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<mods:titleid="A6B8D0722F263B8CA5A53F6BE799E51C">A new species of Speonemadus from Portugal, with the revision of the escalerai-group (Coleoptera, Leiodidae)</mods:title>
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;
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<dateid="FFECC9E5666BAB5DB5D16337FE8A3D39"box="[189,348,1626,1653]"pageId="14"pageNumber="15"value="1922-05-16">16 May 1922</date>
(
<bibRefCitationid="EFC392D4666BAB5DB4006337FE313D39"author="Blas M."box="[364,487,1626,1653]"firstAuthor="Blas"journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona"pageId="14"pageNumber="15"refId="ref11559"refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona."title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)"type="book"year="1979">Blas 1979</bibRefCitation>
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<bibRefCitationid="EFC392D4666BAB5DB72C6337FCB03D39"author="Giachino P. M. & Vailati D."box="[576,870,1626,1653]"firstAuthor="Giachino"journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana"pageId="14"pageNumber="15"pagination="1 - 314"part="18"refId="ref12035"refString="Giachino P. M. & Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana " 18: 1 - 314."title="Revisione degli Anemadinae"type="journal article"year="1993">Giachino & Vailati 1993</bibRefCitation>
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); Ronda, Sierra de las Nieves, Cueva del Rejete (
<bibRefCitationid="EFC392D4666BAB5DB74B6313FD1A3DD4"author="Coiffait H."box="[551,716,1661,1688]"firstAuthor="Coiffait"journalOrPublisher="Notes biospeologiques"pageId="14"pageNumber="15"pagination="21 - 24"part="9"refId="ref11682"refString="Coiffait H. 1954. Catopides recoltes en Andalousie. Description de trois formes nouvelles. Notes biospeologiques 9 (1): 21 - 24."title="Catopides recoltes en Andalousie. Description de trois formes nouvelles"type="journal article"year="1954">Coiffait 1954</bibRefCitation>
;
<bibRefCitationid="EFC392D4666BAB5DB7B66313FCA63DD4"author="Ribera"box="[730,880,1662,1688]"firstAuthor="Ribera"pageId="14"pageNumber="15"pagination="123 - 129"refId="ref13567"refString="Ribera C. 1970. Resultados faunisticos de la campana Serrania de Ronda. In: Actas I Congreso Nacional de Espeleologia: 123 - 129. Editorial Ayuntamiento, Barcelona."type="book chapter"year="1970">Ribera 1970</bibRefCitation>
;
<bibRefCitationid="EFC392D4666BAB5DB6126313FC2F3DDB"author="Blas M."box="[894,1017,1661,1688]"firstAuthor="Blas"journalOrPublisher="Congreso Nacional de Espeleologia"pageId="14"pageNumber="15"pagination="157 - 160"part="4"refId="ref11503"refString="Blas M. 1976. Coleopteros cavernicolas del distrito andaluz (Catopidae). Actas 4 Congreso Nacional de Espeleologia: 157 - 160."title="Coleopteros cavernicolas del distrito andaluz (Catopidae). Actas"type="journal article"year="1976">Blas 1976</bibRefCitation>
,
<bibRefCitationid="EFC392D4666BAB5DB16B6313FB923DD4"author="Blas M."box="[1031,1092,1662,1688]"firstAuthor="Blas"journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia"pageId="14"pageNumber="15"pagination="125 - 130"refId="ref11528"refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130."title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)"type="book chapter"year="1977">1977</bibRefCitation>
,
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,
<bibRefCitationid="EFC392D4666BAB5DB1F26313FB0A3DD4"author="Blas M."box="[1182,1244,1662,1688]"firstAuthor="Blas"journalOrPublisher="Memoires de Biospeologie"pageId="14"pageNumber="15"pagination="135 - 140"part="16"refId="ref11653"refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140."title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)"type="journal article"year="1989">1989</bibRefCitation>
;
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;
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); Ronda, Sumidero del Rejete (
<bibRefCitationid="EFC392D4666BAB5DB6B363CCFB8E3DF7"author="Blas M."box="[991,1112,1697,1723]"firstAuthor="Blas"journalOrPublisher="Congreso Nacional de Espeleologia"pageId="14"pageNumber="15"pagination="157 - 160"part="4"refId="ref11503"refString="Blas M. 1976. Coleopteros cavernicolas del distrito andaluz (Catopidae). Actas 4 Congreso Nacional de Espeleologia: 157 - 160."title="Coleopteros cavernicolas del distrito andaluz (Catopidae). Actas"type="journal article"year="1976">Blas 1976</bibRefCitation>
,
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); Sierra de Ronda, Sima E Las Palomas (
<bibRefCitationid="EFC392D4666BAB5DB4D263A9FDEE3D92"author="Blas M."box="[446,568,1732,1758]"firstAuthor="Blas"journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona"pageId="14"pageNumber="15"refId="ref11559"refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona."title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)"type="book"year="1979">Blas 1979</bibRefCitation>
,
<bibRefCitationid="EFC392D4666BAB5DB72863A9FD543D92"author="Blas M."box="[580,642,1732,1758]"firstAuthor="Blas"journalOrPublisher="Memoires de Biospeologie"pageId="14"pageNumber="15"pagination="135 - 140"part="16"refId="ref11653"refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140."title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)"type="journal article"year="1989">1989</bibRefCitation>
;
<bibRefCitationid="EFC392D4666BAB5DB7E263A9FCF63D92"author="Tinaut"box="[654,800,1732,1758]"firstAuthor="Tinaut"pageId="14"pageNumber="15"pagination="3 - 28"refId="ref13778"refString="Tinaut A. 1998. Artropodos terrestres de las cavidades andaluzas. Zoologica Baetica 9: 3 - 28."type="journal article"year="1998">Tinaut 1998</bibRefCitation>
<figureCitationid="1369F3A0666BAB5DB63A6217FC763CD8"box="[854,928,1914,1940]"captionStart="Figs 1 – 5"captionStartId="3.[189,241,1892,1918]"captionTargetBox="[189,1395,1323,1850]"captionTargetId="figure@3.[189,1398,1323,1850]"captionTargetPageId="3"captionText="Figs 1 – 5. Speonemadus of the escalera i- group habitus. 1. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 2. S. angusticollis (Kraatz, 1870) (Rute, Spain). 3. S. bolivari (Jeannel, 1922) (Ardales, Spain). 4. S. breuili (Jeannel, 1922) (Motillas, Spain). 5. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 1 mm."httpUri="https://zenodo.org/record/889348/files/figure.png"pageId="14"pageNumber="15">Fig. 3</figureCitation>
). The Integument light brown, slightly darker on the head. Pubescence fairly long, golden and slightly raised. Head retractable with welldeveloped eyes; pubescence adpressed on its forehead and slightly raised on the clypeus.
ANTENNAE. Long, slender, light brown and extended beyond the basal fourth of elytra; 3rd antennomere slightly longer than the 2nd one; 6th one shorter than 4th and 5th ones, and the latter one as long as the 7th one; 8th almost 1.5 × longer than wide (
<tableCitationid="C6D0DA9E666AAB5CB7E9643FFD0B3A20"box="[645,733,338,364]"captionStart="Table 1"captionStartId="4.[188,257,267,293]"captionTargetBox="[193,1384,385,1378]"captionTargetPageId="4"captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width."httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04"pageId="15"pageNumber="16"tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
(females); with a maximum width towards the middle; basal impressions are not hardly noticeable; side edges regularly arched forward, almost straight and towards the posterior angles that are obtuse and rounded in the vertex (
<taxonomicNameid="4C5294A6666AAB5CB40C665DFD473806"authority="Ana Sofa P.S. Reboleira & Javier Fresneda & José Maria Salgado, 2017"authorityName="Ana Sofa P.S. Reboleira & Javier Fresneda & José Maria Salgado"authorityYear="2017"box="[352,657,816,842]"class="Insecta"family="Leiodidae"genus="Speonemadus"higherTaxonomySource="GBIF"kingdom="Animalia"order="Coleoptera"pageId="15"pageNumber="16"phylum="Arthropoda"rank="species"species="algarvensis"status="sp. nov.">
; basal lamina of median lobe well developed and more than half of the median lobe length; ventral blade of tegmen short, inconsistent and forms the widest part of the aedeagus. In dorsal view, median lobe is bottle-shaped, with converging sides towards the base and narrowed distally; apex tapers and ends in a triangular tip with the sides slightly beaded dorsally (
<figureCitationid="1369F3A0666AAB5CB49A66D1FD82389B"box="[502,596,956,983]"captionStart="Figs 21 – 25"captionStartId="7.[189,241,1930,1956]"captionTargetBox="[252,1351,550,1857]"captionTargetId="figure@7.[189,1398,543,1890]"captionTargetPageId="7"captionText="Figs 21 – 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm."httpUri="https://zenodo.org/record/889354/files/figure.png"pageId="15"pageNumber="16">Fig. 23</figureCitation>
). Parameres robust, only curved forwards; apical parts obliquely truncated on the inner side, where four setae are inserted, three thin, subequal, and a larger one, as well as a thick and well sclerotized small tooth. Inner sac with two longitudinal chains formed by small sclerotized parts and surrounded by two longer chains of fine and sharp spines that come together in the apical region, but do not form a typical sclerotized tooth as in other species.
); first three protarsomeres dilated, being the first almost as wide as the apical tip of the protibia; mesotibia strongly arched and metatibia straight.
Antennae slightly shorter and more robust than in the male, reaching third basal part of elytra; 2nd antennomere is slightly longer than the 3rd one and 5th one longer than 4th and 6th ones; the 8th being clearly longer than wide (
<tableCitationid="C6D0DA9E666AAB5CB4FE6011FE3C3EDA"box="[402,490,1404,1430]"captionStart="Table 1"captionStartId="4.[188,257,267,293]"captionTargetBox="[193,1384,385,1378]"captionTargetPageId="4"captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width."httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04"pageId="15"pageNumber="16"tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
). Elytra elliptical and elongated with notched and serrated apical areas. The sexual dimorphism is clear for all species of this group, as well as the structures in uroventrites 7th and 8th, and spermatheca model.
<emphasisid="B9263337666AAB5CB5D16369FE603D52"bold="true"box="[189,438,1540,1566]"pageId="15"pageNumber="16">Biology and ecology</emphasis>
</heading>
</paragraph>
<paragraphid="8BEDEF25666AAB5CB5D1635FFAEE3D3C"blockId="15.[189,1398,1540,1648]"pageId="15"pageNumber="16">Subterranean species, found in deep areas of scattered caves, under stones or on calcite mantles. Groups of adults and larvae have been found in small accumulations of organic matter, such as bat guano.</paragraph>
is endemic to several karst caves located in a narrow strip within the province of Málaga, between the Serrania de Ronda, at the western limit, and the karst areas at its border with the province of
province are located to the north and east of the massif of the Sierra Nevada (
<figureCitationid="1369F3A0666AAB5CB0646242FA893C05"box="[1288,1375,1839,1865]"captionStart="Fig. 32"captionStartId="18.[189,232,1955,1981]"captionTargetBox="[189,1398,986,1883]"captionTargetId="figure@18.[189,1398,986,1914]"captionTargetPageId="18"captionText="Fig. 32. Distribution of Speonemadus of the escalerai - group: S. algarvensis sp. nov., S. angusticollis (Kraatz, 1870), S. bolivari (Jeannel, 1922), S. breuili (Jeannel, 1922) and S. escalerai (Uhagón, 1898)."httpUri="https://zenodo.org/record/889358/files/figure.png"pageId="15"pageNumber="16">Fig. 32</figureCitation>
