166 lines
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166 lines
20 KiB
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<document id="FCAEC17E830998E402EB3270421D6F1A" ID-CLB-Dataset="68941" ID-DOI="10.5281/zenodo.5184182" ID-GBIF-Dataset="dcb37aba-9f9f-402a-91db-c1c6e310d61c" ID-ISSN="0253-116X" ID-Zenodo-Dep="5184182" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" checkinTime="1628735848729" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2015" docId="03915A153A20FFB9FF0AEE93FB88FA2F" docLanguage="en" docName="LinzerbiolBeitr.47.2.1133-1178.pdf" docOrigin="Linzer biologische Beiträge 47 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Lithocharis bitriangulata Assing 2015, nov.sp." docType="treatment" docVersion="10" lastPageNumber="1165" masterDocId="FFA8226D3A00FF99FFB8EF41FFB6FFD8" masterDocTitle="A revison of the Lithocharis species of the Palaearctic, Oriental and Australian regions (Coleoptera: Staphylinidae: Paederinae: Medonina)" masterLastPageNumber="1178" masterPageNumber="1133" pageNumber="1165" updateTime="1699050958296" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FD604C45BE22C8378F5C3D817789630C">A revison of the Lithocharis species of the Palaearctic, Oriental and Australian regions (Coleoptera: Staphylinidae: Paederinae: Medonina)</mods:title>
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<mods:namePart id="49B890508967FDBF0ADE4F19651F5E76">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="C2BA1838A580C83B652CAFAB7F3F6F6B">text</mods:typeOfResource>
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<mods:title id="3BC0771C28740321C24B3B07B475E02C">Linzer biologische Beiträge</mods:title>
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<mods:part id="1308B0E0FC9516771828686C2F76ED0B">
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<mods:date id="74A476238DFEC63D252E8DFB5664519D">2015</mods:date>
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<mods:number id="242D62EB0F3781125CDCCD4489AF57C9">2015-12-30</mods:number>
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<mods:number id="3FDB7E8473E4248077C0615CC01842D9">2</mods:number>
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<mods:identifier id="948AE0E7C14D1ADCBC7C40A2369629EB" type="CLB-Dataset">68941</mods:identifier>
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<mods:identifier id="1D676465603730E2FA852DA6403EA5B7" type="DOI">10.5281/zenodo.5184182</mods:identifier>
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<treatment id="03915A153A20FFB9FF0AEE93FB88FA2F" ID-DOI="http://doi.org/10.5281/zenodo.6631563" ID-GBIF-Taxon="195832407" ID-Zenodo-Dep="6631563" LSID="urn:lsid:plazi:treatment:03915A153A20FFB9FF0AEE93FB88FA2F" httpUri="http://treatment.plazi.org/id/03915A153A20FFB9FF0AEE93FB88FA2F" lastPageNumber="1165" pageId="32" pageNumber="1165">
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<subSubSection id="C322B8883A20FFB9FF0AEE93FDBAFE30" box="[178,524,466,488]" pageId="32" pageNumber="1165" type="nomenclature">
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<paragraph id="8B87EB033A20FFB9FF0AEE93FDBAFE30" blockId="32.[178,657,466,489]" box="[178,524,466,488]" pageId="32" pageNumber="1165">
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<heading id="D0CF5C6F3A20FFB9FF0AEE93FDBAFE30" box="[178,524,466,488]" fontSize="8" level="2" pageId="32" pageNumber="1165" reason="2">
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<emphasis id="B94C37113A20FFB9FF0AEE93FDBAFE30" bold="true" box="[178,524,466,488]" pageId="32" pageNumber="1165">
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<taxonomicName id="4C3890803A20FFB9FF0AEE93FE0FFE30" ID-CoL="3VK8F" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" box="[178,441,466,488]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1165" phylum="Arthropoda" rank="species" species="bitriangulata" status="sp. nov.">
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<emphasis id="B94C37113A20FFB9FF0AEE93FE0FFE30" bold="true" box="[178,441,466,488]" italics="true" pageId="32" pageNumber="1165">Lithocharis bitriangulata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27F8A6A3A20FFB9FE78EE92FDBAFE30" box="[448,524,467,488]" pageId="32" pageNumber="1165" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C322B8883A20FFB9FDAAEE92FD27FE31" box="[530,657,467,489]" pageId="32" pageNumber="1165" type="description">
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<paragraph id="8B87EB033A20FFB9FDAAEE92FD27FE31" blockId="32.[178,657,466,489]" box="[530,657,467,489]" pageId="32" pageNumber="1165">
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<heading id="D0CF5C6F3A20FFB9FDAAEE92FD27FE31" box="[530,657,467,489]" fontSize="8" level="2" pageId="32" pageNumber="1165" reason="2">
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<emphasis id="B94C37113A20FFB9FDAAEE92FD27FE31" bold="true" box="[530,657,467,489]" pageId="32" pageNumber="1165">
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(
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<figureCitation id="1303F7863A20FFB9FDA1EE92FD3DFE31" box="[537,651,467,489]" captionStart="Figs 90-94" captionStartId="33.[178,218,1272,1294]" captionTargetBox="[186,1033,212,1250]" captionTargetId="figure-1@33.[178,1091,212,1263]" captionTargetPageId="33" captionText="Figs 90-94: Lithocharis bitriangulata nov.sp.: (90) forebody; (91) male sternite VII; (92) male sternite VIII; (93-94) aedeagus in lateral and in ventral view. Scale bars: 90: 1.0 mm; 91-94: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184212" httpUri="https://zenodo.org/record/5184212/files/figure.png" pageId="32" pageNumber="1165">Figs 90-94</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C322B8883A20FFB9FF0AEEB9FB9EFDFD" pageId="32" pageNumber="1165" type="materials_examined">
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<paragraph id="8B87EB033A20FFB9FF0AEEB9FB9EFDFD" blockId="32.[178,1085,504,549]" pageId="32" pageNumber="1165">
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Type material:
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<materialsCitation id="3B50E15E3A20FFB9FE14EEB9FB9EFDFD" ID-GBIF-Occurrence="3805307371" collectingDate="2001-06-01" collectingDateMax="2001-06-04" collectingDateMin="2001-06-01" collectionCode="NHMB" collectorName="Bolm" determinerName="V. Assing" elevation="1200" location="Rana lake" pageId="32" pageNumber="1165" specimenCount="1" stateProvince="Sumatra" typeStatus="holotype">
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<typeStatus id="548355A13A20FFB9FE14EEB9FDB6FDD6" box="[428,512,504,526]" pageId="32" pageNumber="1165" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D3E208A3A20FFB9FDB2EEB9FDABFDD5" box="[522,541,504,525]" pageId="32" pageNumber="1165" type="male">♂</specimenCount>
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: "
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<collectingRegion id="49FC25E13A20FFB9FD8CEEB9FD34FDD5" box="[564,642,504,525]" country="Indonesia" name="Sumatera Utara" pageId="32" pageNumber="1165">Sumatra</collectingRegion>
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(S),
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<locationDeviation id="9A35F2213A20FFB9FD0FEEB9FC9BFDD6" box="[695,813,504,526]" pageId="32" pageNumber="1165">SW coast of</locationDeviation>
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<location id="8EE7BDD83A20FFB9FC8FEEB9FC21FDD6" LSID="urn:lsid:plazi:treatment:03915A153A20FFB9FF0AEE93FB88FA2F:8EE7BDD83A20FFB9FC8FEEB9FC21FDD6" box="[823,919,504,526]" name="Rana lake" pageId="32" pageNumber="1165" stateProvince="Sumatra">Rana lake</location>
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,
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<date id="FF86CDC33A20FFB9FC1CEEB9FB8CFDD5" box="[932,1082,504,525]" pageId="32" pageNumber="1165" value="2001-06-01" valueMax="2001-06-04" valueMin="2001-06-01">
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<collectingDate id="EFC2342B3A20FFB9FC1CEEB9FB8CFDD5" box="[932,1082,504,525]" pageId="32" pageNumber="1165" value="2001-06-01" valueMax="2001-06-04" valueMin="2001-06-01">1.-4. June 2001</collectingDate>
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</date>
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,
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<collectorName id="26CD8ED53A20FFB9FF0AED4FFF52FDFC" box="[178,228,526,548]" pageId="32" pageNumber="1165">Bolm</collectorName>
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lgt.,
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<quantity id="4CC046E63A20FFB9FEABED4EFEEEFDFC" box="[275,344,527,548]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="1165" unit="m" value="1200.0">
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<elevation id="00150C303A20FFB9FEABED4EFEEEFDFC" box="[275,344,527,548]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="1165" unit="m" value="1200.0">1200 m</elevation>
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</quantity>
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/ Holotypus ♂
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<originalDetermination id="A82653FF3A20FFB9FE5FED4FFCB2FDFC" box="[487,772,526,548]" pageId="32" pageNumber="1165">
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<taxonomicName id="4C3890803A20FFB9FE5FED4FFD7BFDFB" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" box="[487,717,526,547]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1165" phylum="Arthropoda" rank="species" species="bitriangulata" status="sp. nov.">
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<emphasis id="B94C37113A20FFB9FE5FED4FFD7BFDFB" box="[487,717,526,547]" italics="true" pageId="32" pageNumber="1165">Lithocharis bitriangulata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27F8A6A3A20FFB9FD6AED4EFCB2FDFC" box="[722,772,527,548]" pageId="32" pageNumber="1165" rank="species">sp. n.</taxonomicNameLabel>
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</originalDetermination>
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det.
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<determinerName id="7E5B07C73A20FFB9FC88ED4EFC3DFDFC" box="[816,907,526,548]" pageId="32" pageNumber="1165">V. Assing</determinerName>
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2013" (
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<collectionCode id="ED2973C63A20FFB9FC6CED4EFBABFDFD" box="[980,1053,527,549]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="32" pageNumber="1165">NHMB</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C322B8883A20FFB9FF0AED73FDA5FDBE" pageId="32" pageNumber="1165" type="etymology">
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<paragraph id="8B87EB033A20FFB9FF0AED73FDA5FDBE" blockId="32.[178,1085,562,614]" pageId="32" pageNumber="1165">Etymology: The specific epithet (adjective) alludes to the pair of triangular apical internal structures of the aedeagus.</paragraph>
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</subSubSection>
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<subSubSection id="C322B8883A20FFB9FF0AED34FC6BFAC1" pageId="32" pageNumber="1165" type="description">
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<paragraph id="8B87EB033A20FFB9FF0AED34FE62FD3B" blockId="32.[178,1085,629,739]" pageId="32" pageNumber="1165">
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Description: Body length
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<quantity id="4CC046E63A20FFB9FDACED34FDD3FD52" box="[532,613,629,650]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="32" pageNumber="1165" unit="mm" value="4.3">4.3 mm</quantity>
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; length of forebody
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<quantity id="4CC046E63A20FFB9FC8FED34FC3EFD52" box="[823,904,629,650]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="32" pageNumber="1165" unit="mm" value="2.3">2.3 mm</quantity>
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. Coloration: head blackish; pronotum dark-reddish; elytra yellowish-brown with indistinctly delimited yellowish margins; abdomen brown, with the margins of the segments yellowish; legs yellowish; antennae reddish.
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</paragraph>
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<paragraph id="8B87EB033A20FFB9FF0AEDB3FD3EFCB8" blockId="32.[178,1085,754,864]" pageId="32" pageNumber="1165">
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Head (
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<figureCitation id="1303F7863A20FFB9FF4FEDB3FEF2FCD0" box="[247,324,754,776]" captionStart="Figs 90-94" captionStartId="33.[178,218,1272,1294]" captionTargetBox="[186,1033,212,1250]" captionTargetId="figure-1@33.[178,1091,212,1263]" captionTargetPageId="33" captionText="Figs 90-94: Lithocharis bitriangulata nov.sp.: (90) forebody; (91) male sternite VII; (92) male sternite VIII; (93-94) aedeagus in lateral and in ventral view. Scale bars: 90: 1.0 mm; 91-94: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184212" httpUri="https://zenodo.org/record/5184212/files/figure.png" pageId="32" pageNumber="1165">Fig. 90</figureCitation>
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) transverse, 1.08 times as broad as long, indistinctly dilated behind eyes; punctation dense and fine; interstices without distinct microsculpture. Eyes large, distinctly longer than postocular portion in dorsal view. Antennae
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<quantity id="4CC046E63A20FFB9FCFAEC6DFC2BFC9A" box="[834,925,812,834]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="32" pageNumber="1165" unit="mm" value="1.45">1.45 mm</quantity>
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long; preapical antennomeres approximately as long as broad.
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</paragraph>
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<paragraph id="8B87EB033A20FFB9FF0AEC31FC7EFC18" blockId="32.[178,1085,879,960]" pageId="32" pageNumber="1165">
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Pronotum (
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<figureCitation id="1303F7863A20FFB9FE9DEC2EFEC7FC5D" box="[293,369,879,901]" captionStart="Figs 90-94" captionStartId="33.[178,218,1272,1294]" captionTargetBox="[186,1033,212,1250]" captionTargetId="figure-1@33.[178,1091,212,1263]" captionTargetPageId="33" captionText="Figs 90-94: Lithocharis bitriangulata nov.sp.: (90) forebody; (91) male sternite VII; (92) male sternite VIII; (93-94) aedeagus in lateral and in ventral view. Scale bars: 90: 1.0 mm; 91-94: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184212" httpUri="https://zenodo.org/record/5184212/files/figure.png" pageId="32" pageNumber="1165">Fig. 90</figureCitation>
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) 1.1 times as wide as long and 1.02 times as broad as head; posterior angles rounded, weakly marked; punctation similar to that of head; midline with indistinctly delimited, narrow impunctate band; interstices without microsculpture.
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</paragraph>
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<paragraph id="8B87EB033A20FFB9FF0AEC8EFC02FBC7" blockId="32.[178,1086,975,1056]" pageId="32" pageNumber="1165">
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Elytra (
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<figureCitation id="1303F7863A20FFB9FF46EC8EFEFFFC3D" box="[254,329,975,997]" captionStart="Figs 90-94" captionStartId="33.[178,218,1272,1294]" captionTargetBox="[186,1033,212,1250]" captionTargetId="figure-1@33.[178,1091,212,1263]" captionTargetPageId="33" captionText="Figs 90-94: Lithocharis bitriangulata nov.sp.: (90) forebody; (91) male sternite VII; (92) male sternite VIII; (93-94) aedeagus in lateral and in ventral view. Scale bars: 90: 1.0 mm; 91-94: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184212" httpUri="https://zenodo.org/record/5184212/files/figure.png" pageId="32" pageNumber="1165">Fig. 90</figureCitation>
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) 1.1 times as long as pronotum; punctation very dense and fine; interstices without microsculpture. Hind wings present. Protarsomeres I-IV distinctly dilated. Metatarsomere I approximately as long as the combined length of II and III.
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</paragraph>
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<paragraph id="8B87EB033A20FFB9FF0AEB6EFCCEFBBA" blockId="32.[178,1085,1071,1122]" pageId="32" pageNumber="1165">Abdomen narrower than elytra; punctation fine and very dense; interstices with distinct microreticulation; posterior margin of tergite VII with palisade fringe.</paragraph>
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<paragraph id="8B87EB033A20FFB9FF0AEB33FC6BFAC1" blockId="32.[178,1086,1137,1306]" pageId="32" pageNumber="1165">
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♂: sternite VII (
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<figureCitation id="1303F7863A20FFB9FEE4EB30FE11FB50" box="[348,423,1137,1160]" captionStart="Figs 90-94" captionStartId="33.[178,218,1272,1294]" captionTargetBox="[186,1033,212,1250]" captionTargetId="figure-1@33.[178,1091,212,1263]" captionTargetPageId="33" captionText="Figs 90-94: Lithocharis bitriangulata nov.sp.: (90) forebody; (91) male sternite VII; (92) male sternite VIII; (93-94) aedeagus in lateral and in ventral view. Scale bars: 90: 1.0 mm; 91-94: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184212" httpUri="https://zenodo.org/record/5184212/files/figure.png" pageId="32" pageNumber="1165">Fig. 91</figureCitation>
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) moderately transverse, posterior margin weakly concave in the middle and with comb of approximately 30 moderately long palisade setae; sternite VIII (
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<figureCitation id="1303F7863A20FFB9FF02EBEDFEB2FB1A" box="[186,260,1196,1218]" captionStart="Figs 90-94" captionStartId="33.[178,218,1272,1294]" captionTargetBox="[186,1033,212,1250]" captionTargetId="figure-1@33.[178,1091,212,1263]" captionTargetPageId="33" captionText="Figs 90-94: Lithocharis bitriangulata nov.sp.: (90) forebody; (91) male sternite VII; (92) male sternite VIII; (93-94) aedeagus in lateral and in ventral view. Scale bars: 90: 1.0 mm; 91-94: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184212" httpUri="https://zenodo.org/record/5184212/files/figure.png" pageId="32" pageNumber="1165">Fig. 92</figureCitation>
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) transverse, extensively without pubescence in postero-median portion, posterior excision large and broadly V-shaped, near middle of margin of this excision with a few long setae; aedeagus (
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<figureCitation id="1303F7863A20FFB9FE2AEBA6FDB4FB25" box="[402,514,1255,1277]" captionStart="Figs 90-94" captionStartId="33.[178,218,1272,1294]" captionTargetBox="[186,1033,212,1250]" captionTargetId="figure-1@33.[178,1091,212,1263]" captionTargetPageId="33" captionText="Figs 90-94: Lithocharis bitriangulata nov.sp.: (90) forebody; (91) male sternite VII; (92) male sternite VIII; (93-94) aedeagus in lateral and in ventral view. Scale bars: 90: 1.0 mm; 91-94: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184212" httpUri="https://zenodo.org/record/5184212/files/figure.png" pageId="32" pageNumber="1165">Figs 93-94</figureCitation>
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) large,
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<quantity id="4CC046E63A20FFB9FDE9EBA6FD1DFB24" box="[593,683,1255,1276]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="32" pageNumber="1165" unit="mm" value="0.92">0.92 mm</quantity>
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long, with strongly sclerotized internal structures of distinctive shapes; dorso-lateral apophyses curved and rather stout.
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</paragraph>
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</subSubSection>
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<subSubSection id="C322B8883A20FFB9FF0AEA68FBA2FA6C" pageId="32" pageNumber="1165" type="discussion">
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<paragraph id="8B87EB033A20FFB9FF0AEA68FBA2FA6C" blockId="32.[178,1085,1321,1460]" pageId="32" pageNumber="1165">
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Comparative notes: Based on the similar external and male sexual characters,
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<taxonomicName id="4C3890803A20FFB9FF5DEA06FE31FA83" authorityName="Assing" authorityYear="2015" box="[229,391,1350,1372]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1165" phylum="Arthropoda" rank="species" species="bitriangulata">
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<emphasis id="B94C37113A20FFB9FF5DEA06FE31FA83" box="[229,391,1350,1372]" italics="true" pageId="32" pageNumber="1165">L. bitriangulata</emphasis>
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</taxonomicName>
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is closely related to
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<taxonomicName id="4C3890803A20FFB9FDE5EA06FD15FA83" authorityName="Kraatz" authorityYear="1859" box="[605,675,1350,1372]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1165" phylum="Arthropoda" rank="species" species="vilis">
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<emphasis id="B94C37113A20FFB9FDE5EA06FD15FA83" box="[605,675,1350,1372]" italics="true" pageId="32" pageNumber="1165">L. vilis</emphasis>
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</taxonomicName>
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. It is distinguished from this species by the less transverse male sternite VII with a differently shaped posterior margin and with a shorter comb with fewer palisade setae, by the different chaetotaxy of the male sternite VIII, and by the slightly smaller aedeagus with internal structures of different shapes.
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</paragraph>
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</subSubSection>
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||
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<subSubSection id="C322B8883A20FFB9FF0AEA82FB88FA2F" pageId="32" pageNumber="1165" type="distribution">
|
||
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<paragraph id="8B87EB033A20FFB9FF0AEA82FB88FA2F" blockId="32.[178,1086,1475,1527]" pageId="32" pageNumber="1165">
|
||
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|
Distribution and natural history: Thetypelocalityissituatedin in the south of
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<collectingRegion id="49FC25E13A20FFB9FEF2EAA0FE17FA2E" box="[330,417,1505,1526]" country="Indonesia" name="Sumatera Utara" pageId="32" pageNumber="1165">Sumatra</collectingRegion>
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, where the
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<typeStatus id="548355A13A20FFB9FDACEAA0FDDBFA2F" box="[532,621,1505,1527]" pageId="32" pageNumber="1165" type="holotype">holotype</typeStatus>
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was collected on, or near, the shore of a lake.
|
||
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</paragraph>
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</subSubSection>
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||
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</treatment>
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||
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</document>
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