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<document id="FAE554324A321BDDD7E2EF9E345E7E3C" ID-CLB-Dataset="8744" ID-DOI="10.11646/zootaxa.4838.2.1" ID-GBIF-Dataset="761fcec7-c79d-4e80-bb2b-23cc9e78e0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403783" ID-ZooBank="0AD43D95-7508-466C-980F-CBEF38CA1198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352386978" checkinUser="plazi" docAuthor="Gompel, Nicolas" docDate="2020" docId="039187D8FFCCBB1AFF1882EDFAEFFD56" docLanguage="en" docName="zootaxa.4838.2.1.pdf" docOrigin="Zootaxa 4838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zarcosia ilonae Gompel 2020, n. sp." docType="treatment" docUuid="48485295-DAF4-46BA-A640-65DA672835E8" docUuidSource="ZooBank" docVersion="8" lastPageNumber="162" masterDocId="FFA8FFA0FFC6BB11FF8F844AFF9DFFFF" masterDocTitle="Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="151" pageNumber="161" updateTime="1698850145988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="59E8BA86AC72F97AE51B049FC12280ED">Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species</mods:title>
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<mods:namePart id="D1BA076FABF2ABB9B8AD8F62E4F43D38">Gompel, Nicolas</mods:namePart>
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<mods:title id="CECF1B9083FDEB48F5FD8A2F511AC67C">Zootaxa</mods:title>
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<mods:date id="74421D79C2336BC0C8260B61A19AC424">2020</mods:date>
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<mods:number id="2E4D5D081E766776939366150FBFEB5E">2020-08-26</mods:number>
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<mods:classification id="380909491538C0BFD39886DED5C7387F">journal article</mods:classification>
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<mods:identifier id="2745ABD193761AACB6B49EA4300910EA" type="DOI">10.11646/zootaxa.4838.2.1</mods:identifier>
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<treatment id="039187D8FFCCBB1AFF1882EDFAEFFD56" ID-DOI="http://doi.org/10.5281/zenodo.4404253" ID-GBIF-Taxon="177012517" ID-Zenodo-Dep="4404253" LSID="urn:lsid:zoobank.org:act:48485295-DAF4-46BA-A640-65DA672835E8" httpUri="http://treatment.plazi.org/id/039187D8FFCCBB1AFF1882EDFAEFFD56" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161">
<subSubSection id="C3226545FFCCBB1BFF1882EDFE00F93E" box="[151,413,1702,1729]" pageId="10" pageNumber="161" type="nomenclature">
<paragraph id="8B8736CEFFCCBB1BFF1882EDFE00F93E" blockId="10.[151,444,1702,1766]" box="[151,413,1702,1729]" pageId="10" pageNumber="161">
<heading id="D0CF81A2FFCCBB1BFF1882EDFE00F93E" bold="true" box="[151,413,1702,1729]" fontSize="11" level="1" pageId="10" pageNumber="161" reason="1">
<emphasis id="B94CEADCFFCCBB1BFF1882EDFE00F93E" bold="true" box="[151,413,1702,1729]" pageId="10" pageNumber="161">
<taxonomicName id="4C384D4DFFCCBB1BFF1882EDFECEF93E" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[151,339,1702,1729]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="ilonae" status="sp. nov.">
<emphasis id="B94CEADCFFCCBB1BFF1882EDFECEF93E" bold="true" box="[151,339,1702,1729]" italics="true" pageId="10" pageNumber="161">Zarcosia ilonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57A7FFCCBB1BFED482EDFE00F93E" box="[347,413,1703,1729]" pageId="10" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFCCBB1BFF188281FE21F919" box="[151,444,1738,1766]" pageId="10" pageNumber="161" type="description">
<paragraph id="8B8736CEFFCCBB1BFF188281FE21F919" blockId="10.[151,444,1702,1766]" box="[151,444,1738,1766]" pageId="10" pageNumber="161">
(
<figureCitation id="13032A4BFFCCBB1BFF108281FE96F91A" box="[159,267,1739,1765]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="10" pageNumber="161">Figures 1</figureCitation>
;
<figureCitation id="13032A4BFFCCBB1BFE978281FEA8F91A" box="[280,309,1739,1765]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="10" pageNumber="161">3c</figureCitation>
;
<figureCitation id="13032A4BFFCCBB1BFECD8281FEFDF91A" box="[322,352,1739,1765]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="10" pageNumber="161">
<quantity id="4CC09B2BFFCCBB1BFECD8281FEFDF91A" box="[322,352,1739,1765]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="10" pageNumber="161" unit="g" value="4.0">4g</quantity>
</figureCitation>
;
<figureCitation id="13032A4BFFCCBB1BFEE38280FE16F91A" box="[364,395,1738,1765]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="10" pageNumber="161">5k</figureCitation>
;
<figureCitation id="13032A4BFFCCBB1BFE188281FE29F919" box="[407,436,1739,1766]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="10" pageNumber="161">6b</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFCCBB1BFF188359FEF9F8AE" pageId="10" pageNumber="161" type="etymology">
<paragraph id="8B8736CEFFCCBB1BFF188359FEF9F8AE" blockId="10.[151,1437,1811,2017]" pageId="10" pageNumber="161">
<emphasis id="B94CEADCFFCCBB1BFF188359FE83F8D2" bold="true" box="[151,286,1811,1837]" pageId="10" pageNumber="161">Etymology.</emphasis>
The species is named after my wife, Ilona Grunwald, to acknowledge her patience and support of my work with insects.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFCCBB1BFF488311FAADF88A" box="[199,1328,1882,1909]" pageId="10" pageNumber="161" type="materials_examined">
<paragraph id="8B8736CEFFCCBB1BFF488311FAADF88A" blockId="10.[151,1437,1811,2017]" box="[199,1328,1882,1909]" pageId="10" pageNumber="161">
<materialsCitation id="3B503C93FFCCBB1BFF488311FAB1F88A" ID-GBIF-Occurrence="3020160303" box="[199,1324,1882,1909]" country="Thailand" latitude="8.866667" location="Tham Wararam" longLatPrecision="1300" longitude="98.65" municipality="Phanom District" pageId="10" pageNumber="161" specimenCount="1" stateProvince="Surat Thani" typeStatus="holotype">
<emphasis id="B94CEADCFFCCBB1BFF488311FEFBF88B" bold="true" box="[199,358,1882,1909]" pageId="10" pageNumber="161">
<typeStatus id="5483886CFFCCBB1BFF488311FE9FF88A" box="[199,258,1883,1909]" pageId="10" pageNumber="161">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32F765EFFCCBB1BFEE38311FE49F88A" box="[364,468,1883,1909]" name="Thailand" pageId="10" pageNumber="161">Thailand</collectingCountry>
:
<collectingRegion id="49FCF82CFFCCBB1BFE508311FD52F88A" box="[479,719,1883,1909]" country="Thailand" name="Surat Thani" pageId="10" pageNumber="161">Surat Thani Province</collectingRegion>
,
<collectingMunicipality id="6BE3ACB4FFCCBB1BFD558311FC09F88A" box="[730,916,1883,1909]" pageId="10" pageNumber="161">Phanom District</collectingMunicipality>
,
<location id="8EE76015FFCCBB1BFC118311FBD2F88A" LSID="urn:lsid:plazi:treatment:039187D8FFCCBB1AFF1882EDFAEFFD56:8EE76015FFCCBB1BFC118311FBD2F88A" box="[926,1103,1883,1909]" country="Thailand" latitude="8.866667" longLatPrecision="1300" longitude="98.65" municipality="Phanom District" name="Tham Wararam" pageId="10" pageNumber="161" stateProvince="Surat Thani">Tham Wararam</location>
,
<geoCoordinate id="EE0C5009FFCCBB1BFBD58310FB22F88A" box="[1114,1215,1882,1909]" degrees="08" direction="north" minutes="52" orientation="latitude" pageId="10" pageNumber="161" precision="925" value="8.866667">08°52’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFCCBB1BFB488311FAB1F88A" box="[1223,1324,1883,1909]" degrees="98" direction="east" minutes="39" orientation="longitude" pageId="10" pageNumber="161" precision="925" value="98.65">98°39’ E</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFCCBB1BFF488334FDD4F866" box="[199,585,1918,1945]" pageId="10" pageNumber="161" type="distribution">
<paragraph id="8B8736CEFFCCBB1BFF488334FDD4F866" blockId="10.[151,1437,1811,2017]" box="[199,585,1918,1945]" pageId="10" pageNumber="161">
<emphasis id="B94CEADCFFCCBB1BFF488334FEFCF867" bold="true" box="[199,353,1918,1944]" pageId="10" pageNumber="161">Distribution.</emphasis>
<collectingCountry id="F32F765EFFCCBB1BFEE88335FE56F866" box="[359,459,1919,1945]" name="Thailand" pageId="10" pageNumber="161">Thailand</collectingCountry>
(
<figureCitation id="13032A4BFFCCBB1BFE568335FDA5F866" box="[473,568,1919,1945]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="10" pageNumber="161">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFCCBB1AFF4883E8FE29FDC2" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161" type="description">
<paragraph id="8B8736CEFFCCBB1BFF4883E8FBFAF81E" blockId="10.[151,1437,1811,2017]" pageId="10" pageNumber="161">
<emphasis id="B94CEADCFFCCBB1BFF4883E8FEC4F843" bold="true" box="[199,345,1954,1980]" pageId="10" pageNumber="161">Description.</emphasis>
(
<figureCitation id="13032A4BFFCCBB1BFEE683E9FE44F842" box="[361,473,1955,1981]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="10" pageNumber="161">Figure 3c</figureCitation>
). Female unknown. Body length.
<specimenCount id="9D3EFD47FFCCBB1BFCD283E9FCEFF842" box="[861,882,1955,1981]" pageId="10" pageNumber="161" type="male">♂</specimenCount>
<typeStatus id="5483886CFFCCBB1BFCF583E9FC7DF842" box="[890,992,1955,1981]" pageId="10" pageNumber="161" type="holotype">holotype</typeStatus>
:
<quantity id="4CC09B2BFFCCBB1BFC6383E8FBC8F842" box="[1004,1109,1954,1981]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="10" pageNumber="161" unit="mm" value="2.55">2.55 mm</quantity>
. Slender, orange-brown with darker head, covered with very short, pulverulent, white pubescence. Appendages pale.
</paragraph>
<paragraph id="8B8736CEFFCDBB1AFF4884DDFB1AFF2A" blockId="11.[151,1436,151,681]" pageId="11" pageNumber="162">
<emphasis id="B94CEADCFFCDBB1AFF4884DDFE99FF4E" box="[199,260,151,177]" italics="true" pageId="11" pageNumber="162">Head</emphasis>
. Moderately transverse, barely wider than pronotum. Palpi yellow, last article securiform. Finely and densely punctuated. Occiput lined with a fringe of dense and short white hairs posteriorly.
</paragraph>
<paragraph id="8B8736CEFFCDBB1AFF488495FAD1FF06" blockId="11.[151,1436,151,681]" box="[199,1356,222,249]" pageId="11" pageNumber="162">
<emphasis id="B94CEADCFFCDBB1AFF488495FEACFF06" box="[199,305,223,249]" italics="true" pageId="11" pageNumber="162">Antennae</emphasis>
. 1.5 times longer than elytra, entirely yellowish, filiform with antenommeres 3–10 elongated.
</paragraph>
<paragraph id="8B8736CEFFCDBB1AFF488549FEAFFEBE" blockId="11.[151,1436,151,681]" pageId="11" pageNumber="162">
<emphasis id="B94CEADCFFCDBB1AFF488549FEABFEE3" box="[199,310,259,284]" italics="true" pageId="11" pageNumber="162">Pronotum</emphasis>
(
<figureCitation id="13032A4BFFCDBB1AFEC98549FE25FEE1" box="[326,440,259,286]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="11" pageNumber="162">Figure 6b</figureCitation>
). Convex, slightly longer than wide, sides regularly curved, wider, almost gibbose in its anterior third.
</paragraph>
<paragraph id="8B8736CEFFCDBB1AFF488501FD13FE52" blockId="11.[151,1436,151,681]" pageId="11" pageNumber="162">
<emphasis id="B94CEADCFFCDBB1AFF488501FE90FE9A" box="[199,269,331,357]" italics="true" pageId="11" pageNumber="162">Elytra</emphasis>
. Length:
<quantity id="4CC09B2BFFCDBB1AFEF68501FE7DFE9A" box="[377,480,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="162" unit="mm" value="1.8">1.80 mm</quantity>
. Punctation strong and dense, with larger points. Space between points slightly wider than the points themselves. Disc subconvex. Pubescence dense and short, comprised of very short hairs borne from the points and interspersed pulverulent hairs.
</paragraph>
<paragraph id="8B8736CEFFCDBB1AFF4885FDFAA1FE2E" blockId="11.[151,1436,151,681]" box="[199,1340,438,465]" pageId="11" pageNumber="162">
<emphasis id="B94CEADCFFCDBB1AFF4885FDFF61FE2F" box="[199,252,439,464]" italics="true" pageId="11" pageNumber="162">Legs</emphasis>
. Particularly elongated. Foretibia weakly broadened in the middle of their inner side (
<figureCitation id="13032A4BFFCDBB1AFB3185FDFAB2FE2E" box="[1214,1327,438,465]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="11" pageNumber="162">Figure 5k</figureCitation>
).
</paragraph>
<paragraph id="8B8736CEFFCDBB1AFF488591FE29FDC2" blockId="11.[151,1436,151,681]" pageId="11" pageNumber="162">
<emphasis id="B94CEADCFFCDBB1AFF488591FEA9FE0A" box="[199,308,475,501]" italics="true" pageId="11" pageNumber="162">Aedeagus</emphasis>
(
<figureCitation id="13032A4BFFCDBB1AFECD8591FE2FFE0A" box="[322,434,475,501]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="11" pageNumber="162">
Figure
<quantity id="4CC09B2BFFCDBB1AFE1C8591FE2FFE0A" box="[403,434,475,501]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="11" pageNumber="162" unit="g" value="4.0">4g</quantity>
</figureCitation>
). Sturdy, lightly pigmented, sides divergent from base in dorsal view, then largely rounded near apex and ending in a broad stinger. Nearly flat in lateral view, with no kink. Lateral appendages thick and as long as the apical stinger.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFCDBB1AFF48860DFAEFFD56" pageId="11" pageNumber="162" type="materials_examined">
<paragraph id="8B8736CEFFCDBB1AFF48860DFC6BFD7A" blockId="11.[151,1436,151,681]" pageId="11" pageNumber="162">
<emphasis id="B94CEADCFFCDBB1AFF48860DFEE5FD9F" bold="true" box="[199,376,582,609]" pageId="11" pageNumber="162">Type material.</emphasis>
<materialsCitation id="3B503C93FFCDBB1AFE0C860DFC6BFD7A" ID-GBIF-Occurrence="3020160323" collectingDate="1987-07-24" collectionCode="MNHN, A" collectorName="L. Deharveng &amp; Bedos" country="Thailand" county="Phanom District" elevation="250" latitude="8.866667" location="Cave" longLatPrecision="1300" longitude="98.65" municipality="Tham Wararam" pageId="11" pageNumber="162" specimenCount="1" specimenCount-male="1" stateProvince="Surat Thani" typeStatus="holotype">
<typeStatus id="5483886CFFCDBB1AFE0C860DFE76FD9E" box="[387,491,583,609]" pageId="11" pageNumber="162" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D3EFD47FFCDBB1AFE70860DFD8BFD9E" box="[511,534,583,609]" pageId="11" pageNumber="162" type="male">♂</specimenCount>
,
<collectionCode id="ED29AE0BFFCDBB1AFDAA860DFD1DFD9E" box="[549,640,583,609]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="162">MNHN</collectionCode>
).
<emphasis id="B94CEADCFFCDBB1AFD1A860CFC9FFD9F" bold="true" box="[661,770,582,608]" pageId="11" pageNumber="162">
<collectingCountry id="F32F765EFFCDBB1AFD1A860CFC9FFD9F" box="[661,770,582,608]" name="Thailand" pageId="11" pageNumber="162">Thailand</collectingCountry>
</emphasis>
:
<collectingRegion id="49FCF82CFFCDBB1AFC9B860DFB97FD9E" box="[788,1034,583,609]" country="Thailand" name="Surat Thani" pageId="11" pageNumber="162">Surat Thani Province</collectingRegion>
,
<collectingCounty id="62E64E42FFCDBB1AFB96860DFB4BFD9E" box="[1049,1238,583,609]" pageId="11" pageNumber="162">Phanom District</collectingCounty>
,
<collectingMunicipality id="6BE3ACB4FFCDBB1AFB6B860DFA05FD9E" box="[1252,1432,583,609]" pageId="11" pageNumber="162">Tham Wararam</collectingMunicipality>
,
<location id="8EE76015FFCDBB1AFF188621FF4EFD7A" LSID="urn:lsid:plazi:treatment:039187D8FFCCBB1AFF1882EDFAEFFD56:8EE76015FFCDBB1AFF188621FF4EFD7A" box="[151,211,619,645]" country="Thailand" county="Phanom District" latitude="8.866667" longLatPrecision="1300" longitude="98.65" municipality="Tham Wararam" name="Cave" pageId="11" pageNumber="162" stateProvince="Surat Thani">Cave</location>
,
<geoCoordinate id="EE0C5009FFCDBB1AFF508620FED9FD7A" box="[223,324,618,645]" degrees="08" direction="north" minutes="52" orientation="latitude" pageId="11" pageNumber="162" precision="925" value="8.866667">08°52’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFCDBB1AFEC48621FE33FD7A" box="[331,430,619,645]" degrees="98" direction="east" minutes="39" orientation="longitude" pageId="11" pageNumber="162" precision="925" value="98.65">98°39’ E</geoCoordinate>
(
<quantity id="4CC09B2BFFCDBB1AFE328620FD9BFD7A" box="[445,518,618,645]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="162" unit="m" value="250.0">
<elevation id="0015D1FDFFCDBB1AFE328620FD9BFD7A" box="[445,518,618,645]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="162" unit="m" value="250.0">250 m</elevation>
</quantity>
) (
<collectorName id="26CD5318FFCDBB1AFD918621FD24FD7A" box="[542,697,619,645]" pageId="11" pageNumber="162">L. Deharveng</collectorName>
and
<collectionCode id="ED29AE0BFFCDBB1AFD628621FC99FD7A" box="[749,772,619,645]" pageId="11" pageNumber="162">A</collectionCode>
.
<collectorName id="26CD5318FFCDBB1AFC9F8621FCC5FD79" box="[784,856,619,646]" pageId="11" pageNumber="162">Bedos</collectorName>
)
<date id="FF86100EFFCDBB1AFCE98621FC6EFD7A" box="[870,1011,619,645]" pageId="11" pageNumber="162" value="1987-07-24">
<collectingDate id="EFC2E9E6FFCDBB1AFCE98621FC6EFD7A" box="[870,1011,619,645]" pageId="11" pageNumber="162" value="1987-07-24">24.VII.1987</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8736CEFFCDBB1AFF4886C5FAEFFD56" blockId="11.[151,1436,151,681]" box="[199,1394,655,681]" pageId="11" pageNumber="162">
<emphasis id="B94CEADCFFCDBB1AFF4886C5FF62FD56" bold="true" box="[199,255,655,681]" pageId="11" pageNumber="162">Note</emphasis>
. The extremely thin and elongated appendages of species likely represent adaptations to life in caves.
</paragraph>
</subSubSection>
</treatment>
</document>
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