166 lines
15 KiB
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166 lines
15 KiB
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<document ID-DOI="10.1111/j.1096-3642.2004.00121.x" ID-GBIF-Dataset="b6188841-cdc6-42fa-bb29-ac39d113dd72" ID-ISSN="0024-4082" ID-Zenodo-Dep="4687299" checkinTime="1618408847792" checkinUser="carolina" docAuthor="Dzik, Jerzy, Ivantsov, Andrey Yu. & Deulin, Yuriy V." docDate="2004" docId="03DFC827FFE5FFC1FECDF374FC99FC5C" docLanguage="en" docName="ZoolJLinnSoc.142.83-90.pdf" docOrigin="Zoological Journal of the Linnean Society 142 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00121.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.1:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="1" docTitle="Archangeliphausia Dzik & Ivantsov & Deulin 2004, GEN. NOV." docType="treatment" docVersion="6" lastPageNumber="86" masterDocId="FFE6B05FFFE6FFC2FFC2F13BFFE3FF84" masterDocTitle="Oldest shrimp and associated phyllocarid from the Lower Devonian of northern Russia" masterLastPageNumber="90" masterPageNumber="83" pageNumber="86" updateTime="1638552323672" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Oldest shrimp and associated phyllocarid from the Lower Devonian of northern Russia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dzik, Jerzy</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ivantsov, Andrey Yu.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Deulin, Yuriy V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2004</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2004-09-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>142</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>83</mods:start>
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<mods:end>90</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00121.x</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1111/j.1096-3642.2004.00121.x</mods:identifier>
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<mods:identifier type="GBIF-Dataset">b6188841-cdc6-42fa-bb29-ac39d113dd72</mods:identifier>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4687299</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5748183" ID-GBIF-Taxon="190875564" ID-Zenodo-Dep="5748183" LSID="urn:lsid:plazi:treatment:03DFC827FFE5FFC1FECDF374FC99FC5C" httpUri="http://treatment.plazi.org/id/03DFC827FFE5FFC1FECDF374FC99FC5C" lastPageNumber="86" pageId="3" pageNumber="86">
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<subSubSection box="[271,656,591,615]" pageId="3" pageNumber="86" type="nomenclature">
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<paragraph blockId="3.[176,751,495,615]" box="[271,656,591,615]" pageId="3" pageNumber="86">
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<heading box="[271,656,591,615]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="86" reason="2">
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<taxonomicName authority="Dzik & Ivantsov & Deulin, 2004" authorityName="Dzik & Ivantsov & Deulin" authorityYear="2004" box="[271,529,591,615]" class="Malacostraca" family="Anthracophausiidae" genus="Archangeliphausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus" status="GEN. NOV.">ARCHANGELIPHAUSIA</taxonomicName>
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<emphasis bold="true" box="[536,656,591,615]" pageId="3" pageNumber="86">
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<taxonomicNameLabel box="[536,656,591,615]" pageId="3" pageNumber="86" rank="genus">GEN. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[156,525,640,661]" pageId="3" pageNumber="86" type="type_taxon">
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<paragraph blockId="3.[156,525,640,661]" box="[156,525,640,661]" pageId="3" pageNumber="86">
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<emphasis box="[156,431,640,661]" italics="true" pageId="3" pageNumber="86">
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<typeStatus box="[156,209,640,661]" pageId="3" pageNumber="86">Type</typeStatus>
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species:
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<taxonomicName authority="Dzik & Ivantsov & Deulin, 2004" authorityName="Dzik & Ivantsov & Deulin" authorityYear="2004" box="[314,431,640,661]" class="Malacostraca" family="Eocaridacea (awaiting allocation)" genus="Archangeliphausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">A. spinosa</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[440,525,640,661]" pageId="3" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="86" type="diagnosis">
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<paragraph blockId="3.[156,771,685,769]" pageId="3" pageNumber="86">
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<emphasis box="[156,275,686,707]" italics="true" pageId="3" pageNumber="86">Diagnosis:</emphasis>
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A generalized anthracophausiid with abdominal pleura bearing a single small spine on their ventral margin.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="86" type="etymology">
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<paragraph blockId="3.[156,771,793,845]" pageId="3" pageNumber="86">
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<emphasis box="[156,282,793,814]" italics="true" pageId="3" pageNumber="86">Etymology:</emphasis>
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Derived from the latinized name of
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<collectingRegion box="[156,303,823,845]" country="Russia" name="Arkhangel'sk" pageId="3" pageNumber="86">Arkhangelsk</collectingRegion>
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and
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<emphasis box="[362,451,823,844]" italics="true" pageId="3" pageNumber="86">phausis</emphasis>
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(Greek = shine).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="86" type="discussion">
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<paragraph blockId="3.[156,772,869,1903]" pageId="3" pageNumber="86">
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<emphasis box="[156,266,869,890]" italics="true" pageId="3" pageNumber="86">Affinities:</emphasis>
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Brooks (1962) indicated a recessed ‘eye socket’ of the proximal segment of peduncle of antennules as the diagnostic character of his
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<taxonomicName authorityName="BROOKS" authorityYear="1962" class="Malacostraca" family="Anthracophausiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="family">Anthracophausiidae</taxonomicName>
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. This feature is not represented in the new genus, which is quite generalized in this respect and may have been anatomically close to the Devonian eocaridids, as indicated also by the long abdominal pleura. However, the more general aspects of
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<taxonomicName authorityName="Peach" authorityYear="1908" class="Malacostraca" family="Anthracophausiidae" genus="Anthracophausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="86">Anthracophausia</emphasis>
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</taxonomicName>
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listed by Brooks (1962) - weak sclerotization of generally smooth carapace, margins reinforced with an unusually narrow band, a short rostrum, and lateral flattening of the body due to compression - fit well the Russian material.
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</paragraph>
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<paragraph blockId="3.[156,772,869,1903]" pageId="3" pageNumber="86">
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The Carboniferous species of
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<taxonomicName authorityName="Peach" authorityYear="1908" box="[575,771,1268,1289]" class="Malacostraca" family="Anthracophausiidae" genus="Anthracophausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
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<emphasis box="[575,771,1268,1289]" italics="true" pageId="3" pageNumber="86">Anthracophausia</emphasis>
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</taxonomicName>
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reveal dramatically different outlines of abdominal pleura from those in the Russian species. In
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<taxonomicName authority="Brooks, 1962" authorityName="Brooks" authorityYear="1962" class="Malacostraca" family="Anthracophausiidae" genus="Anthracophausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="strongi">
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<emphasis box="[662,771,1330,1351]" italics="true" pageId="3" pageNumber="86">A. strongi</emphasis>
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Brooks, 1962
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</taxonomicName>
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, from the Late Carboniferous Mazon Creek fauna of Illinois, the pleural lobes of the abdominal tergites are broadly rounded. In
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<taxonomicName box="[574,712,1421,1443]" class="Malacostraca" family="Anthracophausiidae" genus="Anthracophausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="dunsiana">
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<emphasis box="[574,712,1421,1443]" italics="true" pageId="3" pageNumber="86">A. dunsiana</emphasis>
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</taxonomicName>
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from the Early Carboniferous Glencartholm Volcanic Beds of Scotland they narrow to form a sharp apex (
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<bibRefCitation author="Schram FR" pageId="3" pageNumber="86" pagination="1 - 129" refId="ref4882" refString="Schram FR. 1979. British Carboniferous Malacostraca. Fieldiana Geology 40: 1 - 129." type="journal article" year="1979">Schram, 1979</bibRefCitation>
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). In the new species, rounded lobes are armed with short spines, which seems to be enough to substantiate its taxonomic distinction. Another possible difference between the Russian form and the Carboniferous species of
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<taxonomicName authorityName="Peach" authorityYear="1908" box="[357,553,1636,1657]" class="Malacostraca" family="Anthracophausiidae" genus="Anthracophausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
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<emphasis box="[357,553,1636,1657]" italics="true" pageId="3" pageNumber="86">Anthracophausia</emphasis>
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</taxonomicName>
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is the increasingly posterior orientation of the abdominal pleural lobes towards the telson (although not easily discernible because of strong flattening of the specimens). In fact, the faint parabolic lines visible on the
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<typeStatus box="[635,737,1759,1780]" pageId="3" pageNumber="86">paratype</typeStatus>
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of
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<taxonomicName authorityName="Brooks" authorityYear="1962" box="[156,265,1790,1811]" class="Malacostraca" family="Anthracophausiidae" genus="Anthracophausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="strongi">
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<emphasis box="[156,265,1790,1811]" italics="true" pageId="3" pageNumber="86">A. strongi</emphasis>
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</taxonomicName>
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(Brooks, 1962; pl. 48: 3) may also indicate a similar shape of the posterior pleural lobes in that species. Some gradient in the shape of pleural lobes is also observable in
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<taxonomicName box="[317,455,1881,1903]" class="Malacostraca" family="Anthracophausiidae" genus="Anthracophausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="dunsiana">
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<emphasis box="[317,455,1881,1903]" italics="true" pageId="3" pageNumber="86">A. dunsiana</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="3.[820,1436,195,984]" pageId="3" pageNumber="86">
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All these distinguishing characters are probably primitive (plesiomorphic) and the new genus is probably transitional between the benthic
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<taxonomicName authorityName="BROOKS" authorityYear="1962" box="[1249,1385,256,278]" class="Malacostraca" family="Eocarididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="86" phylum="Arthropoda" rank="family">Eocarididae</taxonomicName>
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and typical Carboniferous
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<taxonomicName authorityName="BROOKS" authorityYear="1962" box="[1094,1328,287,309]" class="Malacostraca" family="Anthracophausiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="family">Anthracophausiidae</taxonomicName>
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. It may represent the beginning of the lineage of
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<taxonomicName authorityName="Peach" authorityYear="1908" class="Malacostraca" family="Anthracophausiidae" genus="Anthracophausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="86">Anthracophausia</emphasis>
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</taxonomicName>
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. Because of the significant time and morphological distance it seems practical, however, to separate them at the generic level.
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</paragraph>
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<paragraph blockId="3.[820,1436,195,984]" pageId="3" pageNumber="86">
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The
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<taxonomicName authorityName="BROOKS" authorityYear="1962" box="[898,1129,440,462]" class="Malacostraca" family="Anthracophausiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eocaridacea" pageId="3" pageNumber="86" phylum="Arthropoda" rank="family">Anthracophausiidae</taxonomicName>
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probably gave rise to the Recent euphausiaceans, closest to the ancestry of Eumalacostraca among the extant orders (
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<bibRefCitation author="Jarman SN & Nicol S & Elliott NG & McMinn A" pageId="3" pageNumber="86" pagination="26 - 36" refId="ref4641" refString="Jarman SN, Nicol S, Elliott NG, McMinn A. 2000. 28 S rDNA evolution in the Eumalacostraca and the phylogenetic position of krill. Molecular Phylogenetics and Evolution 17: 26 - 36." type="journal article" year="2000">
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Jarman
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<emphasis box="[820,882,532,554]" italics="true" pageId="3" pageNumber="86">et al.,</emphasis>
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2000
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</bibRefCitation>
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). The identified distinction of the anthracophausiids in respect to the euphausiaceans is invariably connected with their basal position in the evolutionary tree: the wide telson, lack of hinge-like connection between the first pleotergite and the carapace, and ventrally extended lobes of the carapace.
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</paragraph>
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<paragraph blockId="3.[820,1436,195,984]" pageId="3" pageNumber="86">
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The Late Carboniferous anthracophausiids are considered near-shore marine filter feeders (
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<bibRefCitation author="Schram FR" pageId="3" pageNumber="86" pagination="126 - 137" refId="ref4922" refString="Schram FR. 1981. Late Paleozoic crustacean communities. Journal of Paleontology 55: 126 - 137." type="journal article" year="1981">Schram, 1981</bibRefCitation>
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). Offshore eumalacostracan communities are inadequately known, with the available evidence restricted to the British late Early Carboniferous, the low diversity community being represented there by schooling species preserved in great numbers of individuals probably as an effect of mass killing (
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<bibRefCitation author="Schram FR" pageId="3" pageNumber="86" pagination="126 - 137" refId="ref4922" refString="Schram FR. 1981. Late Paleozoic crustacean communities. Journal of Paleontology 55: 126 - 137." type="journal article" year="1981">Schram, 1981</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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