treatments-xml/data/9E/6B/78/9E6B7895E289B40220F7952879357EA6.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.248.4106" ID-GBIF-Dataset="605e7744-e499-4617-bb69-6531a82ddbae" ID-PMC="PMC3520135" ID-Pensoft-Pub="1313-2970-248-1" ID-PubMed="23275757" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-248-1" ModsDocOrigin="ZooKeys 248" ModsDocTitle="A revision of the New World species of Gymnoclasiopa Hendel (Diptera, Ephydridae)" checkinTime="1451247880623" checkinUser="pensoft" docAuthor="Mathis, Wayne N. &amp; Zatwarnicki, Tadeusz" docDate="2012" docId="9E6B7895E289B40220F7952879357EA6" docLanguage="en" docName="ZooKeys 248: 1-69" docOrigin="ZooKeys 248" docSource="http://dx.doi.org/10.3897/zookeys.248.4106" docTitle="Gymnoclasiopa bohemanni" docType="treatment" docVersion="3" lastPageNumber="23" masterDocId="23013B478951FFB1FFF7FFACFFBFFF8F" masterDocTitle="A revision of the New World species of Gymnoclasiopa Hendel (Diptera, Ephydridae)" masterLastPageNumber="69" masterPageNumber="1" pageNumber="18" updateTime="1668154988994" updateUser="ExternalLinkService">
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<mods:title>A revision of the New World species of Gymnoclasiopa Hendel (Diptera, Ephydridae)</mods:title>
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<mods:namePart>Mathis, Wayne N.</mods:namePart>
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<mods:namePart>Zatwarnicki, Tadeusz</mods:namePart>
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<mods:date>2012</mods:date>
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<mods:number>248</mods:number>
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<treatment ID-GBIF-Taxon="152039115" LSID="urn:lsid:plazi:treatment:9E6B7895E289B40220F7952879357EA6" httpUri="http://treatment.plazi.org/id/9E6B7895E289B40220F7952879357EA6" lastPageId="22" lastPageNumber="23" pageId="17" pageNumber="18">
<subSubSection pageId="17" pageNumber="18" type="nomenclature">
<paragraph pageId="17" pageNumber="18">
<taxonomicName LSID="http://species-id.net/wiki/Gymnoclasiopa_bohemanni" authority="Becker" class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa bohemanni" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bohemanni">Gymnoclasiopa bohemanni (Becker)</taxonomicName>
Figs 21-24
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="18" type="reference_group">
<paragraph pageId="17" pageNumber="18">
<taxonomicName class="Insecta" family="Ephydridae" genus="Clasiopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clasiopa bohemanni" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bohemanni">Clasiopa bohemanni</taxonomicName>
<bibRefCitation author="Becker, Th" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="50" pageNumber="51" pagination="91 - 276" title="Dipterologische Studien IV. Ephydridae." volume="41" year="1896">Becker 1896</bibRefCitation>
: 159.
</paragraph>
<paragraph pageId="17" pageNumber="18">
<taxonomicName class="Insecta" family="Ephydridae" genus="Discocerina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discocerina bohemanni" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bohemanni">Discocerina bohemanni</taxonomicName>
. 
<bibRefCitation author="Cresson, ET Jr" journalOrPublisher="Transactions of the American Entomological Society" pageId="50" pageNumber="51" pagination="227 - 258" title="Studies in the dipterous family Ephydridae, excluding the North and South American faunas." volume="51" year="1925">Cresson 1925</bibRefCitation>
: 256 [generic combination].
</paragraph>
<paragraph pageId="17" pageNumber="18">
<taxonomicName class="Insecta" family="Ephydridae" genus="Ditrichophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditrichophora bohemanni" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bohemanni">Ditrichophora bohemanni</taxonomicName>
. 
<bibRefCitation author="Hackman, W" journalOrPublisher="Notulae Entomologicae" pageId="50" pageNumber="51" pagination="117 - 162" title="A check list of the Finnish Diptera II. Cyclorrhapha." volume="60" year="1980">Hackman 1980</bibRefCitation>
: 128 [generic combination].
</paragraph>
<paragraph pageId="17" pageNumber="18">
<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa bohemanni" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bohemanni">Gymnoclasiopa bohemanni</taxonomicName>
. 
<bibRefCitation author="Mathis, WN" journalOrPublisher="Memoirs on Entomology, International" pageId="50" pageNumber="51" pagination="1 - 423" title="A world catalog of the shore flies (Diptera: Ephydridae)." volume="4" year="1995">Mathis and Zatwarnicki 1995</bibRefCitation>
: 175 [generic combination; world catalog].
</paragraph>
<paragraph pageId="17" pageNumber="18">
<taxonomicName class="Insecta" family="Ephydridae" genus="Ditrichophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditrichophora (Gymnoclasiopa) montana" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="montana" subGenus="Gymnoclasiopa">Ditrichophora (Gymnoclasiopa) montana</taxonomicName>
<bibRefCitation author="Cresson, ET Jr" journalOrPublisher="Transactions of the American Entomological Society" pageId="50" pageNumber="51" pagination="101 - 128" title="Synopses of North American Ephydridae (Diptera) I. The subfamily Psilopinae, with descriptions of new species." volume="68" year="1942">Cresson 1942</bibRefCitation>
: 120. 
<bibRefCitation author="Wirth, WW" journalOrPublisher="S. Department of Agriculture, Washington, D. C." pageId="50" pageNumber="51" title="Ephydridae. Pp. 734 - 759. In: Stone A, Sabrosky CW, Wirth WW, Foote RH, Coulson JR (Eds) A Catalog of the Diptera of America North of Mexico. Handbook 276, U." year="1965">Wirth 1965</bibRefCitation>
: 739 [Nearctic catalog]. 
<bibRefCitation author="Cole, FR (with the collaboration of E. T. Schlinger)" journalOrPublisher="University of California Press, Berkeley, Los Angeles, and London" pageId="50" pageNumber="51" title="The flies of western North America." year="1969">Cole 1969</bibRefCitation>
: 398 [fauna, western North America]. NEW SYNONYM
</paragraph>
<paragraph pageId="17" pageNumber="18">
<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa montana" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="montana">Gymnoclasiopa montana</taxonomicName>
. 
<bibRefCitation author="Mathis, WN" journalOrPublisher="Memoirs on Entomology, International" pageId="50" pageNumber="51" pagination="1 - 423" title="A world catalog of the shore flies (Diptera: Ephydridae)." volume="4" year="1995">Mathis and Zatwarnicki 1995</bibRefCitation>
: 177 [generic combination; world catalog].
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="diagnosis">
<paragraph pageId="17" pageNumber="18">Diagnosis.</paragraph>
<paragraph lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
This species is distinguished from congeners by the following combination of characters: Small to medium-sized shore flies, body length 1.80-3.35 mm; head and thorax generally microtomentose gray, abdomen subshiny to shiny black. Head: Frons densely but finely microtomentose. Antenna yellowish, sometimes basal flagellomere slightly darkened dorsally; arista bearing 5 dorsal rays. Face relatively flat with antennal grooves inconspicuous; facial microtomentum generally sericeous, bright yellow (males) or grayish yellow (females, grayer in antennal grooves); gena moderately high, gena-to-eye ratio 0.14-0.18. Maxillary palpus yellow to yellowish red. Thorax: Anterolateral area of mesonotum, just mediad of area from postpronotum through notopleuron, densely microtomentose, mostly dull; lateral mesonotal area from and including postpronotum and notopleuron, densely and finely microtomentose; this area of males brown, contrasted with whitish to silvery gray microtomentum of broad, medial portion; same area in female concolorous with medial coloration. Wing of hyaline, not darkened along anterior region; costal ratio 0.0.39-0.50; M vein ratio 0.50-0.59; halter stem yellowish tan, knob yellowish white to white. Forecoxa yellow ventrally; foretibia mostly yellow, sometimes slightly brownish to grayish medially; midtibia mostly yellow to mostly grayish with only apices yellow, but with brownish area around apical 1/3; hindtibia without ventroapical spur, mostly grayish except for apices. Abdomen: Tergites 1-4 sparsely and finely microtomentose medially, subshiny; lateral margins of tergites and tergite 5 shiny black with microtomentum either lacking or sparse. Male terminalia (Figs 21-23): Epandrium in posterior view (Fig. 21) as a broadly formed, inverted U with the base more narrowly formed, dorsal portion more 
<pageBreakToken pageId="18" pageNumber="19" start="start">thinly</pageBreakToken>
developed than lateral arms, lateral arms widespread ventrally, shallowly arched, enlarged ventrally, broadly rounded, setulae more clustered at ventral margin; cercus in posterior view (Fig. 21) elongate, irregularly lunate, dorsal apex very narrow, digitiform, expanded toward ventral apex, ventral apex rounded, both lateral and medial margins arched, setulae more clustered at ventral margin; aedeagus in lateral view (Fig. 23) slipper-like, base shallowly emarginate, tapered very gradually toward apex, apical half nearly parallel sided, apex bluntly rounded, in ventral view (Fig. 22) elongate, expanded laterally from narrow base on basal 1/4, thereafter to apex almost parallel sided, apical margin rounded with tiny notch medially; phallapodeme in lateral view (Fig. 23) more or less irregularly triangular, extension toward hypandrium more elongate than angle towards aedeagal base, in ventral view (Fig. 22) I-shaped, subapical crossbar shorter and basal crossbar, with tapered shoulders, basal crossbar wider, widely Y-shaped, apical margin very shallowly emarginate; ejaculatory apodeme in lateral view robustly comma-shaped, in ventral view obtusely L-shaped; postgonite in lateral view (Fig. 23) with basal half robust, thereafter abruptly tapered to a narrow, slightly tapered, digitiform process, posterior margin with a few setulae, extended process with a single, longer setula, in ventral view (Fig. 22) as an elongate isosceles triangle, tapered gradually toward apex, width of base about half length, angles rounded, lateral and medial margins nearly straight; pregonite in lateral view (Fig. 23) moderately elongate, triangular, width at base almost twice lengths of sides, in ventral view (Fig. 22) lunate with apices pointed; hypandrium in ventral view (Fig. 22) robustly V to U-shaped, lateral margins slightly expanded posteriorly, anterior margin very broadly rounded, posterior margin conspicuously emarginate, widely U-shaped, in lateral view (Fig. 23) narrowly elongate, nearly straight.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="type material">
<paragraph pageId="18" pageNumber="19">Type material.</paragraph>
<paragraph pageId="18" pageNumber="19">
Lectotype female of 
<taxonomicName class="Insecta" family="Ephydridae" genus="Clasiopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clasiopa bohemanni" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bohemanni">Clasiopa bohemanni</taxonomicName>
Becker is labeled &quot;[empty red square]/Type [printed]/Bohemanni Beck [handwritten]/39 [printed]/174 57 [beige; 
<normalizedToken originalValue="“57”">&quot;57&quot;</normalizedToken>
handwritten]/22 64 [beige; 
<normalizedToken originalValue="“64”">&quot;64&quot;</normalizedToken>
handwritten]/LECTOTYPE ♀ 
<taxonomicName class="Insecta" family="Ephydridae" genus="Clasiopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clasiopa bohemanni" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bohemanni">Clasiopa bohemanni</taxonomicName>
Becker by Mathis &amp; Zatwarnicki NRS [red].&quot; The lectotype female is double mounted, is in good condition (anterior margin of right wing slightly broken near middle), and is deposited in the NRS. When 
<bibRefCitation author="Becker, Th" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="50" pageNumber="51" pagination="91 - 276" title="Dipterologische Studien IV. Ephydridae." volume="41" year="1896">Becker (1896</bibRefCitation>
; 159) described this species he noted that &quot;diese neue Art fand ich in 
<normalizedToken originalValue="Bohemann’s">Bohemann's</normalizedToken>
Sammlung.&quot;
</paragraph>
<paragraph pageId="18" pageNumber="19">
The holotype male of 
<taxonomicName class="Insecta" family="Ephydridae" genus="Ditrichophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditrichophora montana" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="montana">Ditrichophora montana</taxonomicName>
Cresson is labeled &quot;GLACIER PARK Avalanche L[a]k[e] [
<geoCoordinate direction="north" orientation="latitude" precision="92" value="48.656666">48°39.4'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-113.785">113°47.1'W</geoCoordinate>
] 14 July 1935[,] A. L. Melander/TYPE Ditrichophora Montana Cress HoloTYPE 6628 [red; all except 
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
handwritten].&quot; The holotype is double mounted (minuten pin in a rectangular card), is in excellent condition, and is deposited in the ANSP (6628).
</paragraph>
<paragraph pageId="18" pageNumber="19">
Type locality. Not given, 
<normalizedToken originalValue="“Bohemann’s">&quot;Bohemann's</normalizedToken>
Sammlung&quot; (=? Sweden).
</paragraph>
</subSubSection>
<subSubSection lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19" type="other specimens examined">
<paragraph pageId="18" pageNumber="19">Other specimens examined.</paragraph>
<paragraph pageId="18" pageNumber="19">
Nearctic. CANADA. ALBERTA. Okotoks, Sheep River Campground (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="50.723335">50°43.4'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-113.971664">113°58.3'W</geoCoordinate>
), 27 Jun 1968, W. W. Wirth (1♂; USNM).
</paragraph>
<paragraph lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
BRITISH COLUMBIA. Emerald Lake, Yoho National Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="51.43833">51°26.3'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-116.541664">116°32.5'W</geoCoordinate>
); 30 Jul 1935, A. L. Melander (1♀; ANSP). Martin Creek, Alaska Highway DC 243 (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="57.288334">57°17.3'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-121.46667">121°28'W</geoCoordinate>
), 13 Aug 1978, P. H. Arnaud, Jr. (1♀; CAS). Pine Pass (37 km NE; highway 97; 
<geoCoordinate direction="north" orientation="latitude" precision="925" value="55.5">55°30'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-122.666664">122°40'W</geoCoordinate>
), 25 Jun 1978, P. H. Arnaud, 
<pageBreakToken pageId="19" pageNumber="20" start="start">Jr</pageBreakToken>
. (3♂, 1♀; CAS). Prophet River Provincial Campground, Alaska Highway DC 221 (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="57.966667">57°58'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-122.78333">122°47'W</geoCoordinate>
), 13 Aug 1978, P. H. Arnaud, Jr. (1♀; CAS). Terrace (52 km SW; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="54.246666">54°14.8'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-130.28667">130°17.2'W</geoCoordinate>
), 5 Jul 1960, J. G. Chillcott (1♂; USNM).
</paragraph>
<paragraph pageId="19" pageNumber="20">
MANITOBA. The Pas (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="53.991665">53°59.5'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-101.253334">101°15.2'W</geoCoordinate>
), 31 Jul 1937, D. G. Denning (1♀; USNM).
</paragraph>
<paragraph pageId="19" pageNumber="20">
NEWFOUNDLAND. Cachrane Pond (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="47.966667">47°58'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-57.22333">57°13.4'W</geoCoordinate>
), 30 Jun 1961, C. P. Alexander (1♀; USNM). Terra Nova National Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="48.53">48°31.8'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-53.928333">53°55.7'W</geoCoordinate>
), 6-8 Jul 1961, C. P. Alexander (3♂; USNM).
</paragraph>
<paragraph pageId="19" pageNumber="20">
ONTARIO. Klotz Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="49.801666">49°48.1'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-85.863335">85°51.8'W</geoCoordinate>
), 5 Jul 1954, A. H. Sturtevant (1♂, 1♀; USNM).
</paragraph>
<paragraph pageId="19" pageNumber="20">
SASKATCHEWAN. Saskatoon, Beaver Creek (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="52.14">52°08.4'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-106.68667">106°41.2'W</geoCoordinate>
), 28 Aug 1955, W. W. Wirth (1♂; USNM).
</paragraph>
<paragraph pageId="19" pageNumber="20">
YUKON TERRITORY. Aishihik River, Alaska Highway DC 996.8 (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.671665">61°40.3'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-137.47166">137°28.3'W</geoCoordinate>
), 7 Aug 1978, P. H. Arnaud, Jr. (2♂, 1♀; CAS).
</paragraph>
<paragraph lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
UNITED STATES. ALASKA. Anchorage: Anchorage (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.218334">61°13.1'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-149.9">149°54'W</geoCoordinate>
; Seward Highway), 5 Aug 1964, K. M. Sommerman (3♂; USNM); Mirror Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.428333">61°25.7'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-149.415">149°24.9'W</geoCoordinate>
), 5 Aug 2002, D. and W. N. Mathis (1♀; USNM). Fairbanks-Northstar: Colorado Creek, Chena Hot Springs (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="65.05334">65°03.2'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-146.04834">146°02.9'W</geoCoordinate>
), 11 Jul 1978, P. H. Arnaud, Jr. (1♂, 1♀; CAS). Juneau: Douglas (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="58.275">58°16.5'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-134.39333">134°23.6'W</geoCoordinate>
), 19 Jun 1954, R. Coleman (1♀; USNM); Gastineau Channel, Thane Road (S Juneau; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="58.281666">58°16.9'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-134.37334">134°22.4'W</geoCoordinate>
), 20-22 Jul 2011, D. and W. N. Mathis (31♂, 9♀; USNM); Juneau, Mendenhall Valley, Riverside Rotary Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="58.38">58°22.8'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-134.58667">134°35.2'W</geoCoordinate>
), 21 Jul 2011, D. and W. N. Mathis (1♂; USNM). Kenai Peninsula: Homer (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="59.646667">59°38.8'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-151.525">151°31.5'W</geoCoordinate>
), 2 Aug 2002, D. and W. N. Mathis (2♂, 1♀; USNM); Kenai Fjord National Park, Exit Glacier (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="60.195">60°11.7'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-149.59666">149°35.8'W</geoCoordinate>
; wetlands), 30 Jul 2002, D. and W. N. Mathis (1♀; USNM); Kenai River, 
<normalizedToken originalValue="Jim’s">Jim's</normalizedToken>
Landing (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="60.481667">60°28.9'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-150.115">150°06.9'W</geoCoordinate>
), 3 Aug 2002, D. and W. N. Mathis (6♂, 1♀; USNM); Ninilchik (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="60.05">60°03'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-151.67">151°40.2'W</geoCoordinate>
; beach), 2 Jul 2006, D. and W. N. Mathis (2♂, 1♀; USNM); Seward (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="60.138332">60°08.3'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-149.38667">149°23.2'W</geoCoordinate>
), 7 Aug 2003, D. and W. N. Mathis (1♀; USNM); Seward (21 km N; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="60.286667">60°17.2'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-149.34166">149°20.5'W</geoCoordinate>
; Snow River), 31 Jul 2002, D. and W. N. Mathis (28♂, 12♀; USNM); Skilak Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="60.43833">60°26.3'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-150.32333">150°19.4'W</geoCoordinate>
), 3 Aug 2002, D. and W. N. Mathis (4♂, ♀; USNM). Matanuska-Susitna: Eklutna (Knik Arm; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.47">61°28.2'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-149.35667">149°21.4'W</geoCoordinate>
), 7 Aug 2002, D. and W. N. Mathis (2♂, 2♀; USNM); Hatcher Pass (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="61.75">61°45'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-149.23167">149°13.9'W</geoCoordinate>
), 6 Aug 2002, D. and W. N. Mathis (1♂; USNM); Knik River (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.463333">61°27.8'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-148.86">148°51.6'W</geoCoordinate>
), 5 Aug 2002, D. and W. N. Mathis (33♂, 7♀; USNM); Little Willow Creek (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.81">61°48.6'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-150.09666">150°05.8'W</geoCoordinate>
; 50 m), 25 Jul 2011, D. and W. N. Mathis (10♂; USNM); Matanuska (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.541668">61°32.5'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-149.23">149°13.8'W</geoCoordinate>
; rotary trap), 28 Apr-21 May 1945, J. C. Chamberlin (1♂, 1♀; USNM); Palmer (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="61.6">61°36'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-149.11333">149°06.8'W</geoCoordinate>
; jeep trap), 7-13 Jul 1964, K. M. Sommerman (2♂, 6♀; USNM); Palmer (Knik River; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.52">61°31.2'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-148.99">148°59.4'W</geoCoordinate>
), 6 Aug 2002, D. and W. N. Mathis (1♀; USNM); Palmer (Matanuska River; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.608334">61°36.5'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-149.06833">149°04.1'W</geoCoordinate>
), 5-16 Aug 2002, 2012, D. and W. N. Mathis (8♂, 4♀; USNM); Sheep Creek (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.97167">61°58.3'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-150.08333">150°05'W</geoCoordinate>
; 55 m), 10 Aug 2011, D. and W.N. Mathis (2♂, 6♀; USNM); Talkeetna 
<pageBreakToken pageId="20" pageNumber="21" start="start">(</pageBreakToken>
Susitna River; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.323334">61°19.4'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-150.12">150°07.2'W</geoCoordinate>
; 120 m), 10 Aug 2011, D. and W.N. Mathis (8♂, 2♀; USNM); Willow Creek (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.768333">61°46.1'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-150.07">150°04.2'W</geoCoordinate>
; 50 m), 26 Jul 2011, D. and W.N. Mathis (1♂; USNM). Southeast Fairbanks Census Area: Dry Creek Campground, Glenn Highway A-192 (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="63.65333">63°39.2'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-144.36333">144°21.8'W</geoCoordinate>
), 3 Aug 1978, P. H. Arnaud, Jr. (1♀; CAS); Gardiner Creek Camp, Alaska Highway DC 1253 (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="62.858334">62°51.5'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-141.46666">141°28'W</geoCoordinate>
), 5 Aug 1978, P. H. Arnaud, Jr. (1♀; CAS); Gerstle River, Alaska Highway DC 1393 (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="64.05666">64°03.4'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-145.135">145°08.1'W</geoCoordinate>
), 9 Jul 1978, P. H. Arnaud, Jr. (2♂; CAS). Valdez-Cordova (Census Area): Glennallen (32 km W; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="62.09833">62°05.9'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-146.09">146°05.4'W</geoCoordinate>
; 665 m), 27 Jul 2011, D. and W. N. Mathis (8♂, 2♀; USNM); Gulkana River (19.3 km N Glenallen; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="62.268333">62°16.1'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-145.385">145°23.1'W</geoCoordinate>
), 27 Jul-7 Aug 2011, 2012, D. and W. N. Mathis (17♂, 6♀; USNM); Klutina River (mile 101; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.953335">61°57.2'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-145.32167">145°19.3'W</geoCoordinate>
; 315 m), 7 Aug 2012, D. and W. N. Mathis (1♂; USNM); Tolsona Creek State Campground, Glenn Highway, A 173 (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="62.065">62°03.9'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-145.99667">145°59.8'W</geoCoordinate>
), 31 Jul 1978, P. H. Arnaud, Jr. (1♂; CAS); Valdez (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.125">61°07.5'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-146.35834">146°21.5'W</geoCoordinate>
; Ruth Pond), 8 Jul 2006, D. and W. N. Mathis (2♂, 1♀; USNM); Valdez, Valdez Glacier Campground (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="61.116665">61°07'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-146.21">146°12.6'W</geoCoordinate>
), 1 Aug 1978, P. H. Arnaud, Jr. (1♂, 1♀; CAS).
</paragraph>
<paragraph lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
COLORADO. Chaffee: Green Timber Gulch, Cottonwood Lake (6 km W; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.76">38°45.6'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-106.33833">106°20.3'W</geoCoordinate>
), 10-12 Jul 1978, T. W. and W. T. Davies (1♂; CAS); Monarch Pass (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.496666">38°29.8'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-106.325">106°19.5'W</geoCoordinate>
; 2440 m), 21 Jun 1940, A. L. Melander (2♂; USNM). Gunnison: Schofield Pass on Gothic Road (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.015">39°0.9'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-107.04667">107°02.8'W</geoCoordinate>
; 2900 m), 
<pageBreakToken pageId="21" pageNumber="22" start="start">1</pageBreakToken>
Aug 1957, A. H. Sturtevant (5♀; USNM). Rio Grande: South Fork (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.67">37°40.2'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-106.64">106°38.4'W</geoCoordinate>
; 2440 m), 20 Jun 1972, W. W. Wirth (1♂, 1♀; USNM). La Plata: Durango (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.275">37°16.5'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-107.88">107°52.8'W</geoCoordinate>
), 10 Aug 1950, A. H. Sturtevant (5♂; USNM). Larimer: Estes Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.376667">40°22.6'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-105.52167">105°31.3'W</geoCoordinate>
), 13 Jul 1934, A. L. Melander (1♀; ANSP); Hidden Valley, Rocky Mountain National Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.393333">40°23.6'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-105.64667">105°38.8'W</geoCoordinate>
), 8 Aug 1934, A. L. Melander (1♂; ANSP); Virginia Dale (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.955">40°57.3'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-105.35">105°21'W</geoCoordinate>
), 27 Jul 1953, R. R. Dreisbach (1♀; USNM).
</paragraph>
<paragraph pageId="21" pageNumber="22">
IDAHO. Latah: Big Meadow Creek Recreation Area (11.25 km N Troy; 
<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.85">46°51'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-116.745">116°44.7'W</geoCoordinate>
; 915 m; sweeping), 7 Aug 1986, W. J. Turner (1♂; WSU).
</paragraph>
<paragraph pageId="21" pageNumber="22">
MONTANA. Flathead: Glacier National Park, Lake McDonald (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="48.583332">48°35'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-113.92667">113°55.6'W</geoCoordinate>
), 13 Jun 1935, A. L. Melander (1♀; ANSP).
</paragraph>
<paragraph pageId="21" pageNumber="22">
NEBRASKA. Dawes: Chadron (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.708332">42°42.5'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-103.01667">103°01'W</geoCoordinate>
), 20 Aug 1950, A. H. Sturtevant (1♂; USNM).
</paragraph>
<paragraph pageId="21" pageNumber="22">
SOUTH DAKOTA. Custer: Sylvan Lakes (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.843334">42°50.6'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-103.56167">103°33.7'W</geoCoordinate>
), 19 Jul 1924 (1♀; ANSP).
</paragraph>
<paragraph pageId="21" pageNumber="22">
UTAH. Carbon: Deadman Canyon (16 km NE Price; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.695">39°41.7'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-110.73333">110°44'W</geoCoordinate>
; 2055 m), 14 Aug 2008, D. and W. N. Mathis (5♂, 4♀; USNM). Garfield: Grand Staircase-Escalante National Monument: spring off highway 12, Henrieville (12 km E; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.613335">37°36.8'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-111.89667">111°53.8'W</geoCoordinate>
; 2005 m), 2-17 Aug 2000, W. N. Mendel, E. C. Green, M. Moody (1♂; BYU). Grand: Harley Dome (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.173332">39°10.4'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-109.13333">109°08'W</geoCoordinate>
), 13 Aug 1958 (1♀; USNM). Salt Lake: Little Cottonwood Canyon (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.578335">40°34.7'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-111.79667">111°47.8'W</geoCoordinate>
), 23 Aug 1940, A. L. Melander (1♀; USNM). Sanpete: Ephraim (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.36">39°21.6'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-111.58667">111°35.2'W</geoCoordinate>
), 18 Aug 1953, A. H. Sturtevant (1♀; USNM).
</paragraph>
<paragraph pageId="21" pageNumber="22">
WASHINGTON. Island: Keystone Ferry (1.6 km E; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="48.168335">48°10.1'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-122.636665">122°38.2'W</geoCoordinate>
), 13 Aug 1977, R. S. &amp; V. L. Zack (1♂; WSU). Pierce: Mount Rainier National Park, Summerland Trail (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="46.866665">46°52.0'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-121.635">121°38.1'W</geoCoordinate>
), 24 Jul 1924, A. L. Melander (1♀; ANSP). Whitman: Steptoe Canyon (16 km SW Colton; 
<geoCoordinate direction="north" orientation="latitude" precision="92" value="46.451668">46°27.1'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-117.206665">117°12.4'W</geoCoordinate>
; Malaise trap with dry ice), 3 Aug 1974, W. J. Turner (1♂; WSU).
</paragraph>
<paragraph pageId="21" pageNumber="22">
WYOMING. Sublette: Slide Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="43.275">43°16.5'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-109.78167">109°46.9'W</geoCoordinate>
), 14 Aug 1951, A. H. Sturtevant (3♂, 2♀; USNM). Teton: Teton Pass (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="43.498333">43°29.9'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.955">110°57.3'W</geoCoordinate>
), 10 Aug 1951, A. H. Sturtevant (2♂; USNM); Yellowstone National Park, Riverside (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="44.47333">44°28.4'N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.84">110°50.4'W</geoCoordinate>
), 4 Aug 1919, A. L. Melander (1♀; ANSP).
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="distribution">
<paragraph pageId="21" pageNumber="22">Distribution</paragraph>
<paragraph pageId="21" pageNumber="22">
(Fig. 24). Nearctic: Canada (Alberta, British Columbia, Manitoba, Newfoundland, Ontario, Quebec, Saskatchewan, Yukon Territory), United States (Alaska, Colorado, Idaho, Montana, Nebraska, South Dakota, Utah, Washington, Wyoming). Palearctic: Austria, Finland, Iceland, Mongolia (Bayan 
<normalizedToken originalValue="Ölgiy">Oelgiy</normalizedToken>
, Khovd), Sweden.
</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="remarks">
<paragraph pageId="21" pageNumber="22">Remarks.</paragraph>
<paragraph pageId="21" pageNumber="22">This species, like many but not all congeners, is sexually dimorphic, especially the coloration of the face and the mesonotal area from the postpronotum to the base of the wing as noted in the diagnosis. These differences, which were not always recognized, probably was a major factor contributing to this species being described more than once.</paragraph>
<paragraph lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
This species is similar to 
<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa pulchella" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pulchella">Gymnoclasiopa pulchella</taxonomicName>
in having yellowish antennae, maxillary palpi, and foretibiae (sometimes brownish apically) but is distinguished from that species 
<pageBreakToken pageId="22" pageNumber="23" start="start">as</pageBreakToken>
follows (also see key): The anterolateral area of the mesonotum, just mediad of the area from the postpronotum through the notopleuron is densely microtomentose, mostly dull; the lateral mesonotal area from and including the postpronotum and the notopleuron is densely and finely microtomentose (this area in males is brown and is contrasted with whitish to silvery gray microtomentum of the broad, medial portion; the same area in females is concolorous with the medial coloration).
</paragraph>
<caption pageId="22" pageNumber="23">
<paragraph pageId="22" pageNumber="23">
Figures 21-23. 
<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa bohemanni" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bohemanni">Gymnoclasiopa bohemanni</taxonomicName>
(Becker) (Canada. Alberta. Okotoks, Sheep River). 21 epandrium and cerci, posterior view 22 internal structures of male terminalia (aedeagus [shaded], phallapodeme, gonite, hypandrium), ventral view 23 same, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<caption pageId="22" pageNumber="23">
<paragraph pageId="22" pageNumber="23">
Figure 24. Distribution map of 
<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa bohemanni" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bohemanni">Gymnoclasiopa bohemanni</taxonomicName>
(Becker).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>