treatments-xml/data/03/C5/89/03C58923FFA96127B69BF9DCFC64FF2B.xml

<document ID-DOI="10.1098/rsos.200092" ID-GBIF-Dataset="1b0ccc7a-833e-4706-bcad-b34c9a7f441a" ID-PMC="PMC7277287" ID-PubMed="32537217" ID-Zenodo-Dep="3831900" approvalRequired="73" approvalRequired_for_illustrations="6" approvalRequired_for_textStreams="66" approvalRequired_for_treatments="1" checkinTime="1589793152131" checkinUser="plazi" docAuthor="Weijola, Valter, Vahtera, Varpu, Koch, André, Schmitz, Andreas &amp; Kraus, Fred" docDate="2020" docId="03C58923FFA96127B69BF9DCFC64FF2B" docLanguage="en" docName="rsos.200092.pdf.imd" docOrigin="Royal Society Open Science 7 (5)" docSource="http://dx.doi.org/10.1098/rsos.200092" docStyle="DocumentStyle{}" docTitle="Varanus bennetti Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="66" masterDocId="FFFCF15BFFBB6130B60FFFFFFF9CFF83" masterDocTitle="Taxonomy of Micronesian monitors (Reptilia: Squamata: Varanus): endemic status of new species argues for caution in pursuing eradication plans" masterLastPageNumber="28" masterPageNumber="1" pageNumber="61" updateTime="1668144117192" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Taxonomy of Micronesian monitors (Reptilia: Squamata: Varanus): endemic status of new species argues for caution in pursuing eradication plans</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Weijola, Valter</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-6907-0619</mods:nameIdentifier>
<mods:affiliation>Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Turku, Finland</mods:affiliation>
<mods:nameIdentifier type="email">vsawei@utu.fi</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vahtera, Varpu</mods:namePart>
<mods:affiliation>0000 - 0002 - 6710 - 6358</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Koch, André</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-2426-1494</mods:nameIdentifier>
<mods:affiliation>Zoological Research Museum Alexander Koenig, Adenauerallee 160, DE- 53113 Bonn, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schmitz, Andreas</mods:namePart>
<mods:affiliation>Natural History Museum of Geneva, Department of Herpetology and Ichthyology, C. P. 6434, CH- 1211 Geneva 6, Switzerland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kraus, Fred</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4194-4959</mods:nameIdentifier>
<mods:affiliation>Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, USA</mods:affiliation>
</mods:name>
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<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-05-13</mods:number>
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<mods:number>7</mods:number>
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<mods:number>5</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3850213" ID-GBIF-Taxon="164167086" ID-Zenodo-Dep="3850213" LSID="urn:lsid:zoobank.org:act:88D49BE5-3AED-4948-95BD-55D322EB57FD" httpUri="http://treatment.plazi.org/id/03C58923FFA96127B69BF9DCFC64FF2B" lastPageId="23" lastPageNumber="66" pageId="18" pageNumber="61">
<emphasis bold="true" pageId="18" pageNumber="61">
<subSubSection box="[148,664,1571,1604]" pageId="18" pageNumber="61" type="nomenclature">
<paragraph blockId="18.[148,664,1571,1604]" box="[148,664,1571,1604]" pageId="18" pageNumber="61">
<heading bold="true" box="[148,664,1571,1604]" fontSize="14" level="3" pageId="18" pageNumber="61" reason="0">
<emphasis bold="true" box="[148,664,1571,1604]" pageId="18" pageNumber="61">
3.5. 
<taxonomicName authority="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus, 2020" authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[202,397,1571,1604]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="61" phylum="Chordata" rank="species" species="bennetti" status="sp. nov.">
<emphasis bold="true" box="[202,397,1571,1604]" italics="true" pageId="18" pageNumber="61">Varanus bennetti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[410,502,1571,1604]" pageId="18" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
<figureCitation box="[515,664,1571,1604]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStart-2="Figure 10" captionStart-3="Figure 11" captionStart-4="Figure 12" captionStartId-0="19.[148,206,1288,1312]" captionStartId-1="20.[148,206,590,614]" captionStartId-2="20.[148,206,1147,1171]" captionStartId-3="21.[148,206,627,651]" captionStartId-4="22.[148,206,1112,1136]" captionTargetBox-0="[375,1016,117,1265]" captionTargetBox-2="[352,1032,670,1123]" captionTargetBox-3="[322,1078,113,604]" captionTargetBox-4="[390,995,113,1089]" captionTargetId-0="figure@19.[376,1017,118,1255]" captionTargetId-2="figure@20.[352,1033,670,1124]" captionTargetId-3="figure@21.[322,1079,113,604]" captionTargetId-4="figure@22.[390,995,113,1090]" captionTargetPageId-0="19" captionTargetPageId-2="20" captionTargetPageId-3="21" captionTargetPageId-4="22" captionText-0="Figure 8. (a) Dorsolateral and ventral aspects of the holotype (USNM 507504) of Varanus bennetti sp. nov. (b) Lateral profile of the head of the holotype (USNM 507504) of Varanus bennetti sp. nov." captionText-1="Figure 9. Varanus bennetti sp. nov., Rock Islands, Palau (photo by Thibaud Aronson)." captionText-2="Figure 10. Varanus bennetti sp. nov., Ngarchelong, Palau (photo by Thibaud Aronson)." captionText-3="Figure 11. Subadult Varanus bennetti sp. nov., Losiep Island, Federated States of Micronesia (photo by James Reardon)." captionText-4="Figure 12. Adult Varanus bennetti sp. nov., Losiep Island, Federated States of Micronesia (photo by James Reardon)." figureDoi-0="http://doi.org/10.5281/zenodo.3831914" figureDoi-2="http://doi.org/10.5281/zenodo.3831916" figureDoi-3="http://doi.org/10.5281/zenodo.3831918" figureDoi-4="http://doi.org/10.5281/zenodo.3831920" httpUri-0="https://zenodo.org/record/3831914/files/figure.png" httpUri-2="https://zenodo.org/record/3831916/files/figure.png" httpUri-3="https://zenodo.org/record/3831918/files/figure.png" httpUri-4="https://zenodo.org/record/3831920/files/figure.png" pageId="18" pageNumber="61">figures 8–12</figureCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="61" type="materials_examined">
<paragraph blockId="18.[148,301,1630,1659]" box="[148,301,1630,1659]" pageId="18" pageNumber="61">
<heading bold="true" box="[148,301,1630,1659]" fontSize="12" level="4" pageId="18" pageNumber="61" reason="0">
<emphasis bold="true" box="[148,301,1630,1659]" pageId="18" pageNumber="61">
3.5.1. 
<typeStatus box="[213,301,1630,1659]" pageId="18" pageNumber="61">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[148,1244,1684,1738]" pageId="18" pageNumber="61">
<materialsCitation ID-GBIF-Occurrence="2620456312" collectingDate="1996-07-31" collectionCode="USNM" collectorName="Ronald Crombie" country="Palau" location="Ngaramasch village" municipality="Ngeaur Island" pageId="18" pageNumber="61" specimenCode="USNM 507504" specimenCount="1">
<specimenCode box="[148,301,1684,1706]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="61" type="Museum">USNM 507504</specimenCode>
(
<figureCitation box="[315,426,1684,1706]" captionStart="Figure 8" captionStartId="19.[148,206,1288,1312]" captionTargetBox="[375,1016,117,1265]" captionTargetId="figure@19.[376,1017,118,1255]" captionTargetPageId="19" captionText="Figure 8. (a) Dorsolateral and ventral aspects of the holotype (USNM 507504) of Varanus bennetti sp. nov. (b) Lateral profile of the head of the holotype (USNM 507504) of Varanus bennetti sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3831914" httpUri="https://zenodo.org/record/3831914/files/figure.png" pageId="18" pageNumber="61">
figure 8 
<emphasis box="[399,409,1684,1705]" italics="true" pageId="18" pageNumber="61">a</emphasis>
, 
<emphasis box="[415,426,1684,1705]" italics="true" pageId="18" pageNumber="61">b</emphasis>
</figureCitation>
), collected by 
<collectorName box="[581,757,1684,1706]" pageId="18" pageNumber="61">Ronald Crombie</collectorName>
, south of 
<location LSID="urn:lsid:plazi:treatment:03C58923FFA96127B69BF9DCFC64FF2B:8EB36EEEFFA96122B551F96BFBAEF929" box="[862,1074,1684,1706]" country="Palau" municipality="Ngeaur Island" name="Ngaramasch village" pageId="18" pageNumber="61">Ngaramasch village</location>
, 
<collectingMunicipality box="[1086,1240,1684,1706]" pageId="18" pageNumber="61">Ngeaur Island</collectingMunicipality>
, 
<collectingCountry box="[148,209,1716,1738]" name="Palau" pageId="18" pageNumber="61">Palau</collectingCountry>
, 
<date box="[221,354,1716,1738]" pageId="18" pageNumber="61" value="1996-07-31">
<collectingDate box="[221,354,1716,1738]" pageId="18" pageNumber="61" value="1996-07-31">
31 July 
<date box="[303,354,1716,1738]" pageId="18" pageNumber="61" value="1996" year="1996">1996</date>
</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph blockId="18.[148,306,1789,1818]" box="[148,306,1789,1818]" pageId="18" pageNumber="61">
<heading bold="true" box="[148,306,1789,1818]" fontSize="12" level="4" pageId="18" pageNumber="61" reason="0">
<emphasis bold="true" box="[148,306,1789,1818]" pageId="18" pageNumber="61">
3.5.2. 
<typeStatus box="[213,306,1789,1818]" pageId="18" pageNumber="61">Paratypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[148,1244,1842,1960]" pageId="18" pageNumber="61">
<materialsCitation ID-GBIF-Occurrence="2620456310" box="[148,621,1842,1864]" collectionCode="USNM" country="Palau" location="Ngeaur Island" pageId="18" pageNumber="61" specimenCode="USNM 514125, 521719" specimenCount="1">
<specimenCode box="[148,301,1842,1864]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="61" type="Museum">USNM 514125</specimenCode>
, 
<specimenCode box="[313,386,1842,1864]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="61" type="Museum">521719</specimenCode>
, 
<location LSID="urn:lsid:plazi:treatment:03C58923FFA96127B69BF9DCFC64FF2B:8EB36EEEFFA96122B782F8CDFDBAF8CB" box="[397,550,1842,1864]" country="Palau" name="Ngeaur Island" pageId="18" pageNumber="61">Ngeaur Island</location>
, 
<collectingCountry box="[561,621,1842,1864]" name="Palau" pageId="18" pageNumber="61">Palau</collectingCountry>
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="2620456307" collectingDate="60-12-25" collectingDateMax="86-01-13" collectingDateMin="60-12-25" collectionCode="USNM, ZMH" country="Palau" location="Losiep Island" municipality="Sarigan Island" pageId="18" pageNumber="61" specimenCode="USNM 495369-70, USNM 212494, AMNH 00624-25, BMNH 98.5, SMF 32808-09, ZMB 17520-22, 7619, R-4727, AMNH 70652-53, USNM 495369" specimenCount="1" stateProvince="Koror">
<specimenCode box="[633,822,1842,1864]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="61" type="Museum">USNM 495369–70</specimenCode>
, 
<collectingCountry box="[833,894,1842,1864]" name="Palau" pageId="18" pageNumber="61">Palau</collectingCountry>
. Commonwealth of the Northern Mariana Islands: 
<collectingMunicipality box="[333,484,1874,1896]" pageId="18" pageNumber="61">Sarigan Island</collectingMunicipality>
(
<specimenCode box="[500,656,1874,1896]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="61" type="Museum">USNM 212494</specimenCode>
); 
<collectingCountry box="[676,1001,1874,1896]" name="Micronesia" pageId="18" pageNumber="61">Federated States of Micronesia</collectingCountry>
: 
<location LSID="urn:lsid:plazi:treatment:03C58923FFA96127B69BF9DCFC64FF2B:8EB36EEEFFA96122B5FAF8ADFB1FF8EB" box="[1013,1155,1874,1896]" country="Palau" municipality="Sarigan Island" name="Losiep Island" pageId="18" pageNumber="61" stateProvince="Koror">Losiep Island</location>
(
<collectionCode box="[1171,1244,1874,1895]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="61" type="Museum">USNM</collectionCode>
<date box="[148,222,1906,1928]" pageId="18" pageNumber="61" value="60-12-25">
<collectingDate box="[148,222,1906,1928]" pageId="18" pageNumber="61" value="60-12-25">122560</collectingDate>
</date>
), Yap Island (
<specimenCode box="[369,554,1906,1928]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="61">AMNH 00624–25</specimenCode>
, 
<specimenCode box="[566,695,1906,1928]" collectionCode="BMNH" pageId="18" pageNumber="61">BMNH 98.5</specimenCode>
.27.1, 
<specimenCode box="[753,905,1906,1928]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="18" pageNumber="61">SMF 32808–09</specimenCode>
, 
<collectionCode box="[917,990,1906,1927]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="61" type="Museum">USNM</collectionCode>
<date box="[998,1071,1906,1928]" pageId="18" pageNumber="61" value="86-01-13">
<collectingDate box="[998,1071,1906,1928]" pageId="18" pageNumber="61" value="86-01-13">130186</collectingDate>
</date>
, 
<specimenCode box="[1083,1240,1906,1928]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="18" pageNumber="61" type="Museum">ZMB 17520–22</specimenCode>
, 
<specimenCode box="[148,195,1938,1960]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="18" pageNumber="61" type="Museum">7619</specimenCode>
and 
<collectionCode box="[247,305,1938,1959]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="18" pageNumber="61" type="Museum">ZMH</collectionCode>
<specimenCode box="[311,385,1938,1960]" collectionCode="R-" pageId="18" pageNumber="61">R-4727</specimenCode>
); 
<collectingCountry box="[400,461,1938,1960]" name="Palau" pageId="18" pageNumber="61">Palau</collectingCountry>
: 
<collectingRegion box="[470,530,1938,1959]" country="Palau" name="Koror" pageId="18" pageNumber="61">Koror</collectingRegion>
Island (
<specimenCode box="[613,796,1938,1960]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="61">AMNH 70652–53</specimenCode>
), Ngcheangel Atoll (
<specimenCode box="[1006,1160,1938,1960]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="61" type="Museum">USNM 495369</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</emphasis>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3831914" ID-Zenodo-Dep="3831914" httpUri="https://zenodo.org/record/3831914/files/figure.png" pageId="19" pageNumber="62" startId="19.[148,206,1288,1312]" targetBox="[375,1016,117,1265]" targetPageId="19">
<paragraph blockId="19.[148,1244,1288,1344]" pageId="19" pageNumber="62">
Figure 8. (
<emphasis bold="true" box="[244,254,1288,1312]" italics="true" pageId="19" pageNumber="62">a</emphasis>
) Dorsolateral and ventral aspects of the holotype (USNM 507504) of 
<taxonomicName authority="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus, 2020" authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[830,967,1288,1312]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="bennetti" status="sp. nov.">
<emphasis bold="true" box="[830,967,1288,1312]" italics="true" pageId="19" pageNumber="62">Varanus bennetti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[974,1039,1288,1312]" pageId="19" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis bold="true" box="[1051,1061,1288,1312]" italics="true" pageId="19" pageNumber="62">b</emphasis>
) Lateral profile of the head of the holotype (USNM 507504) of 
<taxonomicName authority="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus, 2020" authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[503,642,1320,1344]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="bennetti" status="sp. nov.">
<emphasis bold="true" box="[503,642,1320,1344]" italics="true" pageId="19" pageNumber="62">Varanus bennetti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[651,718,1320,1344]" pageId="19" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection pageId="19" pageNumber="62" type="diagnosis">
<paragraph blockId="19.[148,305,1403,1432]" box="[148,305,1403,1432]" pageId="19" pageNumber="62">
<heading bold="true" box="[148,305,1403,1432]" fontSize="12" level="4" pageId="19" pageNumber="62" reason="0">
<emphasis bold="true" box="[148,305,1403,1432]" pageId="19" pageNumber="62">3.5.3. Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[148,1244,1456,1638]" pageId="19" pageNumber="62">
<taxonomicName authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[148,324,1456,1478]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="bennetti">
<emphasis box="[148,324,1456,1478]" italics="true" pageId="19" pageNumber="62">Varanus bennetti</emphasis>
</taxonomicName>
can be distinguished from all other members of 
<taxonomicName box="[920,1071,1457,1478]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="69" phylum="Chordata" rank="subGenus" subGenus="Euprepiosaurus">
<emphasis box="[920,1071,1457,1478]" italics="true" pageId="19" pageNumber="62">Euprepiosaurus</emphasis>
</taxonomicName>
by its unique combination of (i) dorsum black and evenly speckled with yellow scales, sometimes arranged in small groups of yellow scales, (ii) tongue dark blue/grey, (iii) venter cream coloured with pale grey crossbands, (iv) tail exceptionally long (F/SVL mean = 1.76, range = 1.60–1.89), high XY scale counts (148–160), (v) a clear yellow temporal stripe present in about half of the studied specimens, and (vi), in life, peach colouring on the throat.
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="63" pageId="19" pageNumber="62" type="discussion">
<paragraph blockId="19.[148,740,1693,1722]" box="[148,740,1693,1722]" pageId="19" pageNumber="62">
<heading bold="true" box="[148,740,1693,1722]" fontSize="12" level="4" pageId="19" pageNumber="62" reason="0">
<emphasis bold="true" box="[148,740,1693,1722]" pageId="19" pageNumber="62">
3.5.4. Comparisons with other members of 
<taxonomicName box="[595,740,1693,1722]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="69" phylum="Chordata" rank="subGenus" subGenus="Euprepiosaurus">
<emphasis bold="true" box="[595,740,1693,1722]" italics="true" pageId="19" pageNumber="62">Euprepiosaurus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[148,1245,1746,1960]" lastBlockId="20.[148,1244,1213,1491]" lastPageId="20" lastPageNumber="63" pageId="19" pageNumber="62">
<taxonomicName authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[148,318,1746,1768]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="bennetti">
<emphasis box="[148,318,1746,1768]" italics="true" pageId="19" pageNumber="62">Varanus bennetti</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName box="[619,775,1746,1768]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="caerulivirens">
<emphasis box="[619,775,1746,1768]" italics="true" pageId="19" pageNumber="62">V. caerulivirens</emphasis>
</taxonomicName>
, 
<emphasis box="[790,998,1746,1768]" italics="true" pageId="19" pageNumber="62">
<taxonomicName box="[790,867,1746,1768]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="colei">V. colei</taxonomicName>
, 
<taxonomicName baseAuthorityName="Meyer" baseAuthorityYear="1874" box="[880,998,1746,1768]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="doreanus">V. doreanus</taxonomicName>
</emphasis>
, 
<taxonomicName box="[1014,1109,1746,1768]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="finschi">
<emphasis box="[1014,1109,1746,1768]" italics="true" pageId="19" pageNumber="62">V. finschi</emphasis>
</taxonomicName>
, 
<taxonomicName box="[1124,1238,1746,1768]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="jobiensis">
<emphasis box="[1124,1238,1746,1768]" italics="true" pageId="19" pageNumber="62">V. jobiensis</emphasis>
</taxonomicName>
, 
<taxonomicName box="[148,289,1778,1800]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="juxtindicus">
<emphasis box="[148,289,1778,1800]" italics="true" pageId="19" pageNumber="62">V. juxtindicus</emphasis>
</taxonomicName>
, 
<taxonomicName box="[304,411,1778,1800]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="melinus">
<emphasis box="[304,411,1778,1800]" italics="true" pageId="19" pageNumber="62">V. melinus</emphasis>
</taxonomicName>
, 
<taxonomicName box="[426,497,1778,1800]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="obor">
<emphasis box="[426,497,1778,1800]" italics="true" pageId="19" pageNumber="62">V. obor</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Weijola, Donnellan &amp; Lindqvist" authorityYear="2016" box="[512,621,1779,1800]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="semotus">
<emphasis box="[512,621,1779,1800]" italics="true" pageId="19" pageNumber="62">V. semotus</emphasis>
</taxonomicName>
and 
<taxonomicName box="[679,793,1779,1800]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="yuwonoi">
<emphasis box="[679,793,1779,1800]" italics="true" pageId="19" pageNumber="62">V. yuwonoi</emphasis>
</taxonomicName>
by having a fully dark blue/grey tongue rather than a (at least partly) pink or yellow tongue; from 
<taxonomicName box="[795,966,1810,1832]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="cerambonensis">
<emphasis box="[795,966,1810,1832]" italics="true" pageId="19" pageNumber="62">V. cerambonensis</emphasis>
</taxonomicName>
by the absence of dorsal cross-bands and its comparatively longer tail (F/SVL: 
<quantity box="[764,893,1842,1864]" metricMagnitude="-2" metricUnit="m" metricValue="4.4323" metricValueMax="4.8006" metricValueMin="4.064" pageId="19" pageNumber="62" unit="in" value="1.745" valueMax="1.89" valueMin="1.6">1.60–1.89 in</quantity>
<taxonomicName authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[907,1020,1842,1863]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="bennetti">
<emphasis box="[907,1020,1842,1863]" italics="true" pageId="19" pageNumber="62">V. bennetti</emphasis>
</taxonomicName>
versus 
<quantity box="[1114,1244,1842,1864]" metricMagnitude="-2" metricUnit="m" metricValue="3.7211" metricValueMax="4.0894" metricValueMin="3.3528" pageId="19" pageNumber="62" unit="in" value="1.465" valueMax="1.61" valueMin="1.32">1.32–1.61 in</quantity>
<taxonomicName box="[148,320,1874,1896]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="cerambonensis">
<emphasis box="[148,320,1874,1896]" italics="true" pageId="19" pageNumber="62">V. cerambonensis</emphasis>
</taxonomicName>
); from 
<taxonomicName box="[406,526,1874,1896]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="douarrha">
<emphasis box="[406,526,1874,1896]" italics="true" pageId="19" pageNumber="62">V. douarrha</emphasis>
</taxonomicName>
by the absence of dorsal ocelli and its comparatively longer tail (F/SVL = 1.60–1.89 versus 1.32–1.61); from 
<taxonomicName baseAuthorityName="Daudin" baseAuthorityYear="1802" box="[634,739,1906,1928]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="indicus">
<emphasis box="[634,739,1906,1928]" italics="true" pageId="19" pageNumber="62">V. indicus</emphasis>
</taxonomicName>
by the presence of a yellow temporal stripe in much of the population, generally higher scale counts (X: 41–48 versus 
<quantity box="[981,1073,1938,1960]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" metricValueMax="10.668" metricValueMin="7.112" pageId="19" pageNumber="62" unit="in" value="35.0" valueMax="42.0" valueMin="28.0">28–42 in</quantity>
<taxonomicName baseAuthorityName="Daudin" baseAuthorityYear="1802" box="[1085,1188,1938,1959]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="62" phylum="Chordata" rank="species" species="indicus">
<emphasis box="[1085,1188,1938,1959]" italics="true" pageId="19" pageNumber="62">V. indicus</emphasis>
</taxonomicName>
, XY: 148–160 versus 
<quantity box="[314,425,1213,1235]" metricMagnitude="0" metricUnit="m" metricValue="3.3909" metricValueMax="4.013199999999999" metricValueMin="2.7686" pageId="20" pageNumber="63" unit="in" value="133.5" valueMax="158.0" valueMin="109.0">109–158 in</quantity>
<taxonomicName baseAuthorityName="Daudin" baseAuthorityYear="1802" box="[433,532,1213,1235]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="indicus">
<emphasis box="[433,532,1213,1235]" italics="true" pageId="20" pageNumber="63">V. indicus</emphasis>
</taxonomicName>
), longer tail (F/SVL = 1.60–1.89 versus 
<quantity box="[959,1082,1213,1235]" metricMagnitude="-2" metricUnit="m" metricValue="3.7084" metricValueMax="4.318" metricValueMin="3.0987999999999998" pageId="20" pageNumber="63" unit="in" value="1.46" valueMax="1.7" valueMin="1.22">1.22–1.70 in</quantity>
<taxonomicName baseAuthorityName="Daudin" baseAuthorityYear="1802" box="[1089,1188,1213,1235]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="indicus">
<emphasis box="[1089,1188,1213,1235]" italics="true" pageId="20" pageNumber="63">V. indicus</emphasis>
</taxonomicName>
) and dark pigmentation on the lower part of the throat (versus dark pigmentation on throat normally absent in 
<taxonomicName baseAuthorityName="Daudin" baseAuthorityYear="1802" box="[176,276,1277,1299]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="indicus">
<emphasis box="[176,276,1277,1299]" italics="true" pageId="20" pageNumber="63">V. indicus</emphasis>
</taxonomicName>
); from 
<taxonomicName box="[356,494,1277,1299]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="lirungensis">
<emphasis box="[356,494,1277,1299]" italics="true" pageId="20" pageNumber="63">V. lirungensis</emphasis>
</taxonomicName>
by the absence of dorsal cross-bands and lower scale counts (P: 31–40 versus 
<quantity box="[228,321,1309,1331]" metricMagnitude="0" metricUnit="m" metricValue="1.0795" metricValueMax="1.1938" metricValueMin="0.9652" pageId="20" pageNumber="63" unit="in" value="42.5" valueMax="47.0" valueMin="38.0">38–47 in</quantity>
<taxonomicName box="[334,476,1309,1331]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="lirungensis">
<emphasis box="[334,476,1309,1331]" italics="true" pageId="20" pageNumber="63">V. lirungensis</emphasis>
</taxonomicName>
, Q: 54–74 versus 
<quantity box="[685,778,1309,1331]" metricMagnitude="0" metricUnit="m" metricValue="2.1209" metricValueMax="2.2352" metricValueMin="2.0065999999999997" pageId="20" pageNumber="63" unit="in" value="83.5" valueMax="88.0" valueMin="79.0">79–88 in</quantity>
<taxonomicName box="[791,933,1309,1331]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="lirungensis">
<emphasis box="[791,933,1309,1331]" italics="true" pageId="20" pageNumber="63">V. lirungensis</emphasis>
</taxonomicName>
, S: 101–126 versus 
<quantity metricMagnitude="0" metricUnit="m" metricValue="3.6195" metricValueMax="3.8354" metricValueMin="3.4036" pageId="20" pageNumber="63" unit="in" value="142.5" valueMax="151.0" valueMin="134.0">134–151 in</quantity>
<taxonomicName box="[181,324,1341,1363]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="lirungensis">
<emphasis box="[181,324,1341,1363]" italics="true" pageId="20" pageNumber="63">V. lirungensis</emphasis>
</taxonomicName>
); from 
<taxonomicName box="[413,598,1341,1363]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="rainerguentheri">
<emphasis box="[413,598,1341,1363]" italics="true" pageId="20" pageNumber="63">V. rainerguentheri</emphasis>
</taxonomicName>
by its longer tail (F/SVL = 1.60–1.89 versus 
<quantity box="[1115,1244,1341,1363]" metricMagnitude="-2" metricUnit="m" metricValue="3.5941" metricValueMax="3.7338" metricValueMin="3.4544" pageId="20" pageNumber="63" unit="in" value="1.415" valueMax="1.47" valueMin="1.36">1.36–1.47 in</quantity>
<taxonomicName box="[148,328,1373,1395]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="rainerguentheri">
<emphasis box="[148,328,1373,1395]" italics="true" pageId="20" pageNumber="63">V. rainerguentheri</emphasis>
</taxonomicName>
), higher average scale counts of all measured characters (table 4) and the peach color of the throat (versus cream in 
<taxonomicName box="[494,677,1405,1427]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="rainerguentheri">
<emphasis box="[494,677,1405,1427]" italics="true" pageId="20" pageNumber="63">V. rainerguentheri</emphasis>
</taxonomicName>
); and from 
<taxonomicName authorityName="Kishida" authorityYear="1929" box="[817,955,1405,1427]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="tsukamotoi">
<emphasis box="[817,955,1405,1427]" italics="true" pageId="20" pageNumber="63">V. tsukamotoi</emphasis>
</taxonomicName>
by its longer tail (F/SVL 1.60–1.89 (1.76) versus 1.33–1.73 (1.58) in 
<taxonomicName authorityName="Kishida" authorityYear="1929" box="[594,729,1437,1459]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="tsukamotoi">
<emphasis box="[594,729,1437,1459]" italics="true" pageId="20" pageNumber="63">V. tsukamotoi</emphasis>
</taxonomicName>
), higher scale counts of all measured characters (table 4), and the peach colour of the throat of live animals (versus yellow in 
<taxonomicName authorityName="Kishida" authorityYear="1929" box="[975,1110,1469,1491]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="tsukamotoi">
<emphasis box="[975,1110,1469,1491]" italics="true" pageId="20" pageNumber="63">V. tsukamotoi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption box="[148,887,590,614]" pageId="20" pageNumber="63">
<paragraph blockId="20.[148,887,590,614]" box="[148,887,590,614]" pageId="20" pageNumber="63">
Figure 9. 
<taxonomicName authority="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus, 2020" authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[241,380,590,614]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="bennetti" status="sp. nov.">
<emphasis bold="true" box="[241,380,590,614]" italics="true" pageId="20" pageNumber="63">Varanus bennetti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[389,457,590,614]" pageId="20" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
, Rock Islands, Palau (photo by Thibaud Aronson).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3831916" ID-Zenodo-Dep="3831916" box="[148,898,1147,1171]" httpUri="https://zenodo.org/record/3831916/files/figure.png" pageId="20" pageNumber="63" startId="20.[148,206,1147,1171]" targetBox="[352,1032,670,1123]" targetPageId="20">
<paragraph blockId="20.[148,898,1147,1171]" box="[148,898,1147,1171]" pageId="20" pageNumber="63">
Figure 10. 
<taxonomicName authority="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus, 2020" authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[252,392,1147,1171]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="63" phylum="Chordata" rank="species" species="bennetti" status="sp. nov.">
<emphasis bold="true" box="[252,392,1147,1171]" italics="true" pageId="20" pageNumber="63">Varanus bennetti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[400,468,1147,1171]" pageId="20" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
, Ngarchelong, Palau (photo by Thibaud Aronson).
</paragraph>
</caption>
<subSubSection lastPageId="21" lastPageNumber="64" pageId="20" pageNumber="63" type="description">
<paragraph blockId="20.[148,489,1565,1594]" box="[148,489,1565,1594]" pageId="20" pageNumber="63">
<heading bold="true" box="[148,489,1565,1594]" fontSize="12" level="4" pageId="20" pageNumber="63" reason="0">
<emphasis bold="true" box="[148,489,1565,1594]" pageId="20" pageNumber="63">
3.5.5. Description of the 
<typeStatus box="[403,489,1565,1594]" pageId="20" pageNumber="63">holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[148,1244,1618,1960]" pageId="20" pageNumber="63">
Subadult specimen of undetermined sex, total length 
<quantity box="[704,788,1618,1640]" metricMagnitude="-1" metricUnit="m" metricValue="7.75" pageId="20" pageNumber="63" unit="mm" value="775.0">775 mm</quantity>
(
<collectionCode box="[801,849,1618,1639]" pageId="20" pageNumber="63">SVL</collectionCode>
: 
<quantity box="[857,943,1618,1640]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" pageId="20" pageNumber="63" unit="mm" value="275.0">275 mm</quantity>
, F: 
<quantity box="[978,1065,1618,1640]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="63" unit="mm" value="500.0">500 mm</quantity>
). Well preserved, without degradation or loss of keratin layer. There is a 
<quantity box="[729,801,1650,1672]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="63" unit="mm" value="25.0">25 mm</quantity>
long incision on the upper abdomen. Tail muscular, long, slender (F/SVL = 1.82, 38.46 times as long as high at midlength), round at base, becoming increasingly laterally compressed and gaining a double dorsal scale ridge distal to 
<quantity box="[1172,1244,1714,1736]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="20" pageNumber="63" unit="mm" value="45.0">45 mm</quantity>
posterior to vent. Dorsum of trunk and limbs black with scattered yellow scales in groups of one to four. Tail black with yellow marbling, without distinctive cross-bands. Venter cream with 
<emphasis box="[1120,1140,1779,1800]" italics="true" pageId="20" pageNumber="63">ca</emphasis>
20 moreor-less complete grey cross-bands. Throat cream with scattered brown scales laterally and near gular fold. Head various shades of brown, with a yellow post-ocular stripe. Tongue blue dorsally and pink ventrally except for a slightly darker median line. Teeth pointed and recurved.
</paragraph>
<paragraph blockId="20.[148,1244,1618,1960]" lastBlockId="21.[148,1244,710,1212]" lastPageId="21" lastPageNumber="64" pageId="20" pageNumber="63">Nuchal scales on anterior half of neck round to slightly oval, flattened or slightly domed, bordered by row of enlarged granules along lower and sometimes lateral margins. Interstitial skin covered by smaller granules. Nuchal scales on posterior half of neck elongate, keeled. Dorsal scales oval to elongate, keeled, becoming rectangular and increasingly elongate distal to base of tail. Suprabrachials and antebrachials elongate, keeled, surrounded by one or more rows of granules. Suprafemorals and tibials small, elongate, keeled, surrounded by numerous small granules. Subbrachials, antebrachials, femorals and tibials polished, round to slightly oval, with row of larger granules along posterior margin.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3831918" ID-Zenodo-Dep="3831918" box="[148,1183,627,651]" httpUri="https://zenodo.org/record/3831918/files/figure.png" pageId="21" pageNumber="64" startId="21.[148,206,627,651]" targetBox="[322,1078,113,604]" targetPageId="21">
<paragraph blockId="21.[148,1183,627,651]" box="[148,1183,627,651]" pageId="21" pageNumber="64">
Figure 11. Subadult 
<taxonomicName authority="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus, 2020" authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[334,473,627,651]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="64" phylum="Chordata" rank="species" species="bennetti" status="sp. nov.">
<emphasis bold="true" box="[334,473,627,651]" italics="true" pageId="21" pageNumber="64">Varanus bennetti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[482,551,627,651]" pageId="21" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
, Losiep Island, Federated States of Micronesia (photo by James Reardon).
</paragraph>
</caption>
<paragraph blockId="21.[148,1244,710,1212]" pageId="21" pageNumber="64">Infracarpals and infratarsals round, domed, dark brown in centre. Claws dark brown, sharp, recurved. Gulars yellow, enlarged, rectangular or irregular toward the snout, quickly decreasing in size ventrally to level of eyes. Towards the gular fold gulars round or slightly oval, bordered by row of granules along posterior and lateral margins, with one to three darker pits. Chest scales cream or grey, irregular in shape. Ventrals cream or grey, rectangular, with rounded posterior corners and small central keel. Subcaudals cream or yellow with brown anterior margin, rectangular, elongate, with sharp central keel. Lateral caudal scales half as long as subcaudals, elongate rectangular, with central keel and pit at posterior end.</paragraph>
<paragraph blockId="21.[148,1244,710,1212]" pageId="21" pageNumber="64">Occipital scale roundish. Enlarged supraocular scales seven on each side. Four scale rows separate mouth and naris; nine dorsal scales between the nares. Enlarged supralabials 24 on each side. Rostral pentagonal; temporals small and irregular.</paragraph>
<paragraph blockId="21.[148,300,1267,1296]" box="[148,300,1267,1296]" pageId="21" pageNumber="64">
<heading bold="true" box="[148,300,1267,1296]" fontSize="12" level="4" pageId="21" pageNumber="64" reason="0">
<emphasis bold="true" box="[148,300,1267,1296]" pageId="21" pageNumber="64">3.5.6. Scalation</emphasis>
</heading>
</paragraph>
<paragraph blockId="21.[148,963,1321,1343]" box="[148,963,1321,1343]" pageId="21" pageNumber="64">
S 138, XY 158, DOR 173, 
<collectionCode box="[420,434,1321,1342]" name="Tavera, Department of Geology and Geophysics" pageId="21" pageNumber="64">T</collectionCode>
100, VEN 118, X 47, m 109, P 45, Q 90 and 
<collectionCode box="[908,924,1321,1342]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="64">R</collectionCode>
64.
</paragraph>
<paragraph blockId="21.[148,355,1398,1427]" box="[148,355,1398,1427]" pageId="21" pageNumber="64">
<heading bold="true" box="[148,355,1398,1427]" fontSize="12" level="4" pageId="21" pageNumber="64" reason="0">
<emphasis bold="true" box="[148,355,1398,1427]" pageId="21" pageNumber="64">3.5.7. Measurements</emphasis>
</heading>
</paragraph>
<paragraph blockId="21.[148,1155,1452,1474]" box="[148,1155,1452,1474]" pageId="21" pageNumber="64">
SVL 
<quantity box="[200,286,1452,1474]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" pageId="21" pageNumber="64" unit="mm" value="275.0">275 mm</quantity>
, F 
<quantity box="[319,405,1452,1474]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="64" unit="mm" value="500.0">500 mm</quantity>
, TL 
<quantity box="[454,540,1452,1474]" metricMagnitude="-1" metricUnit="m" metricValue="7.75" pageId="21" pageNumber="64" unit="mm" value="775.0">775 mm</quantity>
; A 
<quantity box="[579,671,1452,1474]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" pageId="21" pageNumber="64" unit="mm" value="44.5">44.5 mm</quantity>
, B 
<quantity box="[705,797,1452,1474]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="21" pageNumber="64" unit="mm" value="25.5">25.5 mm</quantity>
, C 
<quantity box="[834,908,1452,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="64" unit="mm" value="17.0">17 mm</quantity>
, G 
<quantity box="[946,1020,1452,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="21" pageNumber="64" unit="mm" value="13.0">13 mm</quantity>
, H 
<quantity box="[1060,1152,1452,1474]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="21" pageNumber="64" unit="mm" value="11.5">11.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="64" type="discussion">
<paragraph blockId="21.[148,403,1530,1559]" box="[148,403,1530,1559]" pageId="21" pageNumber="64">
<heading bold="true" box="[148,403,1530,1559]" fontSize="12" level="4" pageId="21" pageNumber="64" reason="0">
<emphasis bold="true" box="[148,403,1530,1559]" pageId="21" pageNumber="64">3.5.8. Molecular evidence</emphasis>
</heading>
</paragraph>
<paragraph blockId="21.[148,1244,1583,1701]" pageId="21" pageNumber="64">
<taxonomicName authority="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus, 2020" authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[148,314,1583,1605]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="64" phylum="Chordata" rank="species" species="bennetti" status="sp. nov.">
<emphasis box="[148,314,1583,1605]" italics="true" pageId="21" pageNumber="64">Varanus bennetti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[319,402,1584,1605]" pageId="21" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
is resolved as a well-supported monophyletic lineage (JF 99, BS 99, syn 6) in both parsimony and likelihood-based phylogenies. The closest evolutionary relatives of the species are 
<taxonomicName authorityName="Kishida" authorityYear="1929" box="[148,290,1647,1668]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="64" phylum="Chordata" rank="species" species="tsukamotoi">
<emphasis box="[148,290,1647,1668]" italics="true" pageId="21" pageNumber="64">V. tsukamotoi</emphasis>
</taxonomicName>
(1.5/0.3% difference in ND4/16S) and 
<taxonomicName box="[754,898,1647,1668]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="64" phylum="Chordata" rank="species" species="lirungensis">
<emphasis box="[754,898,1647,1668]" italics="true" pageId="21" pageNumber="64">V. lirungensis</emphasis>
</taxonomicName>
(1.4/0.3% difference), but the evolutionary affinities among these three species remain unresolved (
<figureCitation box="[885,980,1679,1701]" captionStart="Figure 3" captionStartId="10.[148,206,1003,1027]" captionTargetBox="[156,1239,114,983]" captionTargetId="figure@10.[155,1240,113,984]" captionTargetPageId="10" captionText="Figure 3. Strict consensus of the nine most-parsimonious trees resulting from the TNT analysis. Jackknife resampling values greater than 50% are shown above, and the numbers of synapomorphies shared by each clade are shown below the nodes in parentheses. Branch lengths represent the number of optimized character-state changes." figureDoi="http://doi.org/10.5281/zenodo.3831906" httpUri="https://zenodo.org/record/3831906/files/figure.png" pageId="21" pageNumber="64">figures 3</figureCitation>
and 
<figureCitation box="[1036,1050,1679,1701]" captionStart="Figure 4" captionStartId="11.[148,206,1714,1738]" captionTargetBox="[156,1239,114,1678]" captionTargetId="figure@11.[155,1240,113,1678]" captionTargetPageId="11" captionText="Figure 4. Optimal maximum-likelihood tree resulting from the RAxML analysis. Bootstrap support values greater than 50% are shown on the nodes. Scale bar corresponds to the mean number of nucleotide substitutions per site." figureDoi="http://doi.org/10.5281/zenodo.3831908" httpUri="https://zenodo.org/record/3831908/files/figure.png" pageId="21" pageNumber="64">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="65" pageId="21" pageNumber="64" type="description">
<paragraph blockId="21.[148,522,1757,1786]" box="[148,522,1757,1786]" pageId="21" pageNumber="64">
<heading bold="true" box="[148,522,1757,1786]" fontSize="12" level="4" pageId="21" pageNumber="64" reason="0">
<emphasis bold="true" box="[148,522,1757,1786]" pageId="21" pageNumber="64">3.5.9. Variation and coloration in life</emphasis>
</heading>
</paragraph>
<paragraph blockId="21.[148,1244,1810,1960]" pageId="21" pageNumber="64">
The light (probably cream coloured to yellow in life) temporal stripe is apparent in only part of the examined material. Some specimens lack tail bands altogether, whereas others show discernible bands. Crombie and Pregill [
<bibRefCitation author="Crombie RI &amp; Pregill GK" box="[480,493,1873,1895]" journalOrPublisher="Herpetol. Monogr." pageId="21" pageNumber="64" pagination="29 - 80" part="13" refId="ref14026" refString="3. Crombie RI, Pregill GK. 1999 A checklist of the herpetofauna of the Palau Islands (Republic of Belau), Oceania. Herpetol. Monogr. 13, 29 - 80. (doi: 10.2307 / 1467060)" title="A checklist of the herpetofauna of the Palau Islands (Republic of Belau), Oceania" type="journal article" year="1999">3</bibRefCitation>
] noted that live animals are black with prominent yellow dorsal rosettes and other irregular markings, and they have a vivid peach-coloured throat (
<figureCitation box="[1058,1191,1906,1928]" captionStart-0="Figure 9" captionStart-1="Figure 10" captionStart-2="Figure 11" captionStart-3="Figure 12" captionStartId-0="20.[148,206,590,614]" captionStartId-1="20.[148,206,1147,1171]" captionStartId-2="21.[148,206,627,651]" captionStartId-3="22.[148,206,1112,1136]" captionTargetBox-1="[352,1032,670,1123]" captionTargetBox-2="[322,1078,113,604]" captionTargetBox-3="[390,995,113,1089]" captionTargetId-1="figure@20.[352,1033,670,1124]" captionTargetId-2="figure@21.[322,1079,113,604]" captionTargetId-3="figure@22.[390,995,113,1090]" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionText-0="Figure 9. Varanus bennetti sp. nov., Rock Islands, Palau (photo by Thibaud Aronson)." captionText-1="Figure 10. Varanus bennetti sp. nov., Ngarchelong, Palau (photo by Thibaud Aronson)." captionText-2="Figure 11. Subadult Varanus bennetti sp. nov., Losiep Island, Federated States of Micronesia (photo by James Reardon)." captionText-3="Figure 12. Adult Varanus bennetti sp. nov., Losiep Island, Federated States of Micronesia (photo by James Reardon)." figureDoi-1="http://doi.org/10.5281/zenodo.3831916" figureDoi-2="http://doi.org/10.5281/zenodo.3831918" figureDoi-3="http://doi.org/10.5281/zenodo.3831920" httpUri-1="https://zenodo.org/record/3831916/files/figure.png" httpUri-2="https://zenodo.org/record/3831918/files/figure.png" httpUri-3="https://zenodo.org/record/3831920/files/figure.png" pageId="21" pageNumber="64">figures 9–12</figureCitation>
, and fig. 
<quantity box="[190,255,1938,1960]" metricMagnitude="0" metricUnit="m" metricValue="4.7752" pageId="21" pageNumber="64" unit="in" value="188.0">188 in</quantity>
[
<bibRefCitation author="Buden DW &amp; Taborosi D." box="[271,285,1937,1960]" journalOrPublisher="Pohnpei, Federated States of Micronesia: Island Research and Education Initiative" pageId="21" pageNumber="64" refId="ref14127" refString="5. Buden DW, Taborosi D. 2016 Reptiles of the Federated States of Micronesia. Pohnpei, Federated States of Micronesia: Island Research and Education Initiative." title="Reptiles of the Federated States of Micronesia" type="book" year="2016">5</bibRefCitation>
]). They also noted the large size (up to 
<quantity box="[717,791,1938,1959]" metricMagnitude="0" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="64" unit="cm" value="180.0">180 cm</quantity>
) attained by 
<taxonomicName authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[935,1044,1938,1959]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="64" phylum="Chordata" rank="species" species="bennetti">
<emphasis box="[935,1044,1938,1959]" italics="true" pageId="21" pageNumber="64">V. bennetti</emphasis>
</taxonomicName>
in 
<collectingCountry box="[1081,1142,1938,1960]" name="Palau" pageId="21" pageNumber="64">Palau</collectingCountry>
, which is
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3831920" ID-Zenodo-Dep="3831920" box="[148,1155,1112,1136]" httpUri="https://zenodo.org/record/3831920/files/figure.png" pageId="22" pageNumber="65" startId="22.[148,206,1112,1136]" targetBox="[390,995,113,1089]" targetPageId="22">
<paragraph blockId="22.[148,1155,1112,1136]" box="[148,1155,1112,1136]" pageId="22" pageNumber="65">
Figure 12. Adult 
<taxonomicName authority="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus, 2020" authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[305,445,1112,1136]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="65" phylum="Chordata" rank="species" species="bennetti" status="sp. nov.">
<emphasis bold="true" box="[305,445,1112,1136]" italics="true" pageId="22" pageNumber="65">Varanus bennetti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[454,523,1112,1136]" pageId="22" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
, Losiep Island, Federated States of Micronesia (photo by James Reardon).
</paragraph>
</caption>
<paragraph blockId="22.[148,1244,1195,1249]" pageId="22" pageNumber="65">
exceptional within the subgenus 
<emphasis box="[504,663,1196,1217]" italics="true" pageId="22" pageNumber="65">
<taxonomicName box="[504,658,1196,1217]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="65" phylum="Chordata" rank="subGenus" subGenus="Euprepiosaurus">Euprepiosaurus</taxonomicName>
.
</emphasis>
Fourteen specimens measured in the field on Sarigan had total lengths of 
<quantity box="[365,477,1227,1249]" metricMagnitude="0" metricUnit="m" metricValue="1.045" metricValueMax="1.42" metricValueMin="0.67" pageId="22" pageNumber="65" unit="cm" value="104.5" valueMax="142.0" valueMin="67.0">67–142 cm</quantity>
, tail/SVL of 1.69–1.88 and weights of 
<quantity box="[888,1002,1227,1249]" metricMagnitude="0" metricUnit="kg" metricValue="1.19" metricValueMax="2.18" metricValueMin="0.2" pageId="22" pageNumber="65" unit="g" value="1190.0" valueMax="2180.0" valueMin="200.0">200–2180 g</quantity>
[
<bibRefCitation author="Vogt S." box="[1017,1046,1226,1248]" journalOrPublisher="CNMI Division of Fish &amp; Wildlife" pageId="22" pageNumber="65" refId="ref16201" refString="54. Vogt S. 2008 Monitor lizard, Varanus indicus, diet on Sarigan Island, Commonwealth of the Northern Mariana Islands. In Technical Report  14 Wildlife and Vegetation Surveys of Sarigan Island April 13 - 25, 2006, pp. 6 - 1 - 6 - 6. CNMI Division of Fish &amp; Wildlife." title="Monitor lizard, Varanus indicus, diet on Sarigan Island, Commonwealth of the Northern Mariana Islands" type="proceedings paper" volumeTitle="Technical Report  14 Wildlife and Vegetation Surveys of Sarigan Island April 13 - 25, 2006" year="2008">54</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="65" type="etymology">
<paragraph blockId="22.[148,327,1304,1333]" box="[148,327,1304,1333]" pageId="22" pageNumber="65">
<heading bold="true" box="[148,327,1304,1333]" fontSize="12" level="4" pageId="22" pageNumber="65" reason="0">
<emphasis bold="true" box="[148,327,1304,1333]" pageId="22" pageNumber="65">3.5.10. Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="22.[148,1244,1357,1443]" pageId="22" pageNumber="65">
The specific epithet is a genitive singular patronym in commemoration of the late Dr Daniel Bennett, 
<date box="[148,258,1389,1411]" pageId="22" pageNumber="65" value="1966" valueMax="2020">1966–2020</date>
, and his life-long commitment to the study and conservation of monitor lizards in Africa and Southeast Asia. As a vernacular name we suggest ‘Bennett’ s long-tailed monitor’.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="65" type="distribution">
<paragraph blockId="22.[148,337,1498,1527]" box="[148,337,1498,1527]" pageId="22" pageNumber="65">
<heading bold="true" box="[148,337,1498,1527]" fontSize="12" level="4" pageId="22" pageNumber="65" reason="0">
<emphasis bold="true" box="[148,337,1498,1527]" pageId="22" pageNumber="65">3.5.11. Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="22.[148,1244,1552,1702]" pageId="22" pageNumber="65">
We have examined specimens of 
<taxonomicName authorityName="Weijola &amp; Vahtera &amp; Koch &amp; Schmitz &amp; Kraus" authorityYear="2020" box="[508,617,1552,1573]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="65" phylum="Chordata" rank="species" species="bennetti">
<emphasis box="[508,617,1552,1573]" italics="true" pageId="22" pageNumber="65">V. bennetti</emphasis>
</taxonomicName>
from 
<collectingRegion box="[685,747,1552,1573]" country="Palau" name="Koror" pageId="22" pageNumber="65">Koror</collectingRegion>
, Ngeaur and Ngcheangel islands in the 
<collectingCountry box="[1185,1244,1552,1574]" name="Palau" pageId="22" pageNumber="65">Palau</collectingCountry>
archipelago, from 
<collectingRegion box="[352,394,1584,1605]" country="Micronesia" name="Yap" pageId="22" pageNumber="65">Yap</collectingRegion>
and Losiep islands in the 
<collectingCountry box="[707,1045,1584,1606]" name="Micronesia" pageId="22" pageNumber="65">Federated States of Micronesia</collectingCountry>
(
<collectingCountry box="[1065,1117,1584,1606]" name="Micronesia" pageId="22" pageNumber="65">FSM</collectingCountry>
), and from Sarigan Island in the 
<collectingCountry box="[382,910,1616,1638]" name="Northern Mariana Islands" pageId="22" pageNumber="65">Commonwealth of the Northern Mariana Islands</collectingCountry>
(
<collectingCountry box="[926,995,1616,1638]" name="Northern Mariana Islands" pageId="22" pageNumber="65">CNMI</collectingCountry>
) (
<figureCitation box="[1017,1102,1616,1638]" captionStart="Figure 1" captionStartId="3.[148,206,929,953]" captionTargetBox="[156,1236,113,906]" captionTargetId="figure@3.[155,1236,113,906]" captionTargetPageId="3" captionText="Figure 1. Map of the Pacific region showing the distribution of Varanus tsukamotoi (white dots), V. bennetti sp. nov. (red dots), V. lirungensis (yellow dot) and V. rainerguentheri (green dot)." figureDoi="http://doi.org/10.5281/zenodo.3831902" httpUri="https://zenodo.org/record/3831902/files/figure.png" pageId="22" pageNumber="65">figure 1</figureCitation>
). Crombie &amp; Pregill [
<bibRefCitation author="Crombie RI &amp; Pregill GK" box="[234,247,1647,1669]" journalOrPublisher="Herpetol. Monogr." pageId="22" pageNumber="65" pagination="29 - 80" part="13" refId="ref14026" refString="3. Crombie RI, Pregill GK. 1999 A checklist of the herpetofauna of the Palau Islands (Republic of Belau), Oceania. Herpetol. Monogr. 13, 29 - 80. (doi: 10.2307 / 1467060)" title="A checklist of the herpetofauna of the Palau Islands (Republic of Belau), Oceania" type="journal article" year="1999">3</bibRefCitation>
] also list this species (as 
<taxonomicName baseAuthorityName="Daudin" baseAuthorityYear="1802" box="[536,735,1648,1670]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="65" phylum="Chordata" rank="species" species="indicus">
<emphasis box="[536,618,1648,1669]" italics="true" pageId="22" pageNumber="65">Varanus</emphasis>
cf. 
<emphasis box="[665,735,1648,1669]" italics="true" pageId="22" pageNumber="65">indicus</emphasis>
</taxonomicName>
) from an additional two islands in the 
<collectingCountry box="[1185,1244,1648,1670]" name="Palau" pageId="22" pageNumber="65">Palau</collectingCountry>
group: Ngeriungs and Babeldaob.
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="66" pageId="22" pageNumber="65" type="biology_ecology">
<paragraph blockId="22.[148,298,1757,1786]" box="[148,298,1757,1786]" pageId="22" pageNumber="65">
<heading bold="true" box="[148,298,1757,1786]" fontSize="12" level="4" pageId="22" pageNumber="65" reason="0">
<emphasis bold="true" box="[148,298,1757,1786]" pageId="22" pageNumber="65">3.5.12. Ecology</emphasis>
</heading>
</paragraph>
<paragraph blockId="22.[148,1244,1809,1960]" pageId="22" pageNumber="65">
Crombie &amp; Pregill [
<bibRefCitation author="Crombie RI &amp; Pregill GK" box="[360,373,1809,1831]" journalOrPublisher="Herpetol. Monogr." pageId="22" pageNumber="65" pagination="29 - 80" part="13" refId="ref14026" refString="3. Crombie RI, Pregill GK. 1999 A checklist of the herpetofauna of the Palau Islands (Republic of Belau), Oceania. Herpetol. Monogr. 13, 29 - 80. (doi: 10.2307 / 1467060)" title="A checklist of the herpetofauna of the Palau Islands (Republic of Belau), Oceania" type="journal article" year="1999">3</bibRefCitation>
] remarked that the monitors on 
<collectingCountry box="[730,789,1810,1832]" name="Palau" pageId="22" pageNumber="65">Palau</collectingCountry>
are decidedly terrestrial and prefer, when possible, to take refuge in terrestrial refuges rather than in trees. On Ngeaur, they are reportedly most common in the rugged limestone interior of the island [
<bibRefCitation author="Crombie RI &amp; Pregill GK" box="[748,762,1873,1895]" journalOrPublisher="Herpetol. Monogr." pageId="22" pageNumber="65" pagination="29 - 80" part="13" refId="ref14026" refString="3. Crombie RI, Pregill GK. 1999 A checklist of the herpetofauna of the Palau Islands (Republic of Belau), Oceania. Herpetol. Monogr. 13, 29 - 80. (doi: 10.2307 / 1467060)" title="A checklist of the herpetofauna of the Palau Islands (Republic of Belau), Oceania" type="journal article" year="1999">3</bibRefCitation>
]. Both features are atypical for species in the 
<taxonomicName baseAuthorityName="Daudin" baseAuthorityYear="1802" box="[148,253,1906,1928]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="65" phylum="Chordata" rank="species" species="indicus">
<emphasis box="[148,253,1906,1928]" italics="true" pageId="22" pageNumber="65">V. indicus</emphasis>
</taxonomicName>
Group, which usually seek refuge in trees and attain their highest densities in coastal habitats [
<bibRefCitation author="Weijola VS-A" box="[246,275,1937,1959]" journalOrPublisher="Biawak" pageId="22" pageNumber="65" pagination="7 - 23" part="4" refId="ref16261" refString="55. Weijola VS-A. 2010 Geographical distribution and habitat use of monitor lizards of the north Moluccas. Biawak 4, 7 - 23. (doi: 10.5281 / zenodo. 194776)" title="Geographical distribution and habitat use of monitor lizards of the north Moluccas" type="journal article" year="2010">55</bibRefCitation>
]. On Sarigan Island, the most common food items found in dissected lizards (
<emphasis box="[1116,1129,1938,1959]" italics="true" pageId="22" pageNumber="65">n</emphasis>
= 16) were
</paragraph>
<paragraph blockId="23.[148,1244,113,168]" pageId="23" pageNumber="66">
rats (
<taxonomicName baseAuthorityName="Peale" baseAuthorityYear="1848" box="[202,346,114,136]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="66" phylum="Chordata" rank="species" species="exulans">
<emphasis box="[202,346,114,136]" italics="true" pageId="23" pageNumber="66">Rattus exulans</emphasis>
</taxonomicName>
), insects and lizards [
<bibRefCitation author="Vogt S." box="[580,609,113,135]" journalOrPublisher="CNMI Division of Fish &amp; Wildlife" pageId="23" pageNumber="66" refId="ref16201" refString="54. Vogt S. 2008 Monitor lizard, Varanus indicus, diet on Sarigan Island, Commonwealth of the Northern Mariana Islands. In Technical Report  14 Wildlife and Vegetation Surveys of Sarigan Island April 13 - 25, 2006, pp. 6 - 1 - 6 - 6. CNMI Division of Fish &amp; Wildlife." title="Monitor lizard, Varanus indicus, diet on Sarigan Island, Commonwealth of the Northern Mariana Islands" type="proceedings paper" volumeTitle="Technical Report  14 Wildlife and Vegetation Surveys of Sarigan Island April 13 - 25, 2006" year="2008">54</bibRefCitation>
]. In addition, that author found a high proportion of males among the specimens examined by him: only four of 
<specimenCount box="[722,863,146,168]" pageId="23" pageNumber="66" type="generic">16 specimens</specimenCount>
were females.
</paragraph>
</subSubSection>
</treatment>
</document>