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161 lines
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<document id="723361B001324F37D92E32B9745A3E37" ID-CLB-Dataset="22102" ID-DOI="10.5852/ejt.2017.322" ID-GBIF-Dataset="2922dbf4-12ec-49d4-ac36-565434419f51" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829305" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589562461714" checkinUser="carolina" docAuthor="Lin, Chien-Hsiang, Brzobohatý, Rostislav, Nolf, Dirk & Girone, Angela" docDate="2017" docId="615996129474CE247A74A070FB06957A" docLanguage="en" docName="ejt-2017-322.pdf.imf" docOrigin="European Journal of Taxonomy 322" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Lampadena gracilis" docType="treatment" docVersion="7" lastPageNumber="19" masterDocId="9D60EE6A9466CE36784FA457FFBC9351" masterDocTitle="Tortonian teleost otoliths from northern Italy: taxonomic synthesis and stratigraphic significance" masterLastPageNumber="44" masterPageNumber="1" pageNumber="19" updateTime="1698839868927" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FA989D286FE85AF2AE9594D78799E380">Tortonian teleost otoliths from northern Italy: taxonomic synthesis and stratigraphic significance</mods:title>
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<mods:namePart id="067FD108FC8661B8BBB1F7384DA6B178">Lin, Chien-Hsiang</mods:namePart>
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<mods:affiliation id="99D8AE670DEB53D21A6260B5A8FC779F">urn: lsid: zoobank. org: author: D 39077 E 1 - 7025 - 4118 - B 258 - C 4 A 896 E 63 E 2 A & Corresponding author: chlin. otolith @ gmail. com; r 97 b 41028 @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="FC8EC3CD400A3F417E25205CA52C3D25" type="email">chlin.otolith@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D509883F414D7C6A216EC2D120F9E7E9">Brzobohatý, Rostislav</mods:namePart>
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<mods:affiliation id="DBBE80FA97E66011E5F008F9BDB336B4">Email: rosta @ sci. muni. cz & urn: lsid: zoobank. org: author: A 354 D 828 - 3 D 07 - 46 F 5 - 9 D 27 - 4 F 9 D 7 B 208 F 12</mods:affiliation>
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<mods:nameIdentifier id="931545C1DC7C1D393ACB674DD7475BE8" type="email">rosta@sci.muni.cz</mods:nameIdentifier>
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<mods:namePart id="072DEA95ACBAED7A875AD2DE3C3DB54F">Nolf, Dirk</mods:namePart>
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<mods:affiliation id="95E2AB929E1B139355E2719CD82AF942">Email: dirk. nolf @ scarlet. be & urn: lsid: zoobank. org: author: 6 BCC 71 A 0 - 1 BEE- 4 BC 0 - BDFC-D 070609 DEFAB</mods:affiliation>
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<mods:nameIdentifier id="8D2E07D7AB9BCEB270C0E01B9154418E" type="email">dirk.nolf@scarlet.be</mods:nameIdentifier>
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<mods:namePart id="26794DA083DF03AD874105223667F549">Girone, Angela</mods:namePart>
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<mods:affiliation id="F45A3C6BFC0E28094A4FFB4F26568DB9">Email: angela. girone @ uniba. it & urn: lsid: zoobank. org: author: B 4212 D 22 - 1 D 1 B- 48 E 9 - 8 ABB-C 1 D 1 E 05573 EF</mods:affiliation>
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<mods:nameIdentifier id="185A9DF0955D5A7FB38800F6105D03E9" type="email">angela.girone@uniba.it</mods:nameIdentifier>
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<mods:title id="CD78E4243E0D2B2D56BAAF884147BBF2">European Journal of Taxonomy</mods:title>
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<mods:date id="826F509F1DF011DE218AA45199A6C9DF">2017</mods:date>
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<mods:number id="0888BE616902EFEDAAF5D11E92239B14">2017-05-23</mods:number>
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<mods:number id="50AE4B4FC20A680736075BDB0B084F2D">322</mods:number>
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<mods:identifier id="B73581F3310748CD958C2BB77A51521F" type="CLB-Dataset">22102</mods:identifier>
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<mods:identifier id="210D879FFD161016EFD9242365787A31" type="DOI">10.5852/ejt.2017.322</mods:identifier>
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<mods:identifier id="B00B6C32EDF52250E22187BAA055C77F" type="ISSN">2118-9773</mods:identifier>
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<treatment id="615996129474CE247A74A070FB06957A" ID-DOI="http://doi.org/10.5281/zenodo.3848327" ID-GBIF-Taxon="164159964" ID-Zenodo-Dep="3848327" LSID="urn:lsid:plazi:treatment:615996129474CE247A74A070FB06957A" httpUri="http://treatment.plazi.org/id/615996129474CE247A74A070FB06957A" lastPageNumber="19" pageId="18" pageNumber="19">
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<subSubSection id="A1EA748F9474CE247A74A070FC449713" box="[571,1016,1063,1090]" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="E94F27049474CE247A74A070FC449713" blockId="18.[571,1016,1063,1090]" box="[571,1016,1063,1090]" pageId="18" pageNumber="19">
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<heading id="B20790689474CE247A74A070FC449713" box="[571,1016,1063,1090]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
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<taxonomicName id="2EF05C879474CE247A74A070FC449713" authority="(Schubert, 1912)" baseAuthorityName="Schubert" baseAuthorityYear="1912" box="[571,1016,1063,1090]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="gracilis">
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<emphasis id="DB84FB169474CE247A74A070FC909713" bold="true" box="[571,812,1063,1090]" italics="true" pageId="18" pageNumber="19">Lampadena gracilis</emphasis>
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(Schubert, 1912)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A1EA748F9474CE2478F2A03EFB06957A" pageId="18" pageNumber="19" type="discussion">
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<paragraph id="E94F27049474CE2478F2A03EFB06957A" blockId="18.[189,1399,1129,1579]" pageId="18" pageNumber="19">
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The otoliths of
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<taxonomicName id="2EF05C879474CE247920A03DFE5B97D2" baseAuthorityName="Schubert" baseAuthorityYear="1912" box="[367,487,1129,1156]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="gracilis">
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<emphasis id="DB84FB169474CE247920A03DFEC397D2" box="[367,383,1130,1155]" italics="true" pageId="18" pageNumber="19">L</emphasis>
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.
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<emphasis id="DB84FB169474CE2479C2A03EFE5B97D2" box="[397,487,1129,1155]" italics="true" pageId="18" pageNumber="19">gracilis</emphasis>
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</taxonomicName>
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are very rare at the investigated localities (
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<figureCitation id="71CB3B819474CE247B95A03EFBA197D5" box="[986,1053,1129,1156]" captionStart="Fig" captionStartId="17.[189,231,1514,1540]" captionTargetBox="[298,1290,265,1505]" captionTargetId="figure@17.[298,1290,265,1505]" captionTargetPageId="17" captionText="Fig. 5. Tortonian fish otoliths from northern Italy. A. Diaphus holti Tåning, 1918, Mondovi, Madonna della Neve (IRSNB P 9732). B. Diaphus metopoclampoides Steurbaut, 1983, Sant’Alosio (IRSNB P 9733). C–E. Diaphus regani Tåning, 1932, Montegibbio (IRSNB P 9734–P 9736). F–G. Diaphus pedemontanus (Robba, 1970), Sant’Alosio (IRSNB P 9737–P 9738). H. Diaphus aff. rafinesquii (Cocco, 1838), Mondovi, Madonna della Neve (IRSNB P 9739). I. Diaphus aff. splendidus (Brauer, 1904), Costa Vescovato (IRSNB P 9740). J. Diaphus cahuzaci Steurbaut, 1979, Mondovi, Madonna della Neve (IRSNB P 9741). K. Lampadena aff. dea Fraser-Brunner, 1949, Torrente Stirone (IRSNB P 9742). L. Lampadena aff. speculigeroides Brzobohatý & Nolf, 1996, Stazzano (IRSNB P 9743). M. Hygophum hygomii (Lütken, 1892), Mondovi, Madonna della Neve (IRSNB P 9744). N–O. Lampadena gracilis (Schubert, 1912); N. Alba, Tanaro (50 m), O. Mondovi, Madonna della Neve (IRSNB P 9745–P 9746). P–Q. Lobianchia gemellarii (Cocco, 1838), Sant’Alosio (IRSNB P 9747–P 9748). R. Hygophum derthonensis (Anfossi & Mosna, 1969), Torrente Stirone (IRSNB P 9749). S–T. Lampanyctus latesulcatus Nolf & Steurbaut, 1983, Sant’Agata Fossili (IRSNB P 9750–P 9751). U. Merluccius cf. merluccius (Linnaeus, 1758), Sant’Agata Fossili (IRSNB P 9752). V. Lobianchia dofleini (Zugmayer, 1911), Costa Vescovato (IRSNB P 9753). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829315" httpUri="https://zenodo.org/record/3829315/files/figure.png" pageId="18" pageNumber="19">Fig. 5</figureCitation>
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N–O). They are characterised by a more ventrally located sulcus, leaving a dorsal area that is wider than the ventral one, and by a prominent postero-dorsal angle. Their rostrum is directed straight anteriorly, and not upward, like in
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<taxonomicName id="2EF05C879474CE2478F2A084FD3F97BC" authority="Fraser-Brunner, 1949" authorityName="Fraser-Brunner" authorityYear="1949" box="[189,643,1235,1261]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="dea">
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<emphasis id="DB84FB169474CE2478F2A084FEC797BC" box="[189,379,1235,1261]" italics="true" pageId="18" pageNumber="19">Lampadena dea</emphasis>
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Fraser-Brunner, 1949
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</taxonomicName>
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(e.g.,
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<figureCitation id="71CB3B819474CE247A9DA084FC8E97BF" box="[722,818,1235,1262]" captionStart="Fig" captionStartId="17.[189,231,1514,1540]" captionTargetBox="[298,1290,265,1505]" captionTargetId="figure@17.[298,1290,265,1505]" captionTargetPageId="17" captionText="Fig. 5. Tortonian fish otoliths from northern Italy. A. Diaphus holti Tåning, 1918, Mondovi, Madonna della Neve (IRSNB P 9732). B. Diaphus metopoclampoides Steurbaut, 1983, Sant’Alosio (IRSNB P 9733). C–E. Diaphus regani Tåning, 1932, Montegibbio (IRSNB P 9734–P 9736). F–G. Diaphus pedemontanus (Robba, 1970), Sant’Alosio (IRSNB P 9737–P 9738). H. Diaphus aff. rafinesquii (Cocco, 1838), Mondovi, Madonna della Neve (IRSNB P 9739). I. Diaphus aff. splendidus (Brauer, 1904), Costa Vescovato (IRSNB P 9740). J. Diaphus cahuzaci Steurbaut, 1979, Mondovi, Madonna della Neve (IRSNB P 9741). K. Lampadena aff. dea Fraser-Brunner, 1949, Torrente Stirone (IRSNB P 9742). L. Lampadena aff. speculigeroides Brzobohatý & Nolf, 1996, Stazzano (IRSNB P 9743). M. Hygophum hygomii (Lütken, 1892), Mondovi, Madonna della Neve (IRSNB P 9744). N–O. Lampadena gracilis (Schubert, 1912); N. Alba, Tanaro (50 m), O. Mondovi, Madonna della Neve (IRSNB P 9745–P 9746). P–Q. Lobianchia gemellarii (Cocco, 1838), Sant’Alosio (IRSNB P 9747–P 9748). R. Hygophum derthonensis (Anfossi & Mosna, 1969), Torrente Stirone (IRSNB P 9749). S–T. Lampanyctus latesulcatus Nolf & Steurbaut, 1983, Sant’Agata Fossili (IRSNB P 9750–P 9751). U. Merluccius cf. merluccius (Linnaeus, 1758), Sant’Agata Fossili (IRSNB P 9752). V. Lobianchia dofleini (Zugmayer, 1911), Costa Vescovato (IRSNB P 9753). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829315" httpUri="https://zenodo.org/record/3829315/files/figure.png" pageId="18" pageNumber="19">Fig. 5K</figureCitation>
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). Recently, based only on otoliths,
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<bibRefCitation id="8D615AF59474CE247C93A084FEA99641" author="Schwarzhans W." pageId="18" pageNumber="19" pagination="151 - 184" refId="ref23703" refString="Schwarzhans W. 2013 b. Otoliths from the Miocene of West Africa, primarily from the Mandorove Formation of Gabon. Palaeo Ichthyologica 13: 151 - 184. https: // doi. org / 10.13140 / 2.1.4576.3200" type="journal article" year="2013">Schwarzhans (2013b</bibRefCitation>
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: p. 160) erected a new fossil genus
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<taxonomicName id="2EF05C879474CE247A83A0A1FC359641" authorityName="Schwarzhans" authorityYear="2013" box="[716,905,1270,1296]" class="Actinopterygii" family="Myctophidae" genus="Paralampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
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<emphasis id="DB84FB169474CE247A83A0A1FC359641" box="[716,905,1270,1296]" italics="true" pageId="18" pageNumber="19">Paralampadena</emphasis>
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</taxonomicName>
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including three fossil species from West Africa as well as
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<taxonomicName id="2EF05C879474CE2479C7A14DFDBD9662" baseAuthorityName="Schubert" baseAuthorityYear="1912" box="[392,513,1305,1332]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="gracilis">
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<emphasis id="DB84FB169474CE2479C7A14DFE249662" box="[392,408,1306,1331]" italics="true" pageId="18" pageNumber="19">L</emphasis>
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.
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<emphasis id="DB84FB169474CE2479E8A14EFDBD9662" box="[423,513,1305,1331]" italics="true" pageId="18" pageNumber="19">gracilis</emphasis>
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</taxonomicName>
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. However, following the diagnostic features described for
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<taxonomicName id="2EF05C879474CE247CFDA14EFAD39662" authorityName="Schwarzhans" authorityYear="2013" box="[1202,1391,1305,1331]" class="Actinopterygii" family="Myctophidae" genus="Paralampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
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<emphasis id="DB84FB169474CE247CFDA14EFAD39662" box="[1202,1391,1305,1331]" italics="true" pageId="18" pageNumber="19">Paralampadena</emphasis>
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</taxonomicName>
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, each criterion is covered by the high variability of extant
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<taxonomicName id="2EF05C879474CE247BD1A16AFB959606" authorityName="Goode & Bean" authorityYear="1893" box="[926,1065,1341,1367]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
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<emphasis id="DB84FB169474CE247BD1A16AFB959606" box="[926,1065,1341,1367]" italics="true" pageId="18" pageNumber="19">Lampadena</emphasis>
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</taxonomicName>
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otoliths. For example, the statement that
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<taxonomicName id="2EF05C879474CE247921A137FD97962B" authorityName="Schwarzhans" authorityYear="2013" box="[366,555,1376,1402]" class="Actinopterygii" family="Myctophidae" genus="Paralampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="DB84FB169474CE247921A137FD97962B" box="[366,555,1376,1402]" italics="true" pageId="18" pageNumber="19">Paralampadena</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
does not exhibit ventral denticles, a postero-dorsal depression, which can be rather deep and angular, thus reducing the posterior rim of the otolith to some extent, and the comparatively long cauda, can all be observed in the otoliths of Recent
|
|||
|
|
<taxonomicName id="2EF05C879474CE247BB6A1F1FACD9690" authority="Parr, 1928" authorityName="Parr" authorityYear="1928" box="[1017,1393,1446,1473]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="anomala">
|
|||
|
|
<emphasis id="DB84FB169474CE247BB6A1F1FB4D9691" box="[1017,1265,1446,1472]" italics="true" pageId="18" pageNumber="19">Lampadena anomala</emphasis>
|
|||
|
|
Parr, 1928
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="2EF05C879474CE2478F2A19CFEBB96B5" authorityName="Fraser-Brunner" authorityYear="1949" box="[189,263,1482,1508]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="dea">
|
|||
|
|
<emphasis id="DB84FB169474CE2478F2A19CFF7196B5" box="[189,205,1483,1508]" italics="true" pageId="18" pageNumber="19">L</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="DB84FB169474CE247892A19DFEBB96B5" box="[221,263,1482,1508]" italics="true" pageId="18" pageNumber="19">dea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="2EF05C879474CE247959A19CFD6096B5" authority="Nafpaktitis & Paxton, 1968" authorityName="Nafpaktitis & Paxton" authorityYear="1968" box="[278,732,1482,1508]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="notialis">
|
|||
|
|
<emphasis id="DB84FB169474CE247959A19CFE3296B5" box="[278,398,1482,1508]" italics="true" pageId="18" pageNumber="19">L. notialis</emphasis>
|
|||
|
|
Nafpaktitis & Paxton, 1968
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="2EF05C879474CE247B53A19CFB7B96B5" authority="Goode & Bean, 1896" authorityName="Goode & Bean" authorityYear="1896" box="[796,1223,1482,1508]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="speculigera">
|
|||
|
|
<emphasis id="DB84FB169474CE247B53A19CFC7896B5" box="[796,964,1482,1508]" italics="true" pageId="18" pageNumber="19">L. speculigera</emphasis>
|
|||
|
|
Goode & Bean, 1896
|
|||
|
|
</taxonomicName>
|
|||
|
|
(see
|
|||
|
|
<bibRefCitation id="8D615AF59474CE247D49A19DFE8B9556" author="Girone A. & Nolf D." pageId="18" pageNumber="19" pagination="493 - 500" refId="ref21925" refString="Girone A. & Nolf D. 2002. Lampadena ionica: a new teleost from the Mediterranean Pleistocene. Rivista Italiana di Paleontologia e Stratigrafia 108 (3): 493 - 500." type="journal article" year="2002">Girone & Nolf 2002</bibRefCitation>
|
|||
|
|
for the otolith iconography of all the Recent
|
|||
|
|
<taxonomicName id="2EF05C879474CE247B18A1BAFC5E9556" authorityName="Goode & Bean" authorityYear="1893" box="[855,994,1517,1543]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="DB84FB169474CE247B18A1BAFC5E9556" box="[855,994,1517,1543]" italics="true" pageId="18" pageNumber="19">Lampadena</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species). Consequently, we retain that there is insufficient evidence for separating
|
|||
|
|
<taxonomicName id="2EF05C879474CE247ABEA246FCD6957B" baseAuthorityName="Schubert" baseAuthorityYear="1912" box="[753,874,1552,1579]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="gracilis">
|
|||
|
|
<emphasis id="DB84FB169474CE247ABEA246FCBD957B" box="[753,769,1553,1578]" italics="true" pageId="18" pageNumber="19">L</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="DB84FB169474CE247B5FA247FCD6957B" box="[784,874,1552,1578]" italics="true" pageId="18" pageNumber="19">gracilis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from the genus
|
|||
|
|
<taxonomicName id="2EF05C879474CE247C66A247FB08957B" authorityName="Goode & Bean" authorityYear="1893" box="[1065,1204,1552,1578]" class="Actinopterygii" family="Myctophidae" genus="Lampadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myctophiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="DB84FB169474CE247C66A247FB08957B" box="[1065,1204,1552,1578]" italics="true" pageId="18" pageNumber="19">Lampadena</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|