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292 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.394.6446" ID-GBIF-Dataset="8534078f-1f99-4b9e-866b-80609e341172" ID-PMC="PMC3978267" ID-Pensoft-Pub="1313-2970-394-1" ID-PubMed="24715784" ID-ZBK="32F5211696ED46C68BF9BE59C09F0B1A" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-394-1" ModsDocOrigin="ZooKeys 394" ModsDocTitle="A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae)" checkinTime="1451246058701" checkinUser="pensoft" docAuthor="Yoshimura, Masashi & Fisher, Brian L." docDate="2014" docId="59BADC10D0382F7920B224B9C60878A3" docLanguage="en" docName="ZooKeys 394: 1-99" docOrigin="ZooKeys 394" docSource="http://dx.doi.org/10.3897/zookeys.394.6446" docTitle="Mystrium eques Yoshimura & Fisher, sp. n." docType="treatment" docUuid="0FF584D9-44A0-44C5-AF5E-BA17BF4ACE7B" docUuidSource="ZooBank" docVersion="4" lastPageNumber="58" masterDocId="FF955F09FFA4FFC3FFC5FFA24014FFB0" masterDocTitle="A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae)" masterLastPageNumber="99" masterPageNumber="1" pageNumber="52" updateTime="1668158113736" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yoshimura, Masashi</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>394</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>99</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.394.6446</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.394.6446</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-394-1</mods:identifier>
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<mods:identifier type="ZBK">32F5211696ED46C68BF9BE59C09F0B1A</mods:identifier>
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<mods:identifier type="ZooBank">32F5211696ED46C68BF9BE59C09F0B1A</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152052265" LSID="urn:lsid:zoobank.org:act:0FF584D9-44A0-44C5-AF5E-BA17BF4ACE7B" httpUri="http://treatment.plazi.org/id/59BADC10D0382F7920B224B9C60878A3" lastPageId="57" lastPageNumber="58" pageId="51" pageNumber="52">
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<subSubSection pageId="51" pageNumber="52" type="nomenclature">
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<paragraph pageId="51" pageNumber="52">
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<taxonomicName LSID="http://zoobank.org/0FF584D9-44A0-44C5-AF5E-BA17BF4ACE7B" authority="Yoshimura & Fisher" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="eques">Mystrium eques Yoshimura & Fisher</taxonomicName>
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<taxonomicNameLabel pageId="51" pageNumber="52">sp. n.</taxonomicNameLabel>
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Figs 18B, 18D, 18F, 21A, 22A, 50A, 51A, 52A, 53A, 54A, 55A, 56A
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</paragraph>
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</subSubSection>
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<subSubSection pageId="51" pageNumber="52" type="holotype">
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<paragraph pageId="51" pageNumber="52">Holotype.</paragraph>
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<paragraph pageId="51" pageNumber="52">Worker: CASENT0418314, BLF04691, MADAGASCAR, Toamasina, Tampolo, Masoala Peninsula, 40.4 km 154° SSE Maroantsetra (-15.73°, 49.96°), 30 m alt., 28.xi.-3.xii.2001, B.L.Fisher & H.J.Ratsirarson leg. [CASC].</paragraph>
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</subSubSection>
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<subSubSection pageId="51" pageNumber="52" type="paratypes">
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<paragraph pageId="51" pageNumber="52">Paratypes.</paragraph>
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<paragraph pageId="51" pageNumber="52">13 workers: CASENT0317390 [CASC], CASENT0317393 [CASC], CASENT0317394 [CASC], CASENT0317395 [CASC], CASENT0317396 [MSNG], CASENT0317397 [MHNG], CASENT0317398 [BMNH], CASENT0317399 [MCZC], CASENT0317443 [CASC], CASENT0418312 [MNHN], CASENT0418313 [CASC], CASENT0418316 [NHMB], CASENT0418318 [ZMHB]; 5 ergatoid queens: CASENT0418311 [CASC], CASENT0418315 [BMNH], CASENT0418317 [MHNG], CASENT0317391 [MCZC], CASENT0317392 [NHMB], with same data as holotype.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52" type="worker">
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<paragraph pageId="51" pageNumber="52">Worker.</paragraph>
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<paragraph pageId="51" pageNumber="52">Description. Measurements: holotype. HL 1.78, HW 1.65, SL 1.23, ML 1.87, HD 1.08, WL 2.01, PnW 0.94, PpW 0.83, PtW 0.85, PtL 0.57, CI 92.6, SI 74.4, MI 113.8, PpI 88.6, PtI 150.4.</paragraph>
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<paragraph pageId="51" pageNumber="52">HL 1.72-2.11, HW 1.59-2, SL 1.26-1.42, ML 1.81-2.18, HD 1.08-1.33, WL 2.07-2.34, PnW 0.92-1.1, PpW 0.83-1.02, PtW 0.83-1.03, PtL 0.54-0.67, CI 91.4-97.3, SI 70.4-78.9, MI 107.4-114.2, PpI 84.7-92.7, PtI 149.5-161.2 (10 specimens measured).</paragraph>
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<paragraph lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52">
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Posterolateral corner of head strongly to moderately expanding posteriorly. Posterior face of vertex forming almost a right angle with dorsal face on median line of head, so that declivity of vertex on lateral part as steep as that on median part. Whole region of vertex finely striated. Eye relatively smaller than that of
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
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. Anterior margin of clypeus strongly convex with long conical setae. Genal tooth of head relatively long, as long as lateral lobe of clypeus. Masticatory margin of mandible almost invisible in full-face view. Width of dorsal surface of mandible on distal portion slightly
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<pageBreakToken pageId="52" pageNumber="53" start="start">wider</pageBreakToken>
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than that on mandibular shaft. Second maxillary palpomere longer than third. First flagellomere (third antennal segment) as long as pedicel (second antennal segment). Strong, deep and thick longitudinal striae regularly impressed on whole central part of pronotal dorsum. Strong, deep, and thick longitudinal striae impressed on lateral surface of pronotum. Mesonotum differentiated from propodeum in dorsal view, length slightly shorter than that of propodeum. Metanotal groove shallowly and gently
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<pageBreakToken pageId="53" pageNumber="54" start="start">impressed</pageBreakToken>
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, and mesonotum higher than pronotum in lateral view. Metapleural gland bulla moderately developed, and propodeal declivity in lateral view almost straight. Petiole widened on posterior 1/4 and gently narrowed anteriorly in dorsal view, anterior margin straight to gently rounded and sometimes edged by thin striae.
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</paragraph>
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<paragraph pageId="53" pageNumber="54">Body color blackish brown. Appendages brighter, and four distal segments of antennal club yellowish.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="ergatoid queen">
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<paragraph pageId="53" pageNumber="54">Ergatoid queen.</paragraph>
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<paragraph pageId="53" pageNumber="54">Description. Measurements: HL 1.48-1.62, HW 1.41-1.57, SL 1.10-1.25, ML 1.38-1.56, HD 0.99-1.09, WL 1.95-2.10, PnW 0.85-0.95, PpW 0.83-0.94, PtW 0.87-0.97, PtL 0.52-0.56, CI 94.0-97.2, SI 77.7-83.9, MI 93.2-107.0, PpI 93.8-101.6, PtI 160.5-172.6 (7 specimens measured).</paragraph>
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<paragraph lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
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<pageBreakToken pageId="54" pageNumber="55" start="start">Wings</pageBreakToken>
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vestigial and forming small but distinct appendages. Wing sclerites undeveloped. Posterolateral corner of head strongly expanding posteriorly, expansion relatively weaker than that of workers. Posterior face of vertex forming almost a right angle with dorsal face on median line of head, so that declivity of vertex on lateral part as steep as on median part. Ventral half of vertex sculptured. Eye small but distinct. Ocelli absent. Anterior margin of clypeus distinctly convex with long to short conical
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<pageBreakToken pageId="55" pageNumber="56" start="start">setae</pageBreakToken>
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. Anterolateral portion of head with short spine. Masticatory margin of mandible almost invisible in full-face view. Dorsal surface of mandible on distal portion slightly wider than that on mandibular shaft. Spatulate or narrow spoon-shaped seta present on basal side of each basal denticle on masticatory margin of mandible. First flagellar segment on antenna about 1.0
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<normalizedToken originalValue="–1.2×">-1.2x</normalizedToken>
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length of pedicel. Setae on pronotum distinctly spatulate, widened distally with sharp or blunt apex. Metapleural gland bulla moderately developed, not expanding to dorsum of propodeal spiracle, so that propodeal declivity in lateral view weakly convex and rounded posteriorly on its ventral 1/3. Petiole relatively long in dorsal view, about 1.0
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<normalizedToken originalValue="–0.8×">-0.8x</normalizedToken>
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length of abdominal segment III.
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</paragraph>
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<paragraph pageId="55" pageNumber="56">Body color blackish to reddish brown.</paragraph>
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</subSubSection>
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<subSubSection pageId="55" pageNumber="56" type="male">
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<paragraph pageId="55" pageNumber="56">Male</paragraph>
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<paragraph pageId="55" pageNumber="56">unknown.</paragraph>
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<caption pageId="55" pageNumber="56">
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<paragraph pageId="55" pageNumber="56">
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Figure 50. Workers of voeltzkowi species group in lateral view. A
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
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sp. n. (CASENT0418314: holotype) B
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium janovitzi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="janovitzi">Mystrium janovitzi</taxonomicName>
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sp. n. (CASENT0482698: holotype) C
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mirror" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mirror">Mystrium mirror</taxonomicName>
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sp. n. (CASENT0429897: holotype) D
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
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(CASENT0496066) E
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium shadow" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="shadow">Mystrium shadow</taxonomicName>
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sp. n. (CASENT0318933: holotype) F
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium voeltzkowi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="voeltzkowi">Mystrium voeltzkowi</taxonomicName>
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(CASENT0002075).
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</paragraph>
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</caption>
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<caption pageId="55" pageNumber="56">
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<paragraph pageId="55" pageNumber="56">
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Figure 51. Head of the workers of voeltzkowi species group in full-face view. A
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
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sp. n. (CASENT0418314: holotype) B
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium janovitzi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="janovitzi">Mystrium janovitzi</taxonomicName>
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sp. n. (CASENT0482698: holotype) C
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mirror" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mirror">Mystrium mirror</taxonomicName>
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sp. n. (CASENT0429897: holotype) D
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
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(CASENT0496066) E
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium shadow" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="shadow">Mystrium shadow</taxonomicName>
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sp. n. (CASENT0318933: holotype) F
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium voeltzkowi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="voeltzkowi">Mystrium voeltzkowi</taxonomicName>
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(CASENT0002075).
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</paragraph>
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</caption>
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<caption pageId="55" pageNumber="56">
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<paragraph pageId="55" pageNumber="56">
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Figure 52. Workers of voeltzkowi species group in dorsal view. A
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
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sp. n. (CASENT0418314: holotype) B
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium janovitzi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="janovitzi">Mystrium janovitzi</taxonomicName>
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sp. n. (CASENT0482698: holotype) C
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mirror" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mirror">Mystrium mirror</taxonomicName>
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sp. n. (CASENT0429897: holotype) D
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
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(CASENT0496066) E
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium shadow" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="shadow">Mystrium shadow</taxonomicName>
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sp. n. (CASENT0318933: holotype) F
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium voeltzkowi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="voeltzkowi">Mystrium voeltzkowi</taxonomicName>
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(CASENT0002075).
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</paragraph>
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</caption>
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<caption pageId="55" pageNumber="56">
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<paragraph pageId="55" pageNumber="56">
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Figure 53. Ergatoid queens of voeltzkowi species group in lateral view. A
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
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sp. n. (CASENT0418317: paratype) B
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium janovitzi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="janovitzi">Mystrium janovitzi</taxonomicName>
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sp. n. (CASENT0482696) C
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mirror" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mirror">Mystrium mirror</taxonomicName>
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sp. n. (CASENT0318940: paratype) D
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
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(CASENT0496064) E
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<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium shadow" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="shadow">Mystrium shadow</taxonomicName>
|
|||
|
|
sp. n. (CASENT0077647: paratype) F
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium voeltzkowi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="voeltzkowi">Mystrium voeltzkowi</taxonomicName>
|
|||
|
|
(CASENT0002092).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption pageId="55" pageNumber="56">
|
|||
|
|
<paragraph pageId="55" pageNumber="56">
|
|||
|
|
Figure 54. Head of the ergatoid queens of voeltzkowi species group in full-face view. A
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
sp. n. (CASENT0418317: paratype) B
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium janovitzi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="janovitzi">Mystrium janovitzi</taxonomicName>
|
|||
|
|
sp. n. (CASENT0482696) C
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mirror" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mirror">Mystrium mirror</taxonomicName>
|
|||
|
|
sp. n. (CASENT0318940: paratype) D
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
(CASENT0496064) E
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium shadow" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="shadow">Mystrium shadow</taxonomicName>
|
|||
|
|
sp. n. (CASENT0077647: paratype) F
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium voeltzkowi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="voeltzkowi">Mystrium voeltzkowi</taxonomicName>
|
|||
|
|
(CASENT0002092).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption pageId="55" pageNumber="56">
|
|||
|
|
<paragraph pageId="55" pageNumber="56">
|
|||
|
|
Figure 55. Ergatoid queens of voeltzkowi species group in dorsal view. A
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
sp. n. (CASENT0418317: paratype) B
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium janovitzi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="janovitzi">Mystrium janovitzi</taxonomicName>
|
|||
|
|
sp. n. (CASENT0482696) C
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mirror" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mirror">Mystrium mirror</taxonomicName>
|
|||
|
|
sp. n. (CASENT0318940: paratype) D
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
(CASENT0496064) E
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium shadow" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="shadow">Mystrium shadow</taxonomicName>
|
|||
|
|
sp. n. (CASENT0077647: paratype) F
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium voeltzkowi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="voeltzkowi">Mystrium voeltzkowi</taxonomicName>
|
|||
|
|
(CASENT0002092).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption pageId="55" pageNumber="56">
|
|||
|
|
<paragraph pageId="55" pageNumber="56">
|
|||
|
|
Figure 56. Distribution maps for voeltzkowi species group. A
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
sp. n. B
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium janovitzi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="janovitzi">Mystrium janovitzi</taxonomicName>
|
|||
|
|
sp. n. C
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mirror" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mirror">Mystrium mirror</taxonomicName>
|
|||
|
|
sp. n. D
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
E
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium shadow" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="shadow">Mystrium shadow</taxonomicName>
|
|||
|
|
sp. n. F
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium voeltzkowi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="voeltzkowi">Mystrium voeltzkowi</taxonomicName>
|
|||
|
|
. Star symbols represent the type locality: the localities for
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium voeltzkowi" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="voeltzkowi">Mystrium voeltzkowi</taxonomicName>
|
|||
|
|
are approximate, which are assumed from the original descriptions.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="56" pageNumber="57" type="etymology">
|
|||
|
|
<paragraph pageId="56" pageNumber="57">
|
|||
|
|
<pageBreakToken pageId="56" pageNumber="57" start="start">Etymology</pageBreakToken>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="56" pageNumber="57">This species name is the Latin word eques, inspired by the distinctly developed clypeus (shield) on the new species. The species epithet is a noun and invariant.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="56" pageNumber="57" type="distribution">
|
|||
|
|
<paragraph pageId="56" pageNumber="57">Distribution.</paragraph>
|
|||
|
|
<paragraph pageId="56" pageNumber="57">MADAGASCAR: as in Figure 56A.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="additional material examined">
|
|||
|
|
<paragraph pageId="56" pageNumber="57">Additional material examined.</paragraph>
|
|||
|
|
<paragraph lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
|
|||
|
|
In addition to the type material, specimens from the following localities were examined in this study: MADAGASCAR. Toamasina. Montagne
|
|||
|
|
<normalizedToken originalValue="d’Akirindro">d'Akirindro</normalizedToken>
|
|||
|
|
7.6 km 341° NNW Ambinanitelo (-15.28833°, 49.54833°), rainforest, 600 m alt.; 5.3 km SSE Ambanizana, Andranobe (-15.66667°, 49.96667°), rainforest, 425 m alt.; Tampolo, Masoala Peninsula, 40.4 km 154° SSE Maroantsetra (-15.73°, 49.96°), rainforest, 30 m alt.; Res. Ambodiriana, 4.8 km 306° Manompana, along Manompana river (-16.67233°, 49.70117°), rainforest, 125 m alt.;
|
|||
|
|
<pageBreakToken pageId="57" pageNumber="58" start="start">
|
|||
|
|
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
|||
|
|
</pageBreakToken>
|
|||
|
|
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
|||
|
|
Ambatovaky, Sandrangato river (-16.77274°, 49.26551°), rainforest, 450 m alt.; Ile Sainte Marie,
|
|||
|
|
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
Kalalao, 9.9 km 34° Ambodifotatra (-16.9225°, 49.88733°), rainforest, 100 m alt.;
|
|||
|
|
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
|||
|
|
Speciale Manombo 24.5 km 228° Farafangana (-23.01583°, 47.719°), rainforest, 30 m alt.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="57" pageNumber="58" type="remarks">
|
|||
|
|
<paragraph pageId="57" pageNumber="58">Remarks.</paragraph>
|
|||
|
|
<paragraph pageId="57" pageNumber="58">
|
|||
|
|
The worker of
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
is easily distinguished from those of other
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">Mystrium</taxonomicName>
|
|||
|
|
species by a combination of the following characters: strong and deep longitudinal sculpture on the dorsal and lateral surface of pronotum (as in Fig. 17B); anteromedial portion of the clypeus strongly convex anteriorly (Fig. 18B); and sharper angle between the posterior and dorsal faces of the vertex on the median line (as in Fig. 16A). The queen of
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
is differentiated from other
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">Mystrium</taxonomicName>
|
|||
|
|
queens by having vestigial wings reduced to small appendages (as in Fig. 25D) with undeveloped wing sclerites (ergatoid), posterior face of the vertex forming approximately a right angle with its dorsal face on the median line (Fig. 21A), and a strongly convex anterior clypeal margin (Fig. 22A).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="57" pageNumber="58">
|
|||
|
|
The worker of
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
is similar to that of
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
, however, the strongly convex anterior clypeal margin separates
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
(Fig. 18B) from
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
(Fig. 18A), in which the anterior margin of the clypeus is straight. The male of
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
is not yet known. Because of the similarity between the workers and queens of
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
, we assume the male of
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
is also similar to that of
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
. The most easily distinguishable character between workers of
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
, the anterior margin of the clypeus (Figs 18A, B), is unfortunately not useful for males because the anterior clypeal margin and the conical setae on it are less developed in males.
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
(Fig. 56A) has a sympatric distribution with
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
(Fig. 56D), so collection locality does not separate these two similar species either. The current male specimens identified as
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
|
|||
|
|
may contain males of
|
|||
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium eques" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="eques">Mystrium eques</taxonomicName>
|
|||
|
|
. Further studies are necessary to resolve this issue.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|