treatments-xml/data/31/FE/63/31FE631E24D607EA5C94870BF77D6620.xml

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.66.684" ID-GBIF-Dataset="35e99d78-af9c-400e-b111-6abc6e1d9b1d" ID-PMC="PMC3088417" ID-Pensoft-Pub="1313-2970-66-1" ID-Pensoft-UUID="AD1FFFAF9122FFEAFF8EEF3BE966F439" ID-PubMed="21594029" ID-Zenodo-Dep="576888" ID-ZooBank="176306EB63424E75AD76C4A82040A002" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-66-1" ModsDocOrigin="ZooKeys 66" ModsDocTitle="Agra, arboreal beetles of Neotropical forests:pusilla group and piranha group systematics and notes on their ways of life (Coleoptera, Carabidae, Lebiini, Agrina)" checkinTime="1451250911506" checkinUser="pensoft" docAuthor="Erwin, Terry L." docDate="2010" docId="31FE631E24D607EA5C94870BF77D6620" docLanguage="en" docName="ZooKeys 66: 1-28" docOrigin="ZooKeys 66" docSource="http://dx.doi.org/10.3897/zookeys.66.684" docTitle="Agra ce Erwin 2010, sp. n." docType="treatment" docUuid="6A25E44D-7927-4FD0-9FD2-9B37E12458D7" docUuidSource="ZooBank" docVersion="7" id="AD1FFFAF9122FFEAFF8EEF3BE966F439" lastPageNumber="16" masterDocId="AD1FFFAF9122FFEAFF8EEF3BE966F439" masterDocTitle="Agra, arboreal beetles of Neotropical forests: pusilla group and piranha group systematics and notes on their ways of life (Coleoptera, Carabidae, Lebiini, Agrina)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="15" updateTime="1732481568501" updateUser="ExternalLinkService">
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<mods:title id="AB4B0DE5EDA41ABBD7B864DCE0760DD0">Agra, arboreal beetles of Neotropical forests: pusilla group and piranha group systematics and notes on their ways of life (Coleoptera, Carabidae, Lebiini, Agrina)</mods:title>
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<mods:namePart id="D1AFF38FD93CE0CEB6B10505F8539F3B">Erwin, Terry L.</mods:namePart>
<mods:affiliation id="D0CAD547EC750F1B4A5A343674999BB4">Washington DC</mods:affiliation>
<mods:nameIdentifier id="9C17CC9AA18DC63CC087C7375E3FB438" type="email">erwint@si.edu</mods:nameIdentifier>
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<mods:classification id="9F784AA9798A66DB86F98D35F0A44B01">journal article</mods:classification>
<mods:identifier id="2B4A65A328C0E9401F9C3C853C4A7D7B" type="DOI">http://dx.doi.org/10.3897/zookeys.66.684</mods:identifier>
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<treatment id="31FE631E24D607EA5C94870BF77D6620" ID-GBIF-Taxon="159360648" LSID="urn:lsid:zoobank.org:act:6A25E44D-7927-4FD0-9FD2-9B37E12458D7" httpUri="http://treatment.plazi.org/id/31FE631E24D607EA5C94870BF77D6620" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" scope_family="Carabidae" scope_order="Coleoptera" scope_subTribe="Agrina" scope_tribe="Lebiini">
<subSubSection id="A00A629D02B7AECB5B0D7125C511380D" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="0F1FB63B0B2C956FA08D5ACB5FD7AB64" pageId="14" pageNumber="15">
<taxonomicName id="307930D085ED6684609B2B41005CC443" ID-CoL="B2JL" LSID="31FE631E-24D6-07EA-5C94-870BF77D6620" authority="Erwin" authorityName="Erwin" authorityYear="2010" class="Insecta" family="Carabidae" genus="Agra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agra ce" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ce" status="sp. n.">Agra ce Erwin</taxonomicName>
<taxonomicNameLabel id="D88F8EF5A143742E1D9CBA3B40AFE7A5" pageId="14" pageNumber="15">sp. n.</taxonomicNameLabel>
<figureCitation id="116237DC32F8937B78F1AEB692BEE967" captionStart="Figure 11" captionStartId="F7" captionText="Figure 11. Distribution map of the species of the piranha group." httpUri="https://binary.pensoft.net/fig/14830" pageId="14" pageNumber="15">Fig. 11</figureCitation>
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</paragraph>
</subSubSection>
<subSubSection id="51E8AD285BE8EDBFB34E3D9A4E78481E" pageId="14" pageNumber="15" type="holotype">
<paragraph id="B15515FDF18E4796CA8F92094DA54872" pageId="14" pageNumber="15">Holotype:</paragraph>
<paragraph id="94016B820DB2A97DE55B44D73364F16D" pageId="14" pageNumber="15">
<emphasis id="1992C541DC5B427688042779552E427A" bold="true" pageId="14" pageNumber="15">
<normalizedToken id="78FB9AD1BC554C43BE1EE52DC8E74A92" originalValue="Perú">Peru</normalizedToken>
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:
</emphasis>
Madre de Dios, 30 air km SW Puerto Maldonado, 205m, &quot;12.8368°S, 69.2933°W,&quot; 10 September 1984 (T.L. Erwin, et al.)(NMNH: ADP 093837, female).
</paragraph>
</subSubSection>
<subSubSection id="D84624EB332DEF8E241D6486EE3AA5B8" pageId="14" pageNumber="15" type="derivation of specific epithet">
<paragraph id="62C86311F791B303F6A095B830DDC7B5" pageId="14" pageNumber="15">Derivation of specific epithet.</paragraph>
<paragraph id="9C81290B12C2645D307A7AB0E3B9C728" pageId="14" pageNumber="15">
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The epithet &quot;
<emphasis id="6143770F66D528E5EC3AD9AC77284BBC" italics="true" pageId="14" pageNumber="15">ce</emphasis>
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&quot; is a combination of pronounceable letters that when joined with the last three letters of the genus name,
<taxonomicName id="CEC425D5B7B773D245E43A61523A33BD" authorityName="Fabricius" authorityYear="1801" class="Insecta" family="Carabidae" genus="Agra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Agra</taxonomicName>
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, spells &quot;grace,&quot; for the Peruvian Ornithologist, Grace Servat, who has shared the lowland Amazon and the high Andes with me for many years, including the known localities of this species.
</paragraph>
</subSubSection>
<subSubSection id="C22F7C13EB76C521B609341B68B8B157" pageId="14" pageNumber="15" type="proposed english vernacular name">
<paragraph id="459934DACF40EBABE7CE2CC49823EB3E" pageId="14" pageNumber="15">Proposed English vernacular name.</paragraph>
<paragraph id="81EC4AA012F9EBE8FE0382A73462E318" pageId="14" pageNumber="15">Graceful Elegant Canopy Beetle.</paragraph>
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</subSubSection>
<subSubSection id="3FF8792E04B77F2C681334DC625D38BC" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="85363FB0F29316C73422CB3AEAC547D6" pageId="14" pageNumber="15">Diagnosis.</paragraph>
<paragraph id="2C83C1DAD3A31B6C679E6BE9868F385F" pageId="14" pageNumber="15">With the attributes of the genus and species-group as described above and elytra and prothorax metallic blue, legs unicolored, frons laterad slightly rugose, prothorax markedly setiferous both laterally and ventrally, and elytra barely constricted at apical third, side margin slightly arcuate.</paragraph>
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</subSubSection>
<subSubSection id="5FCF75196D2D25F8E34BAB7A7A1A5EEC" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="17BCC6E069102E8C97B9E4A5941389CD" pageId="14" pageNumber="15">Description.</paragraph>
<paragraph id="063129051259EEF20F382488E50B1803" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="AEB1081B464B7C10A2396119B3A39436" italics="true" pageId="14" pageNumber="15">Size</emphasis>
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: Small, ABL = 8.04 - 9.21 mm, SBL = 7.09 - 8.07 mm, TW = 2.36 - 2.78 mm.
<emphasis id="C232DBBC6214B3A27FE4DB67F741FF69" italics="true" pageId="14" pageNumber="15">Color:</emphasis>
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Head black with bluish reflection posteriorly, body and legs metallic blue; antennae and mouthparts piceous, scape with slight metallic blue reflections.
<emphasis id="A284FFB06951394C212C33C34D1DA407" italics="true" pageId="14" pageNumber="15">Luster:</emphasis>
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Shiny metallic.
<emphasis id="5F0E412D677A4963FA711F1ABDB6DD00" italics="true" pageId="14" pageNumber="15">Head:</emphasis>
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Labrum moderately elongate and rounded apically. Frons medially raised and smooth, laterally depressed and rugose. Gena almost squared to constricted neck in females. Genae and occiput moderately densely punctate, each puncture setiferous.
<emphasis id="87272C77BB521C34D781C0ED6C612F9B" italics="true" pageId="14" pageNumber="15">Prothorax:</emphasis>
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Slightly broader medially, flared basally; surface with dense and coarse setiferous punctures; lateral elongate callous with single row of setiferous punctures along middle.
<emphasis id="5257E49D65F4DE2365770B17B5AC3FED" italics="true" pageId="14" pageNumber="15">Pterothorax:</emphasis>
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Elytron markedly convex, inte
<pageBreakToken id="90EA3133AD18F55E9239F886C4BBF706" pageId="15" pageNumber="16" start="start">rvals</pageBreakToken>
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moderately costate, interneurs of rows of somewhat laterally ovate punctures, apex truncate, barely lobate, apical dentation asymmetric, lateral tooth small, acute, sutural apex slightly produced, narrowly pointed. Metasternum sparsely setiferous in female.
<emphasis id="6536CE0BA37D94D10EBD4264B1D9DA49" italics="true" pageId="15" pageNumber="16">Legs:</emphasis>
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Normal in female.
<emphasis id="49B01645A639900F80807B218D9B2890" italics="true" pageId="15" pageNumber="16">Abdomen</emphasis>
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: Abdominal sterna III to VII of female moderately and bilaterally setiferous; sternum VII of female barely emarginated, corners rounded.
<emphasis id="7E53B40719EC046429E6798AC98B7C2A" italics="true" pageId="15" pageNumber="16">Male genitalia:</emphasis>
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Unknown.
<emphasis id="96351E531242FD48765484CAE60FAF53" italics="true" pageId="15" pageNumber="16">Female ovipositor:</emphasis>
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Stylomere 2 as in
<taxonomicName id="C0EFB86E9E5D24816B25AAB7FA03E467" authorityName="Erwin" authorityYear="2010" class="Insecta" family="Carabidae" genus="Agra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="piranha">Agra piranha</taxonomicName>
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(
<figureCitation id="AB3C435471FBD4DB96B7C7082132FDB1" captionStart="Figures 89" captionStartId="F5" captionText="Figures 8 - 9. 8 Agra piranha Erwin, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 087440) 9 Agra piranha Erwin, sp. n., female stylomeres (dorsal aspect) (ADP 117227)." httpUri="https://binary.pensoft.net/fig/14828" pageId="15" pageNumber="16">Fig. 9</figureCitation>
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).
</paragraph>
</subSubSection>
<subSubSection id="4C0827AFD9709DF20EA9B9EE1A9B903B" pageId="15" pageNumber="16" type="dispersal potential">
<paragraph id="1D235E92473C53E52D749A4D77767D65" pageId="15" pageNumber="16">Dispersal potential.</paragraph>
<paragraph id="00ACE1161CEDBF3BF46E0B702AD8BCAB" pageId="15" pageNumber="16">These beetles are macropterous and are capable of flight; they are swift and agile runners.</paragraph>
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</subSubSection>
<subSubSection id="59DA397556C276E6005B91331D28DFA4" pageId="15" pageNumber="16" type="way of life">
<paragraph id="05F1BB09CC14A5A0C9F8DBFF156AC07B" pageId="15" pageNumber="16">Way of life.</paragraph>
<paragraph id="D2791353918E9AF48F8D2DB82A7BFB7E" pageId="15" pageNumber="16">
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Adults of other
<taxonomicName id="2B6456029E9EF49D00EA083229EF378F" authorityName="Fabricius" authorityYear="1801" class="Insecta" family="Carabidae" genus="Agra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Agra</taxonomicName>
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species are found in the canopy of rainforest trees; larvae of this genus are found under the bark of the these trees, however they must also roam on the surface, as they have been collected by insecticidal fogging techniques in the very early morning before first light. Members of
<taxonomicName id="E62C7121EB181D3B32E6B246C9B89A5B" authorityName="Erwin" authorityYear="2010" class="Insecta" family="Carabidae" genus="Agra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ce">Agra ce</taxonomicName>
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occur at lowland altitudes in the Amazon Basin. Adults are active in September, the late dry season. The holotype was collected in an Erwin Plot at the type locality; the forest of this plot is designated as a Swamp Forest with internal drainage (
<bibRefCitation id="4EE8364C7A081D589F426075B96D0569" author="Erwin, TL" journalOrPublisher="Revista Peruana de Entomologia" pageId="22" pageNumber="23" pagination="1 - 8" refId="B10" refString="Erwin, TL, 1985. Tambopata Reserved Zone, Madre de Dios, Peru: History and description of the reserve. Revista Peruana de Entomologia 27: 1 - 8" title="Tambopata Reserved Zone, Madre de Dios, Peru: History and description of the reserve." volume="27" year="1985">Erwin 1985</bibRefCitation>
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) and is dominated by the palm
<taxonomicName id="DC4D660206C1FBC336110E1EE857F908" class="Liliopsida" family="Arecaceae" genus="Mauritia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Arecales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="flexuosa">Mauritia flexuosa</taxonomicName>
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L. and the hardwood tree
<taxonomicName id="2FA76A0149172A40C583544A952139FD" authorityName="Burret" authorityYear="1926" class="Magnoliopsida" family="Malvaceae" genus="Lueheopsis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Malvales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="hoehnei">Lueheopsis hoehnei</taxonomicName>
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Burret. The holotype was fogged from the later named species. The paratype was attracted to MV light.
</paragraph>
</subSubSection>
<subSubSection id="45CCABC305D7BB873D35CB10FC4C9718" pageId="15" pageNumber="16" type="other specimens examined">
<paragraph id="FDF7B40160675FA091E3C281BA486B8E" pageId="15" pageNumber="16">Other specimens examined.</paragraph>
<paragraph id="5BA1805522A5638B469F6037364EBB36" pageId="15" pageNumber="16">
<emphasis id="C631A7F5E84CB41A3451A57D23C02B4B" bold="true" pageId="15" pageNumber="16">
<normalizedToken id="88D127406BDC51F1683A8FBF5B4C4D1A" originalValue="Perú">Peru</normalizedToken>
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</emphasis>
: Madre de Dios, Pakitza, Trocha Uno /14, 324m, &quot;11.9352°S, 71.3039°W,&quot; 8 September 1989 (R.A. Faitoute, et al.)(NMNH: BIOLAT 017465, female paratype).
</paragraph>
</subSubSection>
<subSubSection id="2847BBECEFF33140E06F6E6EAAE60588" pageId="15" pageNumber="16" type="geographic distribution">
<paragraph id="DA2BC84998790D00C96969B4A1232C84" pageId="15" pageNumber="16">Geographic distribution.</paragraph>
<paragraph id="E505C31F019D0CFC683B9A2AF3950C59" pageId="15" pageNumber="16">
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(
<figureCitation id="7865CAF32D28BE442BDC080CAD5AE9E7" captionStart="Figure 11" captionStartId="F7" captionText="Figure 11. Distribution map of the species of the piranha group." httpUri="https://binary.pensoft.net/fig/14830" pageId="15" pageNumber="16">Fig. 11</figureCitation>
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). This species is currently known only from two localities in southeastern
<normalizedToken id="54008A44AF2FB9EE66883D60C39CBA9A" originalValue="Perú">Peru</normalizedToken>
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.
</paragraph>
</subSubSection>
</treatment>
</document>