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<document id="D1908BA61F3099B0F73D2BF0394594F0" ID-DOI="10.11646/phytotaxa.218.2.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="13635772" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725332500112" checkinUser="felipe" docAuthor="Shang, Hui, Ma, Qi-Xia &amp; Yan, Yue-Hong" docDate="2015" docId="1E078795FF81FFCEFF46FB30A70DEE60" docLanguage="en" docName="phytotaxa.218.2.5.pdf" docOrigin="Phytotaxa 218 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.218.2.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Dryopteris shiakeana H. Shang &amp; Y. H. Yan 2015, sp. nov." docType="treatment" docVersion="2" lastPageNumber="161" masterDocId="E23EFFEDFF83FFCBFFCEFFC4A62BEF45" masterDocTitle="Dryopteris shiakeana (Dryopteridaceae): A new fern from Danxiashan in Guangdong, China" masterLastPageNumber="162" masterPageNumber="156" pageNumber="158" updateTime="1725333454318" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="36BF049F10ED6A9998DA033307A6CBD0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F00A45D98CBC3E81F1A04D1ED649B1D6">Dryopteris shiakeana (Dryopteridaceae): A new fern from Danxiashan in Guangdong, China</mods:title>
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<mods:name id="B1336E466A6D3C7EC15BC263ACDE41FE" type="personal">
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<mods:namePart id="BF98622AC1A26B40636A69FB28922054">Shang, Hui</mods:namePart>
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<mods:namePart id="6074139D344960261D820436EFA73B5F">Ma, Qi-Xia</mods:namePart>
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<mods:namePart id="4502EB917BDAB11C1C767B752B2B65A9">Yan, Yue-Hong</mods:namePart>
</mods:name>
<mods:typeOfResource id="D2E2FBF0CE7EF6F509CE7B68A31E8F04">text</mods:typeOfResource>
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<mods:title id="9252ED5E94DFDC61462F10BBA5BF9141">Phytotaxa</mods:title>
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<mods:part id="0608A365BE1A6FB7EEDF0BF0E67CED77">
<mods:date id="3F160DC39B4FD01A2BF0E9121D709223">2015</mods:date>
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<mods:number id="449EE5ACEE97A270149419624571EC18">2015-07-02</mods:number>
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<mods:number id="B6E7F9491A23645C1EF09FD24E09ACCD">218</mods:number>
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<mods:classification id="AA0F383143F345D50E052EF06673852A">journal article</mods:classification>
<mods:identifier id="B108B1EFC04D68EBC163E0A255DA1390" type="DOI">10.11646/phytotaxa.218.2.5</mods:identifier>
<mods:identifier id="A9EA566184A3DDFB05F628BDD0D2F952" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="BDD8969AA37D2CE98580CE98DAC8AA7E" type="Zenodo-Dep">13635772</mods:identifier>
</mods:mods>
<treatment id="1E078795FF81FFCEFF46FB30A70DEE60" LSID="urn:lsid:plazi:treatment:1E078795FF81FFCEFF46FB30A70DEE60" httpUri="http://treatment.plazi.org/id/1E078795FF81FFCEFF46FB30A70DEE60" lastPageId="5" lastPageNumber="161" pageId="2" pageNumber="158">
<subSubSection id="DEB46508FF81FFC9FF46FB30A564EA4A" box="[136,847,1268,1295]" pageId="2" pageNumber="158" type="nomenclature">
<paragraph id="96113683FF81FFC9FF46FB30A564EA4A" blockId="2.[136,847,1268,1295]" box="[136,847,1268,1295]" pageId="2" pageNumber="158">
<heading id="CD5981EFFF81FFC9FF46FB30A564EA4A" box="[136,847,1268,1295]" fontSize="11" level="2" pageId="2" pageNumber="158" reason="3">
<taxonomicName id="51AE4D00FF81FFC9FF46FB30A452EA4A" ID-CoL="37XP4" authority="H. Shang &amp;Y. H. Yan" authorityName="H. Shang &amp; Y. H. Yan" authorityYear="2015" box="[136,633,1268,1295]" class="Polypodiopsida" family="Dryopteridaceae" genus="Dryopteris" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="158" phylum="Tracheophyta" rank="species" species="shiakeana" status="sp. nov.">
<emphasis id="A4DAEA91FF81FFC9FF46FB30A755EA4A" bold="true" box="[136,382,1268,1295]" italics="true" pageId="2" pageNumber="158">Dryopteris shiakeana</emphasis>
H. Shang &amp;Y. H. Yan
</taxonomicName>
,
<emphasis id="A4DAEA91FF81FFC9FD4AFB32A4F2EA4A" box="[644,729,1270,1295]" italics="true" pageId="2" pageNumber="158">
<taxonomicNameLabel id="BFE957EAFF81FFC9FD4AFB32A4F2EA4A" box="[644,729,1270,1295]" pageId="2" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<figureCitation id="0E952A06FF81FFC9FD2EFB32A507EA4A" box="[736,812,1270,1295]" captionStart="FIGURE 1" captionStartId="3.[136,229,1749,1770]" captionText="FIGURE 1. Dryopteris shiakeana H. Shang &amp; Y. H. Yan. A. Habit. B. Pinnule. C. Basal petiole scale. D. Sporangia. Drawn by Yunxiao Liu based on the holotype (YYH1205119, CSH)." pageId="2" pageNumber="158">Figs. 1</figureCitation>
,
<figureCitation id="0E952A06FF81FFC9FCF4FB32A560EA4A" box="[826,843,1270,1295]" captionStart="FIGURE 2" captionStartId="4.[136,229,2032,2053]" captionTargetBox="[173,1414,193,2003]" captionTargetId="figure-14@4.[170,1417,190,2006]" captionTargetPageId="4" captionText="FIGURE 2. Morphological comparison between D. dehuaensis (1A–1E) and D. shiakeana (2A–2E). A, B. Lamina. C, D. Pinnules and costa. E. Basal petiole and scales." figureDoi="http://doi.org/10.5281/zenodo.13635774" httpUri="https://zenodo.org/record/13635774/files/figure.png" pageId="2" pageNumber="158">2</figureCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DEB46508FF81FFC9FF46FA84A20CE746" pageId="2" pageNumber="158" type="materials_examined">
<paragraph id="96113683FF81FFC9FF46FA84A776EA3C" blockId="2.[136,1451,1344,1401]" pageId="2" pageNumber="158">
<emphasis id="A4DAEA91FF81FFC9FF46FA84A691EA10" bold="true" box="[136,186,1344,1365]" pageId="2" pageNumber="158">Type</emphasis>
:—
<materialsCitation id="26C63CDEFF81FFC9FF16FA84A771EA3C" collectingDate="2012-05-05" collectionCode="CSH" country="China" latitude="25.020555" location="Danxiashan National Park" longLatPrecision="21" longitude="113.67361" pageId="2" pageNumber="158" specimenCount="2" stateProvince="Guangdong" typeStatus="holotype">
<collectingCountry id="EEB97613FF81FFC9FF16FA84A73DEA10" box="[216,278,1344,1365]" name="China" pageId="2" pageNumber="158">China</collectingCountry>
.
<location id="93716058FF81FFC9FEEFFA84A40FEA10" LSID="urn:lsid:plazi:treatment:1E078795FF81FFCEFF46FB30A70DEE60:93716058FF81FFC9FEEFFA84A40FEA10" box="[289,548,1344,1365]" country="China" latitude="25.020555" longLatPrecision="21" longitude="113.67361" name="Danxiashan National Park" pageId="2" pageNumber="158" stateProvince="Guangdong">Danxiashan National Park</location>
,
<collectingRegion id="546AF861FF81FFC9FDFEFA84A488EA10" box="[560,675,1344,1365]" country="China" name="Guangdong" pageId="2" pageNumber="158">Guangdong</collectingRegion>
.
<geoCoordinate id="F39A5044FF81FFC9FD61FA84A516EA10" box="[687,829,1344,1365]" degrees="25" direction="north" minutes="01" orientation="latitude" pageId="2" pageNumber="158" precision="15" seconds="14" value="25.020555">
25° 01
<emphasis id="A4DAEA91FF81FFC9FD3FFA84A4DDEA10" bold="true" box="[753,758,1344,1365]" italics="true" pageId="2" pageNumber="158">′</emphasis>
14
<emphasis id="A4DAEA91FF81FFC9FCD9FA84A509EA10" bold="true" box="[791,802,1344,1365]" italics="true" pageId="2" pageNumber="158">′′</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="F39A5044FF81FFC9FC84FA84A5CAEA10" box="[842,993,1344,1365]" degrees="113" direction="east" minutes="40" orientation="longitude" pageId="2" pageNumber="158" precision="15" seconds="25" value="113.67361">
113° 40
<emphasis id="A4DAEA91FF81FFC9FC59FA84A5B7EA10" bold="true" box="[919,924,1344,1365]" italics="true" pageId="2" pageNumber="158">′</emphasis>
25
<emphasis id="A4DAEA91FF81FFC9FC73FA84A5E3EA10" bold="true" box="[957,968,1344,1365]" italics="true" pageId="2" pageNumber="158">′′</emphasis>
E
</geoCoordinate>
,
<date id="E2101043FF81FFC9FC23FA84A243EA10" box="[1005,1128,1344,1365]" pageId="2" pageNumber="158" value="2012-05-05">
<collectingDate id="F254E9ABFF81FFC9FC23FA84A243EA10" box="[1005,1128,1344,1365]" pageId="2" pageNumber="158" value="2012-05-05">5 May 2012</collectingDate>
</date>
, YYH1205119 (
<typeStatus id="49158821FF81FFC9FAC4FA84A349EA10" box="[1290,1378,1344,1365]" pageId="2" pageNumber="158" type="holotype">holotype</typeStatus>
,
<collectionCode id="F0BFAE46FF81FFC9FAA0FA84A38FEA10" box="[1390,1444,1344,1365]" country="China" httpUri="http://grbio.org/cool/vr0i-2neq" name="Chenshan Botanical Garden" pageId="2" pageNumber="158" type="Herbarium">CSH</collectionCode>
!;
<typeStatus id="49158821FF81FFC9FF73FAA0A725EA3C" box="[189,270,1380,1401]" pageId="2" pageNumber="158" type="isotype">isotypes</typeStatus>
,
<collectionCode id="F0BFAE46FF81FFC9FED6FAA0A764EA3C" box="[280,335,1380,1401]" country="China" httpUri="http://grbio.org/cool/vr0i-2neq" name="Chenshan Botanical Garden" pageId="2" pageNumber="158" type="Herbarium">CSH</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="96113683FF81FFC9FF46FA6EA231E96A" blockId="2.[136,1452,1450,2051]" pageId="2" pageNumber="158">
<taxonomicName id="51AE4D00FF81FFC9FF46FA6EA750EA86" box="[136,379,1450,1475]" class="Polypodiopsida" family="Dryopteridaceae" genus="Dryopteris" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="158" phylum="Tracheophyta" rank="species" species="shiakeana">
<emphasis id="A4DAEA91FF81FFC9FF46FA6EA750EA86" box="[136,379,1450,1475]" italics="true" pageId="2" pageNumber="158">Dryopteris shiakeana</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="51AE4D00FF81FFC9FD81FA6EA4C5EA86" box="[591,750,1450,1475]" class="Polypodiopsida" family="Dryopteridaceae" genus="Dryopteris" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="158" phylum="Tracheophyta" rank="species" species="dehuaensis">
<emphasis id="A4DAEA91FF81FFC9FD81FA6EA4C5EA86" box="[591,750,1450,1475]" italics="true" pageId="2" pageNumber="158">D. dehuaensis</emphasis>
</taxonomicName>
in general morphology; both species contain exindusiate sori, basiscopic pinnule stalks exceeding
<quantity id="51569B66FF81FFC9FDEFFA0AA477EAA2" box="[545,604,1486,1511]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="158" unit="cm" value="1.0">1 cm</quantity>
, and downside swallow-tailed, elongate basal pinnae. However, the former differs from the latter because by having dense scales on the rachis of laminae and pinnae, dissected pinnules, more prominent veinlets, reddish brown scales on stipes, and toothed segment margins.
</paragraph>
<paragraph id="96113683FF81FFC9FF73F9FEA20CE746" blockId="2.[136,1452,1450,2051]" pageId="2" pageNumber="158">
Terrestrial, evergreen,
<quantity id="51569B66FF81FFC9FE72F9FEA407E916" box="[444,556,1594,1619]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="2" pageNumber="158" unit="cm" value="50.0" valueMax="60.0" valueMin="40.0">40–60 cm</quantity>
tall. Rhizome, recumbent or erect, apex densely covered with linear-lanceolate and lustrous reddish-brown scales. Cespitose, 3–5 fronds per plant. Stipes, sepia and reddish-brown at base,
<quantity id="51569B66FF81FFC9FAF8F99AA38CE932" box="[1334,1447,1630,1655]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="158" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 cm</quantity>
, ca.
<quantity id="51569B66FF81FFC9FF60F946A6C4E9DE" box="[174,239,1666,1691]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="158" unit="mm" value="4.0">4 mm</quantity>
in diameter at the base and densely covered with lanceolate, reddish-brown scales at the base; upward scales becoming small, appressed. Lamina, ovate-lanceolate, approximately
<quantity id="51569B66FF81FFC9FC64F962A236E9FA" box="[938,1053,1702,1727]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="158" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 cm</quantity>
long,
<quantity id="51569B66FF81FFC9FBA4F962A2F6E9FA" box="[1130,1245,1702,1727]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="158" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 cm</quantity>
wide at the base, tripinnate, pinnatifid, and acuminate at the apex, lowest basiscopic pinnule on the lowest pinna distinctly enlarged, directed downward. Pinnae, 8–12 pairs, alternate, lanceolate, stalked; largest basal pinnae,
<quantity id="51569B66FF81FFC9FBB1F92AA2ECE842" box="[1151,1223,1774,1799]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="158" unit="cm" value="15.0">15 cm</quantity>
, with stalks
<quantity id="51569B66FF81FFC9FA9FF92AA383E842" box="[1361,1448,1774,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="158" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
; basiscopic pinnules usually larger than acroscopic ones; lowest basiscopic pinnule on lowest pinna largest. Pinnules, 15–18 pairs, oblong, round-cuneate base forms, apices blunt or subacute, base pinnules pinnatisect; pinnules on upper part from pinnatisect to pinnatilobate; segments closely linked, divided only by a narrow slit, nearly oblong, truncate at the apex, with teeth at the margin. Veins, visible at both sides, prominent abaxially; veinlets, 3–4 pairs each segment, 1- or 2-pinnate, branching, reaching the margin. Lamina, papery, yellowish-green; rachis and pinna rachis densely covered with reddish-brown appressed scales, pinnule rachis and costa covered with brown, bullate scales abaxially. Sori, small and medial, between the midrib and the margin of pinnule, exindusiate.
</paragraph>
</subSubSection>
<caption id="C2D1660BFF80FFC8FF46F911A44BE84B" pageId="3" pageNumber="159" startId="3.[136,229,1749,1770]" targetType="figure">
<paragraph id="96113683FF80FFC8FF46F911A44BE84B" blockId="3.[136,1451,1749,1806]" pageId="3" pageNumber="159">
<emphasis id="A4DAEA91FF80FFC8FF46F911A6D5E9AF" bold="true" box="[136,254,1749,1770]" pageId="3" pageNumber="159">FIGURE 1.</emphasis>
<taxonomicName id="51AE4D00FF80FFC8FECAF911A58FE9AF" authority="H. Shang &amp; Y. H. Yan. A. Habit. B. Pinnule. C." box="[260,932,1749,1770]" class="Polypodiopsida" family="Dryopteridaceae" genus="Dryopteris" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="159" phylum="Tracheophyta" rank="species" species="shiakeana">
<emphasis id="A4DAEA91FF80FFC8FECAF911A7FFE9AF" box="[260,468,1749,1770]" italics="true" pageId="3" pageNumber="159">Dryopteris shiakeana</emphasis>
H. Shang &amp; Y. H. Yan. A. Habit. B. Pinnule. C.
</taxonomicName>
Basal petiole scale. D. Sporangia. Drawn by Yunxiao Liu based on the holotype (YYH1205119, CSH).
</paragraph>
</caption>
<subSubSection id="DEB46508FF80FFCEFF73F8FAA70DEE60" lastPageId="5" lastPageNumber="161" pageId="3" pageNumber="159" type="etymology">
<paragraph id="96113683FF80FFC8FF73F8FAA380E812" blockId="3.[136,1451,1854,2059]" box="[189,1451,1854,1879]" pageId="3" pageNumber="159">
<emphasis id="A4DAEA91FF80FFC8FF73F8FAA763E812" bold="true" box="[189,328,1854,1879]" pageId="3" pageNumber="159">Etymology:</emphasis>
—The epithet is from the name of a famous Chinese geographer Xiake Xu, who first described
</paragraph>
<paragraph id="96113683FF80FFC8FF46F8A6A380E886" blockId="3.[136,1451,1854,2059]" pageId="3" pageNumber="159">
Danxia landform in southern
<collectingCountry id="EEB97613FF80FFC8FE1CF8A6A43EE83E" box="[466,533,1890,1915]" name="China" pageId="3" pageNumber="159">China</collectingCountry>
in the book
<emphasis id="A4DAEA91FF80FFC8FD6FF8A6A556E83E" box="[673,893,1890,1915]" italics="true" pageId="3" pageNumber="159">Travels of Xu Xiake</emphasis>
, which was published in 1642.
<emphasis id="A4DAEA91FF80FFC8FF73F842A706E8DA" bold="true" box="[189,301,1926,1951]" pageId="3" pageNumber="159">Habitats:</emphasis>
—Terrestrial, subtropical evergreen broadleaved forest in Danxiashan National Park.
<emphasis id="A4DAEA91FF80FFC8FF73F86EA4C2E886" bold="true" box="[189,745,1962,1987]" pageId="3" pageNumber="159">
Additional specimens examined (
<typeStatus id="49158821FF80FFC8FD90F86EA4F0E886" box="[606,731,1962,1987]" pageId="3" pageNumber="159" type="paratype">paratypes</typeStatus>
):
</emphasis>
—The same site of
<typeStatus id="49158821FF80FFC8FC2AF86EA262E886" box="[996,1097,1962,1987]" pageId="3" pageNumber="159" type="holotype">holotype</typeStatus>
, YYH13163-1, YYH13163-2,
</paragraph>
<paragraph id="96113683FF80FFC8FF46F80AA380E8A2" blockId="3.[136,1451,1854,2059]" box="[136,1451,1998,2023]" pageId="3" pageNumber="159">YYH13163-3, YYH13163-4, YYH13163-5, YYH13163-6, YYH13163-7, YYH13163-8, YYH13163-9, YYH13163-</paragraph>
<paragraph id="96113683FF80FFC8FF46F836A248E74E" blockId="3.[136,1451,1854,2059]" box="[136,1123,2034,2059]" pageId="3" pageNumber="159">10, YYH13163-11, YYH13163-12, YYH13163-13, YYH13163-14, and YYH13163-15.</paragraph>
<caption id="C2D1660BFF87FFCFFF46F834A7E7E76C" ID-DOI="http://doi.org/10.5281/zenodo.13635774" ID-Zenodo-Dep="13635774" httpUri="https://zenodo.org/record/13635774/files/figure.png" pageId="4" pageNumber="160" startId="4.[136,229,2032,2053]" targetBox="[173,1414,193,2003]" targetPageId="4" targetType="figure">
<paragraph id="96113683FF87FFCFFF46F834A7E7E76C" blockId="4.[136,1452,2032,2089]" pageId="4" pageNumber="160">
<emphasis id="A4DAEA91FF87FFCFFF46F834A6D5E740" bold="true" box="[136,254,2032,2053]" pageId="4" pageNumber="160">FIGURE 2.</emphasis>
Morphological comparison between
<taxonomicName id="51AE4D00FF87FFCFFDA6F834A4D9E740" box="[616,754,2032,2053]" class="Polypodiopsida" family="Dryopteridaceae" genus="Dryopteris" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="160" phylum="Tracheophyta" rank="species" species="dehuaensis">
<emphasis id="A4DAEA91FF87FFCFFDA6F834A4D9E740" box="[616,754,2032,2053]" italics="true" pageId="4" pageNumber="160">D. dehuaensis</emphasis>
</taxonomicName>
(1A–1E) and
<taxonomicName id="51AE4D00FF87FFCFFCB3F834A5D7E740" box="[893,1020,2032,2053]" class="Polypodiopsida" family="Dryopteridaceae" genus="Dryopteris" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="160" phylum="Tracheophyta" rank="species" species="shiakeana">
<emphasis id="A4DAEA91FF87FFCFFCB3F834A5D7E740" box="[893,1020,2032,2053]" italics="true" pageId="4" pageNumber="160">D. shiakeana</emphasis>
</taxonomicName>
(2A–2E). A, B. Lamina. C, D. Pinnules and costa. E. Basal petiole and scales.
</paragraph>
</caption>
<paragraph id="96113683FF86FFCEFF73FF64A70DEE60" blockId="5.[136,1452,160,293]" pageId="5" pageNumber="161">
<emphasis id="A4DAEA91FF86FFCEFF73FF64A46BEFFC" bold="true" box="[189,576,160,185]" pageId="5" pageNumber="161">IUCN Conservation assessment:</emphasis>
—EN(B1ab(iii)). No other individuals were found in other locations. Only ca. 20 individuals were collected in the
<typeStatus id="49158821FF86FFCEFDE6FF00A473EF98" box="[552,600,196,221]" pageId="5" pageNumber="161">type</typeStatus>
locality. Based on restricted geographic range, small populations, and less number of individuals,
<taxonomicName id="51AE4D00FF86FFCEFE40FF2CA409EE44" box="[398,546,232,257]" class="Polypodiopsida" family="Dryopteridaceae" genus="Dryopteris" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="161" phylum="Tracheophyta" rank="species" species="shiakeana">
<emphasis id="A4DAEA91FF86FFCEFE40FF2CA409EE44" box="[398,546,232,257]" italics="true" pageId="5" pageNumber="161">D. shiakeana</emphasis>
</taxonomicName>
should be considered endangered in accordance with the IUCN Red List criteria (
<bibRefCitation id="F23F4B72FF86FFCEFF5EFEC8A731EE60" author="IUCN" box="[144,282,268,293]" pageId="5" pageNumber="161" refId="ref2504" refString="IUCN (2014) Guidelines for using the IUCN red list categories and criteria, version 11. Standards and Petitions Subcommittee of the IUCN Species Survival Commission, Gland, Switzerland and Cambridge, United Kingdom. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 28 April 2015)" type="book" year="2014">IUCN 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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