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<document id="2473DEFBFC2A5CB6A974DF7D68006CB6" ID-CLB-Dataset="29103" ID-DOI="10.11646/zootaxa.4455.2.5" ID-GBIF-Dataset="568a5364-52f2-4feb-b08b-87eb85d4cf77" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457283" ID-ZooBank="8CB311F1-431C-4861-956E-3200329E4F0F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539261916473" checkinUser="plazi" docAuthor="Nada, B. &amp; Ballantyne, L. A." docDate="2018" docId="039887E6FFF37A0D9ECB2492A29B639F" docLanguage="en" docName="zootaxa.4455.2.5.pdf" docOrigin="Zootaxa 4455 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pygoluciola" docType="key" docVersion="7" lastPageNumber="348" masterDocId="FFA1FF9EFFF67A089E5C2657A7076769" masterDocTitle="A new species of Pygoluciola Wittmer with unusual abdominal configuration, from lowland dipterocarp forest in Peninsular Malaysia (Coleoptera: Lampyridae: Luciolinae)" masterLastPageNumber="362" masterPageNumber="343" pageNumber="348" updateTime="1698575593942" updateUser="ExternalLinkService">
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<mods:titleInfo id="D1EF7611B1DF93B72B7B4BD7142F7283">
<mods:title id="602E4938EBDD931C7DACA4A2BE424BFF">A new species of Pygoluciola Wittmer with unusual abdominal configuration, from lowland dipterocarp forest in Peninsular Malaysia (Coleoptera: Lampyridae: Luciolinae)</mods:title>
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<mods:name id="788EB6C652AFB4CDBD06262EBD3A826B" type="personal">
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<mods:namePart id="CC1414EA8367D57D76C478C50487464A">Nada, B.</mods:namePart>
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<mods:name id="5451D95F5585E6EC3D3B7CDF3DA10B13" type="personal">
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<mods:namePart id="5EC3F6F7BCB2629DC52FC2EF636A7193">Ballantyne, L. A.</mods:namePart>
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<mods:title id="70A9A44326EB123FDA8F9435B8328C26">Zootaxa</mods:title>
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<mods:date id="9E750D651CC93385C7EFB3250D0A74B4">2018</mods:date>
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<mods:number id="85E310BC74E67CB6E64520B5849186AA">2018-08-02</mods:number>
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<mods:number id="667D7522F4F8897EF71D7C0028E89385">4455</mods:number>
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<mods:identifier id="191BB71806046B3FF2AE4E89D58C6E73" type="DOI">10.11646/zootaxa.4455.2.5</mods:identifier>
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<treatment id="039887E6FFF37A0D9ECB2492A29B639F" ID-DOI="http://doi.org/10.5281/zenodo.6485263" ID-GBIF-Taxon="148691253" ID-Zenodo-Dep="6485263" LSID="urn:lsid:plazi:treatment:039887E6FFF37A0D9ECB2492A29B639F" httpUri="http://treatment.plazi.org/id/039887E6FFF37A0D9ECB2492A29B639F" lastPageNumber="348" pageId="5" pageNumber="348">
<subSubSection id="C32B657BFFF37A0D9ECB2492A59E6469" pageId="5" pageNumber="348" type="nomenclature">
<paragraph id="8B8E36F0FFF37A0D9ECB2492A5AB65B6" blockId="5.[151,684,709,768]" box="[151,684,709,735]" pageId="5" pageNumber="348">
<heading id="D0C6819CFFF37A0D9ECB2492A5AB65B6" bold="true" box="[151,684,709,735]" fontSize="11" level="1" pageId="5" pageNumber="348" reason="1">
<emphasis id="B945EAE2FFF37A0D9ECB2492A5AB65B6" bold="true" box="[151,684,709,735]" pageId="5" pageNumber="348">
Key to species of
<taxonomicName id="4C314D73FFF37A0D9F312492A6FB65B6" ID-CoL="63QVY" box="[365,508,709,735]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAE2FFF37A0D9F312492A6FB65B6" bold="true" box="[365,508,709,735]" italics="true" pageId="5" pageNumber="348">Pygoluciola</emphasis>
</taxonomicName>
using females
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E36F0FFF37A0D9ECB24B0A59E6469" blockId="5.[151,684,709,768]" box="[151,665,743,768]" pageId="5" pageNumber="348">
Extended from
<bibRefCitation id="EFA04B01FFF37A0D9F1924B0A59E6469" box="[325,665,743,768]" pageId="5" pageNumber="348" refId="ref8493" refString="Ballantyne, L. A. &amp; Lambkin, C. (2006) A phylogenetic reassessment of the rare S. E. Asian firefly genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae). Raffles Bulletin of Zoology, 54 (1), 21 - 48." type="journal article">Ballantyne &amp; Lambkin (2006)</bibRefCitation>
</paragraph>
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<subSubSection id="C32B657BFFF37A0D9ECB257EA29B639F" pageId="5" pageNumber="348" type="key">
<keyStep id="30C52E55FFF37A0D9ECB257EA29C6432" pageId="5" pageNumber="348">
<paragraph id="8B8E36F0FFF37A0D9ECB257EA29B6457" blockId="5.[151,1436,809,1270]" box="[151,1436,809,830]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB257EA29B6457" box="[151,1436,809,830]" pageId="5" pageNumber="348">
1 All tibiae curved.....................................................................
<taxonomicName id="4C314D73FFF37A0D9A93257EA29B6457" ID-CoL="4QP8S" authority="(Ballantyne)" authorityName="Ballantyne" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[1231,1436,809,830]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="guigliae">
<emphasis id="B945EAE2FFF37A0D9A93257EA21A6457" box="[1231,1309,809,830]" italics="true" pageId="5" pageNumber="348">guigliae</emphasis>
(Ballantyne)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF37A0D9ECB2511A29C6432" blockId="5.[151,1436,809,1270]" box="[151,1435,838,859]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB2511A29C6432" box="[151,1435,838,859]" pageId="5" pageNumber="348">- No tibiae curved...................................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E55FFF37A0D9ECB2533A29B6362" pageId="5" pageNumber="348">
<paragraph id="8B8E36F0FFF37A0D9ECB2533A29B64B8" blockId="5.[151,1436,809,1270]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB2533A29B64B8" pageId="5" pageNumber="348">
2. Posterior margin of
<date id="FF8F1030FFF37A0D9FFF2533A6C76410" box="[419,448,868,889]" pageId="5" pageNumber="348">V7</date>
deeply emarginate; bearing a small ridge anterior to median area of deepest emargination (possibility this ridge is an artefact having been seen only on dried pinned specimens);
<date id="FF8F1030FFF37A0D9DED25D6A4C864FF" box="[945,975,897,918]" pageId="5" pageNumber="348">V8</date>
with anteromedian prolongation not any more sclerotised than remainder of
<date id="FF8F1030FFF37A0D9FA225C9A51964DA" box="[510,542,926,947]" pageId="5" pageNumber="348">V8</date>
; T7 with anteromedian area rounded and elevated, lateral areas not flattened (
<bibRefCitation id="EFA04B01FFF37A0D9B4B25C9A6DE64B8" pageId="5" pageNumber="348" refId="ref8435" refString="Ballantyne, L. A. &amp; Lambkin, C. (2001) A new firefly, Luciola (Pygoluciola) kinabalua sp. nov. (Coleoptera: Lampyridae), from Malaysia, with observations on a possible copulation clamp. Raffles Bulletin of Zoology, 49 (2), 363 - 377." type="journal article">Ballantyne &amp; Lambkin 2001 figs 16, 17</bibRefCitation>
)............................................................
<taxonomicName id="4C314D73FFF37A0D9AE125EBA29B64B8" ID-CoL="3WCSR" authority="(Ballantyne)" authorityName="Ballantyne" baseAuthorityName="Ballantyne &amp; Lambkin" baseAuthorityYear="2001" box="[1213,1436,956,977]" class="Insecta" family="Lampyridae" genus="Luciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="kinabalua">
<emphasis id="B945EAE2FFF37A0D9AE125EBA21964B8" box="[1213,1310,956,977]" italics="true" pageId="5" pageNumber="348">kinabalua</emphasis>
(Ballantyne)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF37A0D9ECB258EA29B6362" blockId="5.[151,1436,809,1270]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB258EA29B6362" pageId="5" pageNumber="348">
- Posterior margin of
<date id="FF8F1030FFF37A0D9FFF258EA6C76487" box="[419,448,985,1006]" pageId="5" pageNumber="348">V7</date>
without an anteromedian ridge;
<date id="FF8F1030FFF37A0D9CA7258EA41E6487" box="[763,793,985,1006]" pageId="5" pageNumber="348">V8</date>
with anteromedian prolongation well sclerotised and visibly separated from remainder of
<date id="FF8F1030FFF37A0D9F9025A1A6EB6362" box="[460,492,1014,1035]" pageId="5" pageNumber="348">V7</date>
; T7 without a rounded and elevated anteromedian area, with lateral areas flattened........... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E55FFF37A0D9ECB2243A29C630A" pageId="5" pageNumber="348">
<paragraph id="8B8E36F0FFF37A0D9ECB2243A29B632F" blockId="5.[151,1436,809,1270]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB2243A29B632F" pageId="5" pageNumber="348">
3 Posterolateral areas of
<date id="FF8F1030FFF37A0D9FE02243A6DE6340" box="[444,473,1044,1065]" pageId="5" pageNumber="348">V7</date>
irregularly expanded with aggregation of fat body beneath (
<bibRefCitation id="EFA04B01FFF37A0D9A462243A2606340" box="[1050,1383,1044,1065]" pageId="5" pageNumber="348" refId="ref8493" refString="Ballantyne, L. A. &amp; Lambkin, C. (2006) A phylogenetic reassessment of the rare S. E. Asian firefly genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae). Raffles Bulletin of Zoology, 54 (1), 21 - 48." type="journal article">Ballantyne &amp; Lambkin 2006 fig. 7</bibRefCitation>
)...........................................................................................
<taxonomicName id="4C314D73FFF37A0D9AA32266A29B632F" ID-CoL="4QP8W" authority="Ballantyne" authorityName="Ballantyne" authorityYear="2008" box="[1279,1436,1073,1094]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="B945EAE2FFF37A0D9AA32266A22A632F" box="[1279,1325,1073,1094]" italics="true" pageId="5" pageNumber="348">satoi</emphasis>
Ballantyne
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF37A0D9ECB2219A29C630A" blockId="5.[151,1436,809,1270]" box="[151,1435,1102,1123]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB2219A29C630A" box="[151,1435,1102,1123]" pageId="5" pageNumber="348">
- Posterolateral areas of
<date id="FF8F1030FFF37A0D9FE02219A6DE630A" box="[444,473,1102,1123]" pageId="5" pageNumber="348">V7</date>
not expanded.................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E55FFF37A0D9ECB223BA29B639F" pageId="5" pageNumber="348">
<paragraph id="8B8E36F0FFF37A0D9ECB223BA29B63D5" blockId="5.[151,1436,809,1270]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB223BA29B63D5" pageId="5" pageNumber="348">
4. Ventral surface yellow except for white LOs in
<date id="FF8F1030FFF37A0D9CE7223BA5DC63E8" box="[699,731,1132,1153]" pageId="5" pageNumber="348">V6</date>
, and narrow black line across posterior margin of
<date id="FF8F1030FFF37A0D9A8D223BA3E863E8" box="[1233,1263,1132,1153]" pageId="5" pageNumber="348">V5</date>
(
<figureCitation id="130A2A75FFF37A0D9AA2223BA29063E8" box="[1278,1431,1132,1153]" captionStart="FIGURES 25–28" captionStartId="11.[151,264,1936,1959]" captionTargetBox="[215,1371,193,1877]" captionTargetId="figure@11.[208,1379,193,1916]" captionTargetPageId="11" captionText="FIGURES 25–28. Ventral aspect, mating pairs, male to left (25 JI40 (8), 26 B10 (gap 5-6)). 27, 28 dissected abdomen, anterior end to top of page (B19 (2-3)) 27 ventral; 28 dorsal. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457293/files/figure.png" pageId="5" pageNumber="348">Figs 25, 26, 27</figureCitation>
, 29‾33); bursa plates in form of two very elongate slender projections one directed anteriorly and one posteriorly (
<figureCitation id="130A2A75FFF37A0D9B7C22DFA26E63F7" box="[1312,1385,1160,1182]" captionStart="FIGURES 34–37" captionStartId="13.[151,264,1896,1919]" captionTargetBox="[220,1342,209,1859]" captionTargetId="figure@13.[220,1342,209,1859]" captionTargetPageId="13" captionText="FIGURES 34–37. Female reproductive system (B19 (2-3)). 34 system without ovaries, ovipositor to left and digesting gland to right, from above; 35 as for 34 from side expanded base of spermatheca arrowed; 36 detail bursa and expanded base of spermatheca (arrowed); 37 bursa in area of bursa hooks from above. Figure legend: B = bursa; SDG = spermatophore digesting gland; V = valvifers of the ovipositor. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457295/files/figure.png" pageId="5" pageNumber="348">Figs 34</figureCitation>
‾37)......................................................................................
<taxonomicName id="4C314D73FFF37A0D9AA922F1A24E63D2" ID-CoL="8N8PY" authority="Nada &amp; Ballantyne, 2018" authorityName="Nada &amp; Ballantyne" authorityYear="2018" box="[1269,1353,1190,1211]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="dunguna" status="sp. nov.">
<emphasis id="B945EAE2FFF37A0D9AA922F1A24E63D2" box="[1269,1353,1190,1211]" italics="true" pageId="5" pageNumber="348">dunguna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765799FFF37A0D9B1322F1A29B63D5" box="[1359,1436,1190,1212]" pageId="5" pageNumber="348" rank="species">
<emphasis id="B945EAE2FFF37A0D9B1322F1A29B63D5" bold="true" box="[1359,1436,1190,1212]" pageId="5" pageNumber="348">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF37A0D9ECB2293A29B639F" blockId="5.[151,1436,809,1270]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB2293A29B639F" pageId="5" pageNumber="348">
<emphasis id="B945EAE2FFF37A0D9ECB2293A79863B0" box="[151,159,1220,1241]" italics="true" pageId="5" pageNumber="348">-</emphasis>
Ventral surface brown except for white LOs in
<date id="FF8F1030FFF37A0D9CF32293A5C863B0" box="[687,719,1220,1241]" pageId="5" pageNumber="348">V6</date>
, without narrow black line across posterior margin of
<date id="FF8F1030FFF37A0D9ABF2293A20463B0" box="[1251,1283,1220,1241]" pageId="5" pageNumber="348">V5</date>
; bursa plates in form of two anteriorly directed hooks....................................................
<emphasis id="B945EAE2FFF37A0D9AE522B6A21A639F" box="[1209,1309,1249,1270]" italics="true" pageId="5" pageNumber="348">.. wittmeri</emphasis>
(Ballantyne)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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