF82C6" box="[1311,1432,1964,1991]" pageId="2" pageNumber="397">D. Savelle</collectorName>
, 
<collectorName id="6449C3F1FF824F48FF12E2F6807482EA" box="[151,275,2000,2027]" pageId="2" pageNumber="397">H. Kritzler</collectorName>
, 
<collectorName id="6449C3F1FF824F48FE9AE2F680FE82EA" box="[287,409,2000,2027]" pageId="2" pageNumber="397">D. Garlick</collectorName>
, 1975, 
<geoCoordinate id="AC88C0E0FF824F48FE6EE2F7830E82EA" box="[491,617,2000,2026]" degrees="29.6994" direction="north" orientation="latitude" pageId="2" pageNumber="397" precision="5" value="29.6994">29.6994°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF824F48FDF0E2F7839282EA" box="[629,757,2000,2026]" degrees="84.1836" direction="west" orientation="longitude" pageId="2" pageNumber="397" precision="5" value="-84.1836">84.1836°W</geoCoordinate>
, 
<quantity id="0E440BC2FF824F48FC84E2F6825C82EB" box="[769,827,2001,2027]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="397" unit="m" value="14.0">
<elevation id="42914114FF824F48FC84E2F6825C82EB" box="[769,827,2001,2027]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="397" unit="m" value="14.0">14 m</elevation>
</quantity>
(1, 
<specimenCode id="991A0E5CFF824F48FCECE2F6854E82EB" box="[873,1065,2001,2027]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="397" type="Museum">USNM 1284131</specimenCode>
).
</materialsCitation>
<materialsCitation id="79D4AC7AFF824F49FBB1E2F782A185D5" collectingDate="1975-09-19" collectionCode="MAFLA" collectorName=" D. Savelle &amp; H. Kritzler &amp; D. Garlick" country="United States of America" county="Franklin County" elevation="29" lastPageId="3" lastPageNumber="398" latitude="29.91" location="Cedar Key" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="2" pageNumber="397" specimenCode="USNM 1284136, USNM 1284137, USNM 1284138, USNM 1284139" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<emphasis id="FBC87A35FF824F48FBB1E2F7859D82EB" bold="true" box="[1076,1274,2000,2027]" pageId="2" pageNumber="397">
—off 
<location id="CC63F0FCFF824F48FBFDE2F7859D82EB" LSID="urn:lsid:plazi:treatment:41151731FF824F4CFF12E5B085D1841D:CC63F0FCFF824F48FBFDE2F7859D82EB" box="[1144,1274,2000,2027]" country="United States of America" county="Franklin County" latitude="29.91" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" name="Cedar Key" pageId="2" pageNumber="397" stateProvince="Florida">Cedar Key</location>
</emphasis>
, 
<collectionCode id="AFAD3EE2FF824F48FA8CE2F6840E82EB" box="[1289,1385,2001,2027]" pageId="2" pageNumber="397">MAFLA</collectionCode>
<collectorName id="6449C3F1FF824F48FAF4E2F684FA82EB" box="[1393,1437,2001,2027]" pageId="2" pageNumber="397">Sta.</collectorName>
2317, coll. 
<collectorName id="6449C3F1FF834F49FE96E5B080EE85B1" box="[275,393,151,177]" pageId="3" pageNumber="398">D. Savelle</collectorName>
, 
<collectorName id="6449C3F1FF834F49FE11E5B0836985B1" box="[404,526,151,177]" pageId="3" pageNumber="398">H. Kritzler</collectorName>
, 
<collectorName id="6449C3F1FF834F49FD9DE5B083F785B1" box="[536,656,151,177]" pageId="3" pageNumber="398">D. Garlick</collectorName>
, 
<date id="BD0280E7FF834F49FD1FE5B0824085B1" box="[666,807,150,177]" pageId="3" pageNumber="398" value="1975-09-19">
<collectingDate id="AD46790FFF834F49FD1FE5B0824085B1" box="[666,807,150,177]" pageId="3" pageNumber="398" value="1975-09-19">19 Sep 1975</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF834F49FCB4E5B082AC85B1" box="[817,971,151,177]" degrees="28.933333" direction="north" orientation="latitude" pageId="3" pageNumber="398" precision="1" value="28.933332">28.933333°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF834F49FC50E5B0851585B1" box="[981,1138,151,177]" degrees="84.100000" direction="west" orientation="longitude" pageId="3" pageNumber="398" precision="1" value="-84.1">84.100000°W</geoCoordinate>
, 
<quantity id="0E440BC2FF834F49FBF9E5B085D185B1" box="[1148,1206,151,177]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="398" unit="m" value="29.0">
<elevation id="42914114FF834F49FBF9E5B085D185B1" box="[1148,1206,151,177]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="398" unit="m" value="29.0">29 m</elevation>
</quantity>
, box core (1, 
<specimenCode id="991A0E5CFF834F49FACEE5B0819A85D5" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 1284136</specimenCode>
; 1, 
<specimenCode id="991A0E5CFF834F49FEA2E59C808085D5" box="[295,487,187,213]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 1284137</specimenCode>
; 1, 
<specimenCode id="991A0E5CFF834F49FD94E59C83B685D5" box="[529,721,187,213]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 1284138</specimenCode>
; 1, 
<specimenCode id="991A0E5CFF834F49FD7EE59C82DD85D5" box="[763,954,187,213]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 1284139</specimenCode>
).
</materialsCitation>
<materialsCitation id="79D4AC7AFF834F49FC40E59C850285F9" collectingDate="1977-11" collectionCode="MAFLA" collectorName="Sta." country="United States of America" county="Franklin County" elevation="35" latitude="29.91" location="Crystal River" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 090156" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
— 
<emphasis id="FBC87A35FF834F49FC66E59D85D685D4" bold="true" box="[995,1201,186,212]" pageId="3" pageNumber="398">
off 
<location id="CC63F0FCFF834F49FB88E59D85D685D4" LSID="urn:lsid:plazi:treatment:41151731FF824F4CFF12E5B085D1841D:CC63F0FCFF834F49FB88E59D85D685D4" box="[1037,1201,186,212]" country="United States of America" county="Franklin County" latitude="29.91" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" name="Crystal River" pageId="3" pageNumber="398" stateProvince="Florida">Crystal River</location>
</emphasis>
, 
<collectionCode id="AFAD3EE2FF834F49FB44E59C844685D5" box="[1217,1313,187,213]" pageId="3" pageNumber="398">MAFLA</collectionCode>
<collectorName id="6449C3F1FF834F49FAAFE59C843285D5" box="[1322,1365,187,213]" pageId="3" pageNumber="398">Sta.</collectorName>
2316, 
<date id="BD0280E7FF834F49FF12E5F8806D85F9" box="[151,266,223,249]" pageId="3" pageNumber="398" value="1977-11">
<collectingDate id="AD46790FFF834F49FF12E5F8806D85F9" box="[151,266,223,249]" pageId="3" pageNumber="398" value="1977-11">Nov 1977</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF834F49FE93E5F880D085F9" box="[278,439,223,249]" degrees="28.700000" direction="north" orientation="latitude" pageId="3" pageNumber="398" precision="1" value="28.7">28.700000° N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF834F49FE46E5F8830085F9" box="[451,615,223,249]" degrees="84.333333" direction="west" orientation="longitude" pageId="3" pageNumber="398" precision="1" value="-84.333336">84.333333° W</geoCoordinate>
, 
<quantity id="0E440BC2FF834F49FDF6E5F983CB85F9" box="[627,684,222,249]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="398" unit="m" value="35.0">
<elevation id="42914114FF834F49FDF6E5F983CB85F9" box="[627,684,222,249]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="398" unit="m" value="35.0">35 m</elevation>
</quantity>
(1, 
<specimenCode id="991A0E5CFF834F49FD5FE5F882EF85F9" box="[730,904,222,249]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 090156</specimenCode>
) (parapodia slides).
</materialsCitation>
<materialsCitation id="79D4AC7AFF834F49FBE1E5F882318441" collectingDate="1981-07" collectionCode="SOFLA, BLM, MMS" collectorName="Sta. &amp; Mote Marine Lab For Bureau &amp; Land Management &amp; Minerals Management Service" country="United States of America" county="Franklin County" elevation="54" latitude="29.91" location="Florida" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 090154" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
— 
<emphasis id="FBC87A35FF834F49FB06E5F9846385F8" bold="true" box="[1155,1284,222,248]" pageId="3" pageNumber="398">
off 
<collectingRegion id="0B7868C5FF834F49FB2EE5F9846385F8" box="[1195,1284,222,248]" country="United States of America" name="Florida" pageId="3" pageNumber="398">Florida</collectingRegion>
</emphasis>
, 
<collectionCode id="AFAD3EE2FF834F49FA96E5F8840D85F9" box="[1299,1386,223,249]" pageId="3" pageNumber="398">SOFLA</collectionCode>
<collectorName id="6449C3F1FF834F49FAF4E5F884FA85F9" box="[1393,1437,223,249]" pageId="3" pageNumber="398">Sta.</collectorName>
16, coll. 
<collectorName id="6449C3F1FF834F49FE84E424832B841D" box="[257,588,259,285]" pageId="3" pageNumber="398">Mote Marine Lab For Bureau</collectorName>
of the 
<collectorName id="6449C3F1FF834F49FD1EE4248208841D" box="[667,879,259,285]" pageId="3" pageNumber="398">Land Management</collectorName>
/ 
<collectorName id="6449C3F1FF834F49FCF2E42485A9841D" box="[887,1230,259,285]" pageId="3" pageNumber="398">Minerals Management Service</collectorName>
(
<collectionCode id="AFAD3EE2FF834F49FB58E424847B841D" box="[1245,1308,259,285]" pageId="3" pageNumber="398">BLM</collectionCode>
/ 
<collectionCode id="AFAD3EE2FF834F49FAA0E424840F841D" box="[1317,1384,259,285]" country="0" httpUri="http://grbio.org/cool/1jzf-hb7f" name="Montshire Museum of Science" pageId="3" pageNumber="398">MMS</collectionCode>
), 
<date id="BD0280E7FF834F49FAFEE42481B68441" pageId="3" pageNumber="398" value="1981-07">
<collectingDate id="AD46790FFF834F49FAFEE42481B68441" pageId="3" pageNumber="398" value="1981-07">Jul 1981</collectingDate>
</date>
, 
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, 
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, 
<quantity id="0E440BC2FF834F49FDABE401830E8441" box="[558,617,294,321]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="3" pageNumber="398" unit="m" value="54.0">
<elevation id="42914114FF834F49FDABE401830E8441" box="[558,617,294,321]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="3" pageNumber="398" unit="m" value="54.0">54 m</elevation>
</quantity>
(1, 
<specimenCode id="991A0E5CFF834F49FD1DE400822D8441" box="[664,842,294,321]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 090154</specimenCode>
);
</materialsCitation>
<materialsCitation id="79D4AC7AFF834F49FCDAE40082888465" collectingDate="1981-05" collectionCode="SOFLA, MMS" collectorName="Sta. &amp; Mote Marine Lab For" country="United States of America" county="Franklin County" elevation="24" latitude="29.91" location="Gulf of Mexico" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 090153" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF834F49FCDAE40082D18441" box="[863,950,295,321]" pageId="3" pageNumber="398">SOFLA</collectionCode>
<collectorName id="6449C3F1FF834F49FC3AE400828D8441" box="[959,1002,295,321]" pageId="3" pageNumber="398">Sta.</collectorName>
02, coll. 
<collectorName id="6449C3F1FF834F49FBD2E40084288441" box="[1111,1359,295,321]" pageId="3" pageNumber="398">Mote Marine Lab For</collectorName>
BLM/ 
<collectionCode id="AFAD3EE2FF834F49FF12E46C81BC8465" box="[151,219,331,357]" country="0" httpUri="http://grbio.org/cool/1jzf-hb7f" name="Montshire Museum of Science" pageId="3" pageNumber="398">MMS</collectionCode>
, 
<date id="BD0280E7FF834F49FF6CE46C80068465" box="[233,353,331,357]" pageId="3" pageNumber="398" value="1981-05">
<collectingDate id="AD46790FFF834F49FF6CE46C80068465" box="[233,353,331,357]" pageId="3" pageNumber="398" value="1981-05">May 1981</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF834F49FEEAE46C836E8465" box="[367,521,331,357]" degrees="26.763889" direction="north" orientation="latitude" pageId="3" pageNumber="398" precision="1" value="26.76389">26.763889°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF834F49FD92E46D83D48465" box="[535,691,330,357]" degrees="82.753056" direction="west" orientation="longitude" pageId="3" pageNumber="398" precision="1" value="-82.75306">82.753056°W</geoCoordinate>
, 
<quantity id="0E440BC2FF834F49FD44E46C839A8465" box="[705,765,331,357]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="398" unit="m" value="24.0">
<elevation id="42914114FF834F49FD44E46C839A8465" box="[705,765,331,357]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="398" unit="m" value="24.0">24 m</elevation>
</quantity>
(1, 
<specimenCode id="991A0E5CFF834F49FCAAE46C82848465" box="[815,995,330,357]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 090153</specimenCode>
);
</materialsCitation>
<materialsCitation id="79D4AC7AFF834F49FC7CE46C83E184AD" collectingDate="1984-05" collectionCode="SOFLA, MMS" collectorName="Sta. &amp; Environmental Science &amp; Engineering &amp; Inc." country="United States of America" county="Ecological Research Associates" elevation="18" latitude="29.91" location="Lgl" longLatPrecision="1" longitude="-84.39528" municipality="Ese" pageId="3" pageNumber="398" specimenCode="USNM 112302" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF834F49FC7CE46C85378465" box="[1017,1104,331,357]" pageId="3" pageNumber="398">SOFLA</collectionCode>
<collectorName id="6449C3F1FF834F49FBDFE46C85E28465" box="[1114,1157,331,357]" pageId="3" pageNumber="398">Sta.</collectorName>
46, coll. 
<collectorName id="6449C3F1FF834F49FB73E46C81898489" pageId="3" pageNumber="398">Environmental Science</collectorName>
and 
<collectorName id="6449C3F1FF834F49FEA1E44880D78489" box="[292,432,367,393]" pageId="3" pageNumber="398">Engineering</collectorName>
, 
<collectorName id="6449C3F1FF834F49FE3FE44880858489" box="[442,482,367,393]" pageId="3" pageNumber="398">Inc.</collectorName>
/LGL 
<collectingCounty id="2062DEABFF834F49FDACE44882ED8489" box="[553,906,367,393]" pageId="3" pageNumber="398">Ecological Research Associates</collectingCounty>
(
<collectingMunicipality id="29673C5DFF834F49FC1DE44882A58489" box="[920,962,367,393]" pageId="3" pageNumber="398">Ese</collectingMunicipality>
/ 
<location id="CC63F0FCFF834F49FC4EE44882948489" LSID="urn:lsid:plazi:treatment:41151731FF824F4CFF12E5B085D1841D:CC63F0FCFF834F49FC4EE44882948489" box="[971,1011,367,393]" country="United States of America" county="Ecological Research Associates" latitude="29.91" longLatPrecision="1" longitude="-84.39528" municipality="Ese" name="Lgl" pageId="3" pageNumber="398" stateProvince="Florida">Lgl</location>
) 
<location id="CC63F0FCFF834F49FB84E448854F8489" LSID="urn:lsid:plazi:treatment:41151731FF824F4CFF12E5B085D1841D:CC63F0FCFF834F49FB84E448854F8489" box="[1025,1064,367,393]" country="United States of America" county="Ecological Research Associates" latitude="29.91" longLatPrecision="1" longitude="-84.39528" municipality="Ese" name="For" pageId="3" pageNumber="398" stateProvince="Florida">For</location>
BLM/ 
<collectionCode id="AFAD3EE2FF834F49FBF0E44885D08489" box="[1141,1207,367,393]" country="0" httpUri="http://grbio.org/cool/1jzf-hb7f" name="Montshire Museum of Science" pageId="3" pageNumber="398">MMS</collectionCode>
, 
<date id="BD0280E7FF834F49FB47E44884508489" box="[1218,1335,367,393]" pageId="3" pageNumber="398" value="1984-05">
<collectingDate id="AD46790FFF834F49FB47E44884508489" box="[1218,1335,367,393]" pageId="3" pageNumber="398" value="1984-05">May 1984</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF834F49FAC4E44881DE84AD" degrees="26.0144" direction="north" orientation="latitude" pageId="3" pageNumber="398" precision="5" value="26.0144">26.0144 °N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF834F49FF4DE4B4802E84AD" box="[200,329,403,429]" degrees="82.1319" direction="west" orientation="longitude" pageId="3" pageNumber="398" precision="5" value="-82.1319">82.1319°W</geoCoordinate>
, 
<quantity id="0E440BC2FF834F49FEDDE4B480F384AD" box="[344,404,403,429]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="398" unit="m" value="18.0">
<elevation id="42914114FF834F49FEDDE4B480F384AD" box="[344,404,403,429]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="398" unit="m" value="18.0">18 m</elevation>
</quantity>
(2, 
<specimenCode id="991A0E5CFF834F49FE42E4B4831C84AD" box="[455,635,403,429]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 112302</specimenCode>
);
</materialsCitation>
<materialsCitation id="79D4AC7AFF834F49FD14E4B483D184D1" collectingDate="1977-05" collectionCode="SABP, MMS" collectorName="Sta." country="United States of America" county="Franklin County" elevation="20" latitude="29.91" location="Instruments" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 59369" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF834F49FD14E4B483B184AD" box="[657,726,403,429]" pageId="3" pageNumber="398">SABP</collectionCode>
<collectorName id="6449C3F1FF834F49FD64E4B4826B84AD" box="[737,780,403,429]" pageId="3" pageNumber="398">Sta.</collectorName>
7B, coll. 
<collectingRegion id="0B7868C5FF834F49FC06E4B482A384AD" box="[899,964,403,429]" country="United States of America" name="Texas" pageId="3" pageNumber="398">Texas</collectingRegion>
<location id="CC63F0FCFF834F49FC4BE4B4853584AD" LSID="urn:lsid:plazi:treatment:41151731FF824F4CFF12E5B085D1841D:CC63F0FCFF834F49FC4BE4B4853584AD" box="[974,1106,403,429]" country="United States of America" county="Franklin County" latitude="29.91" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" name="Instruments" pageId="3" pageNumber="398" stateProvince="Texas">Instruments</location>
for BLM/ 
<collectionCode id="AFAD3EE2FF834F49FB4AE4B4847684AD" box="[1231,1297,403,429]" country="0" httpUri="http://grbio.org/cool/1jzf-hb7f" name="Montshire Museum of Science" pageId="3" pageNumber="398">MMS</collectionCode>
, 
<date id="BD0280E7FF834F49FA9AE4B484FF84AD" box="[1311,1432,403,429]" pageId="3" pageNumber="398" value="1977-05">
<collectingDate id="AD46790FFF834F49FA9AE4B484FF84AD" box="[1311,1432,403,429]" pageId="3" pageNumber="398" value="1977-05">May 1977</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF834F49FF12E490807284D1" box="[151,277,439,465]" degrees="29.4664" direction="north" orientation="latitude" pageId="3" pageNumber="398" precision="5" value="29.4664">29.4664°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF834F49FE9BE49180F884D1" box="[286,415,438,465]" degrees="80.9525" direction="west" orientation="longitude" pageId="3" pageNumber="398" precision="5" value="-80.9525">80.9525°W</geoCoordinate>
, 
<quantity id="0E440BC2FF834F49FE2CE490808784D1" box="[425,480,439,465]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="398" unit="m" value="20.0">
<elevation id="42914114FF834F49FE2CE490808784D1" box="[425,480,439,465]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="398" unit="m" value="20.0">20 m</elevation>
</quantity>
(1, 
<specimenCode id="991A0E5CFF834F49FD8CE49083CC84D1" box="[521,683,438,465]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 59369</specimenCode>
);
</materialsCitation>
<materialsCitation id="79D4AC7AFF834F49FD39E490818B84F5" collectingDate="1977-11" collectionCode="MAFLA" collectorName="Sta." country="United States of America" county="Franklin County" elevation="27" latitude="29.91" location="Gulf of Mexico" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 90164" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF834F49FD39E490827B84D1" box="[700,796,439,465]" pageId="3" pageNumber="398">MAFLA</collectionCode>
<collectorName id="6449C3F1FF834F49FCA7E490822A84D1" box="[802,845,439,465]" pageId="3" pageNumber="398">Sta.</collectorName>
2960, 
<date id="BD0280E7FF834F49FC12E490856F84D1" box="[919,1032,439,465]" pageId="3" pageNumber="398" value="1977-11">
<collectingDate id="AD46790FFF834F49FC12E490856F84D1" box="[919,1032,439,465]" pageId="3" pageNumber="398" value="1977-11">Nov 1977</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF834F49FB94E491851584D1" box="[1041,1138,438,465]" degrees="25.67" direction="north" orientation="latitude" pageId="3" pageNumber="398" precision="555" value="25.67">25.67°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF834F49FBF9E490858684D1" box="[1148,1249,439,465]" degrees="82.33" direction="west" orientation="longitude" pageId="3" pageNumber="398" precision="555" value="-82.33">82.33°W</geoCoordinate>
, 
<quantity id="0E440BC2FF834F49FB6EE490844584D1" box="[1259,1314,439,465]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="398" unit="m" value="27.0">
<elevation id="42914114FF834F49FB6EE490844584D1" box="[1259,1314,439,465]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="398" unit="m" value="27.0">27 m</elevation>
</quantity>
(1, 
<specimenCode id="991A0E5CFF834F49FACEE490818784F5" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 90164</specimenCode>
).
</materialsCitation>
<materialsCitation id="79D4AC7AFF834F49FF6EE4FC819D8719" collectingDate="1977-08" collectionCode="MAFLA" collectorName="Sta." country="United States of America" county="Franklin County" elevation="19" latitude="29.91" location="St. Petersburg" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 090155" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
— 
<emphasis id="FBC87A35FF834F49FE8FE4FD80B884F5" bold="true" box="[266,479,474,501]" pageId="3" pageNumber="398">
off 
<location id="CC63F0FCFF834F49FEB6E4FD80B884F5" LSID="urn:lsid:plazi:treatment:41151731FF824F4CFF12E5B085D1841D:CC63F0FCFF834F49FEB6E4FD80B884F5" box="[307,479,474,501]" country="United States of America" county="Franklin County" latitude="29.91" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" name="St. Petersburg" pageId="3" pageNumber="398" stateProvince="Florida">St. Petersburg</location>
</emphasis>
, 
<collectionCode id="AFAD3EE2FF834F49FE6AE4FC832884F5" box="[495,591,475,501]" pageId="3" pageNumber="398">MAFLA</collectionCode>
<collectorName id="6449C3F1FF834F49FDDDE4FC83E484F5" box="[600,643,475,501]" pageId="3" pageNumber="398">Sta.</collectorName>
2207, coll. 
<date id="BD0280E7FF834F49FC8EE4FC821984F5" box="[779,894,475,501]" pageId="3" pageNumber="398" value="1977-08">
<collectingDate id="AD46790FFF834F49FC8EE4FC821984F5" box="[779,894,475,501]" pageId="3" pageNumber="398" value="1977-08">Aug 1977</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF834F49FC0EE4FD854284F5" box="[907,1061,474,501]" degrees="27.950000" direction="north" orientation="latitude" pageId="3" pageNumber="398" precision="1" value="27.95">27.950000°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF834F49FBB4E4FD85A984F5" box="[1073,1230,474,501]" degrees="83.150000" direction="west" orientation="longitude" pageId="3" pageNumber="398" precision="1" value="-83.15">83.150000°W</geoCoordinate>
, 
<quantity id="0E440BC2FF834F49FB5EE4FC847084F5" box="[1243,1303,475,501]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="398" unit="m" value="19.0">
<elevation id="42914114FF834F49FB5EE4FC847084F5" box="[1243,1303,475,501]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="398" unit="m" value="19.0">19 m</elevation>
</quantity>
, (1, 
<specimenCode id="991A0E5CFF834F49FACEE4FC81898719" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 090155</specimenCode>
).
</materialsCitation>
<materialsCitation id="79D4AC7AFF834F49FF7FE4D98549873D" collectingDate="1975-07" collectionCode="MAFLA" collectorName="Sta. &amp; D. Savelle &amp; H. Kritzler &amp; D. Garlick" country="United States of America" county="Franklin County" elevation="35" latitude="29.91" location="Apalachicola Bay" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 1284140, USNM 1284141, USNM 1284670" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
— 
<emphasis id="FBC87A35FF834F49FE9DE4D9836C8719" bold="true" box="[280,523,510,537]" pageId="3" pageNumber="398">
off 
<location id="CC63F0FCFF834F49FEB9E4D9836C8719" LSID="urn:lsid:plazi:treatment:41151731FF824F4CFF12E5B085D1841D:CC63F0FCFF834F49FEB9E4D9836C8719" box="[316,523,510,537]" country="United States of America" county="Franklin County" latitude="29.91" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" name="Apalachicola Bay" pageId="3" pageNumber="398" stateProvince="Florida">Apalachicola Bay</location>
</emphasis>
, 
<collectionCode id="AFAD3EE2FF834F49FD92E4D8831B8719" box="[535,636,511,537]" pageId="3" pageNumber="398">MAFLA</collectionCode>
, 
<collectorName id="6449C3F1FF834F49FD00E4D883D78719" box="[645,688,511,537]" pageId="3" pageNumber="398">Sta.</collectorName>
2316, coll. 
<collectorName id="6449C3F1FF834F49FCAAE4D882C28719" box="[815,933,511,537]" pageId="3" pageNumber="398">D. Savelle</collectorName>
, 
<collectorName id="6449C3F1FF834F49FC2BE4D885408719" box="[942,1063,511,537]" pageId="3" pageNumber="398">H. Kritzler</collectorName>
, 
<collectorName id="6449C3F1FF834F49FBB4E4D885CF8719" box="[1073,1192,511,537]" pageId="3" pageNumber="398">D. Garlick</collectorName>
, 
<date id="BD0280E7FF834F49FB34E4D884758719" box="[1201,1298,510,537]" pageId="3" pageNumber="398" value="1975-07">
<collectingDate id="AD46790FFF834F49FB34E4D884758719" box="[1201,1298,510,537]" pageId="3" pageNumber="398" value="1975-07">Jul 1975</collectingDate>
</date>
, 
<geoCoordinate id="AC88C0E0FF834F49FA9EE4D884FF8719" box="[1307,1432,511,537]" degrees="28.7003" direction="north" orientation="latitude" pageId="3" pageNumber="398" precision="5" value="28.7003">28.7003°N</geoCoordinate>
, 
<geoCoordinate id="AC88C0E0FF834F49FF12E704807F873D" box="[151,280,547,573]" degrees="84.3336" direction="west" orientation="longitude" pageId="3" pageNumber="398" precision="5" value="-84.3336">84.3336°W</geoCoordinate>
, 
<quantity id="0E440BC2FF834F49FEA1E7058007873D" box="[292,352,546,573]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="398" unit="m" value="35.0">
<elevation id="42914114FF834F49FEA1E7058007873D" box="[292,352,546,573]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="398" unit="m" value="35.0">35 m</elevation>
</quantity>
, (2, 
<specimenCode id="991A0E5CFF834F49FE16E7048335873D" box="[403,594,547,573]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 1284140</specimenCode>
; 1, 
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; 1, 
<specimenCode id="991A0E5CFF834F49FCE7E7048545873D" box="[866,1058,547,573]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 1284670</specimenCode>
);
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<materialsCitation id="79D4AC7AFF834F49FBB3E70483C28761" collectingDate="1974-06" collectionCode="MAFLA" collectorName="Sta" country="United States of America" county="Franklin County" elevation="19" latitude="29.91" location="Gulf of Mexico" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 1284677" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF834F49FBB3E70485FC873D" box="[1078,1179,547,573]" pageId="3" pageNumber="398">MAFLA</collectionCode>
, 
<collectorName id="6449C3F1FF834F49FB22E70485AC873D" box="[1191,1227,547,573]" pageId="3" pageNumber="398">Sta</collectorName>
48, coll. DG., 
<date id="BD0280E7FF834F49FAF0E70481B58761" pageId="3" pageNumber="398" value="1974-06">
<collectingDate id="AD46790FFF834F49FAF0E70481B58761" pageId="3" pageNumber="398" value="1974-06">Jun 1974</collectingDate>
</date>
, 
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, 
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(2, 
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);
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<materialsCitation id="79D4AC7AFF834F49FD28E760806F8785" collectingDate="1974-06" collectionCode="MAFLA" collectorName="Sta" country="United States of America" county="Franklin County" elevation="19" latitude="29.91" location="Gulf of Mexico" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 1284679" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF834F49FD28E76082758761" box="[685,786,583,609]" pageId="3" pageNumber="398">MAFLA</collectionCode>
, 
<collectorName id="6449C3F1FF834F49FC98E76082268761" box="[797,833,583,609]" pageId="3" pageNumber="398">Sta</collectorName>
42, coll. DG., 
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</date>
, 
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, 
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(1, 
<specimenCode id="991A0E5CFF834F49FACEE760819B8785" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 1284679</specimenCode>
).
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<materialsCitation id="79D4AC7AFF834F49FE82E74C80CE87A9" collectingDate="1975-07" collectionCode="MAFLA" collectorName=" D. Bishof &amp; D. Savelle" country="United States of America" county="Franklin County" elevation="19" latitude="29.91" location="Clear Water" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 1284671" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
— 
<emphasis id="FBC87A35FF834F49FEA3E74D80848785" bold="true" box="[294,483,618,645]" pageId="3" pageNumber="398">
off 
<location id="CC63F0FCFF834F49FECBE74D80848785" LSID="urn:lsid:plazi:treatment:41151731FF824F4CFF12E5B085D1841D:CC63F0FCFF834F49FECBE74D80848785" box="[334,483,618,645]" country="United States of America" county="Franklin County" latitude="29.91" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" name="Clear Water" pageId="3" pageNumber="398" stateProvince="Florida">Clear Water</location>
</emphasis>
, 
<collectionCode id="AFAD3EE2FF834F49FE74E74C83318785" box="[497,598,619,645]" pageId="3" pageNumber="398">MAFLA</collectionCode>
, 
<collectorName id="6449C3F1FF834F49FDE7E74C83E98785" box="[610,654,619,645]" pageId="3" pageNumber="398">Sta.</collectorName>
2207, coll. 
<collectorName id="6449C3F1FF834F49FC90E74C82E18785" box="[789,902,619,645]" pageId="3" pageNumber="398">D. Bishof</collectorName>
, 
<collectorName id="6449C3F1FF834F49FC17E74C856D8785" box="[914,1034,619,645]" pageId="3" pageNumber="398">D. Savelle</collectorName>
, 
<date id="BD0280E7FF834F49FB93E74C851D8785" box="[1046,1146,618,645]" pageId="3" pageNumber="398" value="1975-07">
<collectingDate id="AD46790FFF834F49FB93E74C851D8785" box="[1046,1146,618,645]" pageId="3" pageNumber="398" value="1975-07">Jul 1975</collectingDate>
</date>
, 
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, 
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, 
<quantity id="0E440BC2FF834F49FA05E74C81C887A9" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="398" unit="m" value="19.0">
<elevation id="42914114FF834F49FA05E74C81C887A9" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="398" unit="m" value="19.0">19 m</elevation>
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, (1, 
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<materialsCitation id="79D4AC7AFF834F49FE35E7A8803887CD" collectingDate="1975-07" collectionCode="MAFLA" collectorName=" D. Bishof &amp; D. Savelle" country="United States of America" county="Franklin County" elevation="34" latitude="29.91" location="Gulf of Mexico" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 1284672" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<collectionCode id="AFAD3EE2FF834F49FE35E7A8837287A9" box="[432,533,655,681]" pageId="3" pageNumber="398">MAFLA</collectionCode>
, 
<collectorName id="6449C3F1FF834F49FD9AE7A8832D87A9" box="[543,586,655,681]" pageId="3" pageNumber="398">Sta.</collectorName>
2209, coll. 
<collectorName id="6449C3F1FF834F49FD49E7A8825B87A9" box="[716,828,655,681]" pageId="3" pageNumber="398">D. Bishof</collectorName>
, 
<collectorName id="6449C3F1FF834F49FCC3E7A882DB87A9" box="[838,956,655,681]" pageId="3" pageNumber="398">D. Savelle</collectorName>
, 
<date id="BD0280E7FF834F49FC42E7A8854E87A9" box="[967,1065,654,681]" pageId="3" pageNumber="398" value="1975-07">
<collectingDate id="AD46790FFF834F49FC42E7A8854E87A9" box="[967,1065,654,681]" pageId="3" pageNumber="398" value="1975-07">Jul 1975</collectingDate>
</date>
, 
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, 
<geoCoordinate id="AC88C0E0FF834F49FB28E7A9844987A9" box="[1197,1326,654,681]" degrees="83.5667" direction="west" orientation="longitude" pageId="3" pageNumber="398" precision="5" value="-83.5667">83.5667°W</geoCoordinate>
, 
<quantity id="0E440BC2FF834F49FABCE7A8841487A9" box="[1337,1395,655,681]" metricMagnitude="1" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="398" unit="m" value="34.0">
<elevation id="42914114FF834F49FABCE7A8841487A9" box="[1337,1395,655,681]" metricMagnitude="1" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="398" unit="m" value="34.0">34 m</elevation>
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, (3, 
<specimenCode id="991A0E5CFF834F49FF12E794803487CD" box="[151,339,691,717]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="398" type="Museum">USNM 1284672</specimenCode>
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<materialsCitation id="79D4AC7AFF834F49FEDAE794833287F1" collectingDate="1975-05" collectionCode="MAFLA" collectorName=" D. Bishof &amp; D. Savelle" country="United States of America" county="Franklin County" elevation="53" latitude="29.91" location="Charlotte Harbor" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 1284678" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
— 
<emphasis id="FBC87A35FF834F49FEF8E795831087CD" bold="true" box="[381,631,690,717]" pageId="3" pageNumber="398">
off 
<location id="CC63F0FCFF834F49FE21E795831087CD" LSID="urn:lsid:plazi:treatment:41151731FF824F4CFF12E5B085D1841D:CC63F0FCFF834F49FE21E795831087CD" box="[420,631,690,717]" country="United States of America" county="Franklin County" latitude="29.91" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" name="Charlotte Harbor" pageId="3" pageNumber="398" stateProvince="Florida">Charlotte Harbor</location>
</emphasis>
, 
<collectionCode id="AFAD3EE2FF834F49FD01E794838E87CD" box="[644,745,691,717]" pageId="3" pageNumber="398">MAFLA</collectionCode>
, 
<collectorName id="6449C3F1FF834F49FD71E794827887CD" box="[756,799,691,717]" pageId="3" pageNumber="398">Sta.</collectorName>
2104, coll. 
<collectorName id="6449C3F1FF834F49FC24E794857687CD" box="[929,1041,691,717]" pageId="3" pageNumber="398">D. Bishof</collectorName>
, 
<collectorName id="6449C3F1FF834F49FB9EE79485F687CD" box="[1051,1169,691,717]" pageId="3" pageNumber="398">D. Savelle</collectorName>
, 
<date id="BD0280E7FF834F49FB19E794847787CD" box="[1180,1296,690,717]" pageId="3" pageNumber="398" value="1975-05">
<collectingDate id="AD46790FFF834F49FB19E794847787CD" box="[1180,1296,690,717]" pageId="3" pageNumber="398" value="1975-05">May 1975</collectingDate>
</date>
, 
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, 
<geoCoordinate id="AC88C0E0FF834F49FF12E7F0807E87F1" box="[151,281,727,753]" degrees="83.3833" direction="west" orientation="longitude" pageId="3" pageNumber="398" precision="5" value="-83.3833">83.3833°W</geoCoordinate>
, 
<quantity id="0E440BC2FF834F49FEA6E7F1803A87F1" box="[291,349,726,753]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="398" unit="m" value="53.0">
<elevation id="42914114FF834F49FEA6E7F1803A87F1" box="[291,349,726,753]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="398" unit="m" value="53.0">53 m</elevation>
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, (1, 
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).
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<materialsCitation id="79D4AC7AFF834F49FDD1E7F082438615" collectingDate="1975-05" collectionCode="MAFLA" collectorName=" D. Bishof &amp; D. Savelle" country="United States of America" county="Franklin County" elevation="11" latitude="29.91" location="Sanibel Island" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" pageId="3" pageNumber="398" specimenCode="USNM 1284683" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
— 
<emphasis id="FBC87A35FF834F49FDF7E7F1822387F1" bold="true" box="[626,836,726,753]" pageId="3" pageNumber="398">
off 
<location id="CC63F0FCFF834F49FD1CE7F1822387F1" LSID="urn:lsid:plazi:treatment:41151731FF824F4CFF12E5B085D1841D:CC63F0FCFF834F49FD1CE7F1822387F1" box="[665,836,726,753]" country="United States of America" county="Franklin County" latitude="29.91" longLatPrecision="1" longitude="-84.39528" municipality="Alligator Harbor" name="Sanibel Island" pageId="3" pageNumber="398" stateProvince="Florida">Sanibel Island</location>
</emphasis>
, 
<collectionCode id="AFAD3EE2FF834F49FCD4E7F082D187F1" box="[849,950,727,753]" pageId="3" pageNumber="398">MAFLA</collectionCode>
, 
<collectorName id="6449C3F1FF834F49FC45E7F0828C87F1" box="[960,1003,727,753]" pageId="3" pageNumber="398">Sta.</collectorName>
2101, coll. 
<collectorName id="6449C3F1FF834F49FBEBE7F085BA87F1" box="[1134,1245,727,753]" pageId="3" pageNumber="398">D. Bishof</collectorName>
, 
<collectorName id="6449C3F1FF834F49FB62E7F0843987F1" box="[1255,1374,727,753]" pageId="3" pageNumber="398">D. Savelle</collectorName>
, 
<date id="BD0280E7FF834F49FAECE7F081B58615" pageId="3" pageNumber="398" value="1975-05">
<collectingDate id="AD46790FFF834F49FAECE7F081B58615" pageId="3" pageNumber="398" value="1975-05">May 1975</collectingDate>
</date>
, 
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, 
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, 
<quantity id="0E440BC2FF834F49FE77E7DC834B8615" box="[498,556,763,789]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="398" unit="m" value="11.0">
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, (1, 
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="81A6F5ACFF834F49FF42E63885558681" pageId="3" pageNumber="398" type="description">
<paragraph id="C903A627FF834F49FF42E63885558681" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="398">
<emphasis id="FBC87A35FF834F49FF42E63880118639" bold="true" box="[199,374,799,825]" pageId="3" pageNumber="398">Measurements</emphasis>
. 
<typeStatus id="16071885FF834F49FE03E63880898639" box="[390,494,799,825]" pageId="3" pageNumber="398" type="holotype">Holotype</typeStatus>
is fragmented in three parts. The most anterior 
<quantity id="0E440BC2FF834F49FB90E639851C8639" box="[1045,1147,798,825]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="398" unit="mm" value="0.75">0.75 mm</quantity>
long, 
<quantity id="0E440BC2FF834F49FB40E638846F8639" box="[1221,1288,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="398" unit="mm" value="3.0">3 mm</quantity>
wide, for 27 chaetigers. 
<bibRefCitation id="AD2DDBD6FF834F49FE9DE66480AA865D" author="Hartman, O." box="[280,461,834,861]" pageId="3" pageNumber="398" pagination="7 - 124" refId="ref8904" refString="Hartman, O. (1951) The littoral marine annelids of the Gulf of Mexico. Publications of the Institute of Marine Science, University of Texas, 2, 7 - 124, 27 pls." type="journal article" year="1951">Hartman (1951)</bibRefCitation>
and 
<bibRefCitation id="AD2DDBD6FF834F49FD83E66483C6865D" author="Taylor, J. L." box="[518,673,835,861]" pageId="3" pageNumber="398" pagination="1 - 38" refId="ref9638" refString="Taylor, J. L. (1984) Chapter 1 Orbiniidae. In: Uebelacker, J. M. &amp; Johnson, P. G. (Eds.), Taxonomic guide to the polychaete of the Northern Gulf of Mexico. Vol. I. Barry A. Vittor &amp; Associates, Inc., Inc. Mobile, Alabama, pp. 1 - 38." type="book chapter" year="1984">Taylor (1984)</bibRefCitation>
reported animals reaching 
<quantity id="0E440BC2FF834F49FC5DE664854F865D" box="[984,1064,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="398" unit="mm" value="30.0">30 mm</quantity>
length and 
<quantity id="0E440BC2FF834F49FB2AE6658596865D" box="[1199,1265,834,861]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="398" unit="mm" value="5.0">5 mm</quantity>
width for over 100 segments for specimens from the same localities and sampling events studied.
</paragraph>
</subSubSection>
<subSubSection id="81A6F5ACFF834F49FF42E6AD8349817D" pageId="3" pageNumber="398" type="diagnosis">
<paragraph id="C903A627FF834F49FF42E6AD8349817D" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="398">
<emphasis id="FBC87A35FF834F49FF42E6AD805D86A4" bold="true" box="[199,314,906,932]" pageId="3" pageNumber="398">Diagnosis</emphasis>
. Spatulate to slightly indented prostomium. Multiple dorsal sensory organs grouped ten per side in a one row anteriorly, increasing to two on posterior thoracic segments, almost crossing midline; abdominal sensory organs forming two distinct groups irregularly arranged, reduced in number (four to five). Thoracic notopodial lobes may be acute-to-rounded-tipped and with median restriction. Thoracic neuropodial lobe with an upper subtriangular papilla. Thoracic neurochaetae with 5–6 transverse, posterior rows of numerous subuluncini; a transverse, anterior row of about 20 crenulate capillaries; an inferior, anterior, oblique row of about 20 hooded uncini and an inferior, posterior, oblique row of capillaries.
</paragraph>
</subSubSection>
<subSubSection id="81A6F5ACFF834F4EFF42E1A180CC83ED" lastPageId="4" lastPageNumber="399" pageId="3" pageNumber="398" type="description">
<paragraph id="C903A627FF834F49FF42E1A1829681E9" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="398">
<emphasis id="FBC87A35FF834F49FF42E1A1800A81A0" bold="true" box="[199,365,1158,1184]" pageId="3" pageNumber="398">Redescription</emphasis>
. 
<typeStatus id="16071885FF834F49FEFEE1A0808481A1" box="[379,483,1159,1185]" pageId="3" pageNumber="398" type="holotype">Holotype</typeStatus>
medium-sized specimen. Color in alcohol pale yellow. Thorax and abdomen distinct, with parapodia inserted laterally in thorax and dorsally in abdomen. Most abdominal branchiae lacking. Ventrally thorax smooth, but abdomen tri-annulate with central ring wider than others.
</paragraph>
<paragraph id="C903A627FF834F49FF42E1D4855F8055" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="398">
Prostomium long, spatulate, slightly indented in front end; no eyespots; nuchal organs present at between posterior end of prostomium and anterior end of peristomium (
<figureCitation id="5187BAA2FF834F49FCF6E03082968031" box="[883,1009,1303,1329]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Fig. 1A–B</figureCitation>
). Peristomium achaetous, as short as following rings. Mouth opening with striated lips; proboscis wide, lobed (
<figureCitation id="5187BAA2FF834F49FC57E01C854C8055" box="[978,1067,1339,1365]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Fig. 1D</figureCitation>
).
</paragraph>
<paragraph id="C903A627FF834F49FF42E07880DC832D" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="398">
Branchiae from chaetiger 6 onward (
<figureCitation id="5187BAA2FF834F49FDF1E07883B58079" box="[628,722,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Figs 1B</figureCitation>
, 
<figureCitation id="5187BAA2FF834F49FD67E078826F8079" box="[738,776,1375,1401]" captionStart="FIGURE 3" captionStartId="5.[151,250,1297,1321]" captionTargetBox="[152,1436,274,1274]" captionTargetId="figure-78@5.[151,1436,273,1274]" captionTargetPageId="5" captionText="FIGURE 3. Naineris bicornis (USNM 1284137).A, anterior end; B, abdominal segments, dorsal view, multiple dorsal organs; C, thoracic neurochaetae; D, abdominal neurochaetae. a—acicula, cc—crenulate capillaries, do—dorsal organs, pe—peristomium, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 400 μm; B: 100 μm.; C–D: 20 μm)." figureDoi="http://doi.org/10.5281/zenodo.13234909" httpUri="https://zenodo.org/record/13234909/files/figure.png" pageId="3" pageNumber="398">3A</figureCitation>
), lanceolate, densely ciliated (
<figureCitation id="5187BAA2FF834F49FBEDE07885878079" box="[1128,1248,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Fig. 1E–F</figureCitation>
); first branchiae approximately ⅓ size of abdominal branchiae. Dorsal sensory organs present from chaetiger 12 (
<figureCitation id="5187BAA2FF834F49FB6CE0A48421809D" box="[1257,1350,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Figs 1B</figureCitation>
, 
<figureCitation id="5187BAA2FF834F49FAD1E0A484F0809D" box="[1364,1431,1411,1437]" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="3" pageNumber="398">2B–C</figureCitation>
, 
<figureCitation id="5187BAA2FF834F49FF12E08081DD80C1" box="[151,186,1447,1473]" captionStart="FIGURE 3" captionStartId="5.[151,250,1297,1321]" captionTargetBox="[152,1436,274,1274]" captionTargetId="figure-78@5.[151,1436,273,1274]" captionTargetPageId="5" captionText="FIGURE 3. Naineris bicornis (USNM 1284137).A, anterior end; B, abdominal segments, dorsal view, multiple dorsal organs; C, thoracic neurochaetae; D, abdominal neurochaetae. a—acicula, cc—crenulate capillaries, do—dorsal organs, pe—peristomium, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 400 μm; B: 100 μm.; C–D: 20 μm)." figureDoi="http://doi.org/10.5281/zenodo.13234909" httpUri="https://zenodo.org/record/13234909/files/figure.png" pageId="3" pageNumber="398">3B</figureCitation>
), multiple per segment, oval-shaped; most anterior grouped ten per side forming one row, increasing to two on posterior thoracic segments, almost crossing midline; abdominal sensory organs forming two distinct groups irregularly disposed, reduced in number (four to five). Dorsal crest represented by a straight, consistent, flat curve, low and ciliated (
<figureCitation id="5187BAA2FF834F49FEDCE33480C9832D" box="[345,430,1555,1581]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Fig. 1F</figureCitation>
).
</paragraph>
<paragraph id="C903A627FF834F49FF42E31082E98205" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="398">
Thorax slightly depressed (
<figureCitation id="5187BAA2FF834F49FD8BE310830C8351" box="[526,619,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Fig. 1E</figureCitation>
), both fragments with 45 chaetigers (~3 transitional segments with intermediate characteristics between thoracic and abdominal segments) (
<figureCitation id="5187BAA2FF834F49FC44E37C855E8375" box="[961,1081,1627,1653]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Fig. 1B–C</figureCitation>
). Parapodia biramous. Thoracic notopodial postchaetal lobes slightly constricted, lanceolate, with elongated tips (
<figureCitation id="5187BAA2FF834F49FB91E358850B8399" box="[1044,1132,1663,1689]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Figs 1E</figureCitation>
, 
<figureCitation id="5187BAA2FF834F49FBF3E35885FA8399" box="[1142,1181,1663,1689]" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="3" pageNumber="398">2D</figureCitation>
). Thoracic neuropodial postchaetal lobes represented by a flange with an upper papilla, subtriangular (
<figureCitation id="5187BAA2FF834F49FBA8E38485C983BD" box="[1069,1198,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Figs 1C, E</figureCitation>
, 
<figureCitation id="5187BAA2FF834F49FB38E384846E83BD" box="[1213,1289,1699,1725]" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="3" pageNumber="398">2A, D</figureCitation>
). Abdominal notopodia with postchaetal lobes hardly constricted, pyriform. Abdominal neuropodial lobes are represented by a flange with a triangular lobe, shorter than thoracic (
<figureCitation id="5187BAA2FF834F49FD52E3CC82358205" box="[727,850,1771,1797]" captionStart="FIGURE 1" captionStartId="2.[151,250,1498,1522]" captionTargetBox="[151,1435,333,1472]" captionTargetId="figure-61@2.[151,1436,332,1472]" captionTargetPageId="2" captionText="FIGURE 1. Naineris bicornis. Holotype (LACM-AHF Poly 674). A, anterior end, dorsal view; B, thorax, dorsal view; C, thoracic parapodia, lateral view; D, thorax, ventral view; E, transverse section of thoracic segment, posterior view; F, transverse section of abdominal segment, posterior view; G, abdominal parapodia, lateral view. br—branchia, dc—dorsal crest, do—dorsal organs, NeL—neuropodial lobe, NoL—notopodial lobe, on—nuchal organ, pa—papilla, pb—proboscis (Scales: A–F: 1 mm; G: 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234905" httpUri="https://zenodo.org/record/13234905/files/figure.png" pageId="3" pageNumber="398">Figs 1F–G</figureCitation>
, 
<figureCitation id="5187BAA2FF834F49FCDBE3CC82E68205" box="[862,897,1771,1797]" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="3" pageNumber="398">2E</figureCitation>
).
</paragraph>
<paragraph id="C903A627FF834F49FF42E228806382DD" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="398">
Thoracic notochaetae with 20–25 crenulate capillaries in two bundles (
<figureCitation id="5187BAA2FF834F49FC6CE22885238229" box="[1001,1092,1807,1833]" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="3" pageNumber="398">Fig. 2D</figureCitation>
). Abdominal notochaetae with about 20 capillaries and a few furcate chaetae each with unequal tines (
<figureCitation id="5187BAA2FF834F49FC46E214857B824D" box="[963,1052,1843,1869]" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="3" pageNumber="398">Fig. 2E</figureCitation>
). Thoracic neurochaetae with 5–6 transverse, posterior rows of numerous subuluncini; a transverse, anterior row of about 20 crenulate capillaries; an inferior, anterior, oblique row of about 20 hooded uncini and an inferior, posterior, oblique row of capillaries (
<figureCitation id="5187BAA2FF834F49FAEFE25C81DB82B9" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="3" pageNumber="398">Figs 2D</figureCitation>
, 
<figureCitation id="5187BAA2FF834F49FF4DE2B8818B82B9" box="[200,236,1951,1977]" captionStart="FIGURE 3" captionStartId="5.[151,250,1297,1321]" captionTargetBox="[152,1436,274,1274]" captionTargetId="figure-78@5.[151,1436,273,1274]" captionTargetPageId="5" captionText="FIGURE 3. Naineris bicornis (USNM 1284137).A, anterior end; B, abdominal segments, dorsal view, multiple dorsal organs; C, thoracic neurochaetae; D, abdominal neurochaetae. a—acicula, cc—crenulate capillaries, do—dorsal organs, pe—peristomium, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 400 μm; B: 100 μm.; C–D: 20 μm)." figureDoi="http://doi.org/10.5281/zenodo.13234909" httpUri="https://zenodo.org/record/13234909/files/figure.png" pageId="3" pageNumber="398">3C</figureCitation>
). Abdominal neurochaetae include three acicular spines (
<figureCitation id="5187BAA2FF834F49FCF5E2B882AF82B9" box="[880,968,1951,1977]" captionStart="FIGURE 3" captionStartId="5.[151,250,1297,1321]" captionTargetBox="[152,1436,274,1274]" captionTargetId="figure-78@5.[151,1436,273,1274]" captionTargetPageId="5" captionText="FIGURE 3. Naineris bicornis (USNM 1284137).A, anterior end; B, abdominal segments, dorsal view, multiple dorsal organs; C, thoracic neurochaetae; D, abdominal neurochaetae. a—acicula, cc—crenulate capillaries, do—dorsal organs, pe—peristomium, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 400 μm; B: 100 μm.; C–D: 20 μm)." figureDoi="http://doi.org/10.5281/zenodo.13234909" httpUri="https://zenodo.org/record/13234909/files/figure.png" pageId="3" pageNumber="398">Fig. 3D</figureCitation>
) and 5–8 crenulate capillaries intermixed (
<figureCitation id="5187BAA2FF834F49FF1AE2E4819082DD" box="[159,247,1987,2013]" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="3" pageNumber="398">Fig. 2E</figureCitation>
).
</paragraph>
<caption id="9DC3F6AFFF844F4EFF12E0DB85B28381" ID-DOI="http://doi.org/10.5281/zenodo.13234907" ID-Zenodo-Dep="13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="4" pageNumber="399" startId="4.[151,250,1532,1556]" targetBox="[152,1426,181,1507]" targetPageId="4" targetType="figure">
<paragraph id="C903A627FF844F4EFF12E0DB85B28381" blockId="4.[151,1436,1532,1665]" pageId="4" pageNumber="399">
<emphasis id="FBC87A35FF844F4EFF12E0DB806C8314" bold="true" box="[151,267,1532,1556]" pageId="4" pageNumber="399">FIGURE 2</emphasis>
. 
<taxonomicName id="0EBCDDA4FF844F4EFE93E0DA80A28314" authorityName="Hartman" authorityYear="1951" box="[278,453,1532,1556]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="4" pageNumber="399" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF844F4EFE93E0DA80A28314" box="[278,453,1532,1556]" italics="true" pageId="4" pageNumber="399">Naineris bicornis</emphasis>
</taxonomicName>
(USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. 
<taxonomicName id="0EBCDDA4FF844F4EFE59E30683EA8338" authorityName="Hartman" authorityYear="1951" box="[476,653,1568,1592]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="4" pageNumber="399" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF844F4EFE59E30683EA8338" box="[476,653,1568,1592]" italics="true" pageId="4" pageNumber="399">Naineris bicornis</emphasis>
</taxonomicName>
(USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm).
</paragraph>
</caption>
<paragraph id="C903A627FF844F4EFF42E3F480CC83ED" blockId="4.[151,1437,1747,2025]" box="[199,427,1747,1773]" pageId="4" pageNumber="399">Pygidium unknown.</paragraph>
</subSubSection>
<subSubSection id="81A6F5ACFF844F4CFF42E3D0809A85D5" lastPageId="6" lastPageNumber="401" pageId="4" pageNumber="399" type="discussion">
<paragraph id="C903A627FF844F4EFF42E3D08328827D" blockId="4.[151,1437,1747,2025]" pageId="4" pageNumber="399">
<emphasis id="FBC87A35FF844F4EFF42E3D080538211" bold="true" box="[199,308,1783,1809]" pageId="4" pageNumber="399">Remarks</emphasis>
: 
<taxonomicName id="0EBCDDA4FF844F4EFEC6E3DF83618211" authorityName="Hartman" authorityYear="1951" box="[323,518,1783,1809]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="4" pageNumber="399" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF844F4EFEC6E3DF83618211" box="[323,518,1783,1809]" italics="true" pageId="4" pageNumber="399">Naineris bicornis</emphasis>
</taxonomicName>
is mainly distinguished by its slightly indented prostomium. Nevertheless, care is required when using the shape of the prostomium as the main diagnostic character. The shape of the prostomium is as it appears in the 
<typeStatus id="16071885FF844F4EFEE9E21880A98259" box="[364,462,1855,1881]" pageId="4" pageNumber="399" type="holotype">holotype</typeStatus>
is only seen occasionally and tends to be spatulate in specimens from other collections, probably due to fixation (
<figureCitation id="5187BAA2FF844F4EFE31E2448376827D" box="[436,529,1891,1917]" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="4" pageNumber="399">Figs 2A</figureCitation>
, 
<figureCitation id="5187BAA2FF844F4EFD98E2448325827D" box="[541,578,1891,1917]" captionStart="FIGURE 3" captionStartId="5.[151,250,1297,1321]" captionTargetBox="[152,1436,274,1274]" captionTargetId="figure-78@5.[151,1436,273,1274]" captionTargetPageId="5" captionText="FIGURE 3. Naineris bicornis (USNM 1284137).A, anterior end; B, abdominal segments, dorsal view, multiple dorsal organs; C, thoracic neurochaetae; D, abdominal neurochaetae. a—acicula, cc—crenulate capillaries, do—dorsal organs, pe—peristomium, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 400 μm; B: 100 μm.; C–D: 20 μm)." figureDoi="http://doi.org/10.5281/zenodo.13234909" httpUri="https://zenodo.org/record/13234909/files/figure.png" pageId="4" pageNumber="399">3A</figureCitation>
).
</paragraph>
<paragraph id="C903A627FF844F4FFF42E2A0840885F9" blockId="4.[151,1437,1747,2025]" lastBlockId="5.[151,1437,151,249]" lastPageId="5" lastPageNumber="400" pageId="4" pageNumber="399">
Paired, dark segmental spots at bases of branchiae were not observed in the 
<typeStatus id="16071885FF844F4EFBCBE2A085D782A1" box="[1102,1200,1927,1953]" pageId="4" pageNumber="399" type="holotype">holotype</typeStatus>
but were present in chaetigers 
<quantity id="0E440BC2FF844F4EFE94E28C803D82C5" box="[273,346,1963,1990]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="4" pageNumber="399" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">5–6 in</quantity>
the additional specimens examined from the type locality or vicinity. They seem to correspond to internal structures that are transparent through the epidermis and must not be confused with the dorsal sensory organs (
<figureCitation id="5187BAA2FF854F4FFF75E5B0800B85B1" box="[240,364,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="5" pageNumber="400">Fig. 2A–B</figureCitation>
). These dark, segmental spots were also present in the juveniles of 
<taxonomicName id="0EBCDDA4FF854F4FFBDDE5B085BD85B1" authorityName="Hartman" authorityYear="1957" box="[1112,1242,151,177]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="australis">
<emphasis id="FBC87A35FF854F4FFBDDE5B085BD85B1" box="[1112,1242,151,177]" italics="true" pageId="5" pageNumber="400">N. australis</emphasis>
</taxonomicName>
, the most similar species (see 
<bibRefCitation id="AD2DDBD6FF854F4FFEA3E59C836685D5" author="Zhadan, A. &amp; Stupnikova, A. &amp; Neretina, T." box="[294,513,186,213]" pageId="5" pageNumber="400" pagination="773 - 801" refId="ref9916" refString="Zhadan, A., Stupnikova, A. &amp; Neretina, T. (2015) Orbiniidae (Annelida: Errantia) from Lizard Island, Great Barrier Reef, Australia with notes on orbiniid phylogeny. Zootaxa, 4019 (1), 773 - 801. https: // doi. org / 10.11646 / zootaxa. 4019.1.27" type="journal article" year="2015">
Zhadan 
<emphasis id="FBC87A35FF854F4FFE06E59C80D185D5" box="[387,438,187,213]" italics="true" pageId="5" pageNumber="400">et al</emphasis>
. 2015
</bibRefCitation>
, fig. 12D). In addition, dorsal sensory organs were not originally described in 
<taxonomicName id="0EBCDDA4FF854F4FFA06E59C819485F9" authorityName="Hartman" authorityYear="1951" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF854F4FFA06E59C819485F9" italics="true" pageId="5" pageNumber="400">N. bicornis</emphasis>
</taxonomicName>
. As in 
<taxonomicName id="0EBCDDA4FF854F4FFEC7E5F880A285F9" authorityName="Hartman" authorityYear="1957" box="[322,453,223,249]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="australis">
<emphasis id="FBC87A35FF854F4FFEC7E5F880A285F9" box="[322,453,223,249]" italics="true" pageId="5" pageNumber="400">N. australis</emphasis>
</taxonomicName>
, they are multiple (
<figureCitation id="5187BAA2FF854F4FFD25E5F8827B85F9" box="[672,796,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1532,1556]" captionTargetBox="[152,1426,181,1507]" captionTargetId="figure-21@4.[151,1436,181,1508]" captionTargetPageId="4" captionText="FIGURE 2. Naineris bicornis (USNM 1284134) (Shirlastain stained).A, anterior end, dorsal view; B, thorax, dorsal view; C, last thoracic segments, dorsal view. Naineris bicornis (USNM 090156). D, thoracic chaetiger; E, abdominal chaetiger. a—acicular spines, br—branchiae, cc—crenulate capillaries, do—dorsal organs, NoL—Abdominal notopodial lobe, pa—neuropodial papilla, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 0.5 mm; B–C: 0.25 mm; D–E: 0.25 mm)." figureDoi="http://doi.org/10.5281/zenodo.13234907" httpUri="https://zenodo.org/record/13234907/files/figure.png" pageId="5" pageNumber="400">Figs 2B–C</figureCitation>
, 
<figureCitation id="5187BAA2FF854F4FFCADE5F8822E85F9" box="[808,841,223,249]" captionStart="FIGURE 3" captionStartId="5.[151,250,1297,1321]" captionTargetBox="[152,1436,274,1274]" captionTargetId="figure-78@5.[151,1436,273,1274]" captionTargetPageId="5" captionText="FIGURE 3. Naineris bicornis (USNM 1284137).A, anterior end; B, abdominal segments, dorsal view, multiple dorsal organs; C, thoracic neurochaetae; D, abdominal neurochaetae. a—acicula, cc—crenulate capillaries, do—dorsal organs, pe—peristomium, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 400 μm; B: 100 μm.; C–D: 20 μm)." figureDoi="http://doi.org/10.5281/zenodo.13234909" httpUri="https://zenodo.org/record/13234909/files/figure.png" pageId="5" pageNumber="400">3B</figureCitation>
) and difficult to see in old, preserved specimens.
</paragraph>
<caption id="9DC3F6AFFF854F4FFF12E036852F8072" ID-DOI="http://doi.org/10.5281/zenodo.13234909" ID-Zenodo-Dep="13234909" httpUri="https://zenodo.org/record/13234909/files/figure.png" pageId="5" pageNumber="400" startId="5.[151,250,1297,1321]" targetBox="[152,1436,274,1274]" targetPageId="5" targetType="figure">
<paragraph id="C903A627FF854F4FFF12E036852F8072" blockId="5.[151,1437,1297,1394]" pageId="5" pageNumber="400">
<emphasis id="FBC87A35FF854F4FFF12E036806C8029" bold="true" box="[151,267,1297,1321]" pageId="5" pageNumber="400">FIGURE 3</emphasis>
. 
<taxonomicName id="0EBCDDA4FF854F4FFE93E03580A3802A" authorityName="Hartman" authorityYear="1951" box="[278,452,1298,1322]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF854F4FFE93E03580A3802A" box="[278,452,1298,1322]" italics="true" pageId="5" pageNumber="400">Naineris bicornis</emphasis>
</taxonomicName>
(USNM 1284137). A, anterior end; B, abdominal segments, dorsal view, multiple dorsal organs; C, thoracic neurochaetae; D, abdominal neurochaetae. a—acicula, cc—crenulate capillaries, do—dorsal organs, pe—peristomium, pr—prostomium, su—subuluncini, u—uncini (Scales: A: 400 μm; B: 100 μm.; C–D: 20 μm).
</paragraph>
</caption>
<paragraph id="C903A627FF854F4FFF42E0E3840383DA" blockId="5.[151,1437,1476,2042]" pageId="5" pageNumber="400">
The chaetal arrangements of the thoracic neuropodia of 
<taxonomicName id="0EBCDDA4FF854F4FFCC2E0E282A280DE" authorityName="Hartman" authorityYear="1951" box="[839,965,1476,1502]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF854F4FFCC2E0E282A280DE" box="[839,965,1476,1502]" italics="true" pageId="5" pageNumber="400">N. bicornis</emphasis>
</taxonomicName>
, 
<taxonomicName id="0EBCDDA4FF854F4FFC51E0E2853E80DE" authorityName="Hartman" authorityYear="1957" box="[980,1113,1476,1502]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="australis">
<emphasis id="FBC87A35FF854F4FFC51E0E2853E80DE" box="[980,1113,1476,1502]" italics="true" pageId="5" pageNumber="400">N. australis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="0EBCDDA4FF854F4FFB1CE0E2847580DE" authorityName="Álvarez &amp; Haggin" authorityYear="2024" box="[1177,1298,1476,1502]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="elegans" status="sp. nov.">
<emphasis id="FBC87A35FF854F4FFB1CE0E2847580DE" box="[1177,1298,1476,1502]" italics="true" pageId="5" pageNumber="400">N. elegans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0FBC74EFF854F4FFA9FE0E3841580DE" box="[1306,1394,1476,1502]" pageId="5" pageNumber="400" rank="species">sp. nov.</taxonomicNameLabel>
are all similar. It includes the subuluncini, crenulate capillaries, and hooded uncini. The characters distinguishing 
<taxonomicName id="0EBCDDA4FF854F4FFF12E32A80708326" authorityName="Hartman" authorityYear="1951" box="[151,279,1548,1574]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF854F4FFF12E32A80708326" box="[151,279,1548,1574]" italics="true" pageId="5" pageNumber="400">N. bicornis</emphasis>
</taxonomicName>
from other similar species is based mainly on the features of thoracic notopodial lobes, distribution of multiple dorsal organs, geographic distribution and bathymetry. Thoracic notopodial lobes may be acute-to-rounded-tipped and possess a constriction in the middle, as in 
<taxonomicName id="0EBCDDA4FF854F4FFCE2E3728281836E" authorityName="Hartman" authorityYear="1951" box="[871,998,1620,1646]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF854F4FFCE2E3728281836E" box="[871,998,1620,1646]" italics="true" pageId="5" pageNumber="400">N. bicornis</emphasis>
</taxonomicName>
. Likewise, dorsal organs they may be grouped in rows or be irregularly distributed and may cross the midline depending on the species. Concerning their distribution, 
<taxonomicName id="0EBCDDA4FF854F4FFEA3E3BA808D83B6" authorityName="Hartman" authorityYear="1951" box="[294,490,1692,1718]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF854F4FFEA3E3BA808D83B6" box="[294,490,1692,1718]" italics="true" pageId="5" pageNumber="400">Naineris bicornis</emphasis>
</taxonomicName>
is distributed mostly in the Tropical Atlantic, whereas the other species are from the Pacific. Finally, 
<taxonomicName id="0EBCDDA4FF854F4FFECBE3E680AC83DA" authorityName="Hartman" authorityYear="1951" box="[334,459,1728,1754]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF854F4FFECBE3E680AC83DA" box="[334,459,1728,1754]" italics="true" pageId="5" pageNumber="400">N. bicornis</emphasis>
</taxonomicName>
occurs at continental shelf depths, whereas the other similar species are intertidal.
</paragraph>
<paragraph id="C903A627FF854F4FFF42E3C380BC82B2" blockId="5.[151,1437,1476,2042]" pageId="5" pageNumber="400">
Recent molecular data have shown that 
<taxonomicName id="0EBCDDA4FF854F4FFDFAE3C2839C83FE" authorityName="Hartman" authorityYear="1951" box="[639,763,1764,1790]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF854F4FFDFAE3C2839C83FE" box="[639,763,1764,1790]" italics="true" pageId="5" pageNumber="400">N. bicornis</emphasis>
</taxonomicName>
is a species complex, and at least two species occur in 
<collectingCountry id="B1ABE6B7FF854F4FFADDE3C384FB83FE" box="[1368,1436,1764,1790]" name="Brazil" pageId="5" pageNumber="400">Brazil</collectingCountry>
(RA unpublished data). Molecular characterization of specimens from the 
<typeStatus id="16071885FF854F4FFC5FE22F856D8222" box="[986,1034,1800,1826]" pageId="5" pageNumber="400">type</typeStatus>
locality and Brazilian records of 
<taxonomicName id="0EBCDDA4FF854F4FFA06E22E81948246" authorityName="Hartman" authorityYear="1951" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF854F4FFA06E22E81948246" italics="true" pageId="5" pageNumber="400">N. bicornis</emphasis>
</taxonomicName>
are necessary, but they are outside the scope of this study. This species was recorded only once in 
<collectingCountry id="B1ABE6B7FF854F4FFADDE20B84FB8246" box="[1368,1436,1836,1862]" name="Brazil" pageId="5" pageNumber="400">Brazil</collectingCountry>
(
<bibRefCitation id="AD2DDBD6FF854F4FFF1AE2778053826A" author="Nonato, E. F." box="[159,308,1872,1898]" pageId="5" pageNumber="400" refId="ref9281" refString="Nonato, E. F. (1981) Contribuicao ao conhecimento dos anelideos poliquetas bentonicos da Plataforma Continental Brasileira, entre Cabo Frio e o Arroio Chui. Tese de Livre Docencia, Instituto Oceanografico, Universidade de Sao Paulo, 248 pp." type="book" year="1981">Nonato 1981</bibRefCitation>
) and was not abundant in the area (
<bibRefCitation id="AD2DDBD6FF854F4FFD49E27782CE826A" author="Amaral, A. C. Z. &amp; Nallin, S. A. H. &amp; Steiner, T. M. &amp; Forroni, T. O. &amp; Gomes-Filho, D. &amp; Araujo, G. R. &amp; Freitas, R. &amp; Costa, C. A. O. &amp; Ruta, C. &amp; Gomes, K. R. E. &amp; Bonaldo, R. O." box="[716,937,1872,1898]" pageId="5" pageNumber="400" refId="ref8022" refString="Amaral, A. C. Z., Nallin, S. A. H., Steiner, T. M., Forroni, T. O., Gomes-Filho, D., Araujo, G. R., Freitas, R., Costa, C. A. O., Ruta, C., Gomes, K. R. E. &amp; Bonaldo, R. O. (2022) Catalogo das especies de Annelida &quot; Polychaeta &quot; do Brasil. Available from: https: // intranet. ib. unicamp. br / intranet / polychaeta / _ apresentacao. php (accessed 10 March 2024)" type="url" year="2022">
Amaral 
<emphasis id="FBC87A35FF854F4FFCAEE276823A826A" box="[811,861,1872,1898]" italics="true" pageId="5" pageNumber="400">et al</emphasis>
. 2022
</bibRefCitation>
). Recent sampling along the Brazilian coast resulted in 
<specimenCount id="DFBA6DAEFF854F4FFE9DE25380A4828E" box="[280,451,1908,1934]" count="2" pageId="5" pageNumber="400" type="generic">two specimens</specimenCount>
, which were similar to 
<taxonomicName id="0EBCDDA4FF854F4FFD57E2528236828E" authorityName="Hartman" authorityYear="1951" box="[722,849,1908,1934]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF854F4FFD57E2528236828E" box="[722,849,1908,1934]" italics="true" pageId="5" pageNumber="400">N. bicornis</emphasis>
</taxonomicName>
, but with consistent molecular differences between them (RA unpublished data).
</paragraph>
<paragraph id="C903A627FF854F4CFF42E29B809A85D5" blockId="5.[151,1437,1476,2042]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="401" pageId="5" pageNumber="400">
We also examined the 
<typeStatus id="16071885FF854F4FFE65E29B832582D6" box="[480,578,1980,2006]" pageId="5" pageNumber="400" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="0EBCDDA4FF854F4FFDF0E29A85B482D7" authority="Hartmann-Schroder, 1965" authorityName="Hartmann-Schroder" authorityYear="1965" box="[629,1235,1980,2007]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="5" pageNumber="400" phylum="Annelida" rank="subSpecies" species="bicornis" subSpecies="minuta">
<emphasis id="FBC87A35FF854F4FFDF0E29A82FC82D6" box="[629,923,1980,2006]" italics="true" pageId="5" pageNumber="400">Naineris bicornis minuta</emphasis>
<bibRefCitation id="AD2DDBD6FF854F4FFC2CE29B85B482D7" author="Hartmann-Schroder, G." box="[937,1235,1980,2007]" pageId="5" pageNumber="400" pagination="81 - 161" refId="ref9016" refString="Hartmann-Schroder, G. (1965) Zur kenntnis der eulitoralen polychaetenfauna von Hawaii, Palmyra und Samoa. Abhandlungen und Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg, 9, 81 - 161." type="journal article" year="1965">Hartmann-Schröder, 1965</bibRefCitation>
</taxonomicName>
(ZMH P-14869, type locality: Oahu, 
<collectingRegion id="0B7868C5FF854F4FFE07E2C780BF82FA" box="[386,472,2016,2042]" country="United States of America" name="Hawaii" pageId="5" pageNumber="400">Hawaii</collectingRegion>
). Because of the juvenile morphology of the 
<typeStatus id="16071885FF854F4FFC5BE2C7852482FA" box="[990,1091,2016,2042]" pageId="5" pageNumber="400" type="holotype">holotype</typeStatus>
, it was impossible to find any morphological differences with 
<taxonomicName id="0EBCDDA4FF864F4CFD87E5B083E785B1" authorityName="Hartman" authorityYear="1951" box="[514,640,151,177]" class="Polychaeta" family="Orbiniidae" genus="Naineris" kingdom="Animalia" pageId="6" pageNumber="401" phylum="Annelida" rank="species" species="bicornis">
<emphasis id="FBC87A35FF864F4CFD87E5B083E785B1" box="[514,640,151,177]" italics="true" pageId="6" pageNumber="401">N. bicornis</emphasis>
</taxonomicName>
. Further studies covering different oceans are necessary to clarify the species remaining in this group.
</paragraph>
</subSubSection>
<subSubSection id="81A6F5ACFF864F4CFF42E5F985D1841D" pageId="6" pageNumber="401" type="distribution">
<paragraph id="C903A627FF864F4CFF42E5F985D1841D" blockId="6.[151,1436,151,285]" pageId="6" pageNumber="401">
<emphasis id="FBC87A35FF864F4CFF42E5F9803E85F8" bold="true" box="[199,345,222,248]" pageId="6" pageNumber="401">Distribution</emphasis>
. Warm Temperate Northwest Atlantic province: Northern Gulf of 
<collectingRegion id="0B7868C5FF864F4CFBBBE5F885F385F9" box="[1086,1172,223,249]" country="Mexico" name="Mexico" pageId="6" pageNumber="401">Mexico</collectingRegion>
ecoregion and Tropical 
<collectingRegion id="0B7868C5FF864F4CFF12E4248048841D" box="[151,303,259,285]" country="Sri Lanka" name="North Western" pageId="6" pageNumber="401">Northwestern</collectingRegion>
Atlantic province: Floridian and Southern Gulf of 
<collectingRegion id="0B7868C5FF864F4CFCECE42482D8841D" box="[873,959,259,285]" country="Mexico" name="Mexico" pageId="6" pageNumber="401">Mexico</collectingRegion>
ecoregions, 
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11– 
<specimenCount id="DFBA6DAEFF864F4CFBF1E42585D5841D" box="[1140,1202,258,285]" count="54" pageId="6" pageNumber="401" type="generic">54 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>