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96 KiB
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442 lines
96 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1151.97514" ID-Pensoft-Pub="1313-2970-1151-67" ID-Pensoft-UUID="0B3CEF4A954955478878AD15C022DCD5" ID-ZooBank="5FEDE55DC9AF47D591259F1758AE2A18" ModsDocID="1313-2970-1151-67" checkinTime="1677749880156" checkinUser="pensoft" docAuthor="Willemse, Luc, Tilmans, Jos, Kotitsa, Nefeli, Trichas, Apostolos, Heller, Klaus-Gerhard, Chobanov, Dragan & Ode, Baudewijn" docDate="2023" docId="1E2F5C7D3A9852D286E7ED67E87F7BEB" docLanguage="en" docName="ZooKeys 1151: 67-158" docOrigin="ZooKeys 1151" docPubDate="2023-03-01" docSource="http://dx.doi.org/10.3897/zookeys.1151.97514" docTitle="Eupholidoptera latens Willemse & Kruseman 1976" docType="treatment" docVersion="2" id="0B3CEF4A954955478878AD15C022DCD5" lastPageNumber="67" masterDocId="0B3CEF4A954955478878AD15C022DCD5" masterDocTitle="A review of Eupholidoptera (Orthoptera, Tettigoniidae) from Crete, Gavdos, Gavdopoula, and Andikithira" masterLastPageNumber="158" masterPageNumber="67" pageNumber="67" updateTime="1677750187672" updateUser="ExternalLinkService">
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<mods:title>A review of Eupholidoptera (Orthoptera, Tettigoniidae) from Crete, Gavdos, Gavdopoula, and Andikithira</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Willemse, Luc</mods:namePart>
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<mods:affiliation>Naturalis, PO Box 9517, 2300, RA Leiden, Netherlands</mods:affiliation>
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<mods:nameIdentifier type="email">luc.willemse@naturalis.nl</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tilmans, Jos</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0127-6191</mods:nameIdentifier>
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<mods:affiliation>Herkenboscher Strasse 33, 41849, Wassenberg-Rothenbach, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kotitsa, Nefeli</mods:namePart>
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<mods:affiliation>Natural History Museum Crete, University of Crete, P. O. Box 2208, 71409, Heraklion, Crete, Greece & Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel blvd., 1000, Sofia, Bulgaria</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Trichas, Apostolos</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7917-5262</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum Crete, University of Crete, P. O. Box 2208, 71409, Heraklion, Crete, Greece</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Heller, Klaus-Gerhard</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3331-3228</mods:nameIdentifier>
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<mods:affiliation>Triesdorf Bahnhof 8, 91732, Merkendorf, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Chobanov, Dragan</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1642-0363</mods:nameIdentifier>
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<mods:affiliation>Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel blvd., 1000, Sofia, Bulgaria</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ode, Baudewijn</mods:namePart>
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<mods:date>2023</mods:date>
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<mods:number>2023-03-01</mods:number>
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<mods:number>1151</mods:number>
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<mods:start>67</mods:start>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1151.97514</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:1E2F5C7D3A9852D286E7ED67E87F7BEB" httpUri="http://treatment.plazi.org/id/1E2F5C7D3A9852D286E7ED67E87F7BEB" lastPageNumber="67" pageId="0" pageNumber="67">
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<taxonomicName ID-CoL="3CMKN" LSID="1E2F5C7D-3A98-52D2-86E7-ED67E87F7BEB" authority="Willemse & Kruseman, 1976" authorityName="Willemse & Kruseman" authorityYear="1976" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera latens" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="latens">Eupholidoptera latens Willemse & Kruseman, 1976</taxonomicName>
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<figureCitation captionStart="Figures 11–24" captionStartId="F5" captionText="Figures 11 - 24. Colour pattern of head, frontal view 11 Eupholidoptera smyrnensis ♀ Makrigiannis RMNH 5014918 12 Eupholidoptera gemellata ♂ Mt. Idhi RMNH. INS 1141843 13 Eupholidoptera pallipes ♂ paratype Mt. Lefka RMNH. INS 1105313 14 Eupholidoptera cretica ♂ Mt. Lefka, Omalos RMNH. INS 114838 15 Eupholidoptera latens ♀ Mt. Lefka, Kalergi CT 1987.047.04 16 Eupholidoptera giuliae ♂ Argoules CT 1995.011.18 17 Eupholidoptera francisae sp. nov. ♀ allotype Andikithira CT 2002.004.11 18 Eupholidoptera annamariae ♀ Kato Zakros CT 1995.020.03 19 Eupholidoptera astyla ♀ Ano Viannos CT 1987.024.03 20 Eupholidoptera feri ♂ holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 21 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH. 5014906 22 Eupholidoptera forcipata ♂ Mt. Idhi CT 1987.044.03 23 Eupholidoptera marietheresae sp. nov. ♂ paratype Mt. Dikti, FC 1606 CT 2000.096.01 24 Eupholidoptera jacquelinae ♂ holotype Gavdos CT 2000.005.02. Scale bars: 5 mm." figureDoi="10.3897/zookeys.1151.97514.figures11-24" httpUri="https://binary.pensoft.net/fig/817935" pageId="0" pageNumber="67">Figs 15</figureCitation>
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<figureCitation captionStart="Figures 25–38" captionStartId="F6" captionText="Figures 25 - 38. Colour pattern of pronotum, dorsal view 25 Eupholidoptera smyrnensis ♂ Makrigiannis RMNH. INS 152909 26 Eupholidoptera gemellata ♂ Mt. Idhi, Amariou RMNH. INS 1141843 27 Eupholidoptera pallipes ♂ paratype Mt. Lefka RMNH. INS 1105313 28 Eupholidoptera cretica ♂ Mt. Lefka, Omalos RMNH. INS 1141838 29 Eupholidoptera latens ♂ Rhodopos CH 8236 30 Eupholidoptera giuliae Chora Sfakion ♂ CT 2000.014.10 31 Eupholidoptera francisae sp. nov. ♂ holotype Andikithira CT 2002.004.04 32 Eupholidoptera annamariae ♂ Kato Zakros CT 2000.030.04 33 Eupholidoptera astyla ♂ Krotos RMNH. INS 1141819 34 Eupholidoptera feri ♂ holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 35 Eupholidoptera mariannae ♀ Kalavros RMNH. 5086974 36 Eupholidoptera forcipata ♂ Mt. Idhi CT 1987.044.03 37 Eupholidoptera marietheresae sp. nov. ♂ holotype Mt. Dikti RMNH. INS 1141850 38 Eupholidoptera jacquelinae ♂ holotype Gavdos CT 2000.005.02. Scale bars: 5 mm." figureDoi="10.3897/zookeys.1151.97514.figures25-38" httpUri="https://binary.pensoft.net/fig/817936" pageId="0" pageNumber="67">, 29</figureCitation>
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<figureCitation captionStart="Figures 39–52" captionStartId="F7" captionText="Figures 39 - 52. Female subgenital plate in ventral view 39 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 40 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 41 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 42 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 43 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 44 Eupholidoptera giuliae Argoules CT 1995.011.03 45 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 46 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 47 Eupholidoptera astyla Krotos RMNH. INS 1141820 48 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 49 Eupholidoptera mariannae Kalavros RMNH 5014912 50 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 51 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 52 Eupholidoptera jacquelinae Gavdos allotype CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures39-52" httpUri="https://binary.pensoft.net/fig/817937" pageId="0" pageNumber="67">, 43</figureCitation>
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<figureCitation captionStart="Figures 53–66" captionStartId="F8" captionText="Figures 53 - 66. Female subgenital plate in lateral view 53 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 54 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 55 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 56 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 57 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 58 Eupholidoptera giuliae Argoules CT 1995.011.03 59 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 60 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 61 Eupholidoptera astyla Krotos RMNH. INS 1141820 62 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 63 Eupholidoptera mariannae Kalavros RMNH 5014912 64 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 65 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 66 Eupholidoptera jacquelinae allotype Gavdos CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures53-66" httpUri="https://binary.pensoft.net/fig/817938" pageId="0" pageNumber="67">, 57</figureCitation>
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<figureCitation captionStart="Figures 67, 68" captionStartId="F9" captionText="Figures 67, 68. Female subgenital plate in lateral view 67 Eupholidoptera francisae sp. nov. Anidhroi CT 2001.002.07 68 Eupholidoptera latens Prases DC-Ort 000512." figureDoi="10.3897/zookeys.1151.97514.figures67-68" httpUri="https://binary.pensoft.net/fig/817939" pageId="0" pageNumber="67">, 68</figureCitation>
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<figureCitation captionStart="Figures 69–82" captionStartId="F10" captionText="Figures 69 - 82. Male anal tergite in dorsal view 69 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 70 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 71 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 72 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 73 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 74 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 75 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 76 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 77 Eupholidoptera astyla Krotos RMNH. INS 1141819 78 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 79 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 80 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 81 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 82 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures69-82" httpUri="https://binary.pensoft.net/fig/817940" pageId="0" pageNumber="67">, 73</figureCitation>
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<figureCitation captionStart="Figures 83–96" captionStartId="F11" captionText="Figures 83 - 96. Male anal tergite in caudal view 83 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 84 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 85 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 86 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 87 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 88 Eupholidoptera giuliae Chora Sfakion CT 2000.014.01 89 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 90 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 91 Eupholidoptera astyla Krotos RMNH. INS 1141819 92 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 93 Eupholidoptera mariannae Prina FC 17798 RMNH. INS 1141840 94 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 95 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 96 Eupholidoptera jacquelinae paratype Gavdos NHMC 2001.004.12. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures83-96" httpUri="https://binary.pensoft.net/fig/817941" pageId="0" pageNumber="67">, 87</figureCitation>
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<figureCitation captionStart="Figures 97–110" captionStartId="F12" captionText="Figures 97 - 110. Male anal tergite in lateral view 97 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 98 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 99 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 100 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 101 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.01 102 Eupholidoptera giuliae Argoules CT 1995.011.11 103 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 104 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 105 Eupholidoptera astyla Ano Viannos CT 1995.024.01 106 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 107 Eupholidoptera mariannae Kalavros CT 2017.029.01 108 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 109 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 110 Eupholidoptera jacquelinae Gavdopoula CT 1996.019.01. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures97-110" httpUri="https://binary.pensoft.net/fig/817942" pageId="0" pageNumber="67">, 101</figureCitation>
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<figureCitation captionStart="Figures 111–125" captionStartId="F13" captionText="Figures 111 - 125. Male cercus in dorsal view 111 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 112 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 113 Eupholidoptera pallipes Mt. Lefka paratype RMNH. INS 1105314 114 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 115 Eupholidoptera latens Xiloskalo IBER DC-Ort 000558 116 Eupholidoptera giuliae Chora Sfakion CT 2000.014.02 117 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 118 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 119 Eupholidoptera annamariae Xerokampos, male with anomalous cercus, IBER DC-Ort 000565 120 Eupholidoptera astyla Ierapetra MfN s. n. 121 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 122 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 123 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 124 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 125 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures111-125" httpUri="https://binary.pensoft.net/fig/817943" pageId="0" pageNumber="67">, 115</figureCitation>
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<figureCitation captionStart="Figures 126–139" captionStartId="F14" captionText="Figures 126 - 139. Male cercus in lateral view 126 Eupholidoptera smyrnensis Makrigiannis RMNH. 5087053 127 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 128 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 129 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 130 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 131 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 132 Eupholidoptera francisae paratype sp. nov. Andikithira CT 2002.004.08 133 Eupholidoptera annamariae Kato Zakros CT 1995.020.02 134 Eupholidoptera astyla paratype Ierapetra MfN s. n. 135 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 136 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 137 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 138 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 139 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures126-139" httpUri="https://binary.pensoft.net/fig/817944" pageId="0" pageNumber="67">, 130</figureCitation>
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<figureCitation captionStart="Figures 140–153" captionStartId="F15" captionText="Figures 140 - 153. Male subgenital plate in ventral view 140 Eupholidoptera smyrnensis Lagkada RMNH 5106270 141 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 142 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 143 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 144 Eupholidoptera latens Mt. Lefka Kalergi CT 1987.047.02 145 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 146 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 147 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 148 Eupholidoptera astyla Krotos RMNH. INS 1141819 149 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 150 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 151 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 152 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 153 Eupholidoptera jacquelinae holotype Gavdos 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures140-153" httpUri="https://binary.pensoft.net/fig/817945" pageId="0" pageNumber="67">, 144</figureCitation>
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<figureCitation captionStart="Figures 154–167" captionStartId="F16" captionText="Figures 154 - 167. Male subgenital plate in lateral view 154 Eupholidoptera smyrnensis Lagkada RMNH. INS 152909 155 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 156 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 157 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 (RMNH. INS 1141838 158 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 159 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 160 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 161 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 162 Eupholidoptera astyla Krotos RMNH 5086992 163 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 164 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 165 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 166 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 167 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures154-167" httpUri="https://binary.pensoft.net/fig/817946" pageId="0" pageNumber="67">, 158</figureCitation>
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<figureCitation captionStart="Figures 168–181" captionStartId="F17" captionText="Figures 168 - 181. Male stylus in lateral view 168 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 169 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 170 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 171 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 172 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 173 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 174 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 175 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 176 Eupholidoptera astyla Krotos RMNH. INS 1141819 177 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 178 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 179 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 180 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 181 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures168-181" httpUri="https://binary.pensoft.net/fig/817947" pageId="0" pageNumber="67">, 172</figureCitation>
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<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">, 186</figureCitation>
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<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">, 187</figureCitation>
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<figureCitation captionStart="Figures 198–212" captionStartId="F19" captionText="Figures 198 - 212. Male titillator in lateral view 198 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 199 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 200 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 201 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 202 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 203 Eupholidoptera latens Rhodopos CH 8236 204 Eupholidoptera giuliae Argoules CT 1995.011.11 205 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 206 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 207 Eupholidoptera astyla Krotos RMNH. INS 1141819 208 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 209 Eupholidoptera mariannae Kalavros CT 2017.029.01 210 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 211 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 212 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures198-212" httpUri="https://binary.pensoft.net/fig/817949" pageId="0" pageNumber="67">, 202</figureCitation>
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<figureCitation captionStart="Figures 198–212" captionStartId="F19" captionText="Figures 198 - 212. Male titillator in lateral view 198 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 199 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 200 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 201 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 202 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 203 Eupholidoptera latens Rhodopos CH 8236 204 Eupholidoptera giuliae Argoules CT 1995.011.11 205 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 206 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 207 Eupholidoptera astyla Krotos RMNH. INS 1141819 208 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 209 Eupholidoptera mariannae Kalavros CT 2017.029.01 210 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 211 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 212 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures198-212" httpUri="https://binary.pensoft.net/fig/817949" pageId="0" pageNumber="67">, 203</figureCitation>
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<figureCitation captionStart="Figures 247–253" captionStartId="F27" captionText="Figures 247 - 253. Field images Eupholidoptera spp. Crete 247 Eupholidoptera giuliae ♂ Prasies RMNH 5087051 248 Eupholidoptera giuliae ♀ Rethymnon RMNH 5014921 249 Eupholidoptera latens ♂ Kolympari RMNH 553681 250 Eupholidoptera mariannae ♂ Kalavros RMNH 5014907 251 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH 5014906 252 Eupholidoptera pallipes ♀ below Pakhnes 253 Eupholidoptera smyrnensis ♂ Makrigiannis RMNH 5087053." figureDoi="10.3897/zookeys.1151.97514.figures247-253" httpUri="https://binary.pensoft.net/fig/817957" pageId="0" pageNumber="67">, 249</figureCitation>
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<figureCitation captionStart="Figures 254, 255" captionStartId="F28" captionText="Figures 254, 255. Distribution maps Eupholidoptera Crete and adjacent islands 254 lowland species 255 highland species." figureDoi="10.3897/zookeys.1151.97514.figures254-255" httpUri="https://binary.pensoft.net/fig/817958" pageId="0" pageNumber="67">, 254</figureCitation>
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<figureCitation captionStart="Figure 256" captionStartId="F29" captionText="Figure 256. Bayesian inference phylogenetic tree of Eupholidoptera of Crete, Andikithira and Gavdos, including two outgroups and congenerics from the Greek and Turkish mainlands, based on a 1684 bp concatenated alignment of the NADH 2 + ITS fragments. Node values at branches show node support with BI posterior probabilities. Different colours correspond to the different morphological species. See Suppl. material 4 for details of the specimens and Suppl. material 5 for details of NADH 2 and IT IS sequences." figureDoi="10.3897/zookeys.1151.97514.figure256" httpUri="https://binary.pensoft.net/fig/817959" pageId="0" pageNumber="67">, 256</figureCitation>
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<figureCitation captionStart="Figure 257" captionStartId="F30" captionText="Figure 257. The Eupholidoptera giuliae - Eupholidoptera latens - Eupholidoptera francisae complex: geographic distribution of five morphological entities in the Eupholidoptera giuliae - Eupholidoptera latens - Eupholidoptera francisae complex: (1) eastern Chania + western Rethimno (E. giuliae), (2) northeastern Chania including Akrotiri peninsula (atypical E. giuliae), (3) northern and central Chania (E. latens), (4) western and southwestern Chania (E. francisae), (5) Andikithira (E. francisae)." figureDoi="10.3897/zookeys.1151.97514.figure257" httpUri="https://binary.pensoft.net/fig/817960" pageId="0" pageNumber="67">, 257</figureCitation>
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<figureCitation captionStart="Figure 258" captionStartId="F31" captionText="Figure 258. Distribution results genetic analysis: geographic pattern of analysis of genetic data of Eupholidoptera populations in western Crete. yellow dots: E. francisae, purple dots: E. latens, blue dots: E. giuliae, light blue dots: atypical E. giuliae." figureDoi="10.3897/zookeys.1151.97514.figure258" httpUri="https://binary.pensoft.net/fig/817961" pageId="0" pageNumber="67">, 258</figureCitation>
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<figureCitation captionStart="Figure 259" captionStartId="F32" captionText="Figure 259. Mean syllable duration in recorded species from Crete (incl. Gavdos and Andikithira). See Suppl. material 3 for measurements per recorded specimen." figureDoi="10.3897/zookeys.1151.97514.figure259" httpUri="https://binary.pensoft.net/fig/817962" pageId="0" pageNumber="67">, 259</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="67" type="reference_group">
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<paragraph pageId="0" pageNumber="67">
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<taxonomicName authorityName="Willemse & Kruseman" authorityYear="1976" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera latens" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="latens">Eupholidoptera latens</taxonomicName>
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Willemse & Kruseman, 1976: 134.
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</paragraph>
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<paragraph pageId="0" pageNumber="67">
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<taxonomicName authorityName="Willemse & Kruseman" authorityYear="1976" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera latens" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="latens">Eupholidoptera latens</taxonomicName>
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Morphological description.
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<bibRefCitation author="Willemse, F" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="67" pagination="123 - 164" refId="B77" refString="Willemse, F, Kruseman, G, 1976. Orthopteroidea of Crete. Tijdschrift voor Entomologie 119: 123 - 164" title="Orthopteroidea of Crete." volume="119" year="1976">Willemse and Kruseman 1976</bibRefCitation>
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: 134, 135.
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</paragraph>
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<paragraph pageId="0" pageNumber="67">
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<taxonomicName authorityName="Willemse & Kruseman" authorityYear="1976" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera latens" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="latens">Eupholidoptera latens</taxonomicName>
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Bioacoustics.
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2156.1.1" author="Ciplak, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="1 - 77" refId="B12" refString="Ciplak, B, Heller, K-G, Willemse, F, 2009. Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song. Zootaxa 2156 (1): 1 - 77, DOI: https://doi.org/10.11646/zootaxa.2156.1.1" title="Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song." url="https://doi.org/10.11646/zootaxa.2156.1.1" volume="2156" year="2009">
|
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<normalizedToken originalValue="Çiplak">Ciplak</normalizedToken>
|
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et al. 2009
|
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</bibRefCitation>
|
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|
: 27, 51, 54, 55.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="67" type="examined specimens">
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<paragraph pageId="0" pageNumber="67">Examined specimens.</paragraph>
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<paragraph pageId="0" pageNumber="67">
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<emphasis bold="true" pageId="0" pageNumber="67">Holotype, allotype</emphasis>
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, 4 ♂, 3f (
|
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<emphasis bold="true" pageId="0" pageNumber="67">paratypes</emphasis>
|
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); 12 ♂, 7 ♀ (for details see Suppl. material 2).
|
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</paragraph>
|
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="67" type="diagnostics features">
|
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<paragraph pageId="0" pageNumber="67">Diagnostics features.</paragraph>
|
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<paragraph pageId="0" pageNumber="67">
|
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|
Frons (Fig.
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<figureCitation captionStart="Figures 11–24" captionStartId="F5" captionText="Figures 11 - 24. Colour pattern of head, frontal view 11 Eupholidoptera smyrnensis ♀ Makrigiannis RMNH 5014918 12 Eupholidoptera gemellata ♂ Mt. Idhi RMNH. INS 1141843 13 Eupholidoptera pallipes ♂ paratype Mt. Lefka RMNH. INS 1105313 14 Eupholidoptera cretica ♂ Mt. Lefka, Omalos RMNH. INS 114838 15 Eupholidoptera latens ♀ Mt. Lefka, Kalergi CT 1987.047.04 16 Eupholidoptera giuliae ♂ Argoules CT 1995.011.18 17 Eupholidoptera francisae sp. nov. ♀ allotype Andikithira CT 2002.004.11 18 Eupholidoptera annamariae ♀ Kato Zakros CT 1995.020.03 19 Eupholidoptera astyla ♀ Ano Viannos CT 1987.024.03 20 Eupholidoptera feri ♂ holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 21 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH. 5014906 22 Eupholidoptera forcipata ♂ Mt. Idhi CT 1987.044.03 23 Eupholidoptera marietheresae sp. nov. ♂ paratype Mt. Dikti, FC 1606 CT 2000.096.01 24 Eupholidoptera jacquelinae ♂ holotype Gavdos CT 2000.005.02. Scale bars: 5 mm." figureDoi="10.3897/zookeys.1151.97514.figures11-24" httpUri="https://binary.pensoft.net/fig/817935" pageId="0" pageNumber="67">15</figureCitation>
|
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) pale with black dots; pronotal disc (Fig.
|
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<figureCitation captionStart="Figures 25–38" captionStartId="F6" captionText="Figures 25 - 38. Colour pattern of pronotum, dorsal view 25 Eupholidoptera smyrnensis ♂ Makrigiannis RMNH. INS 152909 26 Eupholidoptera gemellata ♂ Mt. Idhi, Amariou RMNH. INS 1141843 27 Eupholidoptera pallipes ♂ paratype Mt. Lefka RMNH. INS 1105313 28 Eupholidoptera cretica ♂ Mt. Lefka, Omalos RMNH. INS 1141838 29 Eupholidoptera latens ♂ Rhodopos CH 8236 30 Eupholidoptera giuliae Chora Sfakion ♂ CT 2000.014.10 31 Eupholidoptera francisae sp. nov. ♂ holotype Andikithira CT 2002.004.04 32 Eupholidoptera annamariae ♂ Kato Zakros CT 2000.030.04 33 Eupholidoptera astyla ♂ Krotos RMNH. INS 1141819 34 Eupholidoptera feri ♂ holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 35 Eupholidoptera mariannae ♀ Kalavros RMNH. 5086974 36 Eupholidoptera forcipata ♂ Mt. Idhi CT 1987.044.03 37 Eupholidoptera marietheresae sp. nov. ♂ holotype Mt. Dikti RMNH. INS 1141850 38 Eupholidoptera jacquelinae ♂ holotype Gavdos CT 2000.005.02. Scale bars: 5 mm." figureDoi="10.3897/zookeys.1151.97514.figures25-38" httpUri="https://binary.pensoft.net/fig/817936" pageId="0" pageNumber="67">29</figureCitation>
|
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) rarely completely pale, mostly frontal half with more or less well defined black central patch, border with pale rear half. V-shaped. Male (Fig.
|
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<figureCitation captionStart="Figures 247–253" captionStartId="F27" captionText="Figures 247 - 253. Field images Eupholidoptera spp. Crete 247 Eupholidoptera giuliae ♂ Prasies RMNH 5087051 248 Eupholidoptera giuliae ♀ Rethymnon RMNH 5014921 249 Eupholidoptera latens ♂ Kolympari RMNH 553681 250 Eupholidoptera mariannae ♂ Kalavros RMNH 5014907 251 Eupholidoptera mariannae ♀ Ag. Ioannis RMNH 5014906 252 Eupholidoptera pallipes ♀ below Pakhnes 253 Eupholidoptera smyrnensis ♂ Makrigiannis RMNH 5087053." figureDoi="10.3897/zookeys.1151.97514.figures247-253" httpUri="https://binary.pensoft.net/fig/817957" pageId="0" pageNumber="67">249</figureCitation>
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) - stridulatory file with 108-123 teeth (including proximal and distal ones), density of teeth in middle two thirds of the file 26-31 teeth per mm; anal tergite (Figs
|
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<figureCitation captionStart="Figures 69–82" captionStartId="F10" captionText="Figures 69 - 82. Male anal tergite in dorsal view 69 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 70 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 71 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 72 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 73 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 74 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 75 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 76 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 77 Eupholidoptera astyla Krotos RMNH. INS 1141819 78 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 79 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 80 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 81 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 82 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures69-82" httpUri="https://binary.pensoft.net/fig/817940" pageId="0" pageNumber="67">73</figureCitation>
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,
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<figureCitation captionStart="Figures 83–96" captionStartId="F11" captionText="Figures 83 - 96. Male anal tergite in caudal view 83 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 84 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 85 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 86 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 87 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 88 Eupholidoptera giuliae Chora Sfakion CT 2000.014.01 89 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 90 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 91 Eupholidoptera astyla Krotos RMNH. INS 1141819 92 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 93 Eupholidoptera mariannae Prina FC 17798 RMNH. INS 1141840 94 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 95 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 96 Eupholidoptera jacquelinae paratype Gavdos NHMC 2001.004.12. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures83-96" httpUri="https://binary.pensoft.net/fig/817941" pageId="0" pageNumber="67">87</figureCitation>
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,
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|
<figureCitation captionStart="Figures 97–110" captionStartId="F12" captionText="Figures 97 - 110. Male anal tergite in lateral view 97 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 98 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 99 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 100 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 101 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.01 102 Eupholidoptera giuliae Argoules CT 1995.011.11 103 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 104 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 105 Eupholidoptera astyla Ano Viannos CT 1995.024.01 106 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 107 Eupholidoptera mariannae Kalavros CT 2017.029.01 108 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 109 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 110 Eupholidoptera jacquelinae Gavdopoula CT 1996.019.01. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures97-110" httpUri="https://binary.pensoft.net/fig/817942" pageId="0" pageNumber="67">101</figureCitation>
|
|||
|
|
) centrally depressed, distally bend downward, forming two lobes with teeth like apex pointing downward, separated by wide excision; cerci (Figs
|
|||
|
|
<figureCitation captionStart="Figures 111–125" captionStartId="F13" captionText="Figures 111 - 125. Male cercus in dorsal view 111 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 112 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 113 Eupholidoptera pallipes Mt. Lefka paratype RMNH. INS 1105314 114 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 115 Eupholidoptera latens Xiloskalo IBER DC-Ort 000558 116 Eupholidoptera giuliae Chora Sfakion CT 2000.014.02 117 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 118 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 119 Eupholidoptera annamariae Xerokampos, male with anomalous cercus, IBER DC-Ort 000565 120 Eupholidoptera astyla Ierapetra MfN s. n. 121 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 122 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 123 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 124 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 125 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures111-125" httpUri="https://binary.pensoft.net/fig/817943" pageId="0" pageNumber="67">115</figureCitation>
|
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|
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,
|
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|
|
<figureCitation captionStart="Figures 126–139" captionStartId="F14" captionText="Figures 126 - 139. Male cercus in lateral view 126 Eupholidoptera smyrnensis Makrigiannis RMNH. 5087053 127 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 128 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 129 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 130 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 131 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 132 Eupholidoptera francisae paratype sp. nov. Andikithira CT 2002.004.08 133 Eupholidoptera annamariae Kato Zakros CT 1995.020.02 134 Eupholidoptera astyla paratype Ierapetra MfN s. n. 135 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 136 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 137 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 138 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 139 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures126-139" httpUri="https://binary.pensoft.net/fig/817944" pageId="0" pageNumber="67">130</figureCitation>
|
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|
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) unarmed, 6-7
|
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|
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<normalizedToken originalValue="×">x</normalizedToken>
|
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|
|
longer than wide, in basal half conical, in apical half cylindrical, slightly curved inward in basal half, in profile straight to weakly upturned; subgenital plate (Figs
|
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|
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<figureCitation captionStart="Figures 140–153" captionStartId="F15" captionText="Figures 140 - 153. Male subgenital plate in ventral view 140 Eupholidoptera smyrnensis Lagkada RMNH 5106270 141 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 142 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 143 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 144 Eupholidoptera latens Mt. Lefka Kalergi CT 1987.047.02 145 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 146 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 147 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 148 Eupholidoptera astyla Krotos RMNH. INS 1141819 149 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 150 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 151 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 152 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 153 Eupholidoptera jacquelinae holotype Gavdos 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures140-153" httpUri="https://binary.pensoft.net/fig/817945" pageId="0" pageNumber="67">144</figureCitation>
|
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,
|
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<figureCitation captionStart="Figures 154–167" captionStartId="F16" captionText="Figures 154 - 167. Male subgenital plate in lateral view 154 Eupholidoptera smyrnensis Lagkada RMNH. INS 152909 155 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 156 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 157 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 (RMNH. INS 1141838 158 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 159 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 160 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 161 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 162 Eupholidoptera astyla Krotos RMNH 5086992 163 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 164 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 165 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 166 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 167 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures154-167" httpUri="https://binary.pensoft.net/fig/817946" pageId="0" pageNumber="67">158</figureCitation>
|
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|
|
) longer than wide, widest in proximal third, tapering and distinctly narrower in distal two thirds, sides rimmed in proximal third, in profile pointing backward, tip apical lobes rounded, spineless, with narrow median excision along one quarter of length its proximal half narrower; styli (Fig.
|
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|
|
<figureCitation captionStart="Figures 168–181" captionStartId="F17" captionText="Figures 168 - 181. Male stylus in lateral view 168 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 169 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 170 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 171 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 172 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 173 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 174 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 175 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 176 Eupholidoptera astyla Krotos RMNH. INS 1141819 177 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 178 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 179 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 180 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 181 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures168-181" httpUri="https://binary.pensoft.net/fig/817947" pageId="0" pageNumber="67">172</figureCitation>
|
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|
|
) short, one quarter as long as cerci, 2-3
|
|||
|
|
<normalizedToken originalValue="×">x</normalizedToken>
|
|||
|
|
longer than wide, round or flattened, inserted at ventral side of apical lobes, proximal of tip, pointing downward; titillator (Figs
|
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|
|
<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">186</figureCitation>
|
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,
|
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|
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<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">187</figureCitation>
|
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,
|
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|
|
<figureCitation captionStart="Figures 198–212" captionStartId="F19" captionText="Figures 198 - 212. Male titillator in lateral view 198 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 199 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 200 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 201 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 202 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 203 Eupholidoptera latens Rhodopos CH 8236 204 Eupholidoptera giuliae Argoules CT 1995.011.11 205 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 206 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 207 Eupholidoptera astyla Krotos RMNH. INS 1141819 208 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 209 Eupholidoptera mariannae Kalavros CT 2017.029.01 210 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 211 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 212 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures198-212" httpUri="https://binary.pensoft.net/fig/817949" pageId="0" pageNumber="67">202</figureCitation>
|
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|
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,
|
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|
|
<figureCitation captionStart="Figures 198–212" captionStartId="F19" captionText="Figures 198 - 212. Male titillator in lateral view 198 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 199 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 200 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 201 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 202 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 203 Eupholidoptera latens Rhodopos CH 8236 204 Eupholidoptera giuliae Argoules CT 1995.011.11 205 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 206 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 207 Eupholidoptera astyla Krotos RMNH. INS 1141819 208 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 209 Eupholidoptera mariannae Kalavros CT 2017.029.01 210 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 211 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 212 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures198-212" httpUri="https://binary.pensoft.net/fig/817949" pageId="0" pageNumber="67">203</figureCitation>
|
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|
) symmetrical, apical arms fused, parallel to slightly widening toward apical third inflated, diverging into two straight diverging hook-like teeth, in profile basal two thirds parallel, in second third swollen forming wide angle with two weakly upward curved hook-like teeth. Female - subgenital plate (Figs
|
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|
|
<figureCitation captionStart="Figures 39–52" captionStartId="F7" captionText="Figures 39 - 52. Female subgenital plate in ventral view 39 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 40 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 41 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 42 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 43 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 44 Eupholidoptera giuliae Argoules CT 1995.011.03 45 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 46 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 47 Eupholidoptera astyla Krotos RMNH. INS 1141820 48 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 49 Eupholidoptera mariannae Kalavros RMNH 5014912 50 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 51 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 52 Eupholidoptera jacquelinae Gavdos allotype CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures39-52" httpUri="https://binary.pensoft.net/fig/817937" pageId="0" pageNumber="67">43</figureCitation>
|
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,
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<figureCitation captionStart="Figures 53–66" captionStartId="F8" captionText="Figures 53 - 66. Female subgenital plate in lateral view 53 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 54 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 55 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 56 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 57 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 58 Eupholidoptera giuliae Argoules CT 1995.011.03 59 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 60 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 61 Eupholidoptera astyla Krotos RMNH. INS 1141820 62 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 63 Eupholidoptera mariannae Kalavros RMNH 5014912 64 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 65 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 66 Eupholidoptera jacquelinae allotype Gavdos CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures53-66" httpUri="https://binary.pensoft.net/fig/817938" pageId="0" pageNumber="67">57</figureCitation>
|
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) generally slightly wider than long, proximally with two concavities separated by keel, tip apical lobes rectangular, rounded, with slit-like to narrow V-shaped excision reaching one third to halfway, in profile rhomboid, upper distal angle rectangularly rounded.
|
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|
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</paragraph>
|
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="67" type="description">
|
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<paragraph pageId="0" pageNumber="67">Measurements.</paragraph>
|
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<paragraph pageId="0" pageNumber="67">
|
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|
|
See Tables
|
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|
|
<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Measurements (in mm), ratios, and biometrics of male Eupholidoptera." httpUri="http://table.plazi.org/id/8B3AFDE00956AD4F11FF9ECA2A2FA78F" pageId="0" pageNumber="67" tableUuid="8B3AFDE00956AD4F11FF9ECA2A2FA78F">6</tableCitation>
|
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,
|
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|
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<tableCitation captionStart="Table 7" captionStartId="T7" captionText="Table 7. Measurements (in mm), ratios, and biometrics of female Eupholidoptera." httpUri="http://table.plazi.org/id/3ABD8C0FD5A9E22855CC92BF15EE30E0" pageId="0" pageNumber="67" tableUuid="3ABD8C0FD5A9E22855CC92BF15EE30E0">7</tableCitation>
|
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|
.
|
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|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
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<subSubSection pageId="0" pageNumber="67" type="bioacoustics">
|
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|
<paragraph pageId="0" pageNumber="67">Bioacoustics.</paragraph>
|
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|
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<paragraph pageId="0" pageNumber="67">
|
|||
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|
Based upon the sound recordings of nine specimens (49 syllables measured), the song of
|
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|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. latens" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="latens">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. latens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, as in all species of
|
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|
|
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
|
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|
|
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera</emphasis>
|
|||
|
|
</taxonomicName>
|
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|
|
, consists of isolated syllables produced in long series with the opening hemisyllable much shorter and weaker than the closing hemisyllable. In
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. latens" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="latens">
|
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|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. latens</emphasis>
|
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|
|
</taxonomicName>
|
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|
|
, the syllable duration is ~ 228 ms, with a syllable rate up to ~ 1/s. Published records (
|
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2156.1.1" author="Ciplak, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="1 - 77" refId="B12" refString="Ciplak, B, Heller, K-G, Willemse, F, 2009. Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song. Zootaxa 2156 (1): 1 - 77, DOI: https://doi.org/10.11646/zootaxa.2156.1.1" title="Review of the genus Eupholidoptera (Orthoptera, Tettigoniidae): Different genitalia, uniform song." url="https://doi.org/10.11646/zootaxa.2156.1.1" volume="2156" year="2009">
|
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|
|
<normalizedToken originalValue="Çiplak">Ciplak</normalizedToken>
|
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|
|
et al. 2009
|
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|
|
</bibRefCitation>
|
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|
|
) show a syllable duration of ~ 161 ms and a syllable repetition rate of ~ 0.5/s at maximum. The song may most likely be confused with the other species of
|
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|
|
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
|
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|
|
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in Crete, except
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. smyrnensis" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="smyrnensis">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. smyrnensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. forcipata" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="forcipata">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. forcipata</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. For details of sound recordings of
|
|||
|
|
<taxonomicName authorityName="Willemse & Kruseman" authorityYear="1976" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera latens" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="latens">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera latens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
see Suppl. material 3.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="67" type="variation">
|
|||
|
|
<paragraph pageId="0" pageNumber="67">Variation.</paragraph>
|
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|
|
<paragraph pageId="0" pageNumber="67">
|
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|
|
<taxonomicName authorityName="Willemse & Kruseman" authorityYear="1976" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera latens" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="latens">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera latens</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
is restricted to the northern and central Chania region in western Crete. The cerci and anal tergite show little variation across its range. Titillators in males from the Rodopou peninsula (Figs
|
|||
|
|
<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">187</figureCitation>
|
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,
|
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|
|
<figureCitation captionStart="Figures 198–212" captionStartId="F19" captionText="Figures 198 - 212. Male titillator in lateral view 198 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 199 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 200 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 201 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 202 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 203 Eupholidoptera latens Rhodopos CH 8236 204 Eupholidoptera giuliae Argoules CT 1995.011.11 205 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 206 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 207 Eupholidoptera astyla Krotos RMNH. INS 1141819 208 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 209 Eupholidoptera mariannae Kalavros CT 2017.029.01 210 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 211 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 212 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures198-212" httpUri="https://binary.pensoft.net/fig/817949" pageId="0" pageNumber="67">203</figureCitation>
|
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|
|
) are relatively slender, with longer apical hooks, being intermediate between
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. latens" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="latens">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. latens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. giuliae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="giuliae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. giuliae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. In males, morphological variation is most pronounced in the subgenital plates. In males from Kolympari, Rodhopos, and Kato Kefalia in central and northern Chania, as well as in the outlier found in Rethimno the subgenital plate is clearly more slender than in the populations of the Lefka area, the transition between the wide basal part and the narrow apical part generally being more distinct. The morphological variation found in
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. latens" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="latens">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. latens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. giuliae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="giuliae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. giuliae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. francisae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="francisae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. francisae</emphasis>
|
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|
|
</taxonomicName>
|
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|
|
sp. nov. is further elaborated in the discussion.
|
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|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="67" type="differential diagnosis">
|
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|
|
<paragraph pageId="0" pageNumber="67">Differential diagnosis.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="67">
|
|||
|
|
Males differ from congenerics in the stout apical arms of the titillator (Figs
|
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|
|
<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">186</figureCitation>
|
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|
|
,
|
|||
|
|
<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">187</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation captionStart="Figures 198–212" captionStartId="F19" captionText="Figures 198 - 212. Male titillator in lateral view 198 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 199 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 200 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 201 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 202 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 203 Eupholidoptera latens Rhodopos CH 8236 204 Eupholidoptera giuliae Argoules CT 1995.011.11 205 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 206 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 207 Eupholidoptera astyla Krotos RMNH. INS 1141819 208 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 209 Eupholidoptera mariannae Kalavros CT 2017.029.01 210 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 211 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 212 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures198-212" httpUri="https://binary.pensoft.net/fig/817949" pageId="0" pageNumber="67">202</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation captionStart="Figures 198–212" captionStartId="F19" captionText="Figures 198 - 212. Male titillator in lateral view 198 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 199 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 200 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 201 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 202 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 203 Eupholidoptera latens Rhodopos CH 8236 204 Eupholidoptera giuliae Argoules CT 1995.011.11 205 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 206 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 207 Eupholidoptera astyla Krotos RMNH. INS 1141819 208 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 209 Eupholidoptera mariannae Kalavros CT 2017.029.01 210 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 211 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 212 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures198-212" httpUri="https://binary.pensoft.net/fig/817949" pageId="0" pageNumber="67">203</figureCitation>
|
|||
|
|
), fused in basal half, separated into two strong, short, parallel or diverging curved hooks in the elongated, in the slender subgenital plate (Figs
|
|||
|
|
<figureCitation captionStart="Figures 140–153" captionStartId="F15" captionText="Figures 140 - 153. Male subgenital plate in ventral view 140 Eupholidoptera smyrnensis Lagkada RMNH 5106270 141 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 142 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 143 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 144 Eupholidoptera latens Mt. Lefka Kalergi CT 1987.047.02 145 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 146 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 147 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 148 Eupholidoptera astyla Krotos RMNH. INS 1141819 149 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 150 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 151 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 152 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 153 Eupholidoptera jacquelinae holotype Gavdos 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures140-153" httpUri="https://binary.pensoft.net/fig/817945" pageId="0" pageNumber="67">144</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation captionStart="Figures 154–167" captionStartId="F16" captionText="Figures 154 - 167. Male subgenital plate in lateral view 154 Eupholidoptera smyrnensis Lagkada RMNH. INS 152909 155 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 156 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 157 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 (RMNH. INS 1141838 158 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 159 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 160 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 161 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 162 Eupholidoptera astyla Krotos RMNH 5086992 163 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 164 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 165 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 166 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 167 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures154-167" httpUri="https://binary.pensoft.net/fig/817946" pageId="0" pageNumber="67">158</figureCitation>
|
|||
|
|
) narrowing at one third of the length, the apical lobes without a teeth at its apex and with short styli (Fig.
|
|||
|
|
<figureCitation captionStart="Figures 168–181" captionStartId="F17" captionText="Figures 168 - 181. Male stylus in lateral view 168 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 169 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 170 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 171 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 172 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 173 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 174 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 175 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 176 Eupholidoptera astyla Krotos RMNH. INS 1141819 177 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 178 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 179 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 180 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 181 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures168-181" httpUri="https://binary.pensoft.net/fig/817947" pageId="0" pageNumber="67">172</figureCitation>
|
|||
|
|
), inserted pre-apically pointing downward, the anal tergite (Figs
|
|||
|
|
<figureCitation captionStart="Figures 69–82" captionStartId="F10" captionText="Figures 69 - 82. Male anal tergite in dorsal view 69 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 70 Eupholidoptera gemellata holotype Mt. Idhi RMNH. INS 1105300 71 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 72 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 73 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 74 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 75 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 76 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 77 Eupholidoptera astyla Krotos RMNH. INS 1141819 78 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 79 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 80 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 81 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 82 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures69-82" httpUri="https://binary.pensoft.net/fig/817940" pageId="0" pageNumber="67">73</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation captionStart="Figures 83–96" captionStartId="F11" captionText="Figures 83 - 96. Male anal tergite in caudal view 83 Eupholidoptera smyrnensis Makrigiannis RMNH. INS 152909 84 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 85 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 86 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 87 Eupholidoptera latens Mt. Lefka, Kalergi CT 2003.009.01 88 Eupholidoptera giuliae Chora Sfakion CT 2000.014.01 89 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 90 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 91 Eupholidoptera astyla Krotos RMNH. INS 1141819 92 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 93 Eupholidoptera mariannae Prina FC 17798 RMNH. INS 1141840 94 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 95 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 96 Eupholidoptera jacquelinae paratype Gavdos NHMC 2001.004.12. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures83-96" httpUri="https://binary.pensoft.net/fig/817941" pageId="0" pageNumber="67">87</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation captionStart="Figures 97–110" captionStartId="F12" captionText="Figures 97 - 110. Male anal tergite in lateral view 97 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 98 Eupholidoptera gemellata Mt. Idhi CT 2000.095.01 99 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 100 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 101 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.01 102 Eupholidoptera giuliae Argoules CT 1995.011.11 103 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.02 104 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 105 Eupholidoptera astyla Ano Viannos CT 1995.024.01 106 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 107 Eupholidoptera mariannae Kalavros CT 2017.029.01 108 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 109 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 110 Eupholidoptera jacquelinae Gavdopoula CT 1996.019.01. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures97-110" httpUri="https://binary.pensoft.net/fig/817942" pageId="0" pageNumber="67">101</figureCitation>
|
|||
|
|
) medially bent downwards forming two widely separated lobes with short pointed tips pointing downward and in the slender, unarmed, weakly inward and upward bent cerci (Figs
|
|||
|
|
<figureCitation captionStart="Figures 111–125" captionStartId="F13" captionText="Figures 111 - 125. Male cercus in dorsal view 111 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 112 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 113 Eupholidoptera pallipes Mt. Lefka paratype RMNH. INS 1105314 114 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 115 Eupholidoptera latens Xiloskalo IBER DC-Ort 000558 116 Eupholidoptera giuliae Chora Sfakion CT 2000.014.02 117 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 118 Eupholidoptera annamariae Kato Zakros CT 2000.030.03 119 Eupholidoptera annamariae Xerokampos, male with anomalous cercus, IBER DC-Ort 000565 120 Eupholidoptera astyla Ierapetra MfN s. n. 121 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 122 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 123 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 124 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 125 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures111-125" httpUri="https://binary.pensoft.net/fig/817943" pageId="0" pageNumber="67">115</figureCitation>
|
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|
|
,
|
|||
|
|
<figureCitation captionStart="Figures 126–139" captionStartId="F14" captionText="Figures 126 - 139. Male cercus in lateral view 126 Eupholidoptera smyrnensis Makrigiannis RMNH. 5087053 127 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 128 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 129 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 130 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 131 Eupholidoptera giuliae Chora Sfakion CT 2000.024.04 132 Eupholidoptera francisae paratype sp. nov. Andikithira CT 2002.004.08 133 Eupholidoptera annamariae Kato Zakros CT 1995.020.02 134 Eupholidoptera astyla paratype Ierapetra MfN s. n. 135 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 136 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 137 Eupholidoptera forcipata Mt. Idhi CT 1987.044.01 138 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 139 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures126-139" httpUri="https://binary.pensoft.net/fig/817944" pageId="0" pageNumber="67">130</figureCitation>
|
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|
|
).
|
|||
|
|
<taxonomicName authorityName="Willemse & Kruseman" authorityYear="1976" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera latens" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="latens">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera latens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
most closely resembles
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. francisae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="francisae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. francisae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov. but differs in the shape of the subgenital plate (compare Fig.
|
|||
|
|
<figureCitation captionStart="Figures 140–153" captionStartId="F15" captionText="Figures 140 - 153. Male subgenital plate in ventral view 140 Eupholidoptera smyrnensis Lagkada RMNH 5106270 141 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 142 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 143 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 144 Eupholidoptera latens Mt. Lefka Kalergi CT 1987.047.02 145 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 146 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 147 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 148 Eupholidoptera astyla Krotos RMNH. INS 1141819 149 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 150 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 151 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 152 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 153 Eupholidoptera jacquelinae holotype Gavdos 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures140-153" httpUri="https://binary.pensoft.net/fig/817945" pageId="0" pageNumber="67">144</figureCitation>
|
|||
|
|
with Fig.
|
|||
|
|
<figureCitation captionStart="Figures 140–153" captionStartId="F15" captionText="Figures 140 - 153. Male subgenital plate in ventral view 140 Eupholidoptera smyrnensis Lagkada RMNH 5106270 141 Eupholidoptera gemellata Mt. Idhi FC 1651 RMNH. INS 1141843 142 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105313 143 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 144 Eupholidoptera latens Mt. Lefka Kalergi CT 1987.047.02 145 Eupholidoptera giuliae Skaloti IBER DC-Ort 000564 146 Eupholidoptera francisae sp. nov. holotype Andikithira CT 2002.004.04 147 Eupholidoptera annamariae Kato Zakros CT 2000.030.04 148 Eupholidoptera astyla Krotos RMNH. INS 1141819 149 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 150 Eupholidoptera mariannae holotype Anatoli RMNH. INS 1105311 151 Eupholidoptera forcipata Mt. Idhi CT 1987.044.03 152 Eupholidoptera marietheresae sp. nov. holotype Mt. Dikti FC 1606 RMNH. INS 1141850 153 Eupholidoptera jacquelinae holotype Gavdos 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures140-153" httpUri="https://binary.pensoft.net/fig/817945" pageId="0" pageNumber="67">146</figureCitation>
|
|||
|
|
), the length-width ratio of styli, the shape of the titillator (compare Figs
|
|||
|
|
<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">186</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">187</figureCitation>
|
|||
|
|
with Fig.
|
|||
|
|
<figureCitation captionStart="Figures 182–197" captionStartId="F18" captionText="Figures 182 - 197. Male titillator in ventral view 182 Eupholidoptera smyrnensis Makrigiannis RMNH 5087053 183 Eupholidoptera gemellata Mt. Idhi Amariou FC 1651 RMNH. INS 1141843 184 Eupholidoptera pallipes paratype Mt. Lefka RMNH. INS 1105314 185 Eupholidoptera cretica Mt. Lefka, Omalos FC 17807 RMNH. INS 1141838 186 Eupholidoptera latens Mt. Lefka Kalergi CT 2003.009.01 187 Eupholidoptera latens Rhodopos CH 8236 188 Eupholidoptera giuliae Argoules CT 1995.011.11 189 Eupholidoptera francisae sp. nov. paratype Andikithira CT 2002.004.08 190 Eupholidoptera annamariae Kato Zakros CT 2000.030.01 191 Eupholidoptera astyla Krotos RMNH. INS 1141819 192 Eupholidoptera astyla Kofinas FC 460 RMNH. INS 1141836 193 Eupholidoptera feri holotype Mt. Dikti, Katharo plain RMNH. INS 1105297 194 Eupholidoptera mariannae paratype Malles CH 2906 A 195 Eupholidoptera forcipata Mt. Idhi CT 2019.047.01 196 Eupholidoptera marietheresae sp. nov. paratype Mt. Dikti FC 1606 CT 2000.096.01 197 Eupholidoptera jacquelinae holotype Gavdos CT 2000.005.02. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures182-197" httpUri="https://binary.pensoft.net/fig/817948" pageId="0" pageNumber="67">189</figureCitation>
|
|||
|
|
), body size and the ratio height-length of the hind femur (see Tables
|
|||
|
|
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Measurements (in mm) and biometrics of male E. latens and E. francisae." httpUri="http://table.plazi.org/id/E8FAC6B421151AA7F436658A4660E555" pageId="0" pageNumber="67" tableUuid="E8FAC6B421151AA7F436658A4660E555">3</tableCitation>
|
|||
|
|
,
|
|||
|
|
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Measurements (in mm) and biometrics of female E. latens and E. francisae." httpUri="http://table.plazi.org/id/FDBD454389BD0FC9F880E417AF4C5F8F" pageId="0" pageNumber="67" tableUuid="FDBD454389BD0FC9F880E417AF4C5F8F">4</tableCitation>
|
|||
|
|
for measurements). Females differ in the subgenital plates (Figs
|
|||
|
|
<figureCitation captionStart="Figures 39–52" captionStartId="F7" captionText="Figures 39 - 52. Female subgenital plate in ventral view 39 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 40 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 41 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 42 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 43 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 44 Eupholidoptera giuliae Argoules CT 1995.011.03 45 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 46 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 47 Eupholidoptera astyla Krotos RMNH. INS 1141820 48 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 49 Eupholidoptera mariannae Kalavros RMNH 5014912 50 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 51 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 52 Eupholidoptera jacquelinae Gavdos allotype CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures39-52" httpUri="https://binary.pensoft.net/fig/817937" pageId="0" pageNumber="67">43</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation captionStart="Figures 53–66" captionStartId="F8" captionText="Figures 53 - 66. Female subgenital plate in lateral view 53 Eupholidoptera smyrnensis Makrigiannis RMNH. 5014918 54 Eupholidoptera gemellata Mt. Idhi, FC 1602 CT 2000.095.02 55 Eupholidoptera pallipes allotype Mt. Lefka RMNH. INS 1105312 56 Eupholidoptera cretica Mt. Lefka, Omalos RMNH. INS 1141837 57 Eupholidoptera latens Mt. Lefka, Kalergi CT 1987.047.04 58 Eupholidoptera giuliae Argoules CT 1995.011.03 59 Eupholidoptera francisae sp. nov. allotype Andikithira CT 2002.004.11 60 Eupholidoptera annamariae Kato Zakros CT 1995.020.03 61 Eupholidoptera astyla Krotos RMNH. INS 1141820 62 Eupholidoptera feri allotype Mt. Dikti, Katharo plain RMNH. INS 1105298 63 Eupholidoptera mariannae Kalavros RMNH 5014912 64 Eupholidoptera forcipata Mt. Idhi, FC 1602 RMNH. INS 1141845 65 Eupholidoptera marietheresae sp. nov. allotype Mt. Dikti RMNH. INS 1141849 66 Eupholidoptera jacquelinae allotype Gavdos CT 2001.004.13. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1151.97514.figures53-66" httpUri="https://binary.pensoft.net/fig/817938" pageId="0" pageNumber="67">57</figureCitation>
|
|||
|
|
) being as long as wide, proximally with two concavities, slit-like to narrow V-shaped excision between the apical lobes reaching halfway. They can be distinguished from females of
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. francisae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="francisae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. francisae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the fact that in ventral view the incision of the hind margin in
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. francisae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="francisae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. francisae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is shaped in the form of a wide V; in profile the apex of the female subgenital plate in
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. latens" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="latens">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. latens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
reaches the distal half of the gonangulum or surpasses it (Fig.
|
|||
|
|
<figureCitation captionStart="Figures 67, 68" captionStartId="F9" captionText="Figures 67, 68. Female subgenital plate in lateral view 67 Eupholidoptera francisae sp. nov. Anidhroi CT 2001.002.07 68 Eupholidoptera latens Prases DC-Ort 000512." figureDoi="10.3897/zookeys.1151.97514.figures67-68" httpUri="https://binary.pensoft.net/fig/817939" pageId="0" pageNumber="67">68</figureCitation>
|
|||
|
|
) while in
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. francisae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="francisae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. francisae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
it does not reach or surpass the proximal half of the gonangulum (Fig.
|
|||
|
|
<figureCitation captionStart="Figures 67, 68" captionStartId="F9" captionText="Figures 67, 68. Female subgenital plate in lateral view 67 Eupholidoptera francisae sp. nov. Anidhroi CT 2001.002.07 68 Eupholidoptera latens Prases DC-Ort 000512." figureDoi="10.3897/zookeys.1151.97514.figures67-68" httpUri="https://binary.pensoft.net/fig/817939" pageId="0" pageNumber="67">67</figureCitation>
|
|||
|
|
). In colouration, the amount of black shown by
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. latens" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="latens">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. latens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is intermediate between overall pale coloured species like
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. cretica" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="cretica">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. cretica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. jacquelinae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="jacquelinae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. jacquelinae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. smyrnensis" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="smyrnensis">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. smyrnensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and dark-coloured species like
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. annamariae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="annamariae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. annamariae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. astyla" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="astyla">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. astyla</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, or
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. mariannae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="mariannae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. mariannae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. For more details differentiating
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. latens" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="latens">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. latens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from other Cretan
|
|||
|
|
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
see Table
|
|||
|
|
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Diagnostic characters and character-states for Eupholidoptera species from Crete, Andikithira, Gavdos, and Gavdopoula." httpUri="http://table.plazi.org/id/DEC408AC569AD02CE9FF184126BFC8DE" pageId="0" pageNumber="67" tableUuid="DEC408AC569AD02CE9FF184126BFC8DE">5</tableCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="67" type="distribution">
|
|||
|
|
<paragraph pageId="0" pageNumber="67">Distribution.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="67">
|
|||
|
|
The species was discovered in 1973 at high altitudes on Mt. Lefka, western Crete and published records from this species originated only from this mountain and its foothills (Lakki). Specimens collected between 2016 and 2019 indicate the species also occurs at low altitudes in the northern and northeastern parts of Chania region (Fig.
|
|||
|
|
<figureCitation captionStart="Figures 254, 255" captionStartId="F28" captionText="Figures 254, 255. Distribution maps Eupholidoptera Crete and adjacent islands 254 lowland species 255 highland species." figureDoi="10.3897/zookeys.1151.97514.figures254-255" httpUri="https://binary.pensoft.net/fig/817958" pageId="0" pageNumber="67">254</figureCitation>
|
|||
|
|
). To the east the distribution area of
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. latens" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="latens">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. latens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
borders to but is separated from
|
|||
|
|
<taxonomicName family="Tettigoniidae" kingdom="Animalia" lsidName="E. giuliae" order="Orthoptera" pageId="0" pageNumber="67" rank="species" species="giuliae">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">E. giuliae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. For a complete list of localities, specimens and repositories see Suppl. material 1.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="67" type="habitat">
|
|||
|
|
<paragraph pageId="0" pageNumber="67">Habitat.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="67">
|
|||
|
|
The species occupies habitats from sea level to alpine regions between 1600 and 1800 m on Mt. Lefka. It hides in low prickly shrublets in the phrygana while at lower altitudes it was also found on shrubs of blackberry (
|
|||
|
|
<taxonomicName class="Insecta" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rubus" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">Rubus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) or gorse (
|
|||
|
|
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Ulex" kingdom="Plantae" lsidName="Ulex" order="Fabales" pageId="0" pageNumber="67" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">Ulex</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="67" type="phenology">
|
|||
|
|
<paragraph pageId="0" pageNumber="67">Phenology.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="67">At low altitudes adults appear around mid-May, at mid-level elevations toward the end of May or early June whereas at high altitudes it may take to the second half of July before the first adults appear.</paragraph>
|
|||
|
|
<caption doi="10.3897/zookeys.1151.97514.figures254-255" httpUri="https://binary.pensoft.net/fig/817958" pageId="0" pageNumber="67" start="Figures 254, 255" startId="F28">
|
|||
|
|
<paragraph pageId="0" pageNumber="67">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="67">Figures 254, 255.</emphasis>
|
|||
|
|
Distribution maps
|
|||
|
|
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupholidoptera" order="Orthoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="67">Eupholidoptera</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Crete and adjacent islands
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="67">254</emphasis>
|
|||
|
|
lowland species
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="67">255</emphasis>
|
|||
|
|
highland species.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|