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<document id="2B35776E4E84CDEFA9D9B1055962D0BE" ID-CLB-Dataset="56366" ID-DOI="10.11646/zootaxa.5306.2.6" ID-GBIF-Dataset="05c564eb-3640-4ffb-903d-06d5219856e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="8058811" ID-ZooBank="7D456B11-BBF6-4816-AB81-5D02ACE02DAD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1687249692376" checkinUser="plazi" docAuthor="Szawaryn, Karol &amp; Czerwiński, Tomasz" docDate="2023" docId="CD03879FFFD5FF84158AF9A21D326DEF" docLanguage="en" docName="zootaxa.5306.2.6.pdf" docOrigin="Zootaxa 5306 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5306.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Merma fisheri Szawaryn &amp; Czerwiński 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="275" masterDocId="313AFFE7FFD3FF8D151DFFDE196E6E02" masterDocTitle="Three new species of Merma Weise (Coleoptera: Coccinellidae: Epilachnini) from Madagascar" masterLastPageNumber="276" masterPageNumber="266" pageNumber="272" updateTime="1687345819786" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9D87EF172864B6D88CB70C438995E6FC">Three new species of Merma Weise (Coleoptera: Coccinellidae: Epilachnini) from Madagascar</mods:title>
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<mods:namePart id="AC72AD69ABCEDA9E1D16B7FB6C60FC43">Szawaryn, Karol</mods:namePart>
<mods:nameIdentifier id="1F10E066D4FE6DB73CF7C01021D18EFA" type="ORCID">0000-0002-9329-4268</mods:nameIdentifier>
<mods:affiliation id="70C9D2464B3C7CFD5D08A48D08D4CDCA">k. szawaryn @ gmail. com; https: // orcid. org / 0000 - 0002 - 9329 - 4268</mods:affiliation>
<mods:nameIdentifier id="93010C1BC7BF90DA6E205BB73377EA3F" type="email">k.szawaryn@gmail.com</mods:nameIdentifier>
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<mods:namePart id="CE2438B86482A3B87FCF161C463F5B55">Czerwiński, Tomasz</mods:namePart>
<mods:nameIdentifier id="97ABB7721780B096FD0350279AC06CDC" type="ORCID">0000-0002-0598-7858</mods:nameIdentifier>
<mods:affiliation id="CC63C8526D3FF5DB6E63D8AD4633B9F3">tczerwinski @ miiz. waw. pl; https: // orcid. org / 0000 - 0002 - 0598 - 7858</mods:affiliation>
<mods:nameIdentifier id="1B17C7E2CC4CE641AC6B3E831F7A29D6" type="email">tczerwinski@miiz.waw.pl</mods:nameIdentifier>
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<mods:title id="1DBBF5C707F1E31BF283CC7143A7ACC3">Zootaxa</mods:title>
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<mods:part id="DF7D8D68DE3B395F268F714B9B5C8DF4">
<mods:date id="F4F376621D9BC7388BF80FF216886D22">2023</mods:date>
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<mods:number id="8F8E569B1F35AB80D79EF0E8269C3294">2023-06-20</mods:number>
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<mods:number id="F48DE651F22516EAA63C90BA8017398C">5306</mods:number>
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<treatment id="CD03879FFFD5FF84158AF9A21D326DEF" ID-DOI="http://doi.org/10.5281/zenodo.8058893" ID-GBIF-Taxon="209696207" ID-Zenodo-Dep="8058893" LSID="urn:lsid:plazi:treatment:CD03879FFFD5FF84158AF9A21D326DEF" httpUri="http://treatment.plazi.org/id/CD03879FFFD5FF84158AF9A21D326DEF" lastPageId="9" lastPageNumber="275" pageId="6" pageNumber="272">
<subSubSection id="0DB06502FFD5FF8B158AF9A218C46897" box="[151,426,1659,1686]" pageId="6" pageNumber="272" type="nomenclature">
<paragraph id="45153689FFD5FF8B158AF9A218C46897" blockId="6.[151,970,1659,1758]" box="[151,426,1659,1686]" pageId="6" pageNumber="272">
<heading id="1E5D81E5FFD5FF8B158AF9A218C46897" bold="true" box="[151,426,1659,1686]" fontSize="11" level="1" pageId="6" pageNumber="272" reason="1">
<emphasis id="77DEEA9BFFD5FF8B158AF9A218C46897" bold="true" box="[151,426,1659,1686]" pageId="6" pageNumber="272">
<taxonomicName id="82AA4D0AFFD5FF8B158AF9A2182B6894" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[151,325,1659,1686]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="272" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
<emphasis id="77DEEA9BFFD5FF8B158AF9A2182B6894" bold="true" box="[151,325,1659,1686]" italics="true" pageId="6" pageNumber="272">Merma fisheri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CED57E0FFD5FF8B1451F9A518C46897" box="[332,426,1659,1685]" pageId="6" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DB06502FFD5FF8B158AF9411AA468DC" pageId="6" pageNumber="272" type="description">
<paragraph id="45153689FFD5FF8B158AF941184A68B8" blockId="6.[151,970,1659,1758]" box="[151,292,1695,1722]" pageId="6" pageNumber="272">
<figureCitation id="DD912A0CFFD5FF8B158AF941186468BB" box="[151,266,1695,1721]" captionStart="FIGURE 1" captionStartId="2.[151,250,1512,1536]" captionTargetBox="[175,1428,187,1482]" captionTargetId="figure-17@2.[153,1434,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Merma species. A–C) M. ankarana sp. nov., male, holotype (MNHN); A) dorsal; B) lateral; C) frontal; D–F) M. antakotako sp. nov., male, paratype (MNHN); D) dorsal; E) lateral; F) frontal; G–I) M. fisheri sp. nov., male, holotype (CAS); G) dorsal; H) lateral; I) frontal." figureDoi="http://doi.org/10.5281/zenodo.8058813" httpUri="https://zenodo.org/record/8058813/files/figure.png" pageId="6" pageNumber="272">Figs 1G–I</figureCitation>
,
<figureCitation id="DD912A0CFFD5FF8B140BF941184A68B8" box="[278,292,1695,1722]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="6" pageNumber="272">5</figureCitation>
</paragraph>
<paragraph id="45153689FFD5FF8B158AF91D1AA468DC" blockId="6.[151,970,1659,1758]" box="[151,970,1731,1758]" pageId="6" pageNumber="272">
<uri id="313B3A8BFFD5FF8B158AF91D1AA468DC" box="[151,970,1731,1758]" pageId="6" pageNumber="272">
urn:lsid:zoobank.org:act:
<uuid id="310C0C5CFFD5FF8B14B1F91D1AA468DC" box="[428,970,1731,1758]" pageId="6" pageNumber="272">8522CA47-C497-4C39-B947-8B0D77A3D26C</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="0DB06502FFD5FF8B158AF8D81C3E69FA" pageId="6" pageNumber="272" type="diagnosis">
<paragraph id="45153689FFD5FF8B158AF8D81C3E69FA" blockId="6.[151,1437,1798,2041]" pageId="6" pageNumber="272">
<emphasis id="77DEEA9BFFD5FF8B158AF8D818636922" bold="true" box="[151,269,1798,1824]" pageId="6" pageNumber="272">Diagnosis.</emphasis>
Coloration of
<taxonomicName id="82AA4D0AFFD5FF8B14B9F8D91B646922" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[420,522,1798,1825]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="272" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
<emphasis id="77DEEA9BFFD5FF8B14B9F8D918D46922" box="[420,442,1799,1824]" italics="true" pageId="6" pageNumber="272">M</emphasis>
.
<emphasis id="77DEEA9BFFD5FF8B14D8F8D81B646922" box="[453,522,1798,1824]" italics="true" pageId="6" pageNumber="272">fisheri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CED57E0FFD5FF8B1712F8D91B0E6923" box="[527,608,1799,1825]" pageId="6" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
is identical to all other Malagasy species of the genus
<taxonomicName id="82AA4D0AFFD5FF8B1189F8D91D8C6922" authorityName="Weise" authorityYear="1898" box="[1172,1250,1799,1824]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="77DEEA9BFFD5FF8B1189F8D91D8C6922" box="[1172,1250,1799,1824]" italics="true" pageId="6" pageNumber="272">Merma</emphasis>
</taxonomicName>
. From
<taxonomicName id="82AA4D0AFFD5FF8B1036F8D91CF26922" baseAuthorityName="Sicard" baseAuthorityYear="1908" box="[1323,1436,1798,1825]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="272" phylum="Arthropoda" rank="species" species="hovana">
<emphasis id="77DEEA9BFFD5FF8B1036F8D91C2F6922" box="[1323,1345,1799,1824]" italics="true" pageId="6" pageNumber="272">M</emphasis>
.
<emphasis id="77DEEA9BFFD5FF8B1050F8D81CF26922" box="[1357,1436,1798,1824]" italics="true" pageId="6" pageNumber="272">hovana</emphasis>
</taxonomicName>
and
<taxonomicName id="82AA4D0AFFD5FF8B15D9F8F518156946" baseAuthorityName="Sicard" baseAuthorityYear="1909" box="[196,379,1834,1861]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="272" phylum="Arthropoda" rank="species" species="quadriguttata">
<emphasis id="77DEEA9BFFD5FF8B15D9F8F519B46946" box="[196,218,1835,1860]" italics="true" pageId="6" pageNumber="272">M</emphasis>
.
<emphasis id="77DEEA9BFFD5FF8B15FAF8F418156946" box="[231,379,1834,1860]" italics="true" pageId="6" pageNumber="272">quadriguttata</emphasis>
</taxonomicName>
it can be easily separated by its larger body size and tarsal claws with additional subquadrate basal tooth.
<taxonomicName id="82AA4D0AFFD5FF8B15C1F8911818696A" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[220,374,1870,1896]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="272" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
<emphasis id="77DEEA9BFFD5FF8B15C1F8911818696A" box="[220,374,1870,1896]" italics="true" pageId="6" pageNumber="272">Merma fisheri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CED57E0FFD5FF8B1460F89118BE696B" box="[381,464,1871,1897]" pageId="6" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
has narrowly explanate but visible lateral elytral margins similar to
<taxonomicName id="82AA4D0AFFD5FF8B11ADF8911C55696A" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[1200,1339,1870,1896]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="272" phylum="Arthropoda" rank="species" species="ankarana" status="sp. nov.">
<emphasis id="77DEEA9BFFD5FF8B11ADF8911C55696A" box="[1200,1339,1870,1896]" italics="true" pageId="6" pageNumber="272">M. ankarana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CED57E0FFD5FF8B105EF8911CF6696B" box="[1347,1432,1871,1897]" pageId="6" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
, while in
<taxonomicName id="82AA4D0AFFD5FF8B15E5F8AD18FA698E" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[248,404,1906,1932]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="272" phylum="Arthropoda" rank="species" species="antakotako" status="sp. nov.">
<emphasis id="77DEEA9BFFD5FF8B15E5F8AD18FA698E" box="[248,404,1906,1932]" italics="true" pageId="6" pageNumber="272">M. antakotako</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CED57E0FFD5FF8B1481F8AD189E698F" box="[412,496,1907,1933]" pageId="6" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
lateral elytral margins are distinctly explanate. Moreover males have broadly rounded apical margin of ventrite 5 while in
<taxonomicName id="82AA4D0AFFD5FF8B1704F8491BDA69B2" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[537,692,1942,1968]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="272" phylum="Arthropoda" rank="species" species="antakotako" status="sp. nov.">
<emphasis id="77DEEA9BFFD5FF8B1704F8491BDA69B2" box="[537,692,1942,1968]" italics="true" pageId="6" pageNumber="272">M. antakotako</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CED57E0FFD5FF8B17A6F8491A6169B3" box="[699,783,1943,1969]" pageId="6" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="82AA4D0AFFD5FF8B165FF8491AA369B2" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[834,973,1942,1968]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="272" phylum="Arthropoda" rank="species" species="ankarana" status="sp. nov.">
<emphasis id="77DEEA9BFFD5FF8B165FF8491AA369B2" box="[834,973,1942,1968]" italics="true" pageId="6" pageNumber="272">M. ankarana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CED57E0FFD5FF8B16C9F8491D4669B3" box="[980,1064,1943,1969]" pageId="6" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
it is emarginate. The most reliable character to separate that species are structures of the male genitalia: penis guide in lateral view with inner margin incised in middle; penis subparallel and broadened apically, with two rows of small denticles, without lateral projection.
</paragraph>
</subSubSection>
<caption id="11D56601FFD4FF8A158AFA2E1B646876" ID-DOI="http://doi.org/10.5281/zenodo.8058825" ID-Zenodo-Dep="8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="7" pageNumber="273" startId="7.[151,250,1520,1544]" targetBox="[151,1436,181,1496]" targetPageId="7" targetType="figure">
<paragraph id="45153689FFD4FF8A158AFA2E1B646876" blockId="7.[151,1436,1520,1653]" pageId="7" pageNumber="273">
<emphasis id="77DEEA9BFFD4FF8A158AFA2E1878680A" bold="true" box="[151,278,1520,1544]" pageId="7" pageNumber="273">FIGURE 5.</emphasis>
<taxonomicName id="82AA4D0AFFD4FF8A1402FA2F18DD680A" authority="Szawaryn &amp; Czerwiński, 2023" authorityName="Szawaryn &amp; Czerwiński" authorityYear="2023" box="[287,435,1520,1544]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="273" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
<emphasis id="77DEEA9BFFD4FF8A1402FA2F18DD680A" box="[287,435,1520,1544]" italics="true" pageId="7" pageNumber="273">Merma fisheri</emphasis>
</taxonomicName>
<emphasis id="77DEEA9BFFD4FF8A14A1FA2E1B61680A" bold="true" box="[444,527,1520,1544]" pageId="7" pageNumber="273">
<taxonomicNameLabel id="6CED57E0FFD4FF8A14A1FA2E1B61680A" box="[444,527,1520,1544]" pageId="7" pageNumber="273" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="77DEEA9BFFD4FF8A1702FA2E1B5F680A" bold="true" box="[543,561,1520,1544]" pageId="7" pageNumber="273">A</emphasis>
,
<emphasis id="77DEEA9BFFD4FF8A175CFA2E1B3D680A" bold="true" box="[577,595,1520,1544]" pageId="7" pageNumber="273">C</emphasis>
,
<emphasis id="77DEEA9BFFD4FF8A177EFA2E1B1A680A" bold="true" box="[611,628,1520,1544]" pageId="7" pageNumber="273">E</emphasis>
,
<emphasis id="77DEEA9BFFD4FF8A1799FA2E1BC3680A" bold="true" box="[644,685,1520,1544]" pageId="7" pageNumber="273">G–I</emphasis>
,
<emphasis id="77DEEA9BFFD4FF8A17A3FA2E1B98680A" bold="true" box="[702,758,1520,1544]" pageId="7" pageNumber="273">K–M</emphasis>
: male, holotype (CAS);
<emphasis id="77DEEA9BFFD4FF8A16E6FA2E1D62680A" bold="true" box="[1019,1036,1520,1544]" pageId="7" pageNumber="273">B</emphasis>
,
<emphasis id="77DEEA9BFFD4FF8A1101FA2E1D40680A" bold="true" box="[1052,1070,1520,1544]" pageId="7" pageNumber="273">D</emphasis>
,
<emphasis id="77DEEA9BFFD4FF8A1123FA2E1D23680A" bold="true" box="[1086,1101,1520,1544]" pageId="7" pageNumber="273">F</emphasis>
,
<emphasis id="77DEEA9BFFD4FF8A1140FA2E1D04680A" bold="true" box="[1117,1130,1520,1544]" pageId="7" pageNumber="273">J</emphasis>
: female, paratype (CAS):
<emphasis id="77DEEA9BFFD4FF8A109CFA2E1CFD680A" bold="true" box="[1409,1427,1520,1544]" pageId="7" pageNumber="273">A</emphasis>
) abdomen, male;
<emphasis id="77DEEA9BFFD4FF8A1459F9CA183B682E" bold="true" box="[324,341,1556,1580]" pageId="7" pageNumber="273">B</emphasis>
) abdomen, female;
<emphasis id="77DEEA9BFFD4FF8A173BF9CA1B56682E" bold="true" box="[550,568,1556,1580]" pageId="7" pageNumber="273">C</emphasis>
) tergite VIII, male;
<emphasis id="77DEEA9BFFD4FF8A1610F9CA1A71682E" bold="true" box="[781,799,1556,1580]" pageId="7" pageNumber="273">D</emphasis>
) tergite VIII, female;
<emphasis id="77DEEA9BFFD4FF8A111AF9CA1D76682E" bold="true" box="[1031,1048,1556,1580]" pageId="7" pageNumber="273">E</emphasis>
) sternite VIII, male;
<emphasis id="77DEEA9BFFD4FF8A11EAF9CA1C68682E" bold="true" box="[1271,1286,1556,1580]" pageId="7" pageNumber="273">F</emphasis>
) sternite VIII, female;
<emphasis id="77DEEA9BFFD4FF8A15F7F9E619906852" bold="true" box="[234,254,1592,1616]" pageId="7" pageNumber="273">G</emphasis>
) penis, lateral;
<emphasis id="77DEEA9BFFD4FF8A1481F9E618DE6852" bold="true" box="[412,432,1592,1616]" pageId="7" pageNumber="273">H</emphasis>
) penis tip, lateral;
<emphasis id="77DEEA9BFFD4FF8A176DF9E61B146852" bold="true" box="[624,634,1592,1616]" pageId="7" pageNumber="273">I</emphasis>
) penis tip, inner;
<emphasis id="77DEEA9BFFD4FF8A1632F9E61A526852" bold="true" box="[815,828,1592,1616]" pageId="7" pageNumber="273">J</emphasis>
) female genitalia;
<emphasis id="77DEEA9BFFD4FF8A16E6F9E61D616852" bold="true" box="[1019,1039,1592,1616]" pageId="7" pageNumber="273">K</emphasis>
) tegmen, inner;
<emphasis id="77DEEA9BFFD4FF8A11AAF9E61DA66852" bold="true" box="[1207,1224,1592,1616]" pageId="7" pageNumber="273">L</emphasis>
) tegmen, lateral;
<emphasis id="77DEEA9BFFD4FF8A1061F9E61CFA6852" bold="true" box="[1404,1428,1592,1616]" pageId="7" pageNumber="273">M</emphasis>
) male abdominal segments IX and X.
</paragraph>
</caption>
<subSubSection id="0DB06502FFD4FF8A15DAF97C189468E2" pageId="7" pageNumber="273" type="etymology">
<paragraph id="45153689FFD4FF8A15DAF97C189468E2" blockId="7.[151,1436,1698,2013]" pageId="7" pageNumber="273">
<emphasis id="77DEEA9BFFD4FF8A15DAF97C182068BE" bold="true" box="[199,334,1698,1724]" pageId="7" pageNumber="273">Etymology.</emphasis>
The new species is named after Bryan L. Fisher, leader of the CAS
<collectingCountry id="3DBD7619FFD4FF8A1159F97C1DA568BE" box="[1092,1227,1698,1724]" name="Madagascar" pageId="7" pageNumber="273">Madagascar</collectingCountry>
project and one of the collectors of the
<typeStatus id="9A11882BFFD4FF8A1461F91918C268E3" box="[380,428,1735,1761]" pageId="7" pageNumber="273">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="0DB06502FFD4FF8415DAF9341B416CF3" lastPageId="9" lastPageNumber="275" pageId="7" pageNumber="273" type="description">
<paragraph id="45153689FFD4FF8A15DAF9341CE56972" blockId="7.[151,1436,1698,2013]" pageId="7" pageNumber="273">
<emphasis id="77DEEA9BFFD4FF8A15DAF93418376906" bold="true" box="[199,345,1770,1796]" pageId="7" pageNumber="273">Description.</emphasis>
Length
<quantity id="82529B6CFFD4FF8A14A9F9341B526907" box="[436,572,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="6.2" metricValueMin="5.1" pageId="7" pageNumber="273" unit="mm" value="5.65" valueMax="6.2" valueMin="5.1">5.1–6.2 mm</quantity>
, width
<quantity id="82529B6CFFD4FF8A1791F9341A7B6907" box="[652,789,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="7" pageNumber="273" unit="mm" value="4.6" valueMax="4.8" valueMin="4.4">4.4–4.8 mm</quantity>
; length/width ratio = 1.17–1.20. Body (
<figureCitation id="DD912A0CFFD4FF8A11D4F9341C596906" box="[1225,1335,1770,1796]" captionStart="FIGURE 1" captionStartId="2.[151,250,1512,1536]" captionTargetBox="[175,1428,187,1482]" captionTargetId="figure-17@2.[153,1434,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Merma species. A–C) M. ankarana sp. nov., male, holotype (MNHN); A) dorsal; B) lateral; C) frontal; D–F) M. antakotako sp. nov., male, paratype (MNHN); D) dorsal; E) lateral; F) frontal; G–I) M. fisheri sp. nov., male, holotype (CAS); G) dorsal; H) lateral; I) frontal." figureDoi="http://doi.org/10.5281/zenodo.8058813" httpUri="https://zenodo.org/record/8058813/files/figure.png" pageId="7" pageNumber="273">Fig. 1G–I</figureCitation>
) roundly oval, strongly convex, dorsum pubescent. Head, mouthparts, legs and abdomen reddish-orange. Pronotum chestnut brown with anterior corners and lateral margins pale orange. Elytra chestnut brown, with two dark yellow or orange maculae on each elytron, anterior macula placed below humeral callus, posterior macula in apical part of elytron.
</paragraph>
<paragraph id="45153689FFD4FF8515DAF8A418D76F43" blockId="7.[151,1436,1698,2013]" lastBlockId="8.[151,1436,150,321]" lastPageId="8" lastPageNumber="274" pageId="7" pageNumber="273">Head transverse, partially concealed in prothorax. Interocular distance about 0.6 times of head width. Inner eye orbits slightly emarginate antero-medially, closest in the middle. Antenna composed of 11 antennomeres, about as long as 0.6 head width; scape large, elongate, slightly swollen; pedicel distinctly narrower than scape, elongate; antennomere 3 elongate, 1.3 times longer than pedicel; antennomeres 4–7 elongate, about 1.5 times as long as its width; antennomere 8 as long as wide; antennomeres 9–11 forming a distinct asymmetrical club. Ventral antennal grooves distinct in anterior part, moderately long and straight, along inner margin of eye. Clypeus short, parallel-sided, its anterior margin emarginate, smooth without groove. Labrum transverse, short, anterior margin slightly emarginate, membranous.</paragraph>
<caption id="11D56601FFDBFF85158AFA661B596BD2" ID-DOI="http://doi.org/10.5281/zenodo.8063852" ID-Zenodo-Dep="8063852" box="[151,567,1464,1489]" httpUri="https://zenodo.org/record/8063852/files/figure.png" pageId="8" pageNumber="274" startId="8.[151,250,1464,1488]" targetBox="[427,1161,351,1435]" targetPageId="8" targetType="figure">
<paragraph id="45153689FFDBFF85158AFA661B596BD2" blockId="8.[151,567,1464,1489]" box="[151,567,1464,1489]" pageId="8" pageNumber="274">
<emphasis id="77DEEA9BFFDBFF85158AFA66187D6BD3" bold="true" box="[151,275,1464,1489]" pageId="8" pageNumber="274">FIGURE 6.</emphasis>
Distribution map of
<taxonomicName id="82AA4D0AFFDBFF8514F5FA671B5F6BD2" authorityName="Weise" authorityYear="1898" box="[488,561,1465,1488]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="77DEEA9BFFDBFF8514F5FA671B5F6BD2" box="[488,561,1465,1488]" italics="true" pageId="8" pageNumber="274">Merma</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="45153689FFDBFF8515DAFA211A986887" blockId="8.[151,1437,1535,2029]" pageId="8" pageNumber="274">Prothorax. Hypomeron smooth, simply / finely punctate. Prosternal process smooth, without carinae, bordered laterally. Prosternum in front of coxa about as long as coxal longitudinal diameter; anterior margin, uniformly arcuate, bordered; posterior margin with bordering line starting from the base of prosternal process, laterally reaching notosternal suture. Surface of prosternum covered with sparse small punctae.</paragraph>
<paragraph id="45153689FFDBFF8515DAF9511C28695F" blockId="8.[151,1437,1535,2029]" pageId="8" pageNumber="274">Pterothorax.Mesoventral process covered with small tubercles along lateral margins, about as wide as mesocoxal diameter; meso-metaventral suture straight. Elytra with punctate of two sizes; lateral margins weakly explanate, clearly visible only in anterior half of its length from above. Epipleuron complete, broad, with distinct depressions to accommodate mid and hind legs in repose, inner margin with broad bordering line present in median and apical part. Mesoventrite covered with sparse and small punctures, with just a few larger in median part. Metaventral postcoxal lines joined on metaventral process forming straight line, recurved and then descending laterally.</paragraph>
<paragraph id="45153689FFDBFF8515DAF8B91C3769EF" blockId="8.[151,1437,1535,2029]" pageId="8" pageNumber="274">Legs short and stout with apices of mid and hind femora not protruding from outer margin of elytral epipleuron. Trochanters angulately produced, with bordering line on their outer surfaces, and cavities on their inner surfaces for receiving tip of tibiae in repose. Protibiae slightly widened, all tibiae on outer edge with oblique carina near apex, without distinct apical spurs. Tarsal claws double with additional subquadrate basal tooth, empodium absent.</paragraph>
<paragraph id="45153689FFDAFF8415DAFF491BB16F67" blockId="9.[151,1437,150,1005]" pageId="9" pageNumber="275">
Abdomen with five ventrites, with ventrite 5 rounded posteriorly in both sexes (
<figureCitation id="DD912A0CFFDAFF84115AFF491DAF6EB3" box="[1095,1217,150,177]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5A, B</figureCitation>
), in female ventrite 5 with transverse impression at base of ventrite. Abdominal postcoxal lines descending, sub-parallel to posterior margin of ventrite 1, recurved shortly laterally. In male apical margin of tergite VIII (
<figureCitation id="DD912A0CFFDAFF841169FF011DA26EFB" box="[1140,1228,222,249]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5C</figureCitation>
) and sternite VIII broadly rounded (
<figureCitation id="DD912A0CFFDAFF84147AFEDD18AC6F1F" box="[359,450,258,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5E</figureCitation>
); apodeme of sternum IX (
<figureCitation id="DD912A0CFFDAFF841619FEDD1A0A6F1F" box="[772,868,258,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5M</figureCitation>
) rod-like. Tergite X transverse, truncate apically (
<figureCitation id="DD912A0CFFDAFF841582FEF9186E6F43" box="[159,256,294,321]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5M</figureCitation>
). In female apical margin of tergite VIII rounded (
<figureCitation id="DD912A0CFFDAFF841621FEF91AF96F43" box="[828,919,294,321]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5D</figureCitation>
), sternite VIII (
<figureCitation id="DD912A0CFFDAFF84115AFEF91DF56F43" box="[1095,1179,294,321]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5F</figureCitation>
) shallowly emarginate medially and with broad, sclerotised lobe anteriorly.
</paragraph>
<paragraph id="45153689FFDAFF8415DAFEB118046C63" blockId="9.[151,1437,150,1005]" pageId="9" pageNumber="275">
Male genitalia (
<figureCitation id="DD912A0CFFDAFF841466FEB11B5E6F8B" box="[379,560,366,393]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5G–I, K, L</figureCitation>
). Tegminal basal piece with a pair of indistinct spines near base of tegminal strut (
<figureCitation id="DD912A0CFFDAFF8415C6FE4D18596FAF" box="[219,311,402,429]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5K</figureCitation>
, arrows); penis guide slightly asymmetrical in inner view, as long as parameres, distal 1/3 of his length strongly tapering into apex; in lateral view (
<figureCitation id="DD912A0CFFDAFF8417C2FE691A5A6FD3" box="[735,820,438,465]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5L</figureCitation>
) outer edge straight with curved tip and with densely setose field in the median part, inner edge smooth. Parameres well developed, rounded apically. Tegminal strut distinctly shorter than penis guide, stout, expanded distally. Penis (
<figureCitation id="DD912A0CFFDAFF841689FE211A836C1B" box="[916,1005,510,537]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5G</figureCitation>
) rod-like, sinuate; penis tip (
<figureCitation id="DD912A0CFFDAFF841027FE211CFA6C1B" box="[1338,1428,510,537]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5H</figureCitation>
) asymmetrical, flattened, in inner view broadened (
<figureCitation id="DD912A0CFFDAFF8417C6FDFD1A456C3F" box="[731,811,546,573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5I</figureCitation>
), with two rows of denticles; penis capsule with arms weakly developed.
</paragraph>
<paragraph id="45153689FFDAFF8415DAFDB51B416CF3" blockId="9.[151,1437,150,1005]" pageId="9" pageNumber="275">
Female genitalia (
<figureCitation id="DD912A0CFFDAFF84148EFDB5188A6C87" box="[403,484,618,645]" captionStart="FIGURE 5" captionStartId="7.[151,250,1520,1544]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-34@7.[151,1436,181,1496]" captionTargetPageId="7" captionText="FIGURE 5. Merma fisheri sp. nov., A, C, E, G–I, K–M: male, holotype (CAS); B, D, F, J: female, paratype (CAS): A) abdomen, male; B) abdomen, female; C) tergite VIII, male; D) tergite VIII, female; E) sternite VIII, male; F) sternite VIII, female; G) penis, lateral; H) penis tip, lateral; I) penis tip, inner; J) female genitalia; K) tegmen, inner; L) tegmen, lateral; M) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.8058825" httpUri="https://zenodo.org/record/8058825/files/figure.png" pageId="9" pageNumber="275">Fig. 5J</figureCitation>
). Coxites rectangular, setose. Styli well developed, elongate, with long apical setae. Bursa copulatrix at base partially sclerotised, simple, not divided, ending with common oviduct. Sperm duct short, originating in the middle of the bursa, dorsally on bursa. Spermatheca lightly sclerotized, simple, vermiform, accessory gland short, membranous.
</paragraph>
</subSubSection>
<subSubSection id="0DB06502FFDAFF8415DAFD241B3E6D17" box="[199,592,762,789]" pageId="9" pageNumber="275" type="distribution">
<paragraph id="45153689FFDAFF8415DAFD241B3E6D17" blockId="9.[151,1437,150,1005]" box="[199,592,762,789]" pageId="9" pageNumber="275">
<emphasis id="77DEEA9BFFDAFF8415DAFD24180F6D16" bold="true" box="[199,353,762,788]" pageId="9" pageNumber="275">Distribution.</emphasis>
<collectingCountry id="3DBD7619FFDAFF84147AFD2518806D17" box="[359,494,763,789]" name="Madagascar" pageId="9" pageNumber="275">Madagascar</collectingCountry>
(
<figureCitation id="DD912A0CFFDAFF8414E3FD251B2C6D17" box="[510,578,763,789]" captionStart="FIGURE 6" captionStartId="8.[151,250,1464,1488]" captionTargetBox="[427,1161,351,1435]" captionTargetId="figure-85@8.[420,1166,345,1441]" captionTargetPageId="8" captionText="FIGURE 6. Distribution map of Merma." figureDoi="http://doi.org/10.5281/zenodo.8063852" httpUri="https://zenodo.org/record/8063852/files/figure.png" pageId="9" pageNumber="275">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0DB06502FFDAFF8415DAFCC11D326DEF" pageId="9" pageNumber="275" type="materials_examined">
<paragraph id="45153689FFDAFF8415DAFCC11D326DEF" blockId="9.[151,1437,150,1005]" pageId="9" pageNumber="275">
<emphasis id="77DEEA9BFFDAFF8415DAFCC118146D3A" bold="true" box="[199,378,798,825]" pageId="9" pageNumber="275">Type material.</emphasis>
<materialsCitation id="F5C23CD4FFDAFF84149BFCC11A136D83" ID-GBIF-Occurrence="4133181308" collectingDate="2001-02-28" collectionCode="CAS" country="Madagascar" elevation="90" location="Foret d'Orangea" pageId="9" pageNumber="275" specimenCode="CAS 3199" specimenCount="1" typeStatus="holotype">
<emphasis id="77DEEA9BFFDAFF84149BFCC118926D3B" bold="true" box="[390,508,799,825]" pageId="9" pageNumber="275">
<typeStatus id="9A11882BFFDAFF84149BFCC118996D3B" box="[390,503,799,825]" pageId="9" pageNumber="275">Holotype</typeStatus>
:
</emphasis>
male,
<collectingCountry id="3DBD7619FFDAFF84174CFCC11BB46D3B" box="[593,730,799,825]" name="Madagascar" pageId="9" pageNumber="275">Madagascar</collectingCountry>
,
<location id="40756052FFDAFF8417F6FCC11AC96D3B" LSID="urn:lsid:plazi:treatment:CD03879FFFD5FF84158AF9A21D326DEF:40756052FFDAFF8417F6FCC11AC96D3B" box="[747,935,799,825]" country="Madagascar" name="Foret d'Orangea" pageId="9" pageNumber="275">Forêt d'Orangea</location>
,
<locationDeviation id="54A72FABFFDAFF8416AAFCC11D8C6D3B" box="[951,1250,799,825]" location="Remena" pageId="9" pageNumber="275">
<quantity id="82529B6CFFDAFF8416AAFCC11D646D3B" box="[951,1034,799,825]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="275" unit="km" value="3.6">3.6 km</quantity>
128° SE Remena
</locationDeviation>
, 22-
<date id="31141049FFDAFF84100BFCC11CF76D3B" box="[1302,1433,799,825]" pageId="9" pageNumber="275" value="2001-02-28">
<collectingDate id="2150E9A1FFDAFF84100BFCC11CF76D3B" box="[1302,1433,799,825]" pageId="9" pageNumber="275" value="2001-02-28">28.02.2001</collectingDate>
</date>
, lat. -12.25889 long. 49.37467,
<quantity id="82529B6CFFDAFF8414EBFC9D1B5E6D5F" box="[502,560,835,861]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="275" unit="m" value="90.0">
<elevation id="CE87D1BAFFDAFF8414EBFC9D1B5E6D5F" box="[502,560,835,861]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="275" unit="m" value="90.0">90 m</elevation>
</quantity>
(a.s.l.), litoral rainforest,
<collectingMethod id="9CEB4E9EFFDAFF841649FC9D1A8D6D5F" box="[852,995,835,861]" pageId="9" pageNumber="275">Malaise trap</collectingMethod>
,
<collectorName id="E85F535FFFDAFF8416EDFC9D1D1C6D5F" box="[1008,1138,835,861]" pageId="9" pageNumber="275">B.L. Fisher</collectorName>
,
<collectorName id="E85F535FFFDAFF841163FC9D1C4D6D5F" box="[1150,1315,835,861]" pageId="9" pageNumber="275">C.E. Griswold</collectorName>
,
<collectorName id="E85F535FFFDAFF841032FC9D18366D83" pageId="9" pageNumber="275">Malagasy Arthropod Team</collectorName>
, collection code BLF03199,
<specimenCode id="150C9EF2FFDAFF8417B1FCB91A486D83" box="[684,806,871,897]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="275" type="Museum">CAS 3199</specimenCode>
(
<collectionCode id="23BBAE4CFFDAFF841627FCB91A186D83" box="[826,886,871,897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="275" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F5C23CD4FFDAFF841690FCB91BFB6DCB" ID-GBIF-Occurrence="4133181307" collectingDate="2003-07-12" collectionCode="CAS" collectorName="de Bekaraoka &amp; B. L. Fisher" country="Madagascar" elevation="150" location="Foret de Bekaraoka" pageId="9" pageNumber="275" specimenCount="1" typeStatus="paratype">
<emphasis id="77DEEA9BFFDAFF841690FCB91D606D83" bold="true" box="[909,1038,871,897]" pageId="9" pageNumber="275">
<typeStatus id="9A11882BFFDAFF841690FCB91D676D83" box="[909,1033,871,897]" pageId="9" pageNumber="275">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="3DBD7619FFDAFF841104FCB91DCC6D83" box="[1049,1186,871,897]" name="Madagascar" pageId="9" pageNumber="275">Madagascar</collectingCountry>
,
<location id="40756052FFDAFF8411ACFCB91CF66D83" LSID="urn:lsid:plazi:treatment:CD03879FFFD5FF84158AF9A21D326DEF:40756052FFDAFF8411ACFCB91CF66D83" box="[1201,1432,871,897]" country="Madagascar" name="Foret de Bekaraoka" pageId="9" pageNumber="275">
Forêt
<collectorName id="E85F535FFFDAFF8411E5FCB91CF66D83" box="[1272,1432,871,897]" pageId="9" pageNumber="275">de Bekaraoka</collectorName>
</location>
,
<locationDeviation id="54A72FABFFDAFF84158AFC5518DC6DA7" box="[151,434,907,933]" location="Daraina" pageId="9" pageNumber="275">
<quantity id="82529B6CFFDAFF84158AFC5519B06DA7" box="[151,222,907,933]" metricMagnitude="3" metricUnit="m" metricValue="6.8" pageId="9" pageNumber="275" unit="km" value="6.8">6.8km</quantity>
60° ENE Daraina
</locationDeviation>
,
<date id="31141049FFDAFF8414A2FC551B516DA7" box="[447,575,907,933]" pageId="9" pageNumber="275" value="2003-07-12">
<collectingDate id="2150E9A1FFDAFF8414A2FC551B516DA7" box="[447,575,907,933]" pageId="9" pageNumber="275" value="2003-07-12">07.12.2003</collectingDate>
</date>
, lat. -13.16667 long. 49.71,
<quantity id="82529B6CFFDAFF841699FC541AA36DA7" box="[900,973,906,933]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="275" unit="m" value="150.0">
<elevation id="CE87D1BAFFDAFF841699FC541AA36DA7" box="[900,973,906,933]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="275" unit="m" value="150.0">150 m</elevation>
</quantity>
(a.s.l.), tropical dry forest,
<collectingMethod id="9CEB4E9EFFDAFF841015FC551CF66DA7" box="[1288,1432,907,933]" pageId="9" pageNumber="275">Malaise trap</collectingMethod>
,
<collectorName id="E85F535FFFDAFF84158AFC7118776DCB" box="[151,281,943,969]" pageId="9" pageNumber="275">B.L. Fisher</collectorName>
, collection code BLF09874,
<collectionCode id="23BBAE4CFFDAFF841742FC711BFB6DCB" box="[607,661,943,969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="275" type="Museum">CAS</collectionCode>
</materialsCitation>
<materialsCitation id="F5C23CD4FFDAFF841780FC711AD76DCB" ID-GBIF-Occurrence="4133181309" box="[669,953,943,969]" collectionCode="CAS" country="Madagascar" pageId="9" pageNumber="275" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
9874 (
<specimenCount id="53ACFD00FFDAFF8417F5FC711A256DCB" box="[744,843,943,969]" pageId="9" pageNumber="275" type="female">1 female</specimenCount>
,
<collectionCode id="23BBAE4CFFDAFF84164AFC711AFC6DCB" box="[855,914,943,969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="275" type="Museum">CAS</collectionCode>
). N
</materialsCitation>
<materialsCitation id="F5C23CD4FFDAFF8416DCFC711D366DEF" ID-GBIF-Occurrence="4133181301" collectingDate="2007-01-06" collectionCode="JVC" collectorName="Ankarana N. P. &amp; P., F. Pavel" country="Madagascar" latitude="-12.969083" location="Mahamatsina" longLatPrecision="1" longitude="49.136196" pageId="9" pageNumber="275" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="3DBD7619FFDAFF8416DCFC711D246DCB" box="[961,1098,943,969]" name="Madagascar" pageId="9" pageNumber="275">Madagascar</collectingCountry>
,
<location id="40756052FFDAFF84114BFC711D836DCB" LSID="urn:lsid:plazi:treatment:CD03879FFFD5FF84158AF9A21D326DEF:40756052FFDAFF84114BFC711D836DCB" box="[1110,1261,943,969]" country="Madagascar" latitude="-12.969083" longLatPrecision="1" longitude="49.136196" name="Mahamatsina" pageId="9" pageNumber="275">Mahamatsina</location>
env.,
<collectorName id="E85F535FFFDAFF841032FC7119A66DEF" pageId="9" pageNumber="275">Ankarana N.P.</collectorName>
, F. Pavel leg, 5-
<date id="31141049FFDAFF841465FC0C18826DEF" box="[376,492,978,1005]" pageId="9" pageNumber="275" value="2007-01-06">
<collectingDate id="2150E9A1FFDAFF841465FC0C18826DEF" box="[376,492,978,1005]" pageId="9" pageNumber="275" value="2007-01-06">6.01.2007</collectingDate>
</date>
<geoCoordinate id="209E504EFFDAFF8414EEFC0D1BFB6DEF" box="[499,661,978,1005]" degrees="12" direction="south" minutes="58" orientation="latitude" pageId="9" pageNumber="275" precision="1" seconds="08.7" value="-12.969083">S 12°58’08.7’’</geoCoordinate>
<geoCoordinate id="209E504EFFDAFF841780FC0D1A2E6DEF" box="[669,832,979,1005]" degrees="49" direction="east" minutes="08" orientation="longitude" pageId="9" pageNumber="275" precision="1" seconds="10.3" value="49.136196">E 49°08’10.3’’</geoCoordinate>
(
<specimenCount id="53ACFD00FFDAFF84164EFC0D1AF16DEF" box="[851,927,979,1005]" pageId="9" pageNumber="275" type="male">1 male</specimenCount>
,
<specimenCount id="53ACFD00FFDAFF8416B7FC0D1D606DEF" box="[938,1038,979,1005]" pageId="9" pageNumber="275" type="female">1 female</specimenCount>
,
<collectionCode id="23BBAE4CFFDAFF841107FC0D1D3E6DEF" box="[1050,1104,979,1005]" pageId="9" pageNumber="275">JVC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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