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269 lines
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<document id="FEAF17A2CC9B44811394E9C44570D7E3" ID-CLB-Dataset="5866" ID-DOI="10.5852/ejt.2021.757.1415" ID-GBIF-Dataset="9950dbcc-5627-479c-b9e7-abe6b8a0ddd1" ID-ISSN="2118-9773" ID-Zenodo-Dep="5047683" ID-ZooBank="F95D404B-0A7C-429F-BC4F-AAEDF9DC870C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625079024571" checkinUser="felipe" docAuthor="Bordera, Santiago & Broad, Gavin R." docDate="2021" docId="03858793FFD9FFABDA17FBE67926FA5B" docLanguage="en" docName="ejt-757_80-101.pdf" docOrigin="European Journal of Taxonomy 757" docSource="http://dx.doi.org/10.5852/ejt.2021.757.1415" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Handaoia mercedensis Bordera 2021, sp. nov." docType="treatment" docUuid="80A6A583-DB23-43D9-AF4A-4D16A6270918" docUuidSource="ZooBank" docVersion="8" lastPageNumber="90" masterDocId="FFBCFFEBFFD1FFA1D83CFFBE781FFF84" masterDocTitle="Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World" masterLastPageNumber="101" masterPageNumber="80" pageNumber="88" updateTime="1699021642505" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="C288F12C0C3AEAF762CAF2354467D313">
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<mods:title id="C147DE973E573EEB48D600B25E64F530">Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World</mods:title>
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<mods:name id="550E05F39213EF9AB855F91671A658DA" type="personal">
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<mods:roleTerm id="371490C80CEC415E30D42E40215EB33F">Author</mods:roleTerm>
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<mods:namePart id="B2270E1F7B54CD3B6F085AE91C055838">Bordera, Santiago</mods:namePart>
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<mods:affiliation id="E107A689745507390C96059881C6FE87">Departamento de Ciencias Ambientales y Recursos Naturales, University of Alicante, San Vicente del Raspeig, E- 03690 Alicante, Spain.</mods:affiliation>
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<mods:nameIdentifier id="33DBF94BA91B7D9D38FF7961F5445634" type="email">s.bordera@ua.es</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="07FD4824E413D372A0F8013E180DE70E">Broad, Gavin R.</mods:namePart>
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<mods:affiliation id="A36E82E2B4AB3F357B350D7B9DA9BBAA">Department of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
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<mods:nameIdentifier id="43D7C7996CFDDD2C6B93223C72F13D4B" type="email">g.broad@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="E8E6B8F2095677AD9175B6710BDBE400">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="7BC490328D158A2B2E449470B85053A9">
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<mods:date id="9854B6E11D09FF0F410852D90DB5DB18">2021</mods:date>
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<mods:number id="01A1E3353A5A37CE4AC2D771513F0DE7">2021-06-30</mods:number>
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<mods:url id="2F8AD16B55A57F189957F5D003C0F8B8">http://dx.doi.org/10.5852/ejt.2021.757.1415</mods:url>
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<treatment id="03858793FFD9FFABDA17FBE67926FA5B" ID-DOI="http://doi.org/10.5281/zenodo.5056070" ID-GBIF-Taxon="183231744" ID-Zenodo-Dep="5056070" LSID="urn:lsid:zoobank.org:act:80A6A583-DB23-43D9-AF4A-4D16A6270918" httpUri="http://treatment.plazi.org/id/03858793FFD9FFABDA17FBE67926FA5B" lastPageId="10" lastPageNumber="90" pageId="8" pageNumber="88">
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<subSubSection id="C336650EFFD9FFA9DA17FBE67C16FBF7" box="[555,1033,1112,1139]" pageId="8" pageNumber="88" type="nomenclature">
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<paragraph id="8B933685FFD9FFA9DA17FBE67C16FBF7" blockId="8.[363,1224,1112,1209]" box="[555,1033,1112,1139]" pageId="8" pageNumber="88">
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<heading id="D0DB81E9FFD9FFA9DA17FBE67C16FBF7" box="[555,1033,1112,1139]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="88" reason="2">
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<taxonomicName id="4C2C4D06FFD9FFA9DA17FBE67BB8FBF6" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[555,935,1112,1139]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="88" phylum="Arthropoda" rank="species" species="mercedensis" status="sp. nov.">
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<emphasis id="B958EA97FFD9FFA9DA17FBE67B5DFBF7" bold="true" box="[555,834,1112,1139]" italics="true" pageId="8" pageNumber="88">Handaoia mercedensis</emphasis>
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Bordera
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</taxonomicName>
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<taxonomicNameLabel id="A26B57ECFFD9FFA9DB93FBE77C16FBF7" box="[943,1033,1113,1139]" pageId="8" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C336650EFFD9FFA9D957FBC27B70FB3D" pageId="8" pageNumber="88" type="description">
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<paragraph id="8B933685FFD9FFA9D957FBC27CD7FB13" blockId="8.[363,1224,1112,1209]" box="[363,1224,1148,1175]" pageId="8" pageNumber="88">
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<uri id="FFBD3A87FFD9FFA9D957FBC27CD7FB13" box="[363,1224,1148,1175]" pageId="8" pageNumber="88">
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urn:lsid:zoobank.org:act:
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<uuid id="FF8A0C50FFD9FFA9DAB7FBC27CD7FB13" box="[651,1224,1148,1175]" pageId="8" pageNumber="88">80A6A583-DB23-43D9-AF4A-4D16A6270918</uuid>
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</uri>
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</paragraph>
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<paragraph id="8B933685FFD9FFA9DAF9FB217B70FB3D" blockId="8.[363,1224,1112,1209]" box="[709,879,1183,1209]" pageId="8" pageNumber="88">
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<figureCitation id="13172A00FFD9FFA9DAF9FB217B3DFB3D" box="[709,802,1183,1209]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="8" pageNumber="88">Figs 1C</figureCitation>
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,
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<figureCitation id="13172A00FFD9FFA9DB13FB217B5FFB3D" box="[815,832,1183,1209]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="8" pageNumber="88">4</figureCitation>
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,
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<figureCitation id="13172A00FFD9FFA9DB70FB217B70FB3D" box="[844,879,1183,1209]" captionStart="Fig" captionStartId="19.[186,229,1963,1989]" captionTargetBox="[189,1398,1195,1923]" captionTargetId="figure-309@19.[189,1398,1195,1923]" captionTargetPageId="19" captionText="Fig. 9. Distribution of New World species of Handaoia Seyrig, 1952. A. North and Central America. B. South America." figureDoi="http://doi.org/10.5281/zenodo.5047703" httpUri="https://zenodo.org/record/5047703/files/figure.png" pageId="8" pageNumber="88">9B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C336650EFFD9FFA9D881FB5E7D16FA32" pageId="8" pageNumber="88" type="diagnosis">
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<paragraph id="8B933685FFD9FFA9D881FB5E7928FB7E" blockId="8.[189,311,1248,1274]" box="[189,311,1248,1274]" pageId="8" pageNumber="88">
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<heading id="D0DB81E9FFD9FFA9D881FB5E7928FB7E" bold="true" box="[189,311,1248,1274]" fontSize="11" level="3" pageId="8" pageNumber="88" reason="3">
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<emphasis id="B958EA97FFD9FFA9D881FB5E7928FB7E" bold="true" box="[189,311,1248,1274]" pageId="8" pageNumber="88">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B933685FFD9FFA9D881FAB17D16FA32" blockId="8.[189,1399,1295,1463]" pageId="8" pageNumber="88">
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<taxonomicName id="4C2C4D06FFD9FFA9D881FAB17A2BFAAD" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[189,564,1295,1321]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="88" phylum="Arthropoda" rank="species" species="mercedensis" status="sp. nov.">
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<emphasis id="B958EA97FFD9FFA9D881FAB179D2FAAD" box="[189,461,1295,1321]" italics="true" pageId="8" pageNumber="88">Handaoia mercedensis</emphasis>
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Bordera
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</taxonomicName>
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<taxonomicNameLabel id="A26B57ECFFD9FFA9DA00FAB17A89FAAD" box="[572,662,1295,1321]" pageId="8" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
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can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum conspicuously elevated, parallelsided with blunt tip (
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<figureCitation id="13172A00FFD9FFA9D990FAEB7A16FAEB" box="[428,521,1365,1391]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="8" pageNumber="88">Fig. 4G</figureCitation>
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, arrow); fore wing with well-defined transverse dark brown band (
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<figureCitation id="13172A00FFD9FFA9DD37FAEB7D76FAEB" box="[1291,1385,1365,1391]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="8" pageNumber="88">Fig. 4A</figureCitation>
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); juxtacoxal carina strong, irregular, joining submetapleural carina at mid length (
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<figureCitation id="13172A00FFD9FFA9DC73FAC67CBCFA17" box="[1103,1187,1400,1427]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="8" pageNumber="88">Fig. 4F</figureCitation>
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, arrow); flagellum dark brown (except white band), with first and second flagellomeres light brown (
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<figureCitation id="13172A00FFD9FFA9DC47FA227CE3FA32" box="[1147,1276,1436,1462]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="8" pageNumber="88">Fig. 4A–B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C336650EFFD9FFA9D881FA6379E2F9CD" pageId="8" pageNumber="88" type="etymology">
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<paragraph id="8B933685FFD9FFA9D881FA63795AFA73" blockId="8.[189,325,1501,1527]" box="[189,325,1501,1527]" pageId="8" pageNumber="88">
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<heading id="D0DB81E9FFD9FFA9D881FA63795AFA73" bold="true" box="[189,325,1501,1527]" fontSize="11" level="3" pageId="8" pageNumber="88" reason="3">
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<emphasis id="B958EA97FFD9FFA9D881FA63795AFA73" bold="true" box="[189,325,1501,1527]" pageId="8" pageNumber="88">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B933685FFD9FFA9D881F9B279E2F9CD" blockId="8.[189,1399,1548,1609]" pageId="8" pageNumber="88">
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The name refers to La Merced, municipality of Chanchamayo in the Department of
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<collectingRegion id="49E8F867FFD9FFA9DCACF9B27CCFF9A2" box="[1168,1232,1548,1574]" country="Peru" name="Junin" pageId="8" pageNumber="88">Junin</collectingRegion>
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(
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<collectingCountry id="F33B7615FFD9FFA9DCE3F9B27D05F9A2" box="[1247,1306,1548,1574]" name="Peru" pageId="8" pageNumber="88">Peru</collectingCountry>
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), where the
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<typeStatus id="54978827FFD9FFA9D8D4F9917950F9CD" box="[232,335,1583,1609]" pageId="8" pageNumber="88" type="holotype">holotype</typeStatus>
|
|||
|
|
was collected.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C336650EFFD9FFA9D881F9CE790AF8C5" pageId="8" pageNumber="88" type="materials_examined">
|
|||
|
|
<paragraph id="8B933685FFD9FFA9D881F9CE79B2F90E" blockId="8.[189,429,1648,1674]" box="[189,429,1648,1674]" pageId="8" pageNumber="88">
|
|||
|
|
<heading id="D0DB81E9FFD9FFA9D881F9CE79B2F90E" bold="true" box="[189,429,1648,1674]" fontSize="11" level="3" pageId="8" pageNumber="88" reason="3">
|
|||
|
|
<emphasis id="B958EA97FFD9FFA9D881F9CE79B2F90E" bold="true" box="[189,429,1648,1674]" pageId="8" pageNumber="88">Material examined</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B933685FFD9FFA9D881F9217A98F93D" blockId="8.[189,647,1695,1721]" box="[189,647,1695,1721]" pageId="8" pageNumber="88">
|
|||
|
|
Known only from the
|
|||
|
|
<typeStatus id="54978827FFD9FFA9D9FEF9217A36F93D" box="[450,553,1695,1721]" pageId="8" pageNumber="88" type="holotype">holotype</typeStatus>
|
|||
|
|
female.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B933685FFD9FFA9D8D6F95E790AF8C5" blockId="8.[189,1398,1760,1857]" pageId="8" pageNumber="88">
|
|||
|
|
<materialsCitation id="3B443CD8FFD9FFA9D8D6F95E790AF8C5" ID-GBIF-Occurrence="3312836303" collectingDate="2009-01-03" collectingDateMax="2009-01-17" collectingDateMin="2009-01-03" collectionCode="CEUA" collectorName="Trampa Malaise" country="Peru" location="La Merced" pageId="8" pageNumber="88" specimenCode="AECID 013484/07" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
|||
|
|
<emphasis id="B958EA97FFD9FFA9D8D6F95E7942F97E" bold="true" box="[234,349,1760,1786]" pageId="8" pageNumber="88">
|
|||
|
|
<typeStatus id="54978827FFD9FFA9D8D6F95E7942F97E" box="[234,349,1760,1786]" pageId="8" pageNumber="88" type="holotype">Holotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
<collectingCountry id="F33B7615FFD9FFA9D881F8BA7917F89A" box="[189,264,1796,1822]" name="Peru" pageId="8" pageNumber="88">PERU</collectingCountry>
|
|||
|
|
•
|
|||
|
|
<specimenCount id="9D2AFD0CFFD9FFA9D914F8BA7920F89A" box="[296,319,1796,1822]" pageId="8" pageNumber="88" type="female">♀</specimenCount>
|
|||
|
|
;
|
|||
|
|
<location id="8EF3605EFFD9FFA9D96DF8BA79C7F899" LSID="urn:lsid:plazi:treatment:03858793FFD9FFABDA17FBE67926FA5B:8EF3605EFFD9FFA9D96DF8BA79C7F899" box="[337,472,1795,1822]" country="Peru" name="La Merced" pageId="8" pageNumber="88">La Merced</location>
|
|||
|
|
,
|
|||
|
|
<location id="8EF3605EFFD9FFA9D9DAF8BD7ADAF89A" LSID="urn:lsid:plazi:treatment:03858793FFD9FFABDA17FBE67926FA5B:8EF3605EFFD9FFA9D9DAF8BD7ADAF89A" box="[486,709,1795,1822]" country="Peru" name="Fundo La Genova" pageId="8" pageNumber="88">Fundo La Génova</location>
|
|||
|
|
;
|
|||
|
|
<collectorName id="26D95353FFD9FFA9DAEFF8BA7B8AF899" box="[723,917,1795,1822]" pageId="8" pageNumber="88">Trampa Malaise</collectorName>
|
|||
|
|
3;
|
|||
|
|
<date id="FF921045FFD9FFA9DBFEF8BA7C61F89A" box="[962,1150,1796,1822]" pageId="8" pageNumber="88" value="2009-01-03" valueMax="2009-01-17" valueMin="2009-01-03">
|
|||
|
|
<collectingDate id="EFD6E9ADFFD9FFA9DBFEF8BA7C61F89A" box="[962,1150,1796,1822]" pageId="8" pageNumber="88" value="2009-01-03" valueMax="2009-01-17" valueMin="2009-01-03">3–17 Jan. 2009</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
;
|
|||
|
|
<specimenCode id="DB8A9EFEFFD9FFA9DCB0F8BA7D6DF89A" box="[1164,1394,1796,1822]" pageId="8" pageNumber="88">AECID 013484/07</specimenCode>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="ED3DAE40FFD9FFA9D881F899790EF8C5" box="[189,273,1831,1857]" pageId="8" pageNumber="88">CEUA</collectionCode>
|
|||
|
|
.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C336650EFFD9FFABD881F8D67926FA5B" lastPageId="10" lastPageNumber="90" pageId="8" pageNumber="88" type="description">
|
|||
|
|
<paragraph id="8B933685FFD9FFA9D881F8D67951F806" blockId="8.[189,334,1896,1922]" box="[189,334,1896,1922]" pageId="8" pageNumber="88">
|
|||
|
|
<heading id="D0DB81E9FFD9FFA9D881F8D67951F806" bold="true" box="[189,334,1896,1922]" fontSize="11" level="3" pageId="8" pageNumber="88" reason="3">
|
|||
|
|
<emphasis id="B958EA97FFD9FFA9D881F8D67951F806" bold="true" box="[189,334,1896,1922]" pageId="8" pageNumber="88">Description</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B933685FFD9FFA9D8D6F8297959F835" blockId="8.[189,1164,1943,2005]" box="[234,326,1943,1969]" pageId="8" pageNumber="88">
|
|||
|
|
<emphasis id="B958EA97FFD9FFA9D8D6F8297959F835" bold="true" box="[234,326,1943,1969]" pageId="8" pageNumber="88">Female</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B933685FFD9FFA9D881F8047C93F850" blockId="8.[189,1164,1943,2005]" box="[189,1164,1978,2005]" pageId="8" pageNumber="88">
|
|||
|
|
MEASUREMENTS. Body length (without ovipositor)
|
|||
|
|
<quantity id="4CD49B60FFD9FFA9DB3BF8047B7BF850" box="[775,868,1978,2004]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="8" pageNumber="88" unit="mm" value="4.3">4.3 mm</quantity>
|
|||
|
|
. Fore wing
|
|||
|
|
<quantity id="4CD49B60FFD9FFA9DBD3F8047C56F850" box="[1007,1097,1978,2004]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="88" unit="mm" value="3.4">3.4 mm</quantity>
|
|||
|
|
long.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF53660DFFD8FFA8D881F8DD7C5BF863" ID-DOI="http://doi.org/10.5281/zenodo.5047693" ID-Zenodo-Dep="5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="9" pageNumber="89" startId="9.[189,232,1891,1917]" targetBox="[189,1398,390,1851]" targetPageId="9">
|
|||
|
|
<paragraph id="8B933685FFD8FFA8D881F8DD7C5BF863" blockId="9.[187,1399,1891,2024]" pageId="9" pageNumber="89">
|
|||
|
|
<emphasis id="B958EA97FFD8FFA8D881F8DD7915F8F9" bold="true" box="[189,266,1891,1917]" pageId="9" pageNumber="89">Fig. 4.</emphasis>
|
|||
|
|
Morphology of
|
|||
|
|
<taxonomicName id="4C2C4D06FFD8FFA8D9FFF8DD7B29F8F9" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[451,822,1891,1917]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="89" phylum="Arthropoda" rank="species" species="mercedensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B958EA97FFD8FFA8D9FFF8DD7ACFF8F9" box="[451,720,1891,1917]" italics="true" pageId="9" pageNumber="89">Handaoia mercedensis</emphasis>
|
|||
|
|
Bordera
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A26B57ECFFD8FFA8DB00F8DA7B88F8FA" box="[828,919,1892,1918]" pageId="9" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, holotype, ♀ (CEUA).
|
|||
|
|
<emphasis id="B958EA97FFD8FFA8DC9DF8DD7CA9F8F9" bold="true" box="[1185,1206,1891,1917]" pageId="9" pageNumber="89">A</emphasis>
|
|||
|
|
. Habitus, lateral view.
|
|||
|
|
<emphasis id="B958EA97FFD8FFA8D93AF8397905F825" bold="true" box="[262,282,1927,1953]" pageId="9" pageNumber="89">B</emphasis>
|
|||
|
|
. Head, front view.
|
|||
|
|
<emphasis id="B958EA97FFD8FFA8DA36F8387A00F824" bold="true" box="[522,543,1926,1952]" pageId="9" pageNumber="89">C</emphasis>
|
|||
|
|
. Head and mesoscutum, dorsal view.
|
|||
|
|
<emphasis id="B958EA97FFD8FFA8DBCDF8397C19F825" bold="true" box="[1009,1030,1927,1953]" pageId="9" pageNumber="89">D</emphasis>
|
|||
|
|
. Head and mesosoma, lateral view.
|
|||
|
|
<emphasis id="B958EA97FFD8FFA8D930F814793FF840" bold="true" box="[268,288,1962,1988]" pageId="9" pageNumber="89">E</emphasis>
|
|||
|
|
. Metasoma, dorsal view.
|
|||
|
|
<emphasis id="B958EA97FFD8FFA8DA55F8147A64F840" bold="true" box="[617,635,1962,1988]" pageId="9" pageNumber="89">F</emphasis>
|
|||
|
|
. Metapleuron (arrow: juxtacoxal carina).
|
|||
|
|
<emphasis id="B958EA97FFD8FFA8DC57F8177C9EF847" bold="true" box="[1131,1153,1961,1987]" pageId="9" pageNumber="89">G</emphasis>
|
|||
|
|
. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8B933685FFDBFFABD881FEB77D02FDB9" blockId="10.[189,1399,265,573]" pageId="10" pageNumber="90">
|
|||
|
|
HEAD. Finely granulate, matt, strongly narrowed behind eyes, occiput very concave centrally in dorsal view (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABD960FE9279A3FEC2" box="[348,444,300,326]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4C</figureCitation>
|
|||
|
|
); gena 0.18 × length of eye in dorsal view; posterior ocellus separated from eye by 1.3× its own maximum diameter; distance between posterior ocelli 0.9 × maximum ocellar diameter (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABD8FAFECC7939FE09" box="[198,294,370,397]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4C</figureCitation>
|
|||
|
|
); occipital carina gently V-shaped medially; malar space about 1.4× basal width of mandible; face slightly swollen medially; clypeus (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABDAA5FE287AEAFE34" box="[665,757,406,432]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4B</figureCitation>
|
|||
|
|
) quite convex, gently granulate, with scattered coarse punctures in apical half, apex slightly rounded, about 2.0× as wide as high; mandible quite tapered to apex, ventral tooth of mandible a little shorter than upper tooth; maxillary palp reaching mid coxa; flagellum widened in middle, tapered towards apex, flagellomeres 8–20 flattened ventrally (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABDD34FE417D76FD9D" box="[1288,1385,511,537]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4A</figureCitation>
|
|||
|
|
); flagellum with 22 segments, length-thickness ratios: 1
|
|||
|
|
<superScript id="7C599BCDFFDBFFABDB0AFD9C7B5EFDB4" attach="left" box="[822,833,546,560]" fontSize="6" pageId="10" pageNumber="90">st</superScript>
|
|||
|
|
segment = 5.0, 2
|
|||
|
|
<superScript id="7C599BCDFFDBFFABDC37FD9C7C03FDB4" attach="left" box="[1035,1052,546,560]" fontSize="6" pageId="10" pageNumber="90">nd</superScript>
|
|||
|
|
= 5.38 and 10
|
|||
|
|
<superScript id="7C599BCDFFDBFFABDCF9FD9C7CCDFDB4" attach="left" box="[1221,1234,546,560]" fontSize="6" pageId="10" pageNumber="90">th</superScript>
|
|||
|
|
= 1.0.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B933685FFDBFFABD881FDDB7D59FC1C" blockId="10.[186,1398,612,920]" pageId="10" pageNumber="90">
|
|||
|
|
MESOSOMA. Finely granulate and matt (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABDA4FFDDA7AEEFDFA" box="[627,753,612,638]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4C–D</figureCitation>
|
|||
|
|
);mesoscutum slightly convex, notauli absent (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABDD31FDDA7D76FDFA" box="[1293,1385,612,638]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4C</figureCitation>
|
|||
|
|
); scutellum moderately convex, without lateral carinae; sternaulus deep, exceeding half the length of mesopleuron (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABD959FD1479DBFD41" box="[357,452,682,709]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4D</figureCitation>
|
|||
|
|
); juxtacoxal carina strong, irregular, joining submetapleural carina at mid length (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABD8FAFD707904FD6C" box="[198,283,718,744]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4F</figureCitation>
|
|||
|
|
, arrow); propodeum (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABDA2BFD707A70FD6C" box="[535,623,718,744]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="10" pageNumber="90">Fig. 1C</figureCitation>
|
|||
|
|
) with strong and conspicuous carinae; area superomedia confluent with area basalis and separated from area petiolaris; inner side of area externa about 4.7× as long as inner side of area dentipara; inner side of area dentipara about 0.3× length of outer side; area spiracularis confluent with area lateralis; posterior transverse carina strong, forming an elevate apophysis joining lateral longitudinal carina, apophysis parallel-sided with blut tip (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABDB86FCE47C08FCF1" box="[954,1047,858,885]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4G</figureCitation>
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, arrow). Length of hind femur about 4.0× its height. Hind wing with vein
|
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<emphasis id="B958EA97FFDBFFABDA88FCC07AF6FC13" box="[692,745,894,919]" italics="true" pageId="10" pageNumber="90">cu-a</emphasis>
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0.5 as long as abscissa of
|
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<emphasis id="B958EA97FFDBFFABDC22FCC07C58FC1C" box="[1054,1095,894,920]" italics="true" pageId="10" pageNumber="90">CU</emphasis>
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between
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<emphasis id="B958EA97FFDBFFABDC84FCC07CCFFC13" box="[1208,1232,894,919]" italics="true" pageId="10" pageNumber="90">M</emphasis>
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and
|
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<emphasis id="B958EA97FFDBFFABDD36FCC07D20FC13" box="[1290,1343,894,919]" italics="true" pageId="10" pageNumber="90">cu-a</emphasis>
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.
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</paragraph>
|
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<paragraph id="8B933685FFDBFFABD881FC7E798FFB0D" blockId="10.[188,1399,959,1162]" pageId="10" pageNumber="90">
|
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METASOMA. Tergite I (
|
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<figureCitation id="13172A00FFDBFFABD9F8FC017A00FC5D" box="[452,543,959,985]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="10" pageNumber="90">Fig. 1C</figureCitation>
|
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) finely granulate, slightly shiny, 1.77 × as long as its maximum width; postpetiole laterally not finely strigose; lateromedian longitudinal carina moderately strong, reaching about 0.7× length of tergite; lateral longitudinal carina complete; tergite II (
|
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<figureCitation id="13172A00FFDBFFABDC0CFBBB7C97FBA4" box="[1072,1160,1029,1056]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4E</figureCitation>
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) shiny, very slightly granulate anteriorly, glabrous, smooth posteriorly, with moderately dense short setae; remaining tergites (
|
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|
|
<figureCitation id="13172A00FFDBFFABD8F9FBF27900FBE2" box="[197,287,1100,1126]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4E</figureCitation>
|
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) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath 0.53 × as long as hind tibia.
|
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</paragraph>
|
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|
|
<paragraph id="8B933685FFDBFFABD881FB0F7D59FAFF" blockId="10.[188,1400,1200,1403]" pageId="10" pageNumber="90">
|
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|
|
COLOUR. Head (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABD944FB0E79E5FB4F" box="[376,506,1200,1227]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4A–D</figureCitation>
|
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|
|
) orange brown, vertical orbits and interocellar area dark brown; palpi dark brown; mandibular teeth black; antenna with scape, pedicel and flagellomeres 1–2 light brown, remaining flagellomeres dark brown, flagellomeres 4–7 and base of 8 dorsally white (
|
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|
|
<figureCitation id="13172A00FFDBFFABDC26FB497C68FA95" box="[1050,1143,1271,1297]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4A</figureCitation>
|
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|
|
). Mesosoma (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABDD2AFB4978EAFAB0" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4A, C–D</figureCitation>
|
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|
|
) entirely orange-brown. Metasoma (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABDA92FAA47B15FAB0" box="[686,778,1306,1332]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4E</figureCitation>
|
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|
|
) except tergite I dark brown, tergite I light brown. Fore and mid legs (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABD9A2FA8379E6FAD3" box="[414,505,1341,1367]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4A</figureCitation>
|
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|
|
) light brown, tarsi fuscous; hind leg dark brown, trochanter and base of tibia white. Wings slightly infuscate, fore wing with well-defined transverse dark brown band (
|
|||
|
|
<figureCitation id="13172A00FFDBFFABDCE5FADE7D26FAFF" box="[1241,1337,1376,1403]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="10" pageNumber="90">Fig. 4A</figureCitation>
|
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|
|
).
|
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|
|
</paragraph>
|
|||
|
|
<paragraph id="8B933685FFDBFFABD8D6FA1C7935FA38" blockId="10.[189,313,1442,1503]" box="[234,298,1442,1468]" pageId="10" pageNumber="90">
|
|||
|
|
<emphasis id="B958EA97FFDBFFABD8D6FA1C7935FA38" bold="true" box="[234,298,1442,1468]" pageId="10" pageNumber="90">Male</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B933685FFDBFFABD881FA7B7926FA5B" blockId="10.[189,313,1442,1503]" box="[189,313,1477,1503]" pageId="10" pageNumber="90">Unknown.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|