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<document id="084A5C9668E44C269814FDFABCBCF124" ID-CLB-Dataset="112235" ID-DOI="10.35929/RSZ.0011" ID-GBIF-Dataset="b650ff17-2f76-4ff7-8d8d-45c94e85fd4e" ID-ISSN="0035-418" ID-Zenodo-Dep="5743355" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638274966629" checkinUser="carolina" docAuthor="Anisyutkin, Leonid N." docDate="2020" docId="E000E77862198F7938F9FB3BFDCEF885" docLanguage="en" docName="RevueSuisseZool.127.1.105-117.pdf" docOrigin="Revue suisse de Zoologie 127 (1)" docSource="http://dx.doi.org/10.35929/rsz.0011" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Poroblattini Roth 1971" docType="treatment" docVersion="8" lastPageNumber="113" masterDocId="1C399F00621E8F703B3FFF8BFFE9FFA7" masterDocTitle="New and little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Muséum d’histoire naturelle de Genève and the Zoological Institute of Saint Petersburg. Part 4" masterLastPageNumber="117" masterPageNumber="105" pageNumber="111" updateTime="1699471431757" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D8E793162CCACF79F6DE955ABCB6C273">New and little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Muséum d’histoire naturelle de Genève and the Zoological Institute of Saint Petersburg. Part 4</mods:title>
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<treatment id="E000E77862198F7938F9FB3BFDCEF885" ID-DOI="http://doi.org/10.5281/zenodo.6960446" ID-GBIF-Taxon="198576791" ID-Zenodo-Dep="6960446" LSID="urn:lsid:plazi:treatment:E000E77862198F7938F9FB3BFDCEF885" httpUri="http://treatment.plazi.org/id/E000E77862198F7938F9FB3BFDCEF885" lastPageId="9" lastPageNumber="113" pageId="7" pageNumber="111">
<subSubSection id="20B305E562198F7738F9FB3BFAFCFB60" box="[966,1301,1200,1223]" pageId="7" pageNumber="111" type="nomenclature">
<paragraph id="6816566E62198F7738F9FB3BFAFCFB60" blockId="7.[966,1301,1200,1223]" box="[966,1301,1200,1223]" pageId="7" pageNumber="111">
<heading id="335EE10262198F7738F9FB3BFAFCFB60" box="[966,1301,1200,1223]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="111" reason="4">
<emphasis id="5ADD8A7C62198F7738F9FB3BFAFCFB60" bold="true" box="[966,1301,1200,1223]" pageId="7" pageNumber="111">
Tribe
<taxonomicName id="AFA92DED62198F773F34FB3BFAFCFB60" ID-CoL="8N9NY" authority="Roth, 1971" authorityName="Roth" authorityYear="1971" box="[1035,1301,1200,1223]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Poroblattini">
Poroblattini
<bibRefCitation id="0C382B9F62198F773FA4FB3BFAFCFB60" author="Roth L. M." box="[1179,1301,1200,1223]" pageId="7" pageNumber="111" pagination="180 - 192" refId="ref8607" refString="Roth L. M. 1971. The male genitalia of Blattaria. VII. Galiblatta, Dryadoblatta, Poroblatta, Colapteroblatta, Nauclidas, Notolampra, Litopeltis, and Cariacasia (Blaberidae: Epilamprinae). Psyche 78: 180 - 192." type="journal article" year="1971">Roth, 1971</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="20B305E562198F773800FB55FAC6FAB6" pageId="7" pageNumber="111" type="reference_group">
<paragraph id="6816566E62198F773800FB55FB7DFB54" blockId="7.[831,1327,1246,1297]" box="[831,1172,1246,1267]" pageId="7" pageNumber="111">
<taxonomicName id="AFA92DED62198F773800FB55FBDDFB54" ID-CoL="8N9N3" authority="Roth, 1971" authorityName="Roth" authorityYear="1971" box="[831,1076,1246,1267]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Notolamprini">
Notolamprini
<bibRefCitation id="0C382B9F62198F7738F8FB55FBDDFB54" author="Roth L. M." box="[967,1076,1246,1267]" pageId="7" pageNumber="111" pagination="180 - 192" refId="ref8607" refString="Roth L. M. 1971. The male genitalia of Blattaria. VII. Galiblatta, Dryadoblatta, Poroblatta, Colapteroblatta, Nauclidas, Notolampra, Litopeltis, and Cariacasia (Blaberidae: Epilamprinae). Psyche 78: 180 - 192." type="journal article" year="1971">Roth, 1971</bibRefCitation>
</taxonomicName>
, syn. nov.
</paragraph>
<paragraph id="6816566E62198F773800FB77FAC6FAB6" blockId="7.[831,1327,1246,1297]" box="[831,1327,1276,1297]" pageId="7" pageNumber="111">
<taxonomicName id="AFA92DED62198F773800FB77FB27FAB6" ID-CoL="8N9KH" authority="Roth &amp; Gutierrez, 1998" authorityName="Roth &amp; Gutierrez" authorityYear="1998" box="[831,1230,1276,1297]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Colapteroblattini">
Colapteroblattini
<bibRefCitation id="0C382B9F62198F7738D7FB77FB27FAB6" author="Roth L. M. &amp; Gutierrez E." box="[1000,1230,1276,1297]" pageId="7" pageNumber="111" pagination="167 - 202" refId="ref8732" refString="Roth L. M., Gutierrez E. 1998. The cockroach genus Colapteroblatta, its synonyms Poroblatta, Acroporoblatta, and Nauclidas, and a new species of Litopeltis (Blattaria: Blaberidae, Epilamprinae). Transactions of the American Entomological Society 124 (3 - 4): 167 - 202." type="journal article" year="1998">Roth &amp; Gutiérrez, 1998</bibRefCitation>
</taxonomicName>
, syn. nov.
</paragraph>
</subSubSection>
<subSubSection id="20B305E562198F773800FAA1FAFEFAE6" box="[831,1303,1322,1345]" pageId="7" pageNumber="111" type="type_taxon">
<paragraph id="6816566E62198F773800FAA1FAFEFAE6" blockId="7.[831,1303,1322,1345]" box="[831,1303,1322,1345]" pageId="7" pageNumber="111">
<emphasis id="5ADD8A7C62198F773800FAA1FC21FAE6" bold="true" box="[831,968,1322,1345]" pageId="7" pageNumber="111">
<typeStatus id="B712E8CC62198F773800FAA1FC9EFAE6" box="[831,887,1322,1345]" pageId="7" pageNumber="111">Type</typeStatus>
genus:
</emphasis>
<taxonomicName id="AFA92DED62198F7738F0FAA1FAFDFAE6" ID-CoL="8MQKD" authority="Hebard, 1919" authorityName="Hebard" authorityYear="1919" box="[975,1300,1322,1345]" class="Insecta" family="Blaberidae" genus="Colapteroblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62198F7738F0FAA1FB91FAE6" box="[975,1144,1322,1345]" italics="true" pageId="7" pageNumber="111">Colapteroblatta</emphasis>
<bibRefCitation id="0C382B9F62198F773F40FAA1FAFDFAE6" author="Hebard M." box="[1151,1300,1322,1345]" pageId="7" pageNumber="111" pagination="89 - 179" refId="ref8368" refString="Hebard M. 1919. Studies in the Dermaptera and Orthoptera of Colombia. Transactions of the American Entomological Society 45: 89 - 179." type="journal article" year="1919">Hebard, 1919</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="20B305E562198F793800FAD2FDCEF885" lastPageId="9" lastPageNumber="113" pageId="7" pageNumber="111" type="discussion">
<paragraph id="6816566E62198F773800FAD2FB78F85B" blockId="7.[831,1437,1369,2077]" pageId="7" pageNumber="111">
<emphasis id="5ADD8A7C62198F773800FAD2FC58FAD7" bold="true" box="[831,945,1369,1392]" pageId="7" pageNumber="111">Remarks:</emphasis>
The tribes
<taxonomicName id="AFA92DED62198F773F7AFAD2FB2CFAD7" authorityName="Roth" authorityYear="1971" box="[1093,1221,1369,1392]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Poroblattini">Poroblattini</taxonomicName>
and
<taxonomicName id="AFA92DED62198F773E34FAD2FA75FAD7" authorityName="Roth" authorityYear="1971" box="[1291,1436,1369,1392]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Notolamprini">Notolamprini</taxonomicName>
were described in the same paper (
<bibRefCitation id="0C382B9F62198F773F8CFAF1FAD9FA36" author="Roth L. M." box="[1203,1328,1402,1425]" pageId="7" pageNumber="111" pagination="180 - 192" refId="ref8607" refString="Roth L. M. 1971. The male genitalia of Blattaria. VII. Galiblatta, Dryadoblatta, Poroblatta, Colapteroblatta, Nauclidas, Notolampra, Litopeltis, and Cariacasia (Blaberidae: Epilamprinae). Psyche 78: 180 - 192." type="journal article" year="1971">Roth, 1971</bibRefCitation>
). The first tribe originally included the genera
<taxonomicName id="AFA92DED62198F773FF4FA10FC9EFA75" authority="Hebard, 1919" authorityName="Hebard" authorityYear="1919" class="Insecta" family="Blaberidae" genus="Poroblatta" higherTaxonomySource="CoL" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62198F773FF4FA10FAD7FA15" box="[1227,1342,1435,1458]" italics="true" pageId="7" pageNumber="111">Poroblatta</emphasis>
<bibRefCitation id="0C382B9F62198F773E78FA10FC9EFA75" author="Hebard M." pageId="7" pageNumber="111" pagination="89 - 179" refId="ref8368" refString="Hebard M. 1919. Studies in the Dermaptera and Orthoptera of Colombia. Transactions of the American Entomological Society 45: 89 - 179." type="journal article" year="1919">Hebard, 1919</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="AFA92DED62198F7738BAFA30FB95FA75" authority="Rehn, 1930" authorityName="Rehn" authorityYear="1930" box="[901,1148,1467,1490]" class="Insecta" family="Blaberidae" genus="Nauclidas" higherTaxonomySource="CoL" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62198F7738BAFA30FC18FA75" box="[901,1009,1467,1490]" italics="true" pageId="7" pageNumber="111">Nauclidas</emphasis>
<bibRefCitation id="0C382B9F62198F7738C4FA30FB95FA75" author="Rehn J. A. G." box="[1019,1148,1467,1490]" pageId="7" pageNumber="111" pagination="19 - 71" refId="ref8545" refString="Rehn J. A. G. 1930. New or little known Neotropical Blattidae. Transactions of the American Entomological Society 56: 19 - 71." type="journal article" year="1930">Rehn, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="AFA92DED62198F773FB5FA30FA71FA75" authority="Hebard, 1927" authorityName="Hebard" authorityYear="1927" box="[1162,1432,1467,1490]" class="Insecta" family="Blaberidae" genus="Galiblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62198F773FB5FA30FB11FA75" box="[1162,1272,1467,1490]" italics="true" pageId="7" pageNumber="111">Galiblatta</emphasis>
<bibRefCitation id="0C382B9F62198F773E3EFA30FA71FA75" author="Hebard M." box="[1281,1432,1467,1490]" pageId="7" pageNumber="111" pagination="135 - 244" refId="ref8394" refString="Hebard M. 1927. The Blattidae of French Guiana. Proceedings of the Academy of Natural Sciences of Philadelphia (for 1926) 78: 135 - 244." type="journal article" year="1927">Hebard, 1927</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="AFA92DED62198F773800FA57FB82FA54" authority="Rehn, 1930" authorityName="Rehn" authorityYear="1930" box="[831,1131,1500,1523]" class="Insecta" family="Blaberidae" genus="Dryadoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62198F773800FA57FC38FA54" box="[831,977,1500,1523]" italics="true" pageId="7" pageNumber="111">Dryadoblatta</emphasis>
<bibRefCitation id="0C382B9F62198F7738DDFA57FB82FA54" author="Rehn J. A. G." box="[994,1131,1500,1523]" pageId="7" pageNumber="111" pagination="19 - 71" refId="ref8545" refString="Rehn J. A. G. 1930. New or little known Neotropical Blattidae. Transactions of the American Entomological Society 56: 19 - 71." type="journal article" year="1930">Rehn, 1930</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="AFA92DED62198F773F87FA57FA8DFA54" authorityName="Hebard" authorityYear="1919" box="[1208,1380,1500,1523]" class="Insecta" family="Blaberidae" genus="Colapteroblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62198F773F87FA57FA8DFA54" box="[1208,1380,1500,1523]" italics="true" pageId="7" pageNumber="111">Colapteroblatta</emphasis>
</taxonomicName>
, the second tribe was monotypic (
<bibRefCitation id="0C382B9F62198F773FABFA77FAFEF9B4" author="Roth L. M." box="[1172,1303,1532,1555]" pageId="7" pageNumber="111" pagination="180 - 192" refId="ref8607" refString="Roth L. M. 1971. The male genitalia of Blattaria. VII. Galiblatta, Dryadoblatta, Poroblatta, Colapteroblatta, Nauclidas, Notolampra, Litopeltis, and Cariacasia (Blaberidae: Epilamprinae). Psyche 78: 180 - 192." type="journal article" year="1971">Roth, 1971</bibRefCitation>
). Later, the genera
<taxonomicName id="AFA92DED62198F7738A0F996FBFDF993" authorityName="Hebard" authorityYear="1919" box="[927,1044,1565,1588]" class="Insecta" family="Blaberidae" genus="Poroblatta" higherTaxonomySource="CoL" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62198F7738A0F996FBFDF993" box="[927,1044,1565,1588]" italics="true" pageId="7" pageNumber="111">Poroblatta</emphasis>
</taxonomicName>
,
<taxonomicName id="AFA92DED62198F773F0DF996FB30F993" authorityName="Hebard" authorityYear="1919" box="[1074,1241,1565,1588]" class="Insecta" family="Blaberidae" genus="Acroporoblatta" higherTaxonomySource="CoL" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62198F773F0DF996FB30F993" box="[1074,1241,1565,1588]" italics="true" pageId="7" pageNumber="111">Acroporoblatta</emphasis>
</taxonomicName>
and
<taxonomicName id="AFA92DED62198F773E11F996FA74F993" authorityName="Rehn" authorityYear="1930" box="[1326,1437,1565,1588]" class="Insecta" family="Blaberidae" genus="Nauclidas" higherTaxonomySource="CoL" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62198F773E11F996FA74F993" box="[1326,1437,1565,1588]" italics="true" pageId="7" pageNumber="111">Nauclidas</emphasis>
</taxonomicName>
were synonymized under
<taxonomicName id="AFA92DED62198F773F40F9B5FAC5F9F2" authorityName="Hebard" authorityYear="1919" box="[1151,1324,1598,1621]" class="Insecta" family="Blaberidae" genus="Colapteroblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62198F773F40F9B5FAC5F9F2" box="[1151,1324,1598,1621]" italics="true" pageId="7" pageNumber="111">Colapteroblatta</emphasis>
</taxonomicName>
and the name
<taxonomicName id="AFA92DED62198F7738BBF9D5FBEAF9D2" authorityName="Roth" authorityYear="1971" box="[900,1027,1630,1653]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Poroblattini">Poroblattini</taxonomicName>
was replaced with
<taxonomicName id="AFA92DED62198F773FE0F9D5FA70F9D2" authorityName="Roth &amp; Gutierrez" authorityYear="1998" box="[1247,1433,1630,1653]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Colapteroblattini">Colapteroblattini</taxonomicName>
: “Because of synonymy this tribe should be called
<taxonomicName id="AFA92DED62198F773800F914FC13F911" authorityName="Roth &amp; Gutierrez" authorityYear="1998" box="[831,1018,1695,1718]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Colapteroblattini">Colapteroblattini</taxonomicName>
” (
<bibRefCitation id="0C382B9F62198F773F20F914FA79F911" author="Roth L. M. &amp; Gutierrez E." box="[1055,1424,1695,1718]" pageId="7" pageNumber="111" pagination="167 - 202" refId="ref8732" refString="Roth L. M., Gutierrez E. 1998. The cockroach genus Colapteroblatta, its synonyms Poroblatta, Acroporoblatta, and Nauclidas, and a new species of Litopeltis (Blattaria: Blaberidae, Epilamprinae). Transactions of the American Entomological Society 124 (3 - 4): 167 - 202." type="journal article" year="1998">Roth &amp; Gutiérrez, 1998: 171</bibRefCitation>
). This replacement is incorrect because of article 40.1. of the International Code of Zoological Nomenclature (
<bibRefCitation id="0C382B9F62198F773895F88AFBC8F89E" author="International Commission on Zoological Nomenclature" pageId="7" pageNumber="111" refId="ref8424" refString="International Commission on Zoological Nomenclature 1999. International Code of Zoological Nomenclature, 4 th Edition. International Trust for Zoological Nomenclature, c / o The Natural History Museum, London, 306 pp." type="book" year="1999">International Commission on Zoological Nomenclature, 1999</bibRefCitation>
): “Validity of family-group names not affected. When the name of a
<typeStatus id="B712E8CC62198F773FDEF8C9FAE6F8FE" box="[1249,1295,1858,1881]" pageId="7" pageNumber="111">type</typeStatus>
genus of a nominal family-group taxon is considered to be a junior synonym of the name of another nominal genus, the family-group name is not to be replaced on that account alone”. Thus I consider
<taxonomicName id="AFA92DED62198F773F77F84FFB14F87C" authorityName="Roth &amp; Gutierrez" authorityYear="1998" box="[1096,1277,1988,2011]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Colapteroblattini">Colapteroblattini</taxonomicName>
<taxonomicNameLabel id="41EE370762198F773E39F84FFA80F87C" box="[1286,1385,1988,2011]" pageId="7" pageNumber="111">syn. nov.</taxonomicNameLabel>
as a junior synonym of
<taxonomicName id="AFA92DED62198F773F34F86EFB64F85B" authorityName="Roth" authorityYear="1971" box="[1035,1165,2021,2044]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Poroblattini">Poroblattini</taxonomicName>
.
</paragraph>
<paragraph id="6816566E62198F793800F78DFDA9FDB2" blockId="7.[831,1437,1369,2077]" lastBlockId="9.[151,757,258,1826]" lastPageId="9" lastPageNumber="113" pageId="7" pageNumber="111">
The tribe
<taxonomicName id="AFA92DED62198F773896F78DFBCEF7BA" authorityName="Roth" authorityYear="1971" box="[937,1063,2054,2077]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="111" phylum="Arthropoda" rank="tribe" tribe="Poroblattini">Poroblattini</taxonomicName>
is based on characters of the male genitalia: “In this tribe the L2d (i.e. apical part of sclerite L2D in present paper) is elongated, curved, sclerotized, tapers slightly toward the tip, and is separated from L2vm (i.e. basal part of sclerite L2D in present paper). Apparently there is no distinct prepuce (i.e. membrane with bristles or teeth or apical sclerite of apical part of sclerite L2D). The R2 has a subapical incision and the shapes of L1 (i.e. right phallomere in present paper) are all basically similar” (
<bibRefCitation id="0C382B9F62178F793ABDFE75FDDAFDB2" author="Roth L. M." box="[386,563,510,533]" pageId="9" pageNumber="113" pagination="180 - 192" refId="ref8607" refString="Roth L. M. 1971. The male genitalia of Blattaria. VII. Galiblatta, Dryadoblatta, Poroblatta, Colapteroblatta, Nauclidas, Notolampra, Litopeltis, and Cariacasia (Blaberidae: Epilamprinae). Psyche 78: 180 - 192." type="journal article" year="1971">Roth, 1971: 181</bibRefCitation>
).
</paragraph>
<caption id="3CD606E662168F783BA8F8CDFCF5F84C" ID-DOI="http://doi.org/10.5281/zenodo.5743367" ID-Zenodo-Dep="5743367" httpUri="https://zenodo.org/record/5743367/files/figure.png" pageId="8" pageNumber="112" startId="8.[151,192,1862,1883]" targetBox="[173,1423,260,1822]" targetPageId="8">
<paragraph id="6816566E62168F783BA8F8CDFCF5F84C" blockId="8.[151,1437,1862,2027]" pageId="8" pageNumber="112">
Figs 25-35.
<taxonomicName id="AFA92DED62168F783A29F8CDFDABF8FC" authority="Roth, 1999" authorityName="Roth" authorityYear="1999" box="[278,578,1862,1883]" class="Insecta" family="Blaberidae" genus="Molytria" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="112" phylum="Arthropoda" rank="species" species="vegranda">
<emphasis id="5ADD8A7C62168F783A29F8CDFE24F8FC" box="[278,461,1862,1883]" italics="true" pageId="8" pageNumber="112">Molytria vegranda</emphasis>
<bibRefCitation id="0C382B9F62168F783AEBF8CDFDABF8FC" author="Roth L. M." box="[468,578,1862,1883]" pageId="8" pageNumber="112" pagination="327 - 364" refId="ref8651" refString="Roth L. M. 1999. New cockroach species, redescriptions, and records, mostly from Australia, and a description of Metanocticola christmasensis gen. nov., sp. nov., from Christmas Island (Blattaria). Records of the Western Australian Museum 19: 327 - 364." type="journal article" year="1999">Roth, 1999</bibRefCitation>
</taxonomicName>
, male. (25-26) Left hind tarsus, ventral (25) and inner (26) view. (27) Abdominal apex, dorsal view. (28) Hypandrium, ventral view. (29) Sclerites L2D, L4U and outlines of caudal part of hypandrium, dorsal view. (30-31) Right phallomere, ventral (30) and dorsal (31) view. (32) Apical part of sclerite L2D, dorsal view. (33) Sclerite L3. (34, 35) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations:
<emphasis id="5ADD8A7C62168F783F9FF816FB2BF815" box="[1184,1218,1949,1970]" italics="true" pageId="8" pageNumber="112">a.s.</emphasis>
,
<emphasis id="5ADD8A7C62168F783FF2F816FAE3F815" box="[1229,1290,1949,1970]" italics="true" pageId="8" pageNumber="112">b.L2D</emphasis>
,
<emphasis id="5ADD8A7C62168F783E2AF816FAA9F815" box="[1301,1344,1949,1970]" italics="true" pageId="8" pageNumber="112">b.L3</emphasis>
,
<emphasis id="5ADD8A7C62168F783E73F816FA7FF815" box="[1356,1430,1949,1970]" italics="true" pageId="8" pageNumber="112">c.p.R1T</emphasis>
,
<emphasis id="5ADD8A7C62168F783A29F832FEDDF869" box="[278,308,1977,1998]" italics="true" pageId="8" pageNumber="112">f.s.</emphasis>
,
<emphasis id="5ADD8A7C62168F783A00F832FE83F869" box="[319,362,1977,1998]" italics="true" pageId="8" pageNumber="112">L4U</emphasis>
,
<emphasis id="5ADD8A7C62168F783A49F832FE78F869" box="[374,401,1977,1998]" italics="true" pageId="8" pageNumber="112">R2</emphasis>
,
<emphasis id="5ADD8A7C62168F783AA1F832FE51F869" box="[414,440,1977,1998]" italics="true" pageId="8" pageNumber="112">R3</emphasis>
,
<emphasis id="5ADD8A7C62168F783AFAF832FE09F869" box="[453,480,1977,1998]" italics="true" pageId="8" pageNumber="112">R4</emphasis>
,
<emphasis id="5ADD8A7C62168F783AD3F832FDEEF869" box="[492,519,1977,1998]" italics="true" pageId="8" pageNumber="112">R5</emphasis>
,
<emphasis id="5ADD8A7C62168F78392BF832FDD9F869" box="[532,560,1977,1998]" italics="true" pageId="8" pageNumber="112">s.t.</emphasis>
,
<emphasis id="5ADD8A7C62168F783903F832FD99F869" box="[572,624,1977,1998]" italics="true" pageId="8" pageNumber="112">scl.a.</emphasis>
- see chapter “abbreviation used in figures”, for details see text. Scale bars 1 mm: a (24-25), b (27-29), c (30-31), d (32), e (33), f (34-35).
</paragraph>
</caption>
<paragraph id="6816566E62178F793BA8FD95FEB5FCF7" blockId="9.[151,757,258,1826]" pageId="9" pageNumber="113">
Roth did not give a clear definition of the tribe
<taxonomicName id="AFA92DED62178F793989FD95FF1CFDF3" authorityName="Roth" authorityYear="1971" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Notolamprini">Notolamprini</taxonomicName>
. It was noted “that the 3 species of
<taxonomicName id="AFA92DED62178F79394CFDB6FD1DFDF3" authorityName="Saussure" authorityYear="1862" box="[627,756,573,596]" class="Insecta" family="Blaberidae" genus="Notolampra" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62178F79394CFDB6FD1DFDF3" box="[627,756,573,596]" italics="true" pageId="9" pageNumber="113">Notolampra</emphasis>
</taxonomicName>
have a markedly convex dorsal surface” (
<bibRefCitation id="0C382B9F62178F79397DFDD6FD05FDD3" author="Roth L. M." box="[578,748,605,628]" pageId="9" pageNumber="113" pagination="180 - 192" refId="ref8607" refString="Roth L. M. 1971. The male genitalia of Blattaria. VII. Galiblatta, Dryadoblatta, Poroblatta, Colapteroblatta, Nauclidas, Notolampra, Litopeltis, and Cariacasia (Blaberidae: Epilamprinae). Psyche 78: 180 - 192." type="journal article" year="1971">Roth, 1971: 181</bibRefCitation>
) and some morphological differences and peculiarities in male genitalia structures: “In
<taxonomicName id="AFA92DED62178F793AF0FD17FDC3FD14" baseAuthorityName="Thunberg" baseAuthorityYear="1826" box="[463,554,668,691]" class="Insecta" family="Blaberidae" genus="Notolampra" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="5ADD8A7C62178F793AF0FD17FE08FD14" box="[463,481,668,691]" italics="true" pageId="9" pageNumber="113">N</emphasis>
.
<emphasis id="5ADD8A7C62178F793AD2FD17FDC3FD14" box="[493,554,668,691]" italics="true" pageId="9" pageNumber="113">gibba</emphasis>
</taxonomicName>
... the L2d is much more robust than the L2d of members of
<taxonomicName id="AFA92DED62178F79397CFD30FD2DFD75" authorityName="Roth" authorityYear="1971" box="[579,708,699,722]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Poroblattini">Poroblattini</taxonomicName>
, and does not taper toward the apex. R1 (i.e. sclerite L
<quantity id="AF51FB8B62178F7939F6FD50FD1DFD55" box="[713,756,731,754]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="113" unit="in" value="3.0">3 in</quantity>
present paper) is long and slender and has a subapical incision; L1 differs in shape from L1 of
<taxonomicName id="AFA92DED62178F793955FC91FD03FC96" authorityName="Roth" authorityYear="1971" box="[618,746,794,817]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Poroblattini">Poroblattini</taxonomicName>
” (
<bibRefCitation id="0C382B9F62178F793BA0FCB2FEB9FCF7" author="Roth L. M." box="[159,336,825,848]" pageId="9" pageNumber="113" pagination="180 - 192" refId="ref8607" refString="Roth L. M. 1971. The male genitalia of Blattaria. VII. Galiblatta, Dryadoblatta, Poroblatta, Colapteroblatta, Nauclidas, Notolampra, Litopeltis, and Cariacasia (Blaberidae: Epilamprinae). Psyche 78: 180 - 192." type="journal article" year="1971">Roth, 1971: 181</bibRefCitation>
).
</paragraph>
<paragraph id="6816566E62178F793BA8FCD2FE40FC69" blockId="9.[151,757,258,1826]" pageId="9" pageNumber="113">
It must be noted that Roth examined the male genitalia structures mounted on slides (
<bibRefCitation id="0C382B9F62178F793AC0FCF3FD6DFC28" author="Roth L. M." box="[511,644,888,911]" pageId="9" pageNumber="113" pagination="180 - 192" refId="ref8607" refString="Roth L. M. 1971. The male genitalia of Blattaria. VII. Galiblatta, Dryadoblatta, Poroblatta, Colapteroblatta, Nauclidas, Notolampra, Litopeltis, and Cariacasia (Blaberidae: Epilamprinae). Psyche 78: 180 - 192." type="journal article" year="1971">Roth, 1971</bibRefCitation>
;
<bibRefCitation id="0C382B9F62178F7939A5FCF3FEA0FC08" author="Roth L. M. &amp; Gutierrez E." pageId="9" pageNumber="113" pagination="167 - 202" refId="ref8732" refString="Roth L. M., Gutierrez E. 1998. The cockroach genus Colapteroblatta, its synonyms Poroblatta, Acroporoblatta, and Nauclidas, and a new species of Litopeltis (Blattaria: Blaberidae, Epilamprinae). Transactions of the American Entomological Society 124 (3 - 4): 167 - 202." type="journal article" year="1998">Roth &amp; Gutiérrez, 1998</bibRefCitation>
). This made it difficult to study fine morphological structures.
</paragraph>
<paragraph id="6816566E62178F793BA8FC5CFD6BFBCB" blockId="9.[151,757,258,1826]" pageId="9" pageNumber="113">
In my opinion,
<taxonomicName id="AFA92DED62178F793A0BFC5CFD2FFC49" authority="Hebard, 1919" authorityName="Hebard" authorityYear="1919" box="[308,710,983,1006]" class="Insecta" family="Blaberidae" genus="Colapteroblatta" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="species" species="compsa">
<emphasis id="5ADD8A7C62178F793A0BFC5CFDDAFC49" box="[308,563,983,1006]" italics="true" pageId="9" pageNumber="113">Colapteroblatta compsa</emphasis>
<bibRefCitation id="0C382B9F62178F793907FC5CFD2FFC49" author="Hebard M." box="[568,710,983,1006]" pageId="9" pageNumber="113" pagination="89 - 179" refId="ref8368" refString="Hebard M. 1919. Studies in the Dermaptera and Orthoptera of Colombia. Transactions of the American Entomological Society 45: 89 - 179." type="journal article" year="1919">Hebard, 1919</bibRefCitation>
</taxonomicName>
(the
<typeStatus id="B712E8CC62178F793BA8FC7CFF2CFBA9" box="[151,197,1015,1038]" pageId="9" pageNumber="113">type</typeStatus>
species of
<taxonomicName id="AFA92DED62178F793A7CFC7CFE05FBA9" authorityName="Hebard" authorityYear="1919" box="[323,492,1015,1038]" class="Insecta" family="Blaberidae" genus="Colapteroblatta" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62178F793A7CFC7CFE05FBA9" box="[323,492,1015,1038]" italics="true" pageId="9" pageNumber="113">Colapteroblatta</emphasis>
</taxonomicName>
) and
<taxonomicName id="AFA92DED62178F793912FC7CFEBDFB8A" authority="(Thunberg, 1826)" baseAuthorityName="Thunberg" baseAuthorityYear="1826" class="Insecta" family="Blaberidae" genus="Notolampra" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="5ADD8A7C62178F793912FC7CFD1DFBA9" box="[557,756,1015,1038]" italics="true" pageId="9" pageNumber="113">Notolampra gibba</emphasis>
(
<bibRefCitation id="0C382B9F62178F793BA1FB9DFEA5FB8A" author="Thunberg C. P." box="[158,332,1046,1069]" pageId="9" pageNumber="113" pagination="275 - 280" refId="ref8923" refString="Thunberg C. P. 1826. Blattarum novae species descriptae. Memoires de L'Academie Imperiale des Sciences de St. Petersbourg 10: 275 - 280." type="journal article" year="1826">Thunberg, 1826</bibRefCitation>
)
</taxonomicName>
(the
<typeStatus id="B712E8CC62178F793AB4FB9DFE50FB8A" box="[395,441,1046,1069]" pageId="9" pageNumber="113">type</typeStatus>
species of
<taxonomicName id="AFA92DED62178F79390EFB9DFD5BFB8A" authorityName="Saussure" authorityYear="1862" box="[561,690,1046,1069]" class="Insecta" family="Blaberidae" genus="Notolampra" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62178F79390EFB9DFD5BFB8A" box="[561,690,1046,1069]" italics="true" pageId="9" pageNumber="113">Notolampra</emphasis>
</taxonomicName>
) have structurally very similar male genitalia. The differences mentioned by
<bibRefCitation id="0C382B9F62178F793A0FFBDEFE58FBCB" author="Roth L. M." box="[304,433,1109,1132]" pageId="9" pageNumber="113" pagination="180 - 192" refId="ref8607" refString="Roth L. M. 1971. The male genitalia of Blattaria. VII. Galiblatta, Dryadoblatta, Poroblatta, Colapteroblatta, Nauclidas, Notolampra, Litopeltis, and Cariacasia (Blaberidae: Epilamprinae). Psyche 78: 180 - 192." type="journal article" year="1971">Roth (1971)</bibRefCitation>
are not substantial.
</paragraph>
<paragraph id="6816566E62178F793BA8FBFEFDA2FB0C" blockId="9.[151,757,258,1826]" pageId="9" pageNumber="113">
The
<typeStatus id="B712E8CC62178F793BF1FBFEFF15FB2B" box="[206,252,1141,1164]" pageId="9" pageNumber="113">type</typeStatus>
species of
<taxonomicName id="AFA92DED62178F793AB7FBFEFDD8FB2B" authorityName="Hebard" authorityYear="1919" box="[392,561,1141,1164]" class="Insecta" family="Blaberidae" genus="Colapteroblatta" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62178F793AB7FBFEFDD8FB2B" box="[392,561,1141,1164]" italics="true" pageId="9" pageNumber="113">Colapteroblatta</emphasis>
</taxonomicName>
and
<taxonomicName id="AFA92DED62178F79394CFBFEFD1DFB2B" authorityName="Saussure" authorityYear="1862" box="[627,756,1141,1164]" class="Insecta" family="Blaberidae" genus="Notolampra" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="5ADD8A7C62178F79394CFBFEFD1DFB2B" box="[627,756,1141,1164]" italics="true" pageId="9" pageNumber="113">Notolampra</emphasis>
</taxonomicName>
have a number of characters in common:
</paragraph>
<paragraph id="6816566E62178F793BA8FB3FFE26FAAD" blockId="9.[151,757,258,1826]" pageId="9" pageNumber="113">1. Structure of tarsi: metatarsus distinctly shorter than other tarsal segments combined, with large euplantula; spines absent.</paragraph>
<paragraph id="6816566E62178F793BA8FA99FD81FACF" blockId="9.[151,757,258,1826]" pageId="9" pageNumber="113">
2. Hypandrium (
<figureCitation id="F0924AEB62178F793A6BFA99FE4AFA8E" box="[340,419,1298,1321]" captionStart="Figs 36-48" captionStartId="10.[151,192,1845,1866]" captionTargetBox="[166,1424,262,1793]" captionTargetId="figure-5@10.[151,1436,254,1799]" captionTargetPageId="10" captionText="Figs 36-48. Notolampra gibba (Thunberg, 1826), male. (36) Facial part of head. (37) Head, pronotum and basal part of left tegmen, dorsal view. (38) Abdominal tergites VIII-IX, dorsal view. (39)Abdominal apex, dorsal view. (40) Paraprocts and adjacent structures, ventral view. (41) Hypandrium, ventral view. (42-43) Right phallomere, dorsal (42) and ventral (43) view. (44) Sclerite L2D, dorsal view. (45-46) Apical part of sclerite L2D, ventral (45) and dorsal (46) view. (47) Sclerite L3. (48) Apex of sclerite L3. Dotted areas show dark colour (37) or membranous parts (38-48). Abbreviations: ap.scl., b.L2D, b.L3, c.p.R1T, R2, R3, R4, R5, tr.l. - see chapter “abbreviation used in figures”, for details see text. Scale bars 1 mm: a (36), b (37), c (38-39), d (40), e (41), f (42-43), g (44), h (45-46), i (47-48)." figureDoi="http://doi.org/10.5281/zenodo.5743369" httpUri="https://zenodo.org/record/5743369/files/figure.png" pageId="9" pageNumber="113">Fig. 41</figureCitation>
cf.
<bibRefCitation id="0C382B9F62178F793AF1FA99FD70FA8E" author="Anisyutkin L. N." box="[462,665,1298,1321]" pageId="9" pageNumber="113" pagination="1 - 16" refId="ref7956" refString="Anisyutkin L. N. 2018 a. Little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Museum d'histoire naturelle de Geneve. Part 3. Revue suisse de Zoologie 125 (1): 1 - 16." type="journal article" year="2018">Anisyutkin, 2018a</bibRefCitation>
: fig. 90) asymmetrical, with membranous area along right side; styli asymmetrical and cylindrical.
</paragraph>
<paragraph id="6816566E62178F793BA8FAFAFD51FA41" blockId="9.[151,757,258,1826]" pageId="9" pageNumber="113">
3. Structure of right phallomere (
<figureCitation id="F0924AEB62178F793900FAFAFD2AFA2F" box="[575,707,1393,1416]" captionStart="Figs 36-48" captionStartId="10.[151,192,1845,1866]" captionTargetBox="[166,1424,262,1793]" captionTargetId="figure-5@10.[151,1436,254,1799]" captionTargetPageId="10" captionText="Figs 36-48. Notolampra gibba (Thunberg, 1826), male. (36) Facial part of head. (37) Head, pronotum and basal part of left tegmen, dorsal view. (38) Abdominal tergites VIII-IX, dorsal view. (39)Abdominal apex, dorsal view. (40) Paraprocts and adjacent structures, ventral view. (41) Hypandrium, ventral view. (42-43) Right phallomere, dorsal (42) and ventral (43) view. (44) Sclerite L2D, dorsal view. (45-46) Apical part of sclerite L2D, ventral (45) and dorsal (46) view. (47) Sclerite L3. (48) Apex of sclerite L3. Dotted areas show dark colour (37) or membranous parts (38-48). Abbreviations: ap.scl., b.L2D, b.L3, c.p.R1T, R2, R3, R4, R5, tr.l. - see chapter “abbreviation used in figures”, for details see text. Scale bars 1 mm: a (36), b (37), c (38-39), d (40), e (41), f (42-43), g (44), h (45-46), i (47-48)." figureDoi="http://doi.org/10.5281/zenodo.5743369" httpUri="https://zenodo.org/record/5743369/files/figure.png" pageId="9" pageNumber="113">Figs 42-43</figureCitation>
cf.
<bibRefCitation id="0C382B9F62178F793B82FA1BFE6CFA00" author="Anisyutkin L. N." box="[189,389,1424,1447]" pageId="9" pageNumber="113" pagination="1 - 16" refId="ref7956" refString="Anisyutkin L. N. 2018 a. Little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Museum d'histoire naturelle de Geneve. Part 3. Revue suisse de Zoologie 125 (1): 1 - 16." type="journal article" year="2018">Anisyutkin, 2018a</bibRefCitation>
: figs 91-92): caudal part of sclerite R1T distinctly enlarged; “upper triangular lobe” present; R3 short and robust, widened caudally.
</paragraph>
<paragraph id="6816566E62178F793BA8FA64FD98F9E2" blockId="9.[151,757,258,1826]" pageId="9" pageNumber="113">
4. Apical part of sclerite L2D developed as flattened, elongated and plate-like sclerite; bristles absent (
<figureCitation id="F0924AEB62178F7939FAF985FF15F9E2" captionStart="Figs 36-48" captionStartId="10.[151,192,1845,1866]" captionTargetBox="[166,1424,262,1793]" captionTargetId="figure-5@10.[151,1436,254,1799]" captionTargetPageId="10" captionText="Figs 36-48. Notolampra gibba (Thunberg, 1826), male. (36) Facial part of head. (37) Head, pronotum and basal part of left tegmen, dorsal view. (38) Abdominal tergites VIII-IX, dorsal view. (39)Abdominal apex, dorsal view. (40) Paraprocts and adjacent structures, ventral view. (41) Hypandrium, ventral view. (42-43) Right phallomere, dorsal (42) and ventral (43) view. (44) Sclerite L2D, dorsal view. (45-46) Apical part of sclerite L2D, ventral (45) and dorsal (46) view. (47) Sclerite L3. (48) Apex of sclerite L3. Dotted areas show dark colour (37) or membranous parts (38-48). Abbreviations: ap.scl., b.L2D, b.L3, c.p.R1T, R2, R3, R4, R5, tr.l. - see chapter “abbreviation used in figures”, for details see text. Scale bars 1 mm: a (36), b (37), c (38-39), d (40), e (41), f (42-43), g (44), h (45-46), i (47-48)." figureDoi="http://doi.org/10.5281/zenodo.5743369" httpUri="https://zenodo.org/record/5743369/files/figure.png" pageId="9" pageNumber="113">Figs 44-46</figureCitation>
cf.
<bibRefCitation id="0C382B9F62178F793A1CF9A5FE05F9E2" author="Anisyutkin L. N." box="[291,492,1582,1605]" pageId="9" pageNumber="113" pagination="1 - 16" refId="ref7956" refString="Anisyutkin L. N. 2018 a. Little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Museum d'histoire naturelle de Geneve. Part 3. Revue suisse de Zoologie 125 (1): 1 - 16." type="journal article" year="2018">Anisyutkin, 2018a</bibRefCitation>
: figs 93-95).
</paragraph>
<paragraph id="6816566E62178F793BA8F9C5FEF4F903" blockId="9.[151,757,258,1826]" pageId="9" pageNumber="113">
Taking into account the aforesaid, I consider the tribe
<taxonomicName id="AFA92DED62178F793BEAF9E6FE8CF923" authorityName="Roth" authorityYear="1971" box="[213,357,1645,1668]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Notolamprini">Notolamprini</taxonomicName>
<taxonomicNameLabel id="41EE370762178F793A4DF9E6FE30F923" box="[370,473,1645,1668]" pageId="9" pageNumber="113">syn. nov.</taxonomicNameLabel>
as a junior synonym of
<taxonomicName id="AFA92DED62178F793BA8F906FEF0F903" authorityName="Roth" authorityYear="1971" box="[151,281,1677,1700]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Poroblattini">Poroblattini</taxonomicName>
.
</paragraph>
<paragraph id="6816566E62178F793BA8F927FDCEF885" blockId="9.[151,757,258,1826]" pageId="9" pageNumber="113">
The features listed above are probably characteristic of the tribe
<taxonomicName id="AFA92DED62178F793BC7F947FE90F944" authorityName="Roth" authorityYear="1971" box="[248,377,1740,1763]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Poroblattini">Poroblattini</taxonomicName>
, but I prefer to postpone proposing a formal revised diagnosis of this tribe due to insufficient knowledge of other epilamprine taxa.
</paragraph>
</subSubSection>
</treatment>
</document>
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