156 lines
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156 lines
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<document id="82AB84D5FE446A73089698FBC8D1CB8C" ID-DOI="10.5281/zenodo.156636" ID-GBIF-Dataset="c9e80f8e-36cc-4c6b-bfb8-bc53428ee8f4" ID-ISSN="11755326" ID-Zenodo-Dep="156636" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459622659579" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2003" docId="36558784E16AAC2BFE80FE17FBA416CD" docLanguage="en" docName="zt00263.pdf" docOrigin="Zootaxa 263" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Haplohyphes yanahuicsa Molineri, 2003, sp. nov." docType="treatment" docVersion="10" lastPageNumber="8" masterDocId="CA6CFFFCE16FAC2CFF88FFF3FFAE132B" masterDocTitle="Haplohyphes (Ephemeroptera: Leptohyphidae), new species and stage descriptions with a key to separate the species of the genus" masterLastPageNumber="11" masterPageNumber="1" pageNumber="6" updateTime="1698203766015" updateUser="plazi">
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<mods:mods id="195F4970AD61C4021D70477417F05B74" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F80CBC99725160D511405ECCA54E3963">
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<mods:title id="F4681FAF1557BF7B9B4C9126328CB359">Haplohyphes (Ephemeroptera: Leptohyphidae), new species and stage descriptions with a key to separate the species of the genus</mods:title>
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<mods:namePart id="AA8132563512E0DFAC661FF656AE2018">Molineri, Carlos</mods:namePart>
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<mods:typeOfResource id="BC22F7365DA0094ACCFE49FEF5B23800">text</mods:typeOfResource>
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<mods:title id="9619A9639407186D4D67A07BE12A5733">Zootaxa</mods:title>
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<mods:part id="42A8202B76B96647B9608133BE2290DC">
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<mods:date id="56547521170CA38D30A1C71A8E284F8E">2003</mods:date>
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<mods:detail id="A10BB0406C4AFA7EDD93BEAE90277966" type="volume">
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<mods:number id="2EBA5C0D8372E39A75BFC3851323F037">263</mods:number>
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<mods:classification id="01803EC27C498B3A1EA8C1504BD194B0">journal article</mods:classification>
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<mods:identifier id="57978A5F2657A732806215D5452EE955" type="DOI">10.5281/zenodo.156636</mods:identifier>
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<mods:identifier id="037CD9C97AA64579E63573DC1B02143B" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="1AF651000DA3E29D309F126D33834F72" type="Zenodo-Dep">156636</mods:identifier>
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<treatment id="36558784E16AAC2BFE80FE17FBA416CD" ID-DOI="http://doi.org/10.5281/zenodo.6275293" ID-GBIF-Taxon="119331951" ID-Zenodo-Dep="6275293" LSID="urn:lsid:plazi:treatment:36558784E16AAC2BFE80FE17FBA416CD" httpUri="http://treatment.plazi.org/id/36558784E16AAC2BFE80FE17FBA416CD" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
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<subSubSection id="F6E66519E16AAC29FE80FE17FD061752" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="BE433692E16AAC29FE80FE17FC3A12D6" blockId="5.[264,916,483,510]" box="[264,916,483,510]" pageId="5" pageNumber="6">
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<heading id="E50B81FEE16AAC29FE80FE17FC3A12D6" bold="true" box="[264,916,483,510]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
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<emphasis id="8C88EA80E16AAC29FE80FE17FC3A12D6" bold="true" box="[264,916,483,510]" pageId="5" pageNumber="6">
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<taxonomicName id="79FC4D11E16AAC29FE80FE17FD9412D5" box="[264,570,484,510]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yanahuicsa" status="sp. nov.">
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<emphasis id="8C88EA80E16AAC29FE80FE17FD9412D5" bold="true" box="[264,570,484,510]" italics="true" pageId="5" pageNumber="6">Haplohyphes yanahuicsa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="97BB57FBE16AAC29FDC9FE10FD3012D6" box="[577,670,483,509]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="26C72A17E16AAC29FD26FE10FC9612D6" box="[686,824,483,509]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="5" pageNumber="6">Figs. 2041</figureCitation>
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,
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<figureCitation id="26C72A17E16AAC29FCCEFE10FC2512D6" box="[838,907,483,509]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="5" pageNumber="6">4647</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="BE433692E16AAC29FE80FDDCFD061752" blockId="5.[264,1324,559,1225]" pageId="5" pageNumber="6">
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Male imago. Length: 4.0
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<quantity id="79049B77E16AAC29FDB2FDDCFD341162" box="[570,666,559,585]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="6" unit="mm" value="4.2">4.2 mm</quantity>
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; fore wings,
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<quantity id="79049B77E16AAC29FCB0FDDCFC691162" box="[824,967,559,585]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="3.9" metricValueMin="3.7" pageId="5" pageNumber="6" unit="mm" value="3.8" valueMax="3.9" valueMin="3.7">3.73.9 mm</quantity>
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. General coloration yellowish white, last segments of abdomen blackish. Head shaded with brownish gray from a transverse line at the median ocellus to posterior margin; antennae yellowish white. Thorax. Pronotum whitish translucent shaded almost completely with brownish gray, meso and metanotum whitish yellow with few very light gray marks; pleurae and sterna whitish yellow translucent shaded with light brownish on pro and mesopleurae and with gray in thoracic sterna. Legs: coxae I and II, fore femora and apex of fore tibiae and fore tarsi shaded with brownish gray, remaining segments of all legs whitish. Wings (
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<figureCitation id="26C72A17E16AAC29FBCCFCB4FB07104A" box="[1092,1193,839,865]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="5" pageNumber="6">Figs. 20</figureCitation>
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ac): membrane of wings hyaline, fore wings shaded with gray on C and Sc area, basally to bullae; longitudinal vein grayish, cross veins paler; hind wings only with two longitudinal veins. Abdomen (
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<figureCitation id="26C72A17E16AAC29FE06FC4CFE4910F2" box="[398,487,959,985]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="5" pageNumber="6">Fig. 46</figureCitation>
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) whitish translucent shaded with blackish gray on terga, shaded very slightly on first segments and becoming darker and more extensive toward rear segments (segments VIIX are much darker than the rest of body). Genitalia (
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<figureCitation id="26C72A17E16AAC29FB92FBFCFB0B1702" box="[1050,1189,1039,1065]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="5" pageNumber="6">Figs. 2124</figureCitation>
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): yellowish white shaded with gray on margins of styliger plate and forceps. Caudal filaments hyaline shaded with gray, except at joining.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6E66519E16AAC2BFEB0FB74FBA416CD" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
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<paragraph id="BE433692E16AAC29FEB0FB74FC1417E2" blockId="5.[264,1324,559,1225]" pageId="5" pageNumber="6">
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Female imago. Unknown, but two mature female nymphs ready to moult (
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<figureCitation id="26C72A17E16AAC29FB41FB74FA8D178A" box="[1225,1315,1159,1185]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="5" pageNumber="6">Fig. 47</figureCitation>
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) show the same characteristic coloration of the male imago.
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</paragraph>
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<caption id="EA83661AE16AAC29FE80F995FEF81400" pageId="5" pageNumber="6">
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<paragraph id="BE433692E16AAC29FE80F995FEF81400" blockId="5.[264,1323,1638,1835]" pageId="5" pageNumber="6">
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<emphasis id="8C88EA80E16AAC29FE80F995FE651555" bold="true" box="[264,459,1638,1662]" pageId="5" pageNumber="6">FIGURES 2041.</emphasis>
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<taxonomicName id="79FC4D11E16AAC29FE5BF994FD4E1555" box="[467,736,1639,1662]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yanahuicsa" status="sp. nov.">
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<emphasis id="8C88EA80E16AAC29FE5BF994FD4E1555" box="[467,736,1639,1662]" italics="true" pageId="5" pageNumber="6">Haplohyphes yanahuicsa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="97BB57FBE16AAC29FD60F995FC931556" box="[744,829,1638,1661]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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, male imago: 20a, fore wing; 20b, hind wing; 20c, idem, detail; 21, male genitalia, v.v.; 22, idem, l.v.; 23ab, penes, details v.v.; 24, detail of forceps, l.v.; nymph: 25, head, d.v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30a, labium, v.v.; 30b, detail of labial palp; 31, labrum, d.v.; 32, hypopharynx, v.v.; 33, left mandible, d.v.; 34, right mandible, d.v.; 35a, maxillae, d.v.; 35b, detail of maxillary palp; 36, gill II, d.v.; 3741, gills II VI, v.v.
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</paragraph>
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</caption>
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<paragraph id="BE433692E168AC2BFEB0FEEFFECD104D" blockId="7.[264,1324,284,1510]" pageId="7" pageNumber="8">
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Nymph (
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<figureCitation id="26C72A17E168AC2BFE2AFEEFFE52121D" box="[418,508,284,310]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="7" pageNumber="8">Fig. 47</figureCitation>
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). Length: body, 5.0
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<quantity id="79049B77E168AC2BFD78FEEFFCE1121D" box="[752,847,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="8" unit="mm" value="5.1">5.1 mm</quantity>
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; mesonotum,
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<quantity id="79049B77E168AC2BFC7FFEEFFBF8121D" box="[1015,1110,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="8" unit="mm" value="1.5">1.5 mm</quantity>
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; hind femora, 1.0 mm. General coloration whitish yellow with blackish marks. Head yellowish white surrounded by thick yellowish setae; shaded with blackish gray on a transversal band between eyes; occipute shaded more slightly with a reticular pattern; antennae whitish translucent; mouth parts whitish. Mouthparts as in
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<figureCitation id="26C72A17E168AC2BFD7BFE4FFCD512FD" box="[755,891,444,470]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="7" pageNumber="8">Figs 3035</figureCitation>
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. Thorax whitish yellow, pronotum shaded extensively with blackish gray, mesonotum shaded more slightly; wing buds shaded with black on basal half of costal margin. Thoracic sterna shaded with gray, metanotum and pleurae without shading. Legs (
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<figureCitation id="26C72A17E168AC2BFD64FDC7FCDD1165" box="[748,883,564,590]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="7" pageNumber="8">Figs. 2628</figureCitation>
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) whitish shaded with gray on coxae I and II; tarsal claws with double row of submarginal denticles near apex (
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<figureCitation id="26C72A17E168AC2BFBF8FDAFFB64115D" box="[1136,1226,604,630]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="7" pageNumber="8">Fig. 29</figureCitation>
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). Abdomen (
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<figureCitation id="26C72A17E168AC2BFEC3FD77FE0F11B5" box="[331,417,644,670]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="7" pageNumber="8">Fig. 47</figureCitation>
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) yellowish white shaded with light gray on terga IIV (sometimes also on V) and shaded with blackish gray on terga VIIX (sometimes also on V); tergum X slightly shaded with gray. Abdominal sterna paler shaded very slightly with gray. Opercullar gill (
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<figureCitation id="26C72A17E168AC2BFE99FD0FFE37103D" box="[273,409,764,790]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="7" pageNumber="8">Figs. 3637</figureCitation>
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) whitish translucent shaded completely with light gray, remaining gills (
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<figureCitation id="26C72A17E168AC2BFB78FD0FFEE31015" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="7" pageNumber="8">Figs. 3841</figureCitation>
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) whitish translucent. Caudal filaments whitish yellow with whorls of setae at each joining.
|
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</paragraph>
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<paragraph id="BE433692E168AC2BFEB0FC87FDA2109D" blockId="7.[264,1324,284,1510]" pageId="7" pageNumber="8">Etymology. Combination of letters including the Quechua voices “yana” (=black) and “huicsa” (=abdomen).</paragraph>
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<paragraph id="BE433692E168AC2BFEB0FC37FEF7178D" blockId="7.[264,1324,284,1510]" pageId="7" pageNumber="8">
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Material.
|
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<typeStatus id="61478830E168AC2BFE24FC37FDB710F5" box="[428,537,964,990]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
|
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male imago from
|
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<collectingCountry id="C6EB7602E168AC2BFD74FC37FCD410F5" box="[764,890,964,990]" name="Bolivia" pageId="7" pageNumber="8">BOLIVIA</collectingCountry>
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: Dept. La Paz, stream between Caranavi and Guanai,
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<geoCoordinate id="DBC85055E168AC2BFE6FFC1FFD29172D" box="[487,647,1004,1030]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="15" value="-15.671667">S 15° 40' 18"</geoCoordinate>
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<geoCoordinate id="DBC85055E168AC2BFD29FC1FFCE2172D" box="[673,844,1004,1030]" direction="west" orientation="longitude" pageId="7" pageNumber="8" precision="15" value="-67.70111">W 67° 42' 04"</geoCoordinate>
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,
|
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<quantity id="79049B77E168AC2BFCECFC1FFC1C172D" box="[868,946,1004,1030]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="m" value="500.0">500 m</quantity>
|
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,
|
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<date id="CA421052E168AC2BFC37FC1FFBE7172D" box="[959,1097,1004,1030]" pageId="7" pageNumber="8" value="2000-11-27">27/XI/2000</date>
|
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, E. Domínguez, C. Molineri y C. Nieto Cols.
|
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|
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<typeStatus id="61478830E168AC2BFDCDFBE7FD121705" box="[581,700,1044,1070]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
|
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:
|
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<specimenCount id="A8FAFD1BE168AC2BFD40FBE7FCB41705" box="[712,794,1044,1070]" pageId="7" pageNumber="8" type="male">3 male</specimenCount>
|
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imagos and
|
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|
|
<specimenCount id="A8FAFD1BE168AC2BFC3DFBE7FBB31705" box="[949,1053,1044,1070]" pageId="7" pageNumber="8" type="female">2 female</specimenCount>
|
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|
mature nymphs, same data as
|
|||
|
|
<typeStatus id="61478830E168AC2BFEE1FBCFFE7D177D" box="[361,467,1084,1110]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
|||
|
|
.
|
|||
|
|
<typeStatus id="61478830E168AC2BFE67FBCFFDF1177D" box="[495,607,1084,1110]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
|
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|
|
, 1
|
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|
|
<typeStatus id="61478830E168AC2BFD02FBCFFD41177D" box="[650,751,1084,1110]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
|
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|
|
male imago and 1
|
|||
|
|
<typeStatus id="61478830E168AC2BFC6EFBCFFBE5177D" box="[998,1099,1084,1110]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
|
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|
|
nymph housed in UMSA; remaining material in IFML. Nymphs and adults are associated by a pharate subimago.
|
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|
|
</paragraph>
|
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|
|
<paragraph id="BE433692E168AC2BFEB0FB47FBA416CD" blockId="7.[264,1324,284,1510]" pageId="7" pageNumber="8">
|
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|
|
Discussion.
|
|||
|
|
<taxonomicName id="79FC4D11E168AC2BFE4EFB47FD4317E6" box="[454,749,1204,1229]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yanahuicsa" status="sp. nov.">
|
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|
|
<emphasis id="8C88EA80E168AC2BFE4EFB47FD4317E6" box="[454,749,1204,1229]" italics="true" pageId="7" pageNumber="8">Haplohyphes yanahuicsa</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="97BB57FBE168AC2BFD7DFB47FCE117E5" box="[757,847,1204,1230]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
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|
|
can be distinguished from the other species of the genus by the following combination of characters. In the imago: 1) hind wings with two longitudinal veins (
|
|||
|
|
<figureCitation id="26C72A17E168AC2BFDEFFAF7FD6C1635" box="[615,706,1284,1310]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="7" pageNumber="8">Fig. 20</figureCitation>
|
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|
c); 2) segment 1 and 2 of forceps similar in length (
|
|||
|
|
<figureCitation id="26C72A17E168AC2BFE99FADFFEC5166D" box="[273,363,1324,1350]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="7" pageNumber="8">Fig. 24</figureCitation>
|
|||
|
|
); 3) penes divided on apical 0.22 (
|
|||
|
|
<figureCitation id="26C72A17E168AC2BFC9CFADFFCD9166D" box="[788,887,1324,1350]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="7" pageNumber="8">Figs. 23</figureCitation>
|
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|
|
ab); 4) membranous lobes of penes rounded (
|
|||
|
|
<figureCitation id="26C72A17E168AC2BFEF3FAA7FE7D1645" box="[379,467,1364,1390]" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="7" pageNumber="8">Fig. 23</figureCitation>
|
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|
|
ab); 5) distal spines of penes shorter than membranous lobes; 6) male fore wing length: 4.0 mm; 7) general coloration pale except apical segments of abdomen blackish (
|
|||
|
|
<figureCitation id="26C72A17E168AC2BFEF2FA57FE7D1695" box="[378,467,1444,1470]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="7" pageNumber="8">Fig. 46</figureCitation>
|
|||
|
|
). In the nymph: 1) markings on head not reaching base of antennae (
|
|||
|
|
<figureCitation id="26C72A17E168AC2BFB73FA57FE8716CD" captionStart="FIGURES 20 41" captionStartId="5.[264,382,1638,1662]" captionText="FIGURES 20 41. Haplohyphes yanahuicsa sp. nov., male imago: 20 a, fore wing; 20 b, hind wing; 20 c, idem, detail; 21, male genitalia, v. v.; 22, idem, l. v.; 23 ab, penes, details v. v.; 24, detail of forceps, l. v.; nymph: 25, head, d. v.; 26, leg III; 27, leg II; 28, leg I; 29, fore tarsal claw; 30 a, labium, v. v.; 30 b, detail of labial palp; 31, labrum, d. v.; 32, hypopharynx, v. v.; 33, left mandible, d. v.; 34, right mandible, d. v.; 35 a, maxillae, d. v.; 35 b, detail of maxillary palp; 36, gill II, d. v.; 37 41, gills II VI, v. v." pageId="7" pageNumber="8">Fig. 25</figureCitation>
|
|||
|
|
); 2) abdomen shaded with blackish on rear segments (
|
|||
|
|
<figureCitation id="26C72A17E168AC2BFC2DFA3FFC5216CD" box="[933,1020,1484,1510]" captionStart="FIGURES 46 - 47" captionStartId="9.[265,381,1437,1461]" captionTargetBox="[406,1181,313,1427]" captionTargetId="figure@9.[408,1181,313,1082]" captionTargetPageId="9" captionText="FIGURES 46 - 47. Haplohyphes yanahuicsa sp. nov.: 46, general view of male imago; 47, general view of nymph." httpUri="https://zenodo.org/record/168751/files/figure.png" pageId="7" pageNumber="8">Fig. 47</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|