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<document id="664249C3EF07AFA757927C6A4BC13BAE" ID-DOI="10.5281/zenodo.189206" ID-GBIF-Dataset="60cbb643-8df2-44b2-b799-b8b1fe7b61a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="189206" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461100215462" checkinUser="plazi" docAuthor="Hervé, Christophe, Roberts, Michael J. &amp; Murphy, John A." docDate="2009" docId="A552823FBF43FFB2FF7E90D51953ED5A" docLanguage="en" docName="zt02171p028.pdf" docOrigin="Zootaxa 2171" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Drassodex heeri Pavesi 1873" docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="596BFA47BF48FFBFFFE990421B59EC7C" masterDocTitle="A taxonomic revision of the genus Drassodex Murphy, 2007 (Araneae: Gnaphosidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="12" updateTime="1698591093210" updateUser="plazi">
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<mods:title id="6A607BDCD714AB3C5D44BBC5D0BB9627">A taxonomic revision of the genus Drassodex Murphy, 2007 (Araneae: Gnaphosidae)</mods:title>
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<mods:namePart id="6F4B13EAE5A857254FD91B201966C637">Hervé, Christophe</mods:namePart>
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<mods:namePart id="FB53A48A51A763AA9BD62137CB8EA7E7">Roberts, Michael J.</mods:namePart>
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<mods:namePart id="EF26475FAD189F5618B1F8A25B91B438">Murphy, John A.</mods:namePart>
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<mods:title id="5B55640369CCFFABBF870F4610CF8A06">Zootaxa</mods:title>
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<mods:part id="1D590BE185AA5C36D0D494D8E6AE9B70">
<mods:date id="378E972BA9FC91F8FC284E6E02AE03FB">2009</mods:date>
<mods:detail id="9259F370BB1A8FA825D956FAF9D7FC3C" type="volume">
<mods:number id="8E37D8AEB2569F996DA4E71A3444219A">2171</mods:number>
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<treatment id="A552823FBF43FFB2FF7E90D51953ED5A" ID-DOI="http://doi.org/10.5281/zenodo.5663958" ID-GBIF-Taxon="119621568" ID-Zenodo-Dep="5663958" LSID="urn:lsid:plazi:treatment:A552823FBF43FFB2FF7E90D51953ED5A" httpUri="http://treatment.plazi.org/id/A552823FBF43FFB2FF7E90D51953ED5A" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<subSubSection id="65E160A2BF43FFB4FF7E90D51A43ECA8" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="2D443329BF43FFB4FF7E90D51956ECCD" blockId="11.[151,527,151,212]" box="[151,527,151,177]" pageId="11" pageNumber="12">
<heading id="760C8445BF43FFB4FF7E90D51956ECCD" bold="true" box="[151,527,151,177]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="EAFB48AABF43FFB4FF7E90D51956ECCD" ID-CoL="37LS7" authority="Pavesi, 1873" authorityName="Pavesi" authorityYear="1873" box="[151,527,151,177]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="1F8FEF3BBF43FFB4FF7E90D51956ECCD" bold="true" box="[151,527,151,177]" pageId="11" pageNumber="12">
<emphasis id="1F8FEF3BBF43FFB4FF7E90D51A02ECCD" bold="true" box="[151,347,151,177]" italics="true" pageId="11" pageNumber="12">Drassodex heeri</emphasis>
(
<bibRefCitation id="496A4ED8BF43FFB4FE8390D5195FECCD" author="Pavesi" box="[362,518,151,177]" pageId="11" pageNumber="27" refString="Pavesi, P. (1873) Catalogo sistematico dei ragni del cantone ticino con la loro distribuzione orizontale e verticale e cenni sulla araneologica elvetica. Annali del Museo civico di storia naturale di Genova, 4, 5 - 215." type="journal article" year="1873">Pavesi, 1873</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="2D443329BF43FFB4FF7E90F81A43ECA8" blockId="11.[151,527,151,212]" box="[151,282,186,212]" pageId="11" pageNumber="12">
<figureCitation id="B5C02FACBF43FFB4FF7E90F81A43ECA8" box="[151,282,186,212]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="11" pageNumber="12">Figs 10–15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="65E160A2BF43FFB4FF7E90BC1AB7ED89" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="2D443329BF43FFB4FF7E90BC1E64ED69" blockId="11.[151,1437,254,501]" box="[151,1341,254,277]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="0DEB1407BF43FFB4FF7E90BC1E64ED69" box="[151,1341,254,277]" pageId="11" pageNumber="12">
<taxonomicName id="EAFB48AABF43FFB4FF7E90BC1AAAED69" authority="Pavesi, 1873: 126" authorityName="Pavesi" authorityPageNumber="126" authorityYear="1873" box="[151,499,254,277]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="1F8FEF3BBF43FFB4FF7E90BC1A75ED69" box="[151,300,254,277]" italics="true" pageId="11" pageNumber="12">Drassus heeri</emphasis>
<treatmentCitation id="AC5A1538BF43FFB4FEDB90BC1AAAED69" author="Pavesi" box="[306,499,254,277]" page="126" pageId="11" pageNumber="12" year="1873">
<bibRefCitation id="496A4ED8BF43FFB4FEDB90BC1AE7ED69" author="Pavesi" box="[306,446,254,277]" pageId="11" pageNumber="27" refString="Pavesi, P. (1873) Catalogo sistematico dei ragni del cantone ticino con la loro distribuzione orizontale e verticale e cenni sulla araneologica elvetica. Annali del Museo civico di storia naturale di Genova, 4, 5 - 215." type="journal article" year="1873">Pavesi, 1873</bibRefCitation>
: 126
</treatmentCitation>
</taxonomicName>
, fig. 4–6 (male and females
<typeStatus id="F2408D8BBF43FFB4FCC890BC18D8ED69" box="[801,897,254,277]" pageId="11" pageNumber="12" type="syntype">syntypes</typeStatus>
, presumably in Pavia,
<collectingCountry id="55EC73B9BF43FFB4FB9A90BC1FF4ED69" box="[1139,1197,254,277]" name="Italy" pageId="11" pageNumber="12">Italia</collectingCountry>
, not located).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2D443329BF43FFB4FF7E915C196BED49" blockId="11.[151,1437,254,501]" box="[151,562,286,309]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="0DEB1407BF43FFB4FF7E915C196BED49" box="[151,562,286,309]" pageId="11" pageNumber="12">
?
<taxonomicName id="EAFB48AABF43FFB4FF40915C1977ED49" authority="Lebert, 1877: 223" authorityName="Lebert" authorityPageNumber="223" authorityYear="1877" box="[169,558,286,309]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="hispanus">
<emphasis id="1F8FEF3BBF43FFB4FF40915C1A3FED49" box="[169,358,286,309]" italics="true" pageId="11" pageNumber="12">Drassus hispanus</emphasis>
<treatmentCitation id="AC5A1538BF43FFB4FE84915C1977ED49" author="Lebert" box="[365,558,286,309]" page="223" pageId="11" pageNumber="12" year="1877">
<bibRefCitation id="496A4ED8BF43FFB4FE84915C1AA0ED49" author="Lebert" box="[365,505,286,309]" pageId="11" pageNumber="27" refString="Lebert H. (1877) Die Spinnen der Schweiz, ihr Bau, ihr Leben, ihre systematische Uebersicht. Neue Denkschriften der schweizerischen naturforschenden Gesellschaft, Zurich, 23, 1 - 321." type="journal article" year="1877">Lebert, 1877</bibRefCitation>
: 223
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2D443329BF43FFB4FF7E917C1868ED29" blockId="11.[151,1437,254,501]" box="[151,817,318,341]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="0DEB1407BF43FFB4FF7E917C1868ED29" box="[151,817,318,341]" pageId="11" pageNumber="12">
?
<taxonomicName id="EAFB48AABF43FFB4FF40917C1939ED29" authority="Bosenberg, 1902: 291" authorityName="Bosenberg" authorityPageNumber="291" authorityYear="1902" box="[169,608,318,341]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="hypocrita">
<emphasis id="1F8FEF3BBF43FFB4FF40917C1A34ED29" box="[169,365,318,341]" italics="true" pageId="11" pageNumber="12">Drassus hypocrita</emphasis>
<treatmentCitation id="AC5A1538BF43FFB4FE9D917C1939ED29" author="Bosenberg" box="[372,608,318,341]" page="291" pageId="11" pageNumber="12" year="1902">
<bibRefCitation id="496A4ED8BF43FFB4FE9D917C1975ED29" author="Bosenberg" box="[372,556,318,341]" pageId="11" pageNumber="27" refString="Bosenberg, W. (1902) Die Spinnen Deutschlands. II - IV. Zoologica, Stuttgart, 14, 97 - 384." type="journal article" year="1902">Bösenberg, 1902</bibRefCitation>
: 291
</treatmentCitation>
</taxonomicName>
, 292, 296, fig. 434.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2D443329BF43FFB4FF7E911C1BAFEDA9" blockId="11.[151,1437,254,501]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="0DEB1407BF43FFB4FF7E911C1BAFEDA9" pageId="11" pageNumber="12">
<taxonomicName id="EAFB48AABF43FFB4FF7E911C1942ED09" authority="Lessert, 1905: 186" authorityName="Lessert" authorityPageNumber="186" authorityYear="1905" box="[151,539,350,373]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="1F8FEF3BBF43FFB4FF7E911C1A1EED09" box="[151,327,350,373]" italics="true" pageId="11" pageNumber="12">Drassodes heeri</emphasis>
<treatmentCitation id="AC5A1538BF43FFB4FEA6911C1942ED09" author="Lessert" box="[335,539,350,373]" page="186" pageId="11" pageNumber="12" year="1905">
<bibRefCitation id="496A4ED8BF43FFB4FEA6911C1ABCED09" author="Lessert" box="[335,485,350,373]" pageId="11" pageNumber="27" refString="Lessert, R. de (1905) Note sur trois especes d'araignees du genre Drassodes. Revue suisse de Zoologie, 13, 185 - 194." type="journal article" year="1905">Lessert, 1905</bibRefCitation>
: 186
</treatmentCitation>
</taxonomicName>
, figs 1–3;
<treatmentCitation id="AC5A1538BF43FFB4FD78911C1809ED09" author="Lessert" box="[657,848,350,373]" page="50" pageId="11" pageNumber="12" year="1910">
<bibRefCitation id="496A4ED8BF43FFB4FD78911C187FED09" author="Lessert" box="[657,806,350,373]" pageId="11" pageNumber="27" refString="Lessert, R. de (1910) Catalogue des invertebres de la Suisse. Fascicule 3. Araignees. Musee d'histoire naturelle de Geneve, Geneve, 635 pp." type="book" year="1910">Lessert, 1910</bibRefCitation>
: 50
</treatmentCitation>
, figs 41–43;
<treatmentCitation id="AC5A1538BF43FFB4FC36911C1FFAED09" author="Simon" box="[991,1187,350,373]" page="133" pageId="11" pageNumber="12" year="1914">
<bibRefCitation id="496A4ED8BF43FFB4FC36911C1F34ED09" author="Simon" box="[991,1133,350,373]" pageId="11" pageNumber="27" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Roret, Paris, 6, 1 - 308." type="journal article" year="1914">Simon, 1914</bibRefCitation>
: 133
</treatmentCitation>
, 207, figs 219–222 (in part);
<treatmentCitation id="AC5A1538BF43FFB4FEE1913C1AB0EDE9" author="Reimoser" box="[264,489,382,405]" page="161" pageId="11" pageNumber="12" year="1919">
<bibRefCitation id="496A4ED8BF43FFB4FEE1913C1AEAEDE9" author="Reimoser" box="[264,435,382,405]" pageId="11" pageNumber="27" refString="Reimoser, E. (1919) Katalog der echten Spinnen (Araneae) des Palaarktischen Gebietes. Abhandlungen der zoologischbotanischen Gesellschaft in Wien, 10 (2), 1 - 280." type="journal article" year="1919">Reimoser, 1919</bibRefCitation>
: 161
</treatmentCitation>
;
<treatmentCitation id="AC5A1538BF43FFB4FE1D913C199AEDE9" author="Roewer" box="[500,707,382,405]" page="386" pageId="11" pageNumber="12" year="1954">
<bibRefCitation id="496A4ED8BF43FFB4FE1D913C19D4EDE9" author="Roewer" box="[500,653,382,405]" pageId="11" pageNumber="27" refString="Roewer, C. F. (1954) Katalog der Araneae von 1758 bis 1940. Bruxelles, 2 (a), 1 - 923." type="journal article" year="1954">Roewer, 1954</bibRefCitation>
: 386
</treatmentCitation>
;
<treatmentCitation id="AC5A1538BF43FFB4FD27913C18FDEDE9" author="Bonnet" box="[718,932,382,405]" page="1566" pageId="11" pageNumber="12" year="1956">
<bibRefCitation id="496A4ED8BF43FFB4FD27913C183BEDE9" author="Bonnet" box="[718,866,382,405]" pageId="11" pageNumber="27" refString="Bonnet, P. (1956) Bibliographia araneorum. Douladoure, Toulouse, 2 (2), 919 - 1926." type="journal article" year="1956">Bonnet, 1956</bibRefCitation>
: 1566
</treatmentCitation>
; Wiehle, 1967: 13, figs 48–53;?
<treatmentCitation id="AC5A1538BF43FFB4FAE0913C1A71EDC9" author="Tyschchenko" page="95" pageId="11" pageNumber="12" year="1971">
<bibRefCitation id="496A4ED8BF43FFB4FAE0913C1A59EDC9" author="Tyschchenko" pageId="11" pageNumber="28" refString="Tyschchenko, V. P. (1971) Opredelitel' paukov evropejskoj casti SSSR. Nauka, Leningrad, 281 pp. Westring, N. (1851) Forteckning ofver de till narvarande tid Kande, i Sverige forekommande Spindlarter, utgorande ett antal af 253, deraf 132 aro nya for svenska Faunan. Goteborgs Kongl. Vetenskaps- och Vitterhets-Samhalles Handlingar, 2, 25 - 62." type="book chapter" year="1971">Tyschchenko, 1971</bibRefCitation>
: 95
</treatmentCitation>
;
<treatmentCitation id="AC5A1538BF43FFB4FEDC91DC1AA7EDC9" author="Grimm" box="[309,510,414,437]" page="109" pageId="11" pageNumber="12" year="1985">
<bibRefCitation id="496A4ED8BF43FFB4FEDC91DC1A90EDC9" author="Grimm" box="[309,457,414,437]" pageId="11" pageNumber="27" refString="Grimm, U. (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF), 26, 1 - 318." type="journal article" year="1985">Grimm, 1985</bibRefCitation>
: 109
</treatmentCitation>
, figs 122, 130–131 (in part);
<treatmentCitation id="AC5A1538BF43FFB4FCD591DC1FD0EDC9" author="Heimer" box="[828,1161,414,437]" page="418" pageId="11" pageNumber="12" year="1991">
<bibRefCitation id="496A4ED8BF43FFB4FCD591DC1F0AEDC9" author="Heimer" box="[828,1107,414,437]" pageId="11" pageNumber="27" refString="Heimer, S. &amp; Nentwig, W. (1991) Spinnen Mitteleuropas: Ein Bestimmungsbuch. Verlag Paul Parey, Berlin, 543 pp." type="book" year="1991">Heimer &amp; Nentwig, 1991</bibRefCitation>
: 418
</treatmentCitation>
, fig. 1104;
<treatmentCitation id="AC5A1538BF43FFB4FAED91DC1BABEDA9" author="Canard" page="190" pageId="11" pageNumber="12" year="2005">
<bibRefCitation id="496A4ED8BF43FFB4FAED91DC1EC1EDC9" author="Canard" box="[1284,1432,414,437]" pageId="11" pageNumber="27" refString="Canard, A. (2005) Catalogue of spiders species from Europe and the mediterranean basin. Parts I &amp; II. Revue Arachnologique, 15 (3), 1 - 255." type="journal article" year="2005">Canard, 2005</bibRefCitation>
: 190
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2D443329BF43FFB4FF7E919C1AB7ED89" blockId="11.[151,1437,254,501]" box="[151,494,478,501]" pageId="11" pageNumber="12">
<taxonomicName id="EAFB48AABF43FFB4FF7E919C1AB2ED89" ID-CoL="37LS7" authority="Platnick, 2008" authorityName="Platnick" authorityYear="2008" box="[151,491,478,501]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="1F8FEF3BBF43FFB4FF7E919C1A1FED89" box="[151,326,478,501]" italics="true" pageId="11" pageNumber="12">Drassodex heeri</emphasis>
<bibRefCitation id="496A4ED8BF43FFB4FEA4919C1AB2ED89" author="Platnick" box="[333,491,478,501]" pageId="11" pageNumber="27" refString="Platnick, N. I. (2008) The world spider catalog, version 9.0 American Museum of Natural History, online at http: // research. amnh. org / entomology / spiders / catalog / index. html (Accessed 15 September 2008)" type="journal article" year="2008">Platnick, 2008</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="65E160A2BF43FFB4FF7E92591949EF38" pageId="11" pageNumber="12" type="discussion">
<paragraph id="2D443329BF43FFB4FF7E92591949EF38" blockId="11.[151,1437,539,1996]" pageId="11" pageNumber="12">
<emphasis id="1F8FEF3BBF43FFB4FF7E92591A4DEE49" bold="true" box="[151,276,539,565]" pageId="11" pageNumber="12">Remarks.</emphasis>
We did not have the opportunity to examine the
<typeStatus id="F2408D8BBF43FFB4FC9A925E188BEE4A" box="[883,978,540,566]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
specimens of
<taxonomicName id="EAFB48AABF43FFB4FB6D925E1FB3EE49" box="[1156,1258,540,565]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="1F8FEF3BBF43FFB4FB6D925E1FB3EE49" box="[1156,1258,540,565]" italics="true" pageId="11" pageNumber="12">D. heeri</emphasis>
</taxonomicName>
described and illustrated by
<bibRefCitation id="496A4ED8BF43FFB4FED292001AB9EE20" author="Pavesi" box="[315,480,578,604]" pageId="11" pageNumber="27" refString="Pavesi, P. (1873) Catalogo sistematico dei ragni del cantone ticino con la loro distribuzione orizontale e verticale e cenni sulla araneologica elvetica. Annali del Museo civico di storia naturale di Genova, 4, 5 - 215." type="journal article" year="1873">Pavesi (1873)</bibRefCitation>
from Ticino (
<collectingCountry id="55EC73B9BF43FFB4FD6F92001842EE20" box="[646,795,578,604]" name="Switzerland" pageId="11" pageNumber="12">Switzerland</collectingCountry>
). Examination of numerous specimens collected from several parts of the Alps allow us to consider the presence of at least three, perhaps four (see Remarks under
<taxonomicName id="EAFB48AABF43FFB4FF7E92D21A49EED5" box="[151,272,656,681]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="drescoi" status="sp. nov.">
<emphasis id="1F8FEF3BBF43FFB4FF7E92D21A49EED5" box="[151,272,656,681]" italics="true" pageId="11" pageNumber="12">D. drescoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04BC5240BF43FFB4FEFE92CD1A2DEED5" box="[279,372,655,681]" pageId="11" pageNumber="12" rank="species">
<emphasis id="1F8FEF3BBF43FFB4FEFE92CD1A2DEED5" bold="true" box="[279,372,655,681]" pageId="11" pageNumber="12">sp. nov.</emphasis>
</taxonomicNameLabel>
), very similar morphospecies, which can agree at first sight with the illustrations of Pavesi (see also figures of
<taxonomicName id="EAFB48AABF43FFB4FE6992F51AA3EEAC" box="[384,506,695,720]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="drescoi" status="sp. nov.">
<emphasis id="1F8FEF3BBF43FFB4FE6992F51AA3EEAC" box="[384,506,695,720]" italics="true" pageId="11" pageNumber="12">D. drescoi</emphasis>
</taxonomicName>
<emphasis id="1F8FEF3BBF43FFB4FDEB92F41939EEAC" bold="true" box="[514,608,694,720]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="04BC5240BF43FFB4FDEB92F41939EEAC" box="[514,608,694,720]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EAFB48AABF43FFB4FD9992F51840EEAC" box="[624,793,694,720]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="drescoi">
<emphasis id="1F8FEF3BBF43FFB4FD9992F519D5EEAC" box="[624,652,695,720]" italics="true" pageId="11" pageNumber="12">D.</emphasis>
aff.
<emphasis id="1F8FEF3BBF43FFB4FD2D92F51840EEAC" box="[708,793,695,720]" italics="true" pageId="11" pageNumber="12">drescoi</emphasis>
</taxonomicName>
and
<taxonomicName id="EAFB48AABF43FFB4FCBD92F5189FEEAC" box="[852,966,695,720]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="simoni" status="sp. nov.">
<emphasis id="1F8FEF3BBF43FFB4FCBD92F5189FEEAC" box="[852,966,695,720]" italics="true" pageId="11" pageNumber="12">D. simoni</emphasis>
</taxonomicName>
<emphasis id="1F8FEF3BBF43FFB4FC2692F41F74EEAC" bold="true" box="[975,1069,694,720]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="04BC5240BF43FFB4FC2692F41F74EEAC" box="[975,1069,694,720]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). One of these morphospecies, represented by the specimens collected in Allgäu (illustrated by Wiehle 1967) and in Graubünden Canton, corresponds better with the descriptions and illustrations of Pavesi. We decide here that these specimens belong to
<taxonomicName id="EAFB48AABF43FFB4FEE593681949EF38" authority="Pavesi." authorityName="Pavesi." box="[268,528,810,836]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="1F8FEF3BBF43FFB4FEE593681AEDEF3F" box="[268,436,810,835]" italics="true" pageId="11" pageNumber="12">D. heeri sensu</emphasis>
Pavesi.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="65E160A2BF43FFB4FF2F93121FB9E850" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="2D443329BF43FFB4FF2F93121FB9E850" blockId="11.[151,1437,539,1996]" pageId="11" pageNumber="12">
<emphasis id="1F8FEF3BBF43FFB4FF2F9312188EEF16" bold="true" box="[198,983,848,874]" pageId="11" pageNumber="12">
Material examined.
<collectingCountry id="55EC73B9BF43FFB4FE31931219EFEF16" box="[472,694,848,874]" name="Switzerland" pageId="11" pageNumber="12">SWITZERLAND</collectingCountry>
. No specific locality:
</emphasis>
1 Ƥ, Nicholson, O. P.-Cambridge collection (
<collectionCode id="4BEAABECBF43FFB4FEF3933A1A32EFEE" box="[282,363,888,914]" pageId="11" pageNumber="12">HECO</collectionCode>
B. 236); 1 Ƥ, 1899, R. J. Pickard-Cambridge, O. P.-Cambridge collection (
<collectionCode id="4BEAABECBF43FFB4FB0B933A1E6DEFEE" box="[1250,1332,888,914]" pageId="11" pageNumber="12">HECO</collectionCode>
B. 236); 1 Ƥ and 1 immature 3, 1906, R. J. Pickard-Cambridge, O. P.-Cambridge collection (
<collectionCode id="4BEAABECBF43FFB4FB3D93DC1E73EFC4" box="[1236,1322,926,952]" pageId="11" pageNumber="12">HECO</collectionCode>
B. 236).
<emphasis id="1F8FEF3BBF43FFB4FF7E93871AFCEFA3" bold="true" box="[151,421,965,991]" pageId="11" pageNumber="12">
<emphasis id="1F8FEF3BBF43FFB4FF7E93871AC2EFA3" bold="true" box="[151,411,965,991]" italics="true" pageId="11" pageNumber="12">Graubünden Canton</emphasis>
:
</emphasis>
1 Ƥ, 1907, R. Lessert, Simon collection n° 23822 (
<collectionCode id="4BEAABECBF43FFB4FBF793871F25EFA3" LSID="urn:lsid:biocol.org:col:34988" box="[1054,1148,965,991]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="12">MNHN</collectionCode>
<collectionCode id="4BEAABECBF43FFB4FB6D93871FF7EFA3" box="[1156,1198,965,991]" pageId="11" pageNumber="12">AR</collectionCode>
9166).
<emphasis id="1F8FEF3BBF43FFB4FAE493871BBAE879" bold="true" pageId="11" pageNumber="12">
<collectingCountry id="55EC73B9BF43FFB4FAE493871ECEEFA3" box="[1293,1431,965,991]" name="Austria" pageId="11" pageNumber="12">AUSTRIA</collectingCountry>
.
<emphasis id="1F8FEF3BBF43FFB4FF7E93AE1B81E87A" bold="true" box="[151,216,1004,1030]" italics="true" pageId="11" pageNumber="12">Tyrol</emphasis>
:
</emphasis>
Sölden, 1 Ƥ, A. W. Pickard-Cambridge, O. P.-Cambridge collection (
<collectionCode id="4BEAABECBF43FFB4FBDA93AE1FDFE87A" box="[1075,1158,1004,1030]" pageId="11" pageNumber="12">HECO</collectionCode>
B. 235).
<emphasis id="1F8FEF3BBF43FFB4FB1393A91BABE850" bold="true" pageId="11" pageNumber="12">
<collectingCountry id="55EC73B9BF43FFB4FB1393A91ECEE879" box="[1274,1431,1003,1029]" name="Germany" pageId="11" pageNumber="12">GERMANY</collectingCountry>
.
<emphasis id="1F8FEF3BBF43FFB4FF7E94501BB1E850" bold="true" box="[151,232,1042,1068]" italics="true" pageId="11" pageNumber="12">Allgäu</emphasis>
:
</emphasis>
2 Ƥ and
<date id="594515E9BF43FFB4FEB794501A8AE850" box="[350,467,1042,1068]" pageId="11" pageNumber="12" value="2000-01-03">
1 3,
<quantity id="EA039ECCBF43FFB4FE7094501A8AE850" box="[409,467,1042,1068]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">2000</quantity>
</date>
–2100 m,
<date id="594515E9BF43FFB4FDA094501847E850" box="[585,798,1042,1068]" pageId="11" pageNumber="12" value="1965-08-11" valueMax="1965-08-13" valueMin="1965-08-11">Aug. 11–13, 1965</date>
, Harms, det. Wiehle (
<collectionCode id="4BEAABECBF43FFB4FBF594501F00E850" LSID="urn:lsid:biocol.org:col:34838" box="[1052,1113,1042,1068]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
22517/13).
</paragraph>
</subSubSection>
<subSubSection id="65E160A2BF43FFB4FF2F947A1F21E947" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="2D443329BF43FFB4FF2F947A1F21E947" blockId="11.[151,1437,539,1996]" pageId="11" pageNumber="12">
<emphasis id="1F8FEF3BBF43FFB4FF2F947A1A1FE82E" bold="true" box="[198,326,1080,1106]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
<taxonomicName id="EAFB48AABF43FFB4FEB9947B1AEBE82E" box="[336,434,1081,1106]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="1F8FEF3BBF43FFB4FEB9947B1AEBE82E" box="[336,434,1081,1106]" italics="true" pageId="11" pageNumber="12">D. heeri</emphasis>
</taxonomicName>
can be associated with
<taxonomicName id="EAFB48AABF43FFB4FD3A947B1816E82E" box="[723,847,1081,1106]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="drescoi" status="sp. nov.">
<emphasis id="1F8FEF3BBF43FFB4FD3A947B1816E82E" box="[723,847,1081,1106]" italics="true" pageId="11" pageNumber="12">D. drescoi</emphasis>
</taxonomicName>
<emphasis id="1F8FEF3BBF43FFB4FCB1947A18E1E82E" bold="true" box="[856,952,1080,1106]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="04BC5240BF43FFB4FCB1947A18E1E82E" box="[856,952,1080,1106]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EAFB48AABF43FFB4FC1F947B1F33E82E" box="[1014,1130,1081,1106]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="simoni" status="sp. nov.">
<emphasis id="1F8FEF3BBF43FFB4FC1F947B1F33E82E" box="[1014,1130,1081,1106]" italics="true" pageId="11" pageNumber="12">D. simoni</emphasis>
</taxonomicName>
<emphasis id="1F8FEF3BBF43FFB4FB9D947A1F8AE82E" bold="true" box="[1140,1235,1080,1106]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="04BC5240BF43FFB4FB9D947A1F8AE82E" box="[1140,1235,1080,1106]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in a complex of species characterised by the presence of 3 or 4 retrolateral spines on tibiae IV, a large embolar coil looped around twice and spiralling upwards along only one axis (
<figureCitation id="B5C02FACBF43FFB4FCA994C418C3E8DC" box="[832,922,1158,1184]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="11" pageNumber="12">Figs 12</figureCitation>
,
<figureCitation id="B5C02FACBF43FFB4FC4194C418ACE8DC" box="[936,1013,1158,1184]" captionStart="FIGURES 20 – 27. 20 – 23" captionStartId="15.[151,269,1890,1914]" captionTargetBox="[168,1417,204,1859]" captionTargetId="figure@15.[162,1424,194,1866]" captionTargetPageId="15" captionText="FIGURES 20 – 27. 20 – 23 Drassodex simoni sp. nov. (Ledoux collection, after Murphy 2007: figs p. 235). 20 – 21 epigyne (20 ventral view, 21 dorsal view); 22 – 23 left male palp (22 ventral view, 23 retrolateral view). 24 – 27 Drassodex drescoi sp. nov. (24 – 25 Dresco collection, MNHN AR 14598, holotype; 26 – 27 MNHN AR 9167). 24 – 25 epigyne (24 ventral view, 25 dorsal view); 26 – 27 left male palp (26 ventral view, 27 retrolateral view)." httpUri="https://zenodo.org/record/189210/files/figure.png" pageId="11" pageNumber="12">22, 26</figureCitation>
), and a spiral-column shaped coiled duct, following internal contour of the membranous coil (
<figureCitation id="B5C02FACBF43FFB4FCD694EF1F79E8BB" box="[831,1056,1197,1223]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="11" pageNumber="12">Figs 11, 15, 17, 19</figureCitation>
,
<figureCitation id="B5C02FACBF43FFB4FBC494EF1F22E8BB" box="[1069,1147,1197,1223]" captionStart="FIGURES 20 – 27. 20 – 23" captionStartId="15.[151,269,1890,1914]" captionTargetBox="[168,1417,204,1859]" captionTargetId="figure@15.[162,1424,194,1866]" captionTargetPageId="15" captionText="FIGURES 20 – 27. 20 – 23 Drassodex simoni sp. nov. (Ledoux collection, after Murphy 2007: figs p. 235). 20 – 21 epigyne (20 ventral view, 21 dorsal view); 22 – 23 left male palp (22 ventral view, 23 retrolateral view). 24 – 27 Drassodex drescoi sp. nov. (24 – 25 Dresco collection, MNHN AR 14598, holotype; 26 – 27 MNHN AR 9167). 24 – 25 epigyne (24 ventral view, 25 dorsal view); 26 – 27 left male palp (26 ventral view, 27 retrolateral view)." httpUri="https://zenodo.org/record/189210/files/figure.png" pageId="11" pageNumber="12">21, 25</figureCitation>
). However,
<taxonomicName id="EAFB48AABF43FFB4FAE394EF1E32E8BA" box="[1290,1387,1197,1222]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="1F8FEF3BBF43FFB4FAE394EF1E32E8BA" box="[1290,1387,1197,1222]" italics="true" pageId="11" pageNumber="12">D. heeri</emphasis>
</taxonomicName>
can be distinguished from these by the shape of the RTA, almost symmetric, with an inferior prong greatly sharpened and as long as the superior prong (
<figureCitation id="B5C02FACBF43FFB4FD4694B8185EE968" box="[687,775,1274,1300]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="11" pageNumber="12">Fig. 13</figureCitation>
), the blunt tip of the embolar coil (
<figureCitation id="B5C02FACBF43FFB4FB4E94B81E69E968" box="[1191,1328,1274,1300]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="11" pageNumber="12">Figs 12–13</figureCitation>
), and the distal part of the membranous coils, extending and projecting forwards (
<figureCitation id="B5C02FACBF43FFB4FC0D95631F32E947" box="[996,1131,1313,1339]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="11" pageNumber="12">Figs 11, 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="65E160A2BF43FFB2FF2F95051993EC83" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="description">
<paragraph id="2D443329BF43FFB4FF2F95051F20EAEB" blockId="11.[151,1437,539,1996]" pageId="11" pageNumber="12">
<emphasis id="1F8FEF3BBF43FFB4FF2F95051A06E91D" bold="true" box="[198,351,1351,1377]" pageId="11" pageNumber="12">Description.</emphasis>
Male dimensions (SMF 22517/13): total length 9.72. Carapace 4.43 long, 3.14 wide. Leg I: femur 3.81, patella 2.06, tibia 3.52, metatarsus 2.97, tarsus 1.92, total 14.28. Leg II: femur 3.57, patella 1.83, tibia 2.84, metatarsus 2.65, tarsus 1.74, total 12.63. Leg III: femur 3.29, patella 1.65, tibia 2.48, metatarsus 2.51, tarsus 1.66, total 11.59. Leg IV: femur 4.19, patella 1.84, tibia 3.52, metatarsus 3.85, tarsus 1.92, total 15.32. Leg spination: femora: I, II d1-1-0; III r0-1(0)-1; IV p0-0-1; tibiae: I
<date id="594515E9BF43FFB4FBFF95A01F68E980" box="[1046,1073,1506,1532]" pageId="11" pageNumber="12">v1</date>
p(0)-2-0; II v0-2-0; III p1-1(1- 1-1),
<date id="594515E9BF43FFB4FF30964B1A4AEA5F" box="[217,275,1545,1571]" pageId="11" pageNumber="12">v2-2</date>
-2, r1-1-1(1-1); IV
<date id="594515E9BF43FFB4FE10964B196CEA5F" box="[505,565,1545,1571]" pageId="11" pageNumber="12">v2-2</date>
(1p)-2, r1-1-1-1; metatarsi: III
<date id="594515E9BF43FFB4FC4F964B1885EA5F" box="[934,988,1545,1571]" pageId="11" pageNumber="12">v2-1</date>
p(2)-2, r1-2-2; IV
<date id="594515E9BF43FFB4FB55964B1FAFEA5F" box="[1212,1270,1545,1571]" pageId="11" pageNumber="12">v2-2</date>
-2. RTA bifid, superior prong short and strongly curved; inferior prong sharp and as long as superior branch, both almost symmetric (
<figureCitation id="B5C02FACBF43FFB4FEC596141AD3EA0C" box="[300,394,1622,1648]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="11" pageNumber="12">Fig. 13</figureCitation>
); median apophysis pointing inwards; embolar coil looped almost twice, spiralling upwards along one axis, without terminal apophysis and blunt at its tip (
<figureCitation id="B5C02FACBF43FFB4FC0A963F1F32EAEB" box="[995,1131,1661,1687]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="11" pageNumber="12">Figs 12–13</figureCitation>
).
</paragraph>
<paragraph id="2D443329BF43FFB2FF2F96E61993EC83" blockId="11.[151,1437,539,1996]" lastBlockId="13.[151,1436,152,294]" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
Female dimensions (SMF 22517/13): total length 10.45–11.36. Carapace 4.95–5.14 long, 3.81–3.86 wide. Leg I: femur 3.95–4.03, patella 2.24–2.27, tibia 3.29–3.29, metatarsus 3.03–3.19, tarsus 1.84–1.90, total 14.46–14.57. Leg II: femur 3.74–3.76, patella 2.14–2.22, tibia 2.93–3.02, metatarsus 2.79–2.79, tarsus 1.74–1.79, total 13.45–13.47. Leg III: femur 3.50–3.50, patella 1.86–2.00, tibia 2.64–2.65, metatarsus 2.72–2.72, tarsus 1.67–1.74, total 12.46–12.54. Leg IV: femur 4.50–4.55, patella 2.06–2.15, tibia 3.86–3.87, metatarsus 3.99–4.07, tarsus 1.99–2.06, total 16.48–16.62. Leg spination: femora: I, II d1-1-0; III r0-1-1; IV p0-0-1; tibiae: II
<date id="594515E9BF43FFB4FE8997CE1A22EBDA" box="[352,379,1932,1958]" pageId="11" pageNumber="12">v1</date>
p(0)-2-0; III
<date id="594515E9BF43FFB4FDFA97CE1912EBDA" box="[531,587,1932,1958]" pageId="11" pageNumber="12">v2-2</date>
-2, r1-1-1(1-1); IV r1-1-1-1; metatarsi: III
<date id="594515E9BF43FFB4FBD397CE1F34EBDA" box="[1082,1133,1932,1958]" pageId="11" pageNumber="12">v2-1</date>
p(0)-2, r1-2-2; IV
<date id="594515E9BF43FFB4FAAF97CE1E26EBDA" box="[1350,1407,1932,1958]" pageId="11" pageNumber="12">v2-2</date>
-2, r1(2)-2-2. Atrium, subtriangular, wider than high (0.37–0.49 wide, 0.32–0.36 high, ratio 1.09–1.38), anterior tongue rectangular and projecting posteriorly (
<figureCitation id="B5C02FACBF45FFB2FD5590DA181CECCE" box="[700,837,152,178]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="13" pageNumber="14">Figs 10, 14</figureCitation>
); copulatory openings narrow, not easily visible in ventral view, clearly separated; membranous coils looped more than twice, with internal contour followed by spiral-column shaped coiled ducts (
<figureCitation id="B5C02FACBF45FFB2FDDF90A719E5EC83" box="[566,700,229,255]" captionStart="FIGURES 10 – 19. 10 – 15" captionStartId="12.[151,269,1802,1826]" captionTargetBox="[171,1414,182,1782]" captionTargetId="figure@12.[160,1427,174,1786]" captionTargetPageId="12" captionText="FIGURES 10 – 19. 10 – 15 Drassodex heeri (Pavesi, 1873) (10 – 13 SMF 22517 / 3; 14 – 15 MNHN AR 9166). 10 – 11, 14 – 15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12 – 13 left male palp (12 ventral view, 13 retrolateral view). 16 – 17 Drassodex simoni sp. nov. (MNHN ARM 034 a, holotype). 16 – 17 epigyne (16 ventral view, 17 dorsal view). 18 – 19 Drassodex aff. drescoi (MNHN AR 9167). 18 – 19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA)." httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="13" pageNumber="14">Figs 11, 15</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="798463A1BF44FFB3FF7E97481B8EEB8E" httpUri="https://zenodo.org/record/189209/files/figure.png" pageId="12" pageNumber="13" targetBox="[171,1414,182,1782]" targetPageId="12">
<paragraph id="2D443329BF44FFB3FF7E97481B8EEB8E" blockId="12.[151,1437,1802,2034]" pageId="12" pageNumber="13">
<emphasis id="1F8FEF3BBF44FFB3FF7E97481A04EB5E" bold="true" box="[151,349,1802,1826]" pageId="12" pageNumber="13">FIGURES 10–19.</emphasis>
10–15
<taxonomicName id="EAFB48AABF44FFB3FE45974919A2EB5E" authority="Pavesi, 1873" authorityName="Pavesi" authorityYear="1873" box="[428,763,1803,1826]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="1F8FEF3BBF44FFB3FE4597491902EB5E" box="[428,603,1803,1826]" italics="true" pageId="12" pageNumber="13">Drassodex heeri</emphasis>
(Pavesi, 1873)
</taxonomicName>
(10–13 SMF 22517/3; 14–15 MNHN AR 9166). 10–11, 14–15 epigyne (10, 14 ventral view; 11, 15 dorsal view); 12–13 left male palp (12 ventral view, 13 retrolateral view). 16–17
<taxonomicName id="EAFB48AABF44FFB3FF7E97121A04EB1B" box="[151,349,1872,1895]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="simoni" status="sp. nov.">
<emphasis id="1F8FEF3BBF44FFB3FF7E97121A04EB1B" box="[151,349,1872,1895]" italics="true" pageId="12" pageNumber="13">Drassodex simoni</emphasis>
</taxonomicName>
<emphasis id="1F8FEF3BBF44FFB3FE8C970D1AE6EB1B" bold="true" box="[357,447,1871,1895]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="04BC5240BF44FFB3FE8C970D1AE6EB1B" box="[357,447,1871,1895]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MNHN ARM 034a, holotype). 16–17 epigyne (16 ventral view, 17 dorsal view). 18–19
<taxonomicName id="EAFB48AABF44FFB3FF7E97311ADEEBF6" box="[151,391,1907,1930]" class="Arachnida" family="Gnaphosidae" genus="Drassodex" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="drescoi">
<emphasis id="1F8FEF3BBF44FFB3FF7E97311A50EBF6" box="[151,265,1907,1930]" italics="true" pageId="12" pageNumber="13">Drassodex</emphasis>
aff.
<emphasis id="1F8FEF3BBF44FFB3FED397311ADEEBF6" box="[314,391,1907,1930]" italics="true" pageId="12" pageNumber="13">drescoi</emphasis>
</taxonomicName>
(MNHN AR 9167). 18–19 epigyne (18 ventral view, 19 dorsal view). (A = atrium; AA = anterior arch; B = bursa; CD = coiled duct; CO = copulatory opening; EC = embolar coil; FLS = flattened lanceolate structure; IP = inferior prong of the RTA; MA = median apophysis; MC = membranous coil; S = socket; SP = superior prong of the RTA).
</paragraph>
</caption>
<subSubSection id="65E160A2BF45FFB2FF2F91491953ED5A" box="[198,522,267,294]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="2D443329BF45FFB2FF2F91491953ED5A" blockId="13.[151,1436,152,294]" box="[198,522,267,294]" pageId="13" pageNumber="14">
<emphasis id="1F8FEF3BBF45FFB2FF2F91491A3EED59" bold="true" box="[198,359,267,293]" pageId="13" pageNumber="14">Distribution.</emphasis>
Central Alps.
</paragraph>
</subSubSection>
</treatment>
</document>
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