treatments-xml/data/03/A3/87/03A387B9FF96FFBFFDD618FAFB29A7E7.xml

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<document id="6891F4BBA10C8051B0E296D505983B8D" ID-DOI="10.5852/ejt.2024.927.2475" ID-ISSN="2118-9773" ID-Zenodo-Dep="10853688" ID-ZooBank="B6E43365-FACA-49BA-8CCD-77E4BF8F0016" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1711115020933" checkinUser="plazi" docAuthor="Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De &amp; Gittenberger, Edmund" docDate="2024" docId="03A387B9FF96FFBFFDD618FAFB29A7E7" docLanguage="en" docName="EJT.2024.927.1-163.pdf" docOrigin="European Journal of Taxonomy 927" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2475/11013" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cochlostoma (Turritus) nanum Cochl." docType="treatment" docVersion="9" lastPageNumber="87" masterDocId="FF9AFFC1FFC4FFE9FFD91D5FFFB5A629" masterDocTitle="Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae)" masterLastPageNumber="163" masterPageNumber="1" pageNumber="83" updateTime="1711726029180" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="071F25D4A7BEA457A72A750F27DEBC74">Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae)</mods:title>
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<mods:namePart id="24FCE9FCBA65B7E5D8D73519FD03EEDE">Zallot, Enrico</mods:namePart>
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<mods:namePart id="71E8373AB122CB500D8E1F968A39FCD2">Kamchev, Panche</mods:namePart>
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<mods:namePart id="7F638D3FBF2739C7F0230F536DCA6573">Mattia, Willy De</mods:namePart>
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<taxonomicName id="4C0A4D2CFF96FFBBFDD618FAFB90A396" ID-CoL="WNNV" authority="(Westerlund, 1879)" authorityName="Cochl." baseAuthorityName="Westerlund" baseAuthorityYear="1879" box="[527,1061,1445,1472]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="82" pageNumber="83" phylum="Mollusca" rank="species" species="nanum" subGenus="Turritus">
<emphasis id="B97EEABDFF96FFBBFDD618FAFD18A3E9" bold="true" box="[527,685,1445,1472]" italics="true" pageId="82" pageNumber="83">Cochlostoma</emphasis>
(
<emphasis id="B97EEABDFF96FFBBFD6718FAFD61A396" bold="true" box="[702,724,1445,1471]" italics="true" pageId="82" pageNumber="83">T.</emphasis>
)
<emphasis id="B97EEABDFF96FFBBFD3C18FAFC8EA396" bold="true" box="[741,827,1445,1471]" italics="true" pageId="82" pageNumber="83">nanum</emphasis>
(
<bibRefCitation id="EF9B4B5EFF96FFBBFC9218FAFBA9A396" author="Westerlund" box="[843,1052,1445,1471]" firstAuthor="Westerlund" pageId="82" pageNumber="83" pagination="156 - 168" refId="ref38386" refString="Westerlund C. A. 1879. Malakozoologische Beitrage. Jahrbucher der deutschen malakozoologischen Gesellschaft 6: 156 - 168." type="journal article" year="1879">Westerlund, 1879</bibRefCitation>
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<figureCitation id="13312A2AFF96FFBBFD8D1895FD1EA3CD" box="[596,683,1482,1508]" captionStart="Fig" captionStartId="83.[189,232,1962,1988]" captionTargetBox="[189,1398,1245,1925]" captionTargetId="figure-319@83.[189,1398,1245,1925]" captionTargetPageId="83" captionText="Fig. 72. Distribution of samples of clade B: red =Cochlostoma (T.) nanum (Westerlund, 1879); cyan =C. (T.) elegans. Black center for the samples with amplified 16S." figureDoi="http://doi.org/10.5281/zenodo.10853846" httpUri="https://zenodo.org/record/10853846/files/figure.png" pageId="82" pageNumber="83">Figs 72</figureCitation>
(red dots), 73AC, 7476
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<treatmentCitation id="0AAB10BEFF96FFBBFF641B50FD01A001" author="Westerlund C. A." box="[189,692,1550,1576]" page="16" pageId="82" pageNumber="83" year="1879">
<taxonomicName id="4C0A4D2CFF96FFBBFF641B50FD01A001" authority="Westerlund, 1879: 167 - 16" authorityName="Westerlund" authorityPageNumber="167 - 16" authorityYear="1879" box="[189,692,1550,1576]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="82" pageNumber="83" phylum="Mollusca" rank="species" species="nanus">
<emphasis id="B97EEABDFF96FFBBFF641B50FECCA001" box="[189,377,1551,1576]" italics="true" pageId="82" pageNumber="83">Pomatias nanus</emphasis>
<bibRefCitation id="EF9B4B5EFF96FFBBFE591B51FD01A001" author="Westerlund" box="[384,692,1550,1576]" firstAuthor="Westerlund" page="167 - 16" pageId="82" pageNumber="83" pagination="156 - 168" refId="ref38386" refString="Westerlund C. A. 1879. Malakozoologische Beitrage. Jahrbucher der deutschen malakozoologischen Gesellschaft 6: 156 - 168." type="journal article" year="1879">Westerlund, 1879: 167-16</bibRefCitation>
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.
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<treatmentCitation id="0AAB10BEFF96FFBBFF641B6BFD3CA067" author="Hirc D." box="[189,649,1587,1614]" page="522" pageId="82" pageNumber="83" year="1881">
<taxonomicName id="4C0A4D2CFF96FFBBFF641B6BFD3CA067" authority="Hirc, 1881: 521 - 522" authorityName="Hirc" authorityPageNumber="521 - 522" authorityYear="1881" box="[189,649,1587,1614]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="82" pageNumber="83" phylum="Mollusca" rank="species" species="clessini">
<emphasis id="B97EEABDFF96FFBBFF641B6BFE3EA064" box="[189,395,1587,1613]" italics="true" pageId="82" pageNumber="83">Pomatias clessini</emphasis>
<bibRefCitation id="EF9B4B5EFF96FFBBFE4B1B6CFD3CA067" author="Hirc" box="[402,649,1587,1614]" firstAuthor="Hirc" page="521 - 522" pageId="82" pageNumber="83" pagination="519 - 530" refId="ref37052" refString="Hirc D. 1881. Die Mollusken-Fauna des liburnischen Karstes. Verhandlungen der kaiserlich-koniglichen Zoologisch-Botanischen Gesellschaft in Wien 30: 519 - 530." type="journal article" year="1881">Hirc, 1881: 521522</bibRefCitation>
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<typeStatus id="54B1880DFF96FFBBFF641B28FEB3A0B8" box="[189,262,1655,1681]" pageId="82" pageNumber="83">Types</typeStatus>
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<emphasis id="B97EEABDFF96FFBBFF641BB5FE3BA12D" bold="true" box="[189,398,1770,1796]" pageId="82" pageNumber="83">Other specimens</emphasis>
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: 5-
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; 2015; AL leg.;
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• 1-
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<geoCoordinate id="EE3E5068FF96FFBBFCE31A01FC7DA150" box="[826,968,1886,1913]" degrees="45.4251" direction="north" orientation="latitude" pageId="82" pageNumber="83" precision="5" value="45.4251">45.4251° N</geoCoordinate>
,
<geoCoordinate id="EE3E5068FF96FFBBFC0E1A00FBD7A150" box="[983,1122,1887,1913]" degrees="14.6162" direction="east" orientation="longitude" pageId="82" pageNumber="83" precision="5" value="14.6162">14.6162° E</geoCoordinate>
; 2006;
<collectorName id="26FF5379FF96FFBBFB661A00FA8CA150" box="[1215,1337,1887,1913]" pageId="82" pageNumber="83">De Mattia</collectorName>
leg.;
<specimenCode id="DBAC9ED4FF96FFBBFF641ADBFE89A1B6" box="[189,316,1924,1951]" collectionCode="WdM" pageId="82" pageNumber="83">WdM5666</specimenCode>
</materialsCitation>
<materialsCitation id="3B623CF2FF96FFBBFE801ADBFAA3A1B6" box="[345,1302,1924,1951]" collectorName="De Mattia" country="Slovenia" latitude="45.5884" location="Mt Sneznik" longLatPrecision="6" longitude="14.4478" pageId="82" pageNumber="83" specimenCode="WdM7005" specimenCount="1">
2-
<location id="8ED56074FF96FFBBFEA21ADBFDB0A1B7" LSID="urn:lsid:plazi:treatment:03A387B9FF96FFBFFDD618FAFB29A7E7:8ED56074FF96FFBBFEA21ADBFDB0A1B7" box="[379,517,1924,1950]" country="Slovenia" latitude="45.5884" longLatPrecision="6" longitude="14.4478" name="Mt Sneznik" pageId="82" pageNumber="83">Mt Snežnik</location>
1764;
<geoCoordinate id="EE3E5068FF96FFBBFD831ADBFD52A1B7" box="[602,743,1924,1951]" degrees="45.5884" direction="north" orientation="latitude" pageId="82" pageNumber="83" precision="5" value="45.5884">45.5884° N</geoCoordinate>
,
<geoCoordinate id="EE3E5068FF96FFBBFD2C1ADBFCCBA1B7" box="[757,894,1924,1950]" degrees="14.4478" direction="east" orientation="longitude" pageId="82" pageNumber="83" precision="5" value="14.4478">14.4478° E</geoCoordinate>
; 2010;
<collectorName id="26FF5379FF96FFBBFC001ADBFBE4A1B7" box="[985,1105,1924,1950]" pageId="82" pageNumber="83">De Mattia</collectorName>
leg.;
<specimenCode id="DBAC9ED4FF96FFBBFB4F1ADBFAA3A1B6" box="[1174,1302,1924,1951]" collectionCode="WdM" pageId="82" pageNumber="83">WdM7005</specimenCode>
</materialsCitation>
<materialsCitation id="3B623CF2FF96FFBBFAEB1ADBFC53A1EA" collectionCode="EZ" collectorName="De Mattia" country="Slovenia" latitude="45.5825" location="Mt Sneznik" longLatPrecision="6" longitude="14.4312" pageId="82" pageNumber="83" specimenCode="EZ0960" specimenCount="1">
3-
<location id="8ED56074FF96FFBBFA8D1ADBFEA9A1EA" LSID="urn:lsid:plazi:treatment:03A387B9FF96FFBFFDD618FAFB29A7E7:8ED56074FF96FFBBFA8D1ADBFEA9A1EA" country="Slovenia" latitude="45.5825" longLatPrecision="6" longitude="14.4312" name="Mt Sneznik" pageId="82" pageNumber="83">Mt Snežnik</location>
1451;
<geoCoordinate id="EE3E5068FF96FFBBFEB01AF6FE47A1ED" box="[361,498,1961,1988]" degrees="45.5825" direction="north" orientation="latitude" pageId="82" pageNumber="83" precision="5" value="45.5825">45.5825° N</geoCoordinate>
,
<geoCoordinate id="EE3E5068FF96FFBBFE251AF6FD37A1ED" box="[508,642,1961,1988]" degrees="14.4312" direction="east" orientation="longitude" pageId="82" pageNumber="83" precision="5" value="14.4312">14.4312° E</geoCoordinate>
; 2010;
<collectorName id="26FF5379FF96FFBBFD0C1AF5FCFCA1EA" box="[725,841,1961,1988]" pageId="82" pageNumber="83">De Mattia</collectorName>
leg.;
<specimenCode id="DBAC9ED4FF96FFBBFC5E1AF6FC53A1EA" box="[903,998,1961,1987]" collectionCode="EZ" pageId="82" pageNumber="83">EZ0960</specimenCode>
</materialsCitation>
<materialsCitation id="3B623CF2FF96FFBBFC221AF6FD6BA1C0" collectionCode="HNHM" collectorName="Slapnik" country="Slovenia" latitude="46.4274" location="Logarska Dolina" longLatPrecision="6" longitude="14.6572" pageId="82" pageNumber="83" specimenCode="HNHM97239" specimenCount="1">
4-
<location id="8ED56074FF96FFBBFBC01AF6FB55A1EA" LSID="urn:lsid:plazi:treatment:03A387B9FF96FFBFFDD618FAFB29A7E7:8ED56074FF96FFBBFBC01AF6FB55A1EA" box="[1049,1248,1961,1987]" country="Slovenia" latitude="46.4274" longLatPrecision="6" longitude="14.6572" name="Logarska Dolina" pageId="82" pageNumber="83">Logarska Dolina</location>
;
<geoCoordinate id="EE3E5068FF96FFBBFB301AF6FAC7A1ED" box="[1257,1394,1961,1988]" degrees="46.4274" direction="north" orientation="latitude" pageId="82" pageNumber="83" precision="5" value="46.4274">46.4274° N</geoCoordinate>
,
<geoCoordinate id="EE3E5068FF96FFBBFF641A91FEF1A1C0" box="[189,324,1998,2025]" degrees="14.6572" direction="east" orientation="longitude" pageId="82" pageNumber="83" precision="5" value="14.6572">14.6572° E</geoCoordinate>
; 2009;
<collectorName id="26FF5379FF96FFBBFE421A91FE40A1C1" box="[411,501,1998,2024]" pageId="82" pageNumber="83">Slapnik</collectorName>
leg.;
<specimenCode id="DBAC9ED4FF96FFBBFDEE1A90FD6BA1C0" box="[567,734,1999,2025]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="82" pageNumber="83">HNHM97239</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB536AFFF97FFBAFF641C56FEE8A70B" blockId="83.[189,349,264,291]" box="[189,349,264,291]" pageId="83" pageNumber="84">
<heading id="D0FD81C3FF97FFBAFF641C56FEE8A70B" bold="true" box="[189,349,264,291]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B97EEABDFF97FFBAFF641C56FEE8A70B" bold="true" box="[189,349,264,291]" pageId="83" pageNumber="84">
<typeStatus id="54B1880DFF97FFBAFF641C56FF4FA70A" box="[189,250,265,291]" pageId="83" pageNumber="84">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFF97FFBAFF1B1C68FE70A75D" blockId="83.[189,1398,311,372]" pageId="83" pageNumber="84">
<materialsCitation id="3B623CF2FF97FFBAFF1B1C68FE74A75D" country="Croatia" location="Mount Risnjak" pageId="83" pageNumber="84" specimenCount="1" typeStatus="holotype">
A generic “Croatia” in the original description. We define as type locality restricted:
<collectingCountry id="F31D763FFF97FFBAFB4D1C68FAA6A778" box="[1172,1299,311,337]" name="Croatia" pageId="83" pageNumber="84">CROATIA</collectingCountry>
<location id="8ED56074FF97FFBAFAF11C67FEA2A75D" LSID="urn:lsid:plazi:treatment:03A387B9FF96FFBFFDD618FAFB29A7E7:8ED56074FF97FFBAFAF11C67FEA2A75D" country="Croatia" name="Mount Risnjak" pageId="83" pageNumber="84">Mount Risnjak</location>
(see remarks)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3106524FF97FFBAFF641CC4FED1A513" pageId="83" pageNumber="84" type="description">
<paragraph id="8BB536AFFF97FFBAFF641CC4FEFBA79C" blockId="83.[189,334,411,437]" box="[189,334,411,437]" pageId="83" pageNumber="84">
<heading id="D0FD81C3FF97FFBAFF641CC4FEFBA79C" bold="true" box="[189,334,411,437]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B97EEABDFF97FFBAFF641CC4FEFBA79C" bold="true" box="[189,334,411,437]" pageId="83" pageNumber="84">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFF97FFBAFF641C95FC60A402" blockId="83.[189,1399,458,555]" pageId="83" pageNumber="84">
<smallCapsWord id="8D53A073FF97FFBAFF641C95FEB4A7CA" baselines="477,478" box="[189,257,458,484]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="83" pageNumber="84">SHELL</smallCapsWord>
. Closely spaced riblets on last part of protoconch. Teleoconch spotless, with weak or very weak ribs almost disappearing on last whorl and approaching aperture. Strong lip with columellar lobe either barely or abruptly inwardly bent, only partially covering umbilicus.
</paragraph>
<paragraph id="8BB536AFFF97FFBAFF641F0CFB3DA4A6" blockId="83.[189,1398,594,656]" pageId="83" pageNumber="84">
<smallCapsWord id="8D53A073FF97FFBAFF641F0CFED9A442" baselines="614,614" box="[189,364,595,621]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="83" pageNumber="84">MEASUREMENTS</smallCapsWord>
.
<specimenCount id="9D0CFD26FF97FFBAFE5C1F0DFE56A444" box="[389,483,594,621]" count="13" pageId="83" pageNumber="84" type="female">13 ♀♀</specimenCount>
: whorls =78, H =
<quantity id="4CF29B4AFF97FFBAFD0A1F0DFCE2A444" box="[723,855,594,621]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" metricValueMax="8.4" metricValueMin="7.0" pageId="83" pageNumber="84" unit="mm" value="7.7" valueMax="8.4" valueMin="7.0">78.4 mm</quantity>
, H/W=2.42.64, roundness =0.10.15, ribs incl.= 5361°, apert. incl.= 1624°, ribs/mm 1
<superScript id="7C7F9BE7FF97FFBAFD051F2AFD52A4AA" attach="left" box="[732,743,629,643]" fontSize="6" pageId="83" pageNumber="84">st</superScript>
wh.= 811, ribs/mm 4
<superScript id="7C7F9BE7FF97FFBAFC2B1F2BFC4AA4AB" attach="left" box="[1010,1023,628,642]" fontSize="6" pageId="83" pageNumber="84">th</superScript>
wh.=817.
</paragraph>
<paragraph id="8BB536AFFF97FFBAFF641FE8FED1A513" blockId="83.[189,1399,695,826]" pageId="83" pageNumber="84">
<smallCapsWord id="8D53A073FF97FFBAFF641FE8FEA1A4E6" baselines="714,714" box="[189,276,695,721]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="83" pageNumber="84">FEMALE</smallCapsWord>
<smallCapsWord id="8D53A073FF97FFBAFEC51FE3FEC6A4E6" baselines="714" box="[284,371,700,719]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genital" pageId="83" pageNumber="84">GENITAL</smallCapsWord>
<smallCapsWord id="8D53A073FF97FFBAFEA51FE3FE65A4E6" baselines="714" box="[380,464,700,719]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="organs" pageId="83" pageNumber="84">ORGANS</smallCapsWord>
. Posterior connection of pedunculus of bursa copulatrix and junction of uterus gland moved far from connection between distal oviduct and pedunculus of bursa copulatrix. Seminal receptacle normally club-shaped, situated on dorsal side of body and with long and undefined basis/ distal oviduct.
</paragraph>
</subSubSection>
<subSubSection id="C3106524FF97FFBFFF641E3DFB29A7E7" lastPageId="86" lastPageNumber="87" pageId="83" pageNumber="84" type="discussion">
<paragraph id="8BB536AFFF97FFBAFF641E3DFE9AA555" blockId="83.[189,303,866,892]" box="[189,303,866,892]" pageId="83" pageNumber="84">
<heading id="D0FD81C3FF97FFBAFF641E3DFE9AA555" bold="true" box="[189,303,866,892]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B97EEABDFF97FFBAFF641E3DFE9AA555" bold="true" box="[189,303,866,892]" pageId="83" pageNumber="84">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFF97FFBFFF641ECFFDF9A76F" blockId="83.[189,1399,912,1185]" lastBlockId="86.[189,1399,265,327]" lastPageId="86" lastPageNumber="87" pageId="83" pageNumber="84">
The
<typeStatus id="54B1880DFF97FFBAFF2F1ECEFE9CA582" box="[246,297,913,939]" pageId="83" pageNumber="84">type</typeStatus>
locality in the original description is rather undefined. We consider Mount Risnjak in the northern part of
<collectingCountry id="F31D763FFF97FFBAFEA21EECFE64A5E4" box="[379,465,947,973]" name="Croatia" pageId="83" pageNumber="84">Croatia</collectingCountry>
as
<typeStatus id="54B1880DFF97FFBAFE2E1EEBFD9FA5E7" box="[503,554,948,974]" pageId="83" pageNumber="84">type</typeStatus>
locality restricted by A.J.
<bibRefCitation id="EF9B4B5EFF97FFBAFC831EEBFB99A5E7" author="Wagner" box="[858,1068,947,974]" firstAuthor="Wagner" page="54" pageId="83" pageNumber="84" pagination="565 - 632" refId="ref38226" refString="Wagner A. J. 1897. Monographie der Gattung Pomatias Studer. Denkschriften der kaiserlichen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse 64: 565 - 632. https: // doi. org / 10.5962 / bhl. title. 10792" type="journal article" year="1897">Wagner (1897: 54</bibRefCitation>
[618]): “Die mir vom Autor übergebenen Originalexemplare stammen ohne nähere Fundortsangabe aus Croatien (ein bestimmter Fundort wird auch sonst vom Autor nirgends angeführt); dieselben erwiesen sich aber als vollkommen identisch mit den mir ebenfalls vorliegenden Originalexemplaren des
<taxonomicName id="4C0A4D2CFF97FFBAFC3D1941FB27A21E" authority="Hirc" authorityName="Hirc" authorityYear="1881" box="[996,1170,1053,1079]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="clessini">
<emphasis id="B97EEABDFF97FFBAFC3D1941FBEDA21E" box="[996,1112,1053,1079]" italics="true" pageId="83" pageNumber="84">P. clessini</emphasis>
Hirc
</taxonomicName>
vom Berge Risnjak in Croatien und dürften wohl auch daher stammen [The original specimens were given to me by the author labeled Croatia without any further information on where they were found (a specific site is also not mentioned anywhere else by the author); but it turned out that they are completely identical to the original specimens of
<taxonomicName id="4C0A4D2CFF92FFBFFE1A1C55FDC1A70A" authority="Hirc" authorityName="Hirc" authorityYear="1881" box="[451,628,265,291]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="86" pageNumber="87" phylum="Mollusca" rank="species" species="clessini">
<emphasis id="B97EEABDFF92FFBFFE1A1C55FD8DA70A" box="[451,568,265,291]" italics="true" pageId="86" pageNumber="87">P. clessini</emphasis>
Hirc
</taxonomicName>
from the Risnjak mountain in
<collectingCountry id="F31D763FFF92FFBFFC061C56FB80A70A" box="[991,1077,265,291]" name="Croatia" pageId="86" pageNumber="87">Croatia</collectingCountry>
that I also have and it may well be that they are from there]”.
</paragraph>
<caption id="DF756627FF97FFBAFF641AF5FBB6A1CE" ID-DOI="http://doi.org/10.5281/zenodo.10853846" ID-Zenodo-Dep="10853846" httpUri="https://zenodo.org/record/10853846/files/figure.png" pageId="83" pageNumber="84" startId="83.[189,232,1962,1988]" targetBox="[189,1398,1245,1925]" targetPageId="83">
<paragraph id="8BB536AFFF97FFBAFF641AF5FBB6A1CE" blockId="83.[189,1399,1962,2024]" pageId="83" pageNumber="84">
<emphasis id="B97EEABDFF97FFBAFF641AF5FE94A1ED" bold="true" box="[189,289,1962,1988]" pageId="83" pageNumber="84">Fig. 72.</emphasis>
Distribution of samples of clade B: red =
<taxonomicName id="4C0A4D2CFF97FFBAFC9B1AF5FAC7A1EC" authority="(Westerlund, 1879)" authorityName="Cochl." baseAuthorityName="Westerlund" baseAuthorityYear="1879" box="[834,1394,1962,1989]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="nanum" subGenus="Turritus">
<emphasis id="B97EEABDFF97FFBAFC9B1AF5FC68A1ED" box="[834,989,1962,1988]" italics="true" pageId="83" pageNumber="84">Cochlostoma</emphasis>
(
<emphasis id="B97EEABDFF97FFBAFC2F1AF4FBB9A1ED" box="[1014,1036,1963,1988]" italics="true" pageId="83" pageNumber="84">T.</emphasis>
)
<emphasis id="B97EEABDFF97FFBAFBFC1AF4FBC0A1ED" box="[1061,1141,1963,1988]" italics="true" pageId="83" pageNumber="84">nanum</emphasis>
(
<bibRefCitation id="EF9B4B5EFF97FFBAFB551AF5FADCA1EC" author="Westerlund" box="[1164,1385,1962,1989]" firstAuthor="Westerlund" pageId="83" pageNumber="84" pagination="156 - 168" refId="ref38386" refString="Westerlund C. A. 1879. Malakozoologische Beitrage. Jahrbucher der deutschen malakozoologischen Gesellschaft 6: 156 - 168." type="journal article" year="1879">Westerlund, 1879</bibRefCitation>
)
</taxonomicName>
; cyan =
<taxonomicName id="4C0A4D2CFF97FFBAFED51A91FE0CA1C1" box="[268,441,1998,2024]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="elegans" subGenus="Turritus">
<emphasis id="B97EEABDFF97FFBAFED51A91FE92A1C1" box="[268,295,1998,2024]" italics="true" pageId="83" pageNumber="84">C.</emphasis>
(
<emphasis id="B97EEABDFF97FFBAFEE11A91FEFBA1CE" box="[312,334,1998,2023]" italics="true" pageId="83" pageNumber="84">T.</emphasis>
)
<emphasis id="B97EEABDFF97FFBAFE861A91FE0CA1C1" box="[351,441,1998,2024]" italics="true" pageId="83" pageNumber="84">elegans</emphasis>
</taxonomicName>
. Black center for the samples with amplified 16S.
</paragraph>
</caption>
<caption id="DF756627FF90FFBDFF641BA5FC9DA1C1" ID-DOI="http://doi.org/10.5281/zenodo.10853848" ID-Zenodo-Dep="10853848" httpUri="https://zenodo.org/record/10853848/files/figure.png" pageId="84" pageNumber="85" startId="84.[189,232,1786,1812]" targetBox="[248,1340,265,1732]" targetPageId="84">
<paragraph id="8BB536AFFF90FFBDFF641BA5FC9DA1C1" blockId="84.[189,1399,1786,2024]" pageId="84" pageNumber="85">
<emphasis id="B97EEABDFF90FFBDFF641BA5FEAFA13D" bold="true" box="[189,282,1786,1812]" pageId="84" pageNumber="85">Fig. 73.</emphasis>
Shell and female genitals of Clade B samples with amplified 16S.
<emphasis id="B97EEABDFF90FFBDFBF61BA5FBF1A13D" bold="true" box="[1071,1092,1786,1812]" pageId="84" pageNumber="85">A</emphasis>
.
<taxonomicName id="4C0A4D2CFF90FFBDFB8A1BA5FE11A111" authority="(Westerlund, 1879)" authorityName="Cochl." baseAuthorityName="Westerlund" baseAuthorityYear="1879" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="84" pageNumber="85" phylum="Mollusca" rank="species" species="nanum" subGenus="Turritus">
<emphasis id="B97EEABDFF90FFBDFB8A1BA5FB5BA13D" box="[1107,1262,1786,1812]" italics="true" pageId="84" pageNumber="85">Cochlostoma</emphasis>
(
<emphasis id="B97EEABDFF90FFBDFAD91BA4FAA3A13D" box="[1280,1302,1787,1812]" italics="true" pageId="84" pageNumber="85">T.</emphasis>
)
<emphasis id="B97EEABDFF90FFBDFAFE1BA4FAC2A13D" box="[1319,1399,1787,1812]" italics="true" pageId="84" pageNumber="85">nanum</emphasis>
(
<bibRefCitation id="EF9B4B5EFF90FFBDFF1C1A41FE2EA111" author="Westerlund" box="[197,411,1822,1848]" firstAuthor="Westerlund" pageId="84" pageNumber="85" pagination="156 - 168" refId="ref38386" refString="Westerlund C. A. 1879. Malakozoologische Beitrage. Jahrbucher der deutschen malakozoologischen Gesellschaft 6: 156 - 168." type="journal article" year="1879">Westerlund, 1879</bibRefCitation>
)
</taxonomicName>
, Tepke, AT (NHMW-109000/AL/00703).
<emphasis id="B97EEABDFF90FFBDFC441A41FC04A111" bold="true" box="[925,945,1822,1848]" pageId="84" pageNumber="85">B</emphasis>
.
<taxonomicName id="4C0A4D2CFF90FFBDFC1B1A41FAF8A111" authority=", Mt Sneznik 1764" authorityName=", Mt Sneznik" authorityYear="1764" box="[962,1357,1822,1848]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="84" pageNumber="85" phylum="Mollusca" rank="species" species="nanum" subGenus="Turritus">
<emphasis id="B97EEABDFF90FFBDFC1B1A41FC68A111" box="[962,989,1822,1848]" italics="true" pageId="84" pageNumber="85">C.</emphasis>
(
<emphasis id="B97EEABDFF90FFBDFC281A40FBB2A111" box="[1009,1031,1823,1848]" italics="true" pageId="84" pageNumber="85">T.</emphasis>
)
<emphasis id="B97EEABDFF90FFBDFBC31A40FBDFA111" box="[1050,1130,1823,1848]" italics="true" pageId="84" pageNumber="85">nanum</emphasis>
, Mt Snežnik 1764
</taxonomicName>
, SI (WdM-7005).
<emphasis id="B97EEABDFF90FFBDFEB21A1EFE35A172" bold="true" box="[363,384,1857,1883]" pageId="84" pageNumber="85">C</emphasis>
.
<taxonomicName id="4C0A4D2CFF90FFBDFE4B1A1EFCABA172" authority=", Mt Risnjak, 1400" authorityName=", Mount Risnjak, HR" authorityYear="1400" box="[402,798,1857,1883]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="84" pageNumber="85" phylum="Mollusca" rank="species" species="nanum" subGenus="Turritus">
<emphasis id="B97EEABDFF90FFBDFE4B1A1EFE18A172" box="[402,429,1857,1883]" italics="true" pageId="84" pageNumber="85">C.</emphasis>
(
<emphasis id="B97EEABDFF90FFBDFE181A1DFE62A172" box="[449,471,1858,1883]" italics="true" pageId="84" pageNumber="85">T.</emphasis>
)
<emphasis id="B97EEABDFF90FFBDFE331A1DFD8FA172" box="[490,570,1858,1883]" italics="true" pageId="84" pageNumber="85">nanum</emphasis>
, Mt Risnjak, 1400
</taxonomicName>
m a.s.l., HR (WdM-5666).
<emphasis id="B97EEABDFF90FFBDFBAA1A1EFB3DA172" bold="true" box="[1139,1160,1857,1883]" pageId="84" pageNumber="85">D</emphasis>
.
<taxonomicName id="4C0A4D2CFF90FFBDFB401A1EFEC3A157" authority="(Wagner, 1906)" authorityName="Cochl." baseAuthorityName="Wagner" baseAuthorityYear="1906" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="84" pageNumber="85" phylum="Mollusca" rank="species" species="mostarense" subGenus="Turritus">
<emphasis id="B97EEABDFF90FFBDFB401A1EFB01A172" box="[1177,1204,1857,1883]" italics="true" pageId="84" pageNumber="85">C.</emphasis>
(
<emphasis id="B97EEABDFF90FFBDFB111A1DFB6BA172" box="[1224,1246,1858,1883]" italics="true" pageId="84" pageNumber="85">T.</emphasis>
)
<emphasis id="B97EEABDFF90FFBDFB2B1A1DFAC2A172" box="[1266,1399,1858,1883]" italics="true" pageId="84" pageNumber="85">mostarense</emphasis>
(
<bibRefCitation id="EF9B4B5EFF90FFBDFF1C1A3BFED9A157" author="Wagner" box="[197,364,1892,1918]" firstAuthor="Wagner" pageId="84" pageNumber="85" pagination="121 - 140" refId="ref38295" refString="Wagner A. J. 1906. Neue Formen und Fundorte der Genera Pomatias Studer und Auritus Westerlund. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 38 (2 - 3): 92 - 101, 121 - 140." type="journal article" year="1906">Wagner, 1906</bibRefCitation>
)
</taxonomicName>
, Mostar, BeH (EZ-1091).
<emphasis id="B97EEABDFF90FFBDFD701A3BFD08A157" bold="true" box="[681,701,1892,1918]" pageId="84" pageNumber="85">E</emphasis>
.
<taxonomicName id="4C0A4D2CFF90FFBDFD131A3AFB86A157" authority="(Clessin, 1879)" authorityName="Orig." authorityYear="1897" baseAuthorityName="Clessin" baseAuthorityYear="1879" box="[714,1075,1892,1919]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="84" pageNumber="85" phylum="Mollusca" rank="species" species="elegans" subGenus="Turritus">
<emphasis id="B97EEABDFF90FFBDFD131A3AFD50A156" box="[714,741,1893,1919]" italics="true" pageId="84" pageNumber="85">C.</emphasis>
(
<emphasis id="B97EEABDFF90FFBDFD2F1A3AFCB9A157" box="[758,780,1893,1918]" italics="true" pageId="84" pageNumber="85">T.</emphasis>
)
<emphasis id="B97EEABDFF90FFBDFCC51A3BFCC3A157" box="[796,886,1892,1918]" italics="true" pageId="84" pageNumber="85">elegans</emphasis>
(
<bibRefCitation id="EF9B4B5EFF90FFBDFC5C1A3BFB9FA157" author="Clessin" box="[901,1066,1892,1918]" firstAuthor="Clessin" pageId="84" pageNumber="85" pagination="116 - 125" refId="ref36680" refString="Clessin S. 1879. Zur Molluskenfauna Croatiens. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 11 (8 - 9): 116 - 125." type="journal article" year="1879">Clessin, 1879</bibRefCitation>
)
</taxonomicName>
, Razbojište near Oltari, HR (WdM-6896).
<emphasis id="B97EEABDFF90FFBDFEBE1AD8FECCA188" bold="true" box="[359,377,1927,1953]" pageId="84" pageNumber="85">F</emphasis>
.
<taxonomicName id="4C0A4D2CFF90FFBDFE5F1AD7FD66A188" authority="(Hirc, 1881)" baseAuthorityName="Hirc" baseAuthorityYear="1881" box="[390,723,1927,1954]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="84" pageNumber="85" phylum="Mollusca" rank="species" species="stossichi" subGenus="Turritus">
<emphasis id="B97EEABDFF90FFBDFE5F1AD7FE14A18B" box="[390,417,1928,1954]" italics="true" pageId="84" pageNumber="85">C.</emphasis>
(
<emphasis id="B97EEABDFF90FFBDFE681AD7FE72A188" box="[433,455,1928,1953]" italics="true" pageId="84" pageNumber="85">T.</emphasis>
)
<emphasis id="B97EEABDFF90FFBDFE0F1AD8FD8EA188" box="[470,571,1927,1953]" italics="true" pageId="84" pageNumber="85">stossichi</emphasis>
(
<bibRefCitation id="EF9B4B5EFF90FFBDFD901AD8FD7FA188" author="Hirc" box="[585,714,1927,1953]" firstAuthor="Hirc" pageId="84" pageNumber="85" pagination="519 - 530" refId="ref37052" refString="Hirc D. 1881. Die Mollusken-Fauna des liburnischen Karstes. Verhandlungen der kaiserlich-koniglichen Zoologisch-Botanischen Gesellschaft in Wien 30: 519 - 530." type="journal article" year="1881">Hirc, 1881</bibRefCitation>
)
</taxonomicName>
, 2 km NNW of Meja, HR (WdM-1240).
<emphasis id="B97EEABDFF90FFBDFB731AD8FB74A188" bold="true" box="[1194,1217,1927,1953]" pageId="84" pageNumber="85">G</emphasis>
.
<taxonomicName id="4C0A4D2CFF90FFBDFB171AD7FEC0A1EC" authority="(Wagner, 1897)" authorityName="Cochl." baseAuthorityName="Wagner" baseAuthorityYear="1897" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="84" pageNumber="85" phylum="Mollusca" rank="species" species="braueri" subGenus="Turritus">
<emphasis id="B97EEABDFF90FFBDFB171AD7FB5CA18B" box="[1230,1257,1928,1954]" italics="true" pageId="84" pageNumber="85">C.</emphasis>
(
<emphasis id="B97EEABDFF90FFBDFB201AD7FABAA188" box="[1273,1295,1928,1953]" italics="true" pageId="84" pageNumber="85">T.</emphasis>
)
<emphasis id="B97EEABDFF90FFBDFAC71AD8FAC3A188" box="[1310,1398,1927,1953]" italics="true" pageId="84" pageNumber="85">braueri</emphasis>
(
<bibRefCitation id="EF9B4B5EFF90FFBDFF1C1AF4FED9A1EC" author="Wagner" box="[197,364,1963,1989]" firstAuthor="Wagner" pageId="84" pageNumber="85" pagination="565 - 632" refId="ref38226" refString="Wagner A. J. 1897. Monographie der Gattung Pomatias Studer. Denkschriften der kaiserlichen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse 64: 565 - 632. https: // doi. org / 10.5962 / bhl. title. 10792" type="journal article" year="1897">Wagner, 1897</bibRefCitation>
)
</taxonomicName>
, Klek Mt, HR (EZ-0068).
<emphasis id="B97EEABDFF90FFBDFD731AF4FD74A1EC" bold="true" box="[682,705,1963,1989]" pageId="84" pageNumber="85">H</emphasis>
. NFS142, Škrčko Jezero, MONT (TxEx-du0044).
<emphasis id="B97EEABDFF90FFBDFACB1AF4FAA8A1EC" bold="true" box="[1298,1309,1963,1989]" pageId="84" pageNumber="85">I</emphasis>
.
<taxonomicName id="4C0A4D2CFF90FFBDFAF51AF4FE92A1C1" authority="Zallot &amp; Kamchev &amp; Schilthuizen &amp; Fehér &amp; Mattia &amp; Gittenberger, 2024" authorityName="Zallot &amp; Kamchev &amp; Schilthuizen &amp; Fehér &amp; Mattia &amp; Gittenberger" authorityYear="2024" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="84" pageNumber="85" phylum="Mollusca" rank="species" species="hallgassi" status="sp. nov." subGenus="Turritus">
<emphasis id="B97EEABDFF90FFBDFAF51AF4FAF2A1EC" box="[1324,1351,1963,1989]" italics="true" pageId="84" pageNumber="85">C.</emphasis>
(
<emphasis id="B97EEABDFF90FFBDFA8E1AF4FAD8A1ED" box="[1367,1389,1963,1988]" italics="true" pageId="84" pageNumber="85">T.</emphasis>
)
<emphasis id="B97EEABDFF90FFBDFF641A91FE92A1C1" box="[189,295,1998,2024]" italics="true" pageId="84" pageNumber="85">hallgassi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C6FF90FFBDFEF71A91FE3EA1C1" box="[302,395,1998,2024]" pageId="84" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
, Monte Petrella 1010, I (EZ-1118).
</paragraph>
</caption>
<caption id="DF756627FF91FFBCFF641A1DFB1EA1A9" ID-DOI="http://doi.org/10.5281/zenodo.10853852" ID-Zenodo-Dep="10853852" httpUri="https://zenodo.org/record/10853852/files/figure.png" pageId="85" pageNumber="86" startId="85.[189,232,1858,1884]" targetBox="[189,1398,265,1809]" targetPageId="85">
<paragraph id="8BB536AFFF91FFBCFF641A1DFB1EA1A9" blockId="85.[189,1399,1858,1920]" pageId="85" pageNumber="86">
<emphasis id="B97EEABDFF91FFBCFF641A1DFEABA175" bold="true" box="[189,286,1858,1884]" pageId="85" pageNumber="86">Fig. 74.</emphasis>
Type of
<taxonomicName id="4C0A4D2CFF91FFBCFE481A1CFD44A174" authority="Hirc, 1881" authorityName="Hirc" authorityYear="1881" box="[401,753,1858,1885]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="85" pageNumber="86" phylum="Mollusca" rank="species" species="clessini">
<emphasis id="B97EEABDFF91FFBCFE481A1CFDD1A175" box="[401,612,1858,1884]" italics="true" pageId="85" pageNumber="86">Pomatias clessini</emphasis>
<bibRefCitation id="EF9B4B5EFF91FFBCFDB61A1DFD44A174" author="Hirc" box="[623,753,1858,1885]" firstAuthor="Hirc" pageId="85" pageNumber="86" pagination="519 - 530" refId="ref37052" refString="Hirc D. 1881. Die Mollusken-Fauna des liburnischen Karstes. Verhandlungen der kaiserlich-koniglichen Zoologisch-Botanischen Gesellschaft in Wien 30: 519 - 530." type="journal article" year="1881">Hirc, 1881</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C0A4D2CFF91FFBCFCD11A1DFA92A174" authority="(Westerlund, 1879)" authorityName="Cochl." baseAuthorityName="Westerlund" baseAuthorityYear="1879" box="[776,1319,1858,1885]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="85" pageNumber="86" phylum="Mollusca" rank="species" species="nanum" subGenus="Turritus">
<emphasis id="B97EEABDFF91FFBCFCD11A1DFC16A175" box="[776,931,1858,1884]" italics="true" pageId="85" pageNumber="86">Cochlostoma</emphasis>
(
<emphasis id="B97EEABDFF91FFBCFC611A1CFC7BA175" box="[952,974,1859,1884]" italics="true" pageId="85" pageNumber="86">T.</emphasis>
)
<emphasis id="B97EEABDFF91FFBCFC3A1A1CFB86A175" box="[995,1075,1859,1884]" italics="true" pageId="85" pageNumber="86">nanum</emphasis>
(
<bibRefCitation id="EF9B4B5EFF91FFBCFB9F1A1DFAABA174" author="Westerlund" box="[1094,1310,1858,1885]" firstAuthor="Westerlund" pageId="85" pageNumber="86" pagination="156 - 168" refId="ref38386" refString="Westerlund C. A. 1879. Malakozoologische Beitrage. Jahrbucher der deutschen malakozoologischen Gesellschaft 6: 156 - 168." type="journal article" year="1879">Westerlund, 1879</bibRefCitation>
)
</taxonomicName>
, ♂, in Wagners collection (NHMW without collection number) from Mount Risnjak, HR.
</paragraph>
</caption>
<paragraph id="8BB536AFFF92FFBFFF641C31FB29A7E7" blockId="86.[189,1399,366,463]" pageId="86" pageNumber="87">
For what is known, the species lives in a few scattered alpine localities not connected by suitable habitats. In the phylogenetic trees the
<taxonomicName id="4C0A4D2CFF92FFBFFDD71CCDFDEBA782" authorityName="Cochl." baseAuthorityName="Westerlund" baseAuthorityYear="1879" box="[526,606,402,427]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="86" pageNumber="87" phylum="Mollusca" rank="species" species="nanum">
<emphasis id="B97EEABDFF92FFBFFDD71CCDFDEBA782" box="[526,606,402,427]" italics="true" pageId="86" pageNumber="87">nanum</emphasis>
</taxonomicName>
samples do not cluster together and the intraspecific p-distances are of the same magnitude of the ones with samples of the other species of the subclade.
</paragraph>
</subSubSection>
</treatment>
</document>