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<mods:title id="31E5A2A6B8D6AB4A395DDF61F8A74481">A morphological, phylogenetic and phylogeographic reappraisal of the land crabs Gecarcinus quadratus De Saussure, 1853, and G. lateralis Fréminville in Guérin 1832 (Decapoda: Gecarcinidae). Are they different species?</mods:title>
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Fréminville in Guérin, 1832
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(
<figureCitation id="13312A3FFFAFFF9E268E2FC100D55198" box="[159,420,180,207]" captionStart-0="FIGURE" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1922,1946]" captionStartId-1="2.[151,250,1928,1952]" captionStartId-2="3.[151,250,1801,1825]" captionTargetBox-0="[151,1436,561,1899]" captionTargetBox-1="[197,1389,181,1904]" captionTargetBox-2="[155,1432,181,1777]" captionTargetId-0="figure-201@1.[151,1436,561,1899]" captionTargetId-1="figure-26@2.[197,1389,181,1904]" captionTargetId-2="figure-17@3.[155,1432,181,1777]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Dorsal view of:A) Gecarcinus lateralis from Río Prieto, Veracruz (CNCR 33942) and B) G. quadratus from Barra de Boca del Cielo, Chiapas (CNCR 34630)." captionText-1="FIGURE 2. Gecarcinus lateralis from Río Prieto, Veracruz (CNCR 33942): A) frontal view, B) ventral vie, C) rows of spines in the dactyli of the pereiopods, D) major chela. Gecarcinus quadratus from Boca del Cielo, Chiapas (CNCR 34630): E) frontal view, F) ventral view, G) rows of spines in the dactyli of the pereiopods, H) major chela." captionText-2="FIGURE 3. Left male first gonopod (G1): Cephalic view, A) G. lateralis from Río Prieto, Veracruz (CNCR 33942); B) G. quadratus from Boca del Cielo, Chiapas (CNCR 34630). Lateral view, C) G. quadratus from Boca del Cielo, Chiapas (CNCR 34630). Ventral surface, D) G. lateralis female from Barra de Sontecomapan, Veracruz (CNCR 33943); E) G. quadratus female from Playa Higuera Blanca, Nayarit (CNCR 34012); F) G. ruricola male from Cuba (MNHN-IU-2017-8392). Frontal view, G) G. ruricola from Cuba (MNHN-IU-2017-8392) (G. ruricola photos modified from Guinot et al., 2018)." figureDoi-0="http://doi.org/10.5281/zenodo.5552005" figureDoi-1="http://doi.org/10.5281/zenodo.5552007" figureDoi-2="http://doi.org/10.5281/zenodo.5552009" httpUri-0="https://zenodo.org/record/5552005/files/figure.png" httpUri-1="https://zenodo.org/record/5552007/files/figure.png" httpUri-2="https://zenodo.org/record/5552009/files/figure.png" pageId="12" pageNumber="227">Figs. 1A; 2A-D; 3A, D</figureCitation>
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<emphasis id="B97EEAA8FFAFFF9E26862F8C00145046" box="[151,357,249,273]" italics="true" pageId="12" pageNumber="227">Gecarcinus lateralis</emphasis>
Fréminville in Guérin 1832
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: 7, pl. 5, figs. 1, 1a–b.
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<taxonomicName id="4C0A4D39FFAFFF9E26862E6D003C5067" authorityName="Guerin" authorityYear="1832" box="[151,333,280,304]" class="Malacostraca" family="Ocypodidae" genus="Ocypoda" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="227" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B97EEAA8FFAFFF9E26862E6D003C5067" box="[151,333,280,304]" italics="true" pageId="12" pageNumber="227">Ocypoda lateralis</emphasis>
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.— 
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<bibRefCitation id="EF9B4B4BFFAFFF9E277A2E6D03225067" author="Freminville, M." box="[363,595,279,304]" pageId="12" pageNumber="227" pagination="213 - 224" refId="ref12257" refString="Freminville, M. (1835) Notice sur les tourlouraux ou crabes de terre des Antilles. Annales des Sciences Naturelles, Zoologie, 2 (2), 213 - 224. [https: // www. biodiversitylibrary. org / item / 111230 # page / 219 / mode / 1 up]" type="journal article" year="1835">Fréminville 1835: 224</bibRefCitation>
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.
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<emphasis id="B97EEAA8FFAFFF9E26862E4200145019" box="[151,357,310,334]" italics="true" pageId="12" pageNumber="227">Gecarcinus lateralis</emphasis>
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.—H. 
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<bibRefCitation id="EF9B4B4BFFAFFF9E27B52E4303D55019" author="Milne-Edwards, H." box="[420,676,310,334]" pageId="12" pageNumber="227" refId="ref14629" refString="Milne-Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant lanatomie, la physiologie et la classification de ces animaux. Vol. 2. Atlas (1834, 1837, 1840). Librairie encyclopedique de Roret, Paris, 532 pp. https: // doi. org / 10.5962 / bhl. title. 6234" type="book" year="1837">Milne-Edwards 1837: 27</bibRefCitation>
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, pl. 18, figs. 1–6. 
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<bibRefCitation id="EF9B4B4BFFAFFF9E254B2E4205365019" author="De Saussure, H." box="[858,1095,310,334]" pageId="12" pageNumber="227" pagination="417 - 496" refId="ref12007" refString="De Saussure, H. (1858) Memoire sur divers Crustaces nouveaux des Antilles et du Mexique. Memoires de la Societe de physique et d'histoire naturelle de Geneve, 14, 417 - 496. [https: // www. biodiversitylibrary. org / item / 41153 # page / 585 / mode / 1 up]" type="journal article" year="1858">De Saussure 1858: 440</bibRefCitation>
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<emphasis id="B97EEAA8FFAFFF9E26862E200003503A" box="[151,370,341,365]" italics="true" pageId="12" pageNumber="227">Geocarcinus lateralis</emphasis>
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.— 
<treatmentCitation id="0AAB10ABFFAFFF9E27812E200335503A" author="Young, C. G." box="[400,580,341,365]" page="239" pageId="12" pageNumber="227" year="1900">
<bibRefCitation id="EF9B4B4BFFAFFF9E27812E200335503A" author="Young, C. G." box="[400,580,341,365]" pageId="12" pageNumber="227" pagination="98 - 99" refId="ref16570" refString="Young, C. G. (1900) The stalk-eyed Crustacea of British Guiana, West Indies and Bermuda. Nature, 64, 98 - 99. https: // doi. org / 10.1038 / 064098 a 0" type="journal article" year="1900">Young 1900: 239</bibRefCitation>
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.
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<emphasis id="B97EEAA8FFAFFF9E26862E01001450DB" box="[151,357,372,396]" italics="true" pageId="12" pageNumber="227">Gecarcinus lateralis</emphasis>
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.— 
<treatmentCitation id="0AAB10ABFFAFFF9E27922E01033A50DC" author="Rathbun, M. J." box="[387,587,371,396]" page="355" pageId="12" pageNumber="227" year="1918">
<bibRefCitation id="EF9B4B4BFFAFFF9E27922E01033A50DC" author="Rathbun, M. J." box="[387,587,371,396]" pageId="12" pageNumber="227" pagination="1 - 461" refId="ref15576" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, 1 - 461. https: // doi. org / 10.5479 / si. 03629236.97. i" type="journal article" year="1918">Rathbun 1918: 355</bibRefCitation>
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. 
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<emphasis id="B97EEAA8FFAFFF9E26862EE6033A50FD" box="[151,587,402,426]" italics="true" pageId="12" pageNumber="227">Gecarcinus (Gecarcinus) lateralis lateralis</emphasis>
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.— 
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<bibRefCitation id="EF9B4B4BFFAFFF9E24782EE7025550FD" author="Turkay, M." box="[617,804,402,426]" pageId="12" pageNumber="227" pagination="333 - 354" refId="ref16252" refString="Turkay, M. (1970) Die Gecarcinidae Amerikas. Miteinem Anhanguber Ucides Rahbun (Crustacea: Decapoda). Senckenbergiana Biologica, 51 (5 / 6), 333 - 354. [https: // research. nhm. org / pdfs / 27369 / 27369. pdf]" type="journal article" year="1970">Türkay 1970: 337</bibRefCitation>
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<taxonomicName id="4C0A4D39FFAFFF9E26862EC40081509E" ID-CoL="3FGCW" authority="Freminville" authorityName="Freminville in Guerin" authorityYear="1832" box="[151,496,433,457]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="227" phylum="Arthropoda" rank="species" species="lateralis" subGenus="Gecarcinus">
<emphasis id="B97EEAA8FFAFFF9E26862EC40081509E" box="[151,496,433,457]" italics="true" pageId="12" pageNumber="227">Gecarcinus (Gecarcinus) lateralis</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="0AAB10ABFFAFFF9E241F2EC403B8509E" author="Turkay, M." box="[526,713,433,457]" page="974" pageId="12" pageNumber="227" year="1973">
<bibRefCitation id="EF9B4B4BFFAFFF9E241F2EC403B8509E" author="Turkay, M." box="[526,713,433,457]" pageId="12" pageNumber="227" pagination="969 - 978" refId="ref16305" refString="Turkay, M. (1973) Bemerkungen zu einigen Landkrabben (Crustacea, Decapoda). Bulletin du Museum National D'Histoire Naturelle, (142), 969 - 978. [https: // decapoda. nhm. org / pdfs / 31464 / 31464. pdf]" type="journal article" year="1973">Türkay 1973: 974</bibRefCitation>
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</paragraph>
<paragraph id="8BB536BAFFAFFF9E26862EA5033250BF" blockId="12.[151,1437,249,611]" box="[151,579,464,488]" pageId="12" pageNumber="227">
<treatmentCitationGroup id="AB1A1194FFAFFF9E26862EA5033250BF" box="[151,579,464,488]" pageId="12" pageNumber="227">
<taxonomicName id="4C0A4D39FFAFFF9E26862EA5001450BF" ID-CoL="3FGCW" authority="Freminville" authorityName="Freminville in Guerin" authorityYear="1832" box="[151,357,464,488]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="227" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B97EEAA8FFAFFF9E26862EA5001450BF" box="[151,357,464,488]" italics="true" pageId="12" pageNumber="227">Gecarcinus lateralis</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="0AAB10ABFFAFFF9E27952EA5034E50BF" author="Powers, L. W." box="[388,575,464,488]" page="139" pageId="12" pageNumber="227" year="1977">
<bibRefCitation id="EF9B4B4BFFAFFF9E27952EA5034E50BF" author="Powers, L. W." box="[388,575,464,488]" pageId="12" pageNumber="227" pagination="3 - 190" refId="ref15426" refString="Powers, L. W. (1977) A catalogue and bibliography to the crabs (Brachyura) of the Gulf of Mexico. Contributions in Marine Science, Supplement, 20, 3 - 190. [https: // research. nhm. org / pdfs / 4119 / 4119. pdf]" type="journal article" year="1977">Powers 1977: 139</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BB536BAFFAFFF9E26862E9A02B65351" blockId="12.[151,1437,249,611]" box="[151,967,494,518]" pageId="12" pageNumber="227">
<treatmentCitationGroup id="AB1A1194FFAFFF9E26862E9A02B65351" box="[151,967,494,518]" pageId="12" pageNumber="227">
<taxonomicName id="4C0A4D39FFAFFF9E26862E9A033A5351" ID-CoL="3FGCW" authority="lateralis" box="[151,587,494,518]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="227" phylum="Arthropoda" rank="subSpecies" species="lateralis" subGenus="Gecarcinus" subSpecies="lateralis">
<emphasis id="B97EEAA8FFAFFF9E26862E9A033A5351" box="[151,587,494,518]" italics="true" pageId="12" pageNumber="227">Gecarcinus (Gecarcinus) lateralis lateralis</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="0AAB10ABFFAFFF9E247B2E9A02B55351" author="von Prahl, H. &amp; Manjarres, G." box="[618,964,494,518]" page="158" pageId="12" pageNumber="227" year="1984">
<bibRefCitation id="EF9B4B4BFFAFFF9E247B2E9A02B55351" author="von Prahl, H. &amp; Manjarres, G." box="[618,964,494,518]" pageId="12" pageNumber="227" pagination="149 - 168" refId="ref16355" refString="von Prahl, H. &amp; Manjarres, G. (1984) Cangrejos Gecarcinidos (Crustacea; Gecarcinidae) de Colombia. Caldasia, 14 (66), 149 - 168. [https: // revistas. unal. edu. co / index. php / cal / article / view / 34905 / 35174]" type="journal article" year="1984">von Prahl &amp; Manjarrés 1984: 158</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BB536BAFFAFFF9E26862D7805585335" blockId="12.[151,1437,249,611]" pageId="12" pageNumber="227">
<treatmentCitationGroup id="AB1A1194FFAFFF9E26862D7804EC5372" box="[151,1437,525,549]" pageId="12" pageNumber="227">
<taxonomicName id="4C0A4D39FFAFFF9E26862D7800145372" ID-CoL="3FGCW" authority="Freminville" authorityName="Freminville in Guerin" authorityYear="1832" box="[151,357,525,549]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="227" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B97EEAA8FFAFFF9E26862D7800145372" box="[151,357,525,549]" italics="true" pageId="12" pageNumber="227">Gecarcinus lateralis</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="0AAB10ABFFAFFF9E27952D78030E5372" author="Abele, L. G. &amp; Kim, W." box="[388,639,525,549]" page="661" pageId="12" pageNumber="227" year="1986">
<bibRefCitation id="EF9B4B4BFFAFFF9E27952D78030E5372" author="Abele, L. G. &amp; Kim, W." box="[388,639,525,549]" pageId="12" pageNumber="227" pagination="1 - 760" refId="ref10796" refString="Abele, L. G. &amp; Kim, W. (1986) An illustrated guide to the marine decapod crustaceans of Florida. Department of Environmental Regulation, State of Florida. Technical Series, 8 (1), 1 - 760. [https: // decapoda. nhm. org / pdfs / 12365 / 12365 - 001. pdf]" type="journal article" year="1986">Abele &amp; Kim 1986: 661</bibRefCitation>
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<treatmentCitation id="0AAB10ABFFAFFF9E24982D7802CB5372" author="Hernandez-Aguilera, J. L. &amp; Toral-Almazan, R. E. &amp; Ruiz-Nuno, J. A." box="[649,954,525,549]" page="92" pageId="12" pageNumber="227" year="1996">
<bibRefCitation id="EF9B4B4BFFAFFF9E24982D7802CB5372" author="Hernandez-Aguilera, J. L. &amp; Toral-Almazan, R. E. &amp; Ruiz-Nuno, J. A." box="[649,954,525,549]" pageId="12" pageNumber="227" refId="ref13019" refString="Hernandez-Aguilera, J. L., Toral-Almazan, R. E. &amp; Ruiz-Nuno, J. A. (1996) Especies catalogadas de Crustaceos estomatopodos y decapodos para el Golfo de Mexico, Rio Bravo, Tamps. A Progreso, Yuc. DGONSM-CONABIO, Mexico, D. F., 132 pp. [http: // www. conabio. gob. mx / institucion / proyectos / resultados / FichapubB 035. pdf]" type="book" year="1996">Hernández-Aguilera 1996: 92</bibRefCitation>
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<treatmentCitation id="0AAB10ABFFAFFF9E25D52D7805B65372" author="Schubart, C. D. &amp; Cannicci, S. &amp; Vannini, M. &amp; Fratini, S." box="[964,1223,525,549]" page="195" pageId="12" pageNumber="227" year="2006">
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Schubart 
<emphasis id="B97EEAA8FFAFFF9E22342D7B05235372" box="[1061,1106,525,549]" italics="true" pageId="12" pageNumber="227">et al</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C3106531FFAFFF9E26862DE500B9550C" pageId="12" pageNumber="227" type="materials_examined">
<paragraph id="8BB536BAFFAFFF9E26862DE500B9550C" blockId="12.[151,1437,656,2015]" pageId="12" pageNumber="227">
<emphasis id="B97EEAA8FFAFFF9E26862DE500F153FC" bold="true" box="[151,384,656,683]" pageId="12" pageNumber="227">Material examined.</emphasis>
<materialsCitation id="3B623CE7FFAFFF9E27992DE402135399" ID-GBIF-Occurrence="3382024312" collectingDate="2014-04-04" collectionCode="CCLEA" collectorName="La Laguna &amp; G. Rodriguez" country="Mexico" location="Boca de Catan" municipality="Municipality San Fernando" pageId="12" pageNumber="227" specimenCode="C86-07182" specimenCount="3" specimenCount-female="2" specimenCount-juvenile="1" stateProvince="Tamaulipas">
<collectingCountry id="F31D762AFFAFFF9E27992DE4008B53FC" box="[392,506,657,683]" name="Mexico" pageId="12" pageNumber="227">MEXICO</collectingCountry>
. 
<collectingRegion id="49CEF858FFAFFF9E24152DE503F853FD" box="[516,649,656,682]" country="Mexico" name="Tamaulipas" pageId="12" pageNumber="227">Tamaulipas</collectingRegion>
: 
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, 
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, 
<collectingMunicipality id="6BD1ACC0FFAFFF9E226D2DE501C45399" pageId="12" pageNumber="227">Municipality San Fernando</collectingMunicipality>
, 
<collectorName id="26FF536CFFAFFF9E26D12DC0004A5398" box="[192,315,693,719]" pageId="12" pageNumber="227">La Laguna</collectorName>
, 
<location id="8ED56061FFAFFF9E27542DC000985398" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E27542DC000985398" box="[325,489,692,719]" country="Mexico" municipality="Municipality San Fernando" name="Boca de Catan" pageId="12" pageNumber="227" stateProvince="Tamaulipas">Boca de Catán</location>
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<date id="FFB4107AFFAFFF9E27E32DC003F65398" box="[498,647,693,719]" pageId="12" pageNumber="227" value="2014-04-04">
<collectingDate id="EFF0E992FFAFFF9E27E32DC003F65398" box="[498,647,693,719]" pageId="12" pageNumber="227" value="2014-04-04">Apr 04, 2014</collectingDate>
</date>
, coll. 
<collectorName id="26FF536CFFAFFF9E24D92DC002135399" box="[712,866,692,719]" pageId="12" pageNumber="227">G. Rodríguez</collectorName>
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; 
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<specimenCount id="9D0CFD33FFAFFF9E257D2DC002BE5399" box="[876,975,692,719]" pageId="12" pageNumber="227" type="female">1 female</specimenCount>
, 
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<location id="8ED56061FFAFFF9E22C32DC1006153A5" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E22C32DC1006153A5" country="Mexico" name="Municipality Soto La Marina" pageId="12" pageNumber="227" stateProvince="Tamaulipas">Municipality Soto La Marina</location>
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<collectorName id="26FF536CFFAFFF9E270A2DAC00F253A4" box="[283,387,729,755]" pageId="12" pageNumber="227">La Pesca</collectorName>
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<collectingDate id="EFF0E992FFAFFF9E279F2DAC035353A4" box="[398,546,729,755]" pageId="12" pageNumber="227" value="2014-10-20">Oct 20, 2014</collectingDate>
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<collectorName id="26FF536CFFAFFF9E24772DAC027153A5" box="[614,768,728,755]" pageId="12" pageNumber="227">G. Rodríguez</collectorName>
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; 
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, 
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jar D, same locality and collector data as previous, 
<date id="FFB4107AFFAFFF9E27FF2D8803FB5241" box="[494,650,764,791]" pageId="12" pageNumber="227" value="2016-09-24">
<collectingDate id="EFF0E992FFAFFF9E27FF2D8803FB5241" box="[494,650,764,791]" pageId="12" pageNumber="227" value="2016-09-24">Sept 24, 2016</collectingDate>
</date>
</materialsCitation>
.– 
<materialsCitation id="3B623CE7FFAFFF9E24B32D8803FE526D" ID-GBIF-Occurrence="3382024314" collectingDate="2015-01-24" collectionCode="CCLEA" collectorName="G. Rodriguez" country="Mexico" location="Barra de Cazones" municipality="Municipality Cazones" pageId="12" pageNumber="227" specimenCode="C86-06601" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave">
<collectingRegion id="49CEF858FFAFFF9E24B32D88027B5240" box="[674,778,765,791]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="12" pageNumber="227">Veracruz</collectingRegion>
: 
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, 
<collectingMunicipality id="6BD1ACC0FFAFFF9E224A2D8904255240" box="[1115,1364,764,791]" pageId="12" pageNumber="227">Municipality Cazones</collectingMunicipality>
, 
<location id="8ED56061FFAFFF9E234F2D88006B526C" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E234F2D88006B526C" country="Mexico" municipality="Municipality Cazones" name="Barra de Cazones" pageId="12" pageNumber="227" stateProvince="Veracruz de Ignacio de la Llave">Barra de Cazones</location>
, 
<date id="FFB4107AFFAFFF9E27352C5400C5526C" box="[292,436,800,827]" pageId="12" pageNumber="227" value="2015-01-24">
<collectingDate id="EFF0E992FFAFFF9E27352C5400C5526C" box="[292,436,800,827]" pageId="12" pageNumber="227" value="2015-01-24">Jan 24, 2015</collectingDate>
</date>
, coll. 
<collectorName id="26FF536CFFAFFF9E27E72C5403FE526D" box="[502,655,800,827]" pageId="12" pageNumber="227">G. Rodríguez</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B623CE7FFAFFF9E248B2C5400D95208" ID-GBIF-Occurrence="3382024301" collectionCode="R" collectorName="Cabada" country="Mexico" location="Rio Prieto" municipality="Municipality Angel" pageId="12" pageNumber="227" specimenCode="CNCR 33942" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Veracruz">
<specimenCount id="9D0CFD33FFAFFF9E248B2C540383526D" box="[666,754,800,827]" pageId="12" pageNumber="227" type="male">2 males</specimenCount>
, 
<specimenCount id="9D0CFD33FFAFFF9E24EC2C540211526D" box="[765,864,800,827]" pageId="12" pageNumber="227" type="female">1 female</specimenCount>
, 
<specimenCode id="DBAC9EC1FFAFFF9E257A2C540577526C" box="[875,1030,801,827]" collectionCode="CNCR" pageId="12" pageNumber="227">CNCR 33942</specimenCode>
, 
<collectingMunicipality id="6BD1ACC0FFAFFF9E22002C55059D526A" box="[1041,1260,800,829]" pageId="12" pageNumber="227">Municipality Ángel</collectingMunicipality>
<collectionCode id="ED1BAE7FFFAFFF9E22E32C540479526C" box="[1266,1288,801,827]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="227">R</collectionCode>
. 
<collectorName id="26FF536CFFAFFF9E23032C550418526D" box="[1298,1385,800,826]" pageId="12" pageNumber="227">Cabada</collectorName>
, 
<location id="8ED56061FFAFFF9E23652C5401AC5209" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E23652C5401AC5209" country="Mexico" municipality="Municipality Angel" name="Rio Prieto" pageId="12" pageNumber="227" stateProvince="Veracruz">Río Prieto</location>
, coll. anonymous
</materialsCitation>
; 
<materialsCitation id="3B623CE7FFAFFF9E27A42C3004E95208" ID-GBIF-Occurrence="3382024325" box="[437,1432,836,863]" collectingDate="2017-04-24" collectionCode="CNCR" country="Mexico" location="Barra de Sontecomapan" municipality="Municipality Catemaco" pageId="12" pageNumber="227" specimenCode="CNCR 33943" specimenCount="2" specimenCount-male="2" stateProvince="Veracruz">
<specimenCount id="9D0CFD33FFAFFF9E27A42C3003615209" box="[437,528,836,863]" pageId="12" pageNumber="227" type="male">2 males</specimenCount>
, 
<specimenCode id="DBAC9EC1FFAFFF9E240C2C3003CA5208" box="[541,699,837,863]" collectionCode="CNCR" pageId="12" pageNumber="227">CNCR 33943</specimenCode>
, 
<collectingMunicipality id="6BD1ACC0FFAFFF9E24D62C3102A55208" box="[711,980,836,863]" pageId="12" pageNumber="227">Municipality Catemaco</collectingMunicipality>
, 
<location id="8ED56061FFAFFF9E25F12C3005835208" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E25F12C3005835208" box="[992,1266,836,863]" country="Mexico" municipality="Municipality Catemaco" name="Barra de Sontecomapan" pageId="12" pageNumber="227" stateProvince="Veracruz">Barra de Sontecomapan</location>
, 
<date id="FFB4107AFFAFFF9E22EC2C3004E95208" box="[1277,1432,837,863]" pageId="12" pageNumber="227" value="2017-04-24">
<collectingDate id="EFF0E992FFAFFF9E22EC2C3004E95208" box="[1277,1432,837,863]" pageId="12" pageNumber="227" value="2017-04-24">Apr 24, 2017</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B623CE7FFAFFF9E26862C1D03E652D5" ID-GBIF-Occurrence="3382024319" box="[151,663,872,899]" collectorName="I. Toledano &amp; J. L. Villalobos" country="Mexico" location="Moreno" pageId="12" pageNumber="227" specimenCount="1" stateProvince="Veracruz">
colls. 
<collectorName id="26FF536CFFAFFF9E26CA2C1C002C52D5" box="[219,349,872,899]" pageId="12" pageNumber="227">I. Toledano</collectorName>
, E. 
<location id="8ED56061FFAFFF9E27982C1C009452D4" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E27982C1C009452D4" box="[393,485,873,899]" country="Mexico" name="Moreno" pageId="12" pageNumber="227" stateProvince="Veracruz">Moreno</location>
, 
<collectorName id="26FF536CFFAFFF9E27E12C1C03E652D5" box="[496,663,872,899]" pageId="12" pageNumber="227">J.L. Villalobos</collectorName>
</materialsCitation>
.– 
<materialsCitation id="3B623CE7FFAFFF9E24A12C1D032C52F1" ID-GBIF-Occurrence="3382024321" collectingDate="2006-10-14" collectionCode="CCLEA" collectorName="G. Rodriguez" country="Mexico" location="Playa Bruja" municipality="Municipality Paraiso" pageId="12" pageNumber="227" specimenCode="C86-06495" specimenCount="1" specimenCount-male="1" stateProvince="Tabasco">
<collectingRegion id="49CEF858FFAFFF9E24A12C1D026152D5" box="[688,784,872,898]" country="Mexico" name="Tabasco" pageId="12" pageNumber="227">Tabasco</collectingRegion>
: 
<specimenCount id="9D0CFD33FFAFFF9E250A2C1C021B52D5" box="[795,874,872,899]" pageId="12" pageNumber="227" type="male">1 male</specimenCount>
, 
<collectionCode id="ED1BAE7FFFAFFF9E25672C1C02A152D4" box="[886,976,873,899]" pageId="12" pageNumber="227">CCLEA</collectionCode>
<specimenCode id="DBAC9EC1FFAFFF9E25C92C1D052952D4" box="[984,1112,872,899]" collectionCode="CCLEA" pageId="12" pageNumber="227">C86-06495</specimenCode>
, 
<collectingMunicipality id="6BD1ACC0FFAFFF9E22742C1D042352D4" box="[1125,1362,872,899]" pageId="12" pageNumber="227">Municipality Paraíso</collectingMunicipality>
, 
<location id="8ED56061FFAFFF9E234F2C1D01A952F1" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E234F2C1D01A952F1" country="Mexico" municipality="Municipality Paraiso" name="Playa Bruja" pageId="12" pageNumber="227" stateProvince="Tabasco">Playa Bruja</location>
, 
<date id="FFB4107AFFAFFF9E26F42CF8000D52F1" box="[229,380,908,935]" pageId="12" pageNumber="227" value="2006-10-14">
<collectingDate id="EFF0E992FFAFFF9E26F42CF8000D52F1" box="[229,380,908,935]" pageId="12" pageNumber="227" value="2006-10-14">Oct 14, 2006</collectingDate>
</date>
, coll. 
<collectorName id="26FF536CFFAFFF9E27D02CF8032C52F1" box="[449,605,908,935]" pageId="12" pageNumber="227">G. Rodríguez</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B623CE7FFAFFF9E24782CF80369529D" ID-GBIF-Occurrence="3382024322" collectingDate="2007-04-06" collectionCode="CCLEA" collectorName="G. Rodriguez." country="Mexico" location="Chiltepec" municipality="Municipality Paraiso" pageId="12" pageNumber="227" specimenCode="C86-06500" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tabasco">
<specimenCount id="9D0CFD33FFAFFF9E24782CF803C852F1" box="[617,697,908,935]" pageId="12" pageNumber="227" type="male">1 male</specimenCount>
, 
<specimenCount id="9D0CFD33FFAFFF9E24D42CF8025A52F1" box="[709,811,908,935]" pageId="12" pageNumber="227" type="female">1 female</specimenCount>
, 
<collectionCode id="ED1BAE7FFFAFFF9E25262CF802E752F0" box="[823,918,909,935]" pageId="12" pageNumber="227">CCLEA</collectionCode>
, 
<specimenCode id="DBAC9EC1FFAFFF9E25B32CF9055252F0" box="[930,1059,908,935]" collectionCode="CCLEA" pageId="12" pageNumber="227">C86-06500</specimenCode>
, 
<collectingMunicipality id="6BD1ACC0FFAFFF9E223E2CF9046C52F0" box="[1071,1309,908,935]" pageId="12" pageNumber="227">Municipality Paraíso</collectingMunicipality>
, 
<location id="8ED56061FFAFFF9E233B2CF904E952F1" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E233B2CF904E952F1" box="[1322,1432,908,934]" country="Mexico" municipality="Municipality Paraiso" name="Chiltepec" pageId="12" pageNumber="227" stateProvince="Tabasco">Chiltepec</location>
, 
<date id="FFB4107AFFAFFF9E26862CC40043529C" box="[151,306,944,971]" pageId="12" pageNumber="227" value="2007-04-06">
<collectingDate id="EFF0E992FFAFFF9E26862CC40043529C" box="[151,306,944,971]" pageId="12" pageNumber="227" value="2007-04-06">Apr 06, 2007</collectingDate>
</date>
, coll. 
<collectorName id="26FF536CFFAFFF9E27692CC40369529D" box="[376,536,944,971]" pageId="12" pageNumber="227">G. Rodríguez.</collectorName>
</materialsCitation>
– 
<materialsCitation id="3B623CE7FFAFFF9E243E2CC5042E529C" ID-GBIF-Occurrence="3382024324" box="[559,1375,944,971]" collectingDate="2017-04-06" collectionCode="CNCR" country="Mexico" location="Cayo Arcas" pageId="12" pageNumber="227" specimenCode="CNCR 34011" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Campeche">
<collectingRegion id="49CEF858FFAFFF9E243E2CC503DB529D" box="[559,682,944,970]" country="Mexico" name="Campeche" pageId="12" pageNumber="227">Campeche</collectingRegion>
: 
<specimenCount id="9D0CFD33FFAFFF9E24A42CC40261529D" box="[693,784,944,971]" pageId="12" pageNumber="227" type="male">2 males</specimenCount>
, 
<specimenCount id="9D0CFD33FFAFFF9E250C2CC402F2529D" box="[797,899,944,971]" pageId="12" pageNumber="227" type="female">1 female</specimenCount>
, 
<specimenCode id="DBAC9EC1FFAFFF9E259E2CC40558529C" box="[911,1065,945,971]" collectionCode="CNCR" pageId="12" pageNumber="227">CNCR 34011</specimenCode>
<location id="8ED56061FFAFFF9E22232CC405C9529C" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E22232CC405C9529C" box="[1074,1208,945,971]" country="Mexico" name="Cayo Arcas" pageId="12" pageNumber="227" stateProvince="Campeche">Cayo Arcas</location>
, 
<date id="FFB4107AFFAFFF9E22D52CC4042E529C" box="[1220,1375,944,971]" pageId="12" pageNumber="227" value="2017-04-06">
<collectingDate id="EFF0E992FFAFFF9E22D52CC4042E529C" box="[1220,1375,944,971]" pageId="12" pageNumber="227" value="2017-04-06">Apr 06, 2017</collectingDate>
</date>
</materialsCitation>
, coll. anonymous; 
<materialsCitation id="3B623CE7FFAFFF9E27382CA000C75545" ID-GBIF-Occurrence="3382024326" collectingDate="2009-08-01" collectionCode="CCY" collectorName="C. Cinthya Delgado." country="Mexico" location="Arrecife Alacranes" municipality="Municipality Progreso" pageId="12" pageNumber="227" specimenCode="YUC-CC-255-11, 001024" specimenCount="1" specimenCount-female="1" stateProvince="Campeche">
<specimenCount id="9D0CFD33FFAFFF9E27382CA000FE52B9" box="[297,399,980,1007]" pageId="12" pageNumber="227" type="female">1 female</specimenCount>
, 
<collectionCode id="ED1BAE7FFFAFFF9E278D2CA000A452B8" box="[412,469,981,1007]" country="Slovakia" httpUri="http://grbio.org/cool/sdab-10ap" name="Culture Collection of Yeasts, Slovak Academy of Sciences, Institute of Chemistry" pageId="12" pageNumber="227">CCY</collectionCode>
<specimenCode id="DBAC9EC1FFAFFF9E27CD2CA103EC52B8" box="[476,669,980,1007]" collectionCode="YUC-CC-" pageId="12" pageNumber="227">YUC-CC-255-11</specimenCode>
- 
<specimenCode id="DBAC9EC1FFAFFF9E24B42CA1038C52B8" box="[677,765,980,1007]" collectionCode="YUC-CC-" pageId="12" pageNumber="227">001024</specimenCode>
, 
<collectingMunicipality id="6BD1ACC0FFAFFF9E251B2CA1057B52B8" box="[778,1034,980,1007]" pageId="12" pageNumber="227">Municipality Progreso</collectingMunicipality>
, 
<location id="8ED56061FFAFFF9E22072CA1059C52B9" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E22072CA1059C52B9" box="[1046,1261,980,1006]" country="Mexico" municipality="Municipality Progreso" name="Arrecife Alacranes" pageId="12" pageNumber="227" stateProvince="Campeche">Arrecife Alacranes</location>
, 
<date id="FFB4107AFFAFFF9E22E92CA004E952B8" box="[1272,1432,981,1007]" pageId="12" pageNumber="227" value="2009-08-01">
<collectingDate id="EFF0E992FFAFFF9E22E92CA004E952B8" box="[1272,1432,981,1007]" pageId="12" pageNumber="227" value="2009-08-01">Aug 01, 2009</collectingDate>
</date>
, coll. 
<collectorName id="26FF536CFFAFFF9E26DE2C8C00C75545" box="[207,438,1016,1043]" pageId="12" pageNumber="227">C. Cinthya Delgado.</collectorName>
</materialsCitation>
– 
<materialsCitation id="3B623CE7FFAFFF9E27DA2C8D030D5560" ID-GBIF-Occurrence="3382024311" collectingDate="2017-03-12" collectionCode="CNCR" collectorName="Municipality Othon P. Blanco &amp; Anonymous" country="Mexico" location="Playa Mahahual" pageId="12" pageNumber="227" specimenCode="CNCR 34010" specimenCount="3" specimenCount-juvenile="2" specimenCount-male="1" stateProvince="Quintana Roo">
<collectingRegion id="49CEF858FFAFFF9E27DA2C8D031D5544" box="[459,620,1016,1043]" country="Mexico" name="Quintana Roo" pageId="12" pageNumber="227">Quintana Roo</collectingRegion>
: 
<specimenCount id="9D0CFD33FFAFFF9E24662C8C03B45545" box="[631,709,1016,1043]" pageId="12" pageNumber="227" type="male">1 male</specimenCount>
, 
<specimenCount id="9D0CFD33FFAFFF9E24C02C8C023F5545" box="[721,846,1016,1043]" pageId="12" pageNumber="227" type="juvenile">2 juveniles</specimenCount>
, 
<specimenCode id="DBAC9EC1FFAFFF9E25482C8C02875544" box="[857,1014,1017,1043]" collectionCode="CNCR" pageId="12" pageNumber="227">CNCR 34010</specimenCode>
, 
<collectorName id="26FF536CFFAFFF9E22102C8D04235545" box="[1025,1362,1016,1043]" pageId="12" pageNumber="227">Municipality Othón P. Blanco</collectorName>
, 
<location id="8ED56061FFAFFF9E234F2C8D007B5561" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E234F2C8D007B5561" country="Mexico" name="Playa Mahahual" pageId="12" pageNumber="227" stateProvince="Quintana Roo">Playa Mahahual</location>
, 
<date id="FFB4107AFFAFFF9E27042B6800DF5560" box="[277,430,1053,1079]" pageId="12" pageNumber="227" value="2017-03-12">
<collectingDate id="EFF0E992FFAFFF9E27042B6800DF5560" box="[277,430,1053,1079]" pageId="12" pageNumber="227" value="2017-03-12">Mar 12, 2017</collectingDate>
</date>
, coll. 
<collectorName id="26FF536CFFAFFF9E27FE2B68030D5560" box="[495,636,1053,1079]" pageId="12" pageNumber="227">Anonymous</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B623CE7FFAFFF9E24972B68018D550C" ID-GBIF-Occurrence="3382024327" collectingDate="2013-03-28" collectionCode="CNCR" country="Mexico" location="Banco Chinchorro" municipality="Isla Cayo Norte Menor" pageId="12" pageNumber="227" specimenCode="CNCR 30834" specimenCount="1" specimenCount-male="1" stateProvince="Quintana Roo">
<specimenCount id="9D0CFD33FFAFFF9E24972B6803A25561" box="[646,723,1052,1079]" pageId="12" pageNumber="227" type="male">1 male</specimenCount>
, 
<specimenCode id="DBAC9EC1FFAFFF9E24CF2B68020B5560" box="[734,890,1053,1079]" collectionCode="CNCR" pageId="12" pageNumber="227">CNCR 30834</specimenCode>
, 
<collectingMunicipality id="6BD1ACC0FFAFFF9E25942B6905F65560" box="[901,1159,1052,1079]" pageId="12" pageNumber="227">Isla Cayo Norte Menor</collectingMunicipality>
, 
<location id="8ED56061FFAFFF9E22832B6804135561" LSID="urn:lsid:plazi:treatment:03A387ACFFAFFF9F26862FE200F75274:8ED56061FFAFFF9E22832B6804135561" box="[1170,1378,1052,1079]" country="Mexico" municipality="Isla Cayo Norte Menor" name="Banco Chinchorro" pageId="12" pageNumber="227" stateProvince="Quintana Roo">Banco Chinchorro</location>
, 
<date id="FFB4107AFFAFFF9E237C2B68018D550C" pageId="12" pageNumber="227" value="2013-03-28">
<collectingDate id="EFF0E992FFAFFF9E237C2B68018D550C" pageId="12" pageNumber="227" value="2013-03-28">Mar 28, 2013</collectingDate>
</date>
</materialsCitation>
, coll. anonymous.
</paragraph>
</subSubSection>
<subSubSection id="C3106531FFAFFF9E26D62B1103CD542D" pageId="12" pageNumber="227" type="diagnosis">
<paragraph id="8BB536BAFFAFFF9E26D62B1103CD542D" blockId="12.[151,1437,656,2015]" pageId="12" pageNumber="227">
<emphasis id="B97EEAA8FFAFFF9E26D62B1100335529" bold="true" box="[199,322,1124,1150]" pageId="12" pageNumber="227">Diagnosis.</emphasis>
Male abdomen triangular; telson campanulate, narrower than sixth abdominal somite, lateral margins slightly marked, tip rounded. Sixth somite with convex lateral margins, forming lateral shoulder (
<figureCitation id="13312A3FFFAFFF9E23252BFD04FE55F4" box="[1332,1423,1160,1187]" captionStart="FIGURE 2" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[197,1389,181,1904]" captionTargetId="figure-26@2.[197,1389,181,1904]" captionTargetPageId="2" captionText="FIGURE 2. Gecarcinus lateralis from Río Prieto, Veracruz (CNCR 33942): A) frontal view, B) ventral vie, C) rows of spines in the dactyli of the pereiopods, D) major chela. Gecarcinus quadratus from Boca del Cielo, Chiapas (CNCR 34630): E) frontal view, F) ventral view, G) rows of spines in the dactyli of the pereiopods, H) major chela." figureDoi="http://doi.org/10.5281/zenodo.5552007" httpUri="https://zenodo.org/record/5552007/files/figure.png" pageId="12" pageNumber="227">Fig. 2B</figureCitation>
). Female abdomen subcircular, completely or almost completely covering thoracic sterna; telson triangular, as wide as posterior margin of sixth somite, lateral margins straight, smooth (
<figureCitation id="13312A3FFFAFFF9E25B32BA5028C55BC" box="[930,1021,1232,1259]" captionStart="FIGURE 3" captionStartId="3.[151,250,1801,1825]" captionTargetBox="[155,1432,181,1777]" captionTargetId="figure-17@3.[155,1432,181,1777]" captionTargetPageId="3" captionText="FIGURE 3. Left male first gonopod (G1): Cephalic view, A) G. lateralis from Río Prieto, Veracruz (CNCR 33942); B) G. quadratus from Boca del Cielo, Chiapas (CNCR 34630). Lateral view, C) G. quadratus from Boca del Cielo, Chiapas (CNCR 34630). Ventral surface, D) G. lateralis female from Barra de Sontecomapan, Veracruz (CNCR 33943); E) G. quadratus female from Playa Higuera Blanca, Nayarit (CNCR 34012); F) G. ruricola male from Cuba (MNHN-IU-2017-8392). Frontal view, G) G. ruricola from Cuba (MNHN-IU-2017-8392) (G. ruricola photos modified from Guinot et al., 2018)." figureDoi="http://doi.org/10.5281/zenodo.5552009" httpUri="https://zenodo.org/record/5552009/files/figure.png" pageId="12" pageNumber="227">Fig. 3D</figureCitation>
). Antero-lateral margins of sternite 3 slightly convex, those of sternite 4 straight, with slight concavity (
<figureCitation id="13312A3FFFAFFF9E259B2B81029F5458" box="[906,1006,1268,1295]" captionStart="FIGURE 2" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[197,1389,181,1904]" captionTargetId="figure-26@2.[197,1389,181,1904]" captionTargetPageId="2" captionText="FIGURE 2. Gecarcinus lateralis from Río Prieto, Veracruz (CNCR 33942): A) frontal view, B) ventral vie, C) rows of spines in the dactyli of the pereiopods, D) major chela. Gecarcinus quadratus from Boca del Cielo, Chiapas (CNCR 34630): E) frontal view, F) ventral view, G) rows of spines in the dactyli of the pereiopods, H) major chela." figureDoi="http://doi.org/10.5281/zenodo.5552007" httpUri="https://zenodo.org/record/5552007/files/figure.png" pageId="12" pageNumber="227">Figs. 2B</figureCitation>
, 
<figureCitation id="13312A3FFFAFFF9E25EA2B8005505458" box="[1019,1057,1269,1295]" captionStart="FIGURE 3" captionStartId="3.[151,250,1801,1825]" captionTargetBox="[155,1432,181,1777]" captionTargetId="figure-17@3.[155,1432,181,1777]" captionTargetPageId="3" captionText="FIGURE 3. Left male first gonopod (G1): Cephalic view, A) G. lateralis from Río Prieto, Veracruz (CNCR 33942); B) G. quadratus from Boca del Cielo, Chiapas (CNCR 34630). Lateral view, C) G. quadratus from Boca del Cielo, Chiapas (CNCR 34630). Ventral surface, D) G. lateralis female from Barra de Sontecomapan, Veracruz (CNCR 33943); E) G. quadratus female from Playa Higuera Blanca, Nayarit (CNCR 34012); F) G. ruricola male from Cuba (MNHN-IU-2017-8392). Frontal view, G) G. ruricola from Cuba (MNHN-IU-2017-8392) (G. ruricola photos modified from Guinot et al., 2018)." figureDoi="http://doi.org/10.5281/zenodo.5552009" httpUri="https://zenodo.org/record/5552009/files/figure.png" pageId="12" pageNumber="227">3D</figureCitation>
). Shallow endostomial ridges delimiting efferent branchial channel, poorly defining this region. Third maxilliped with merus generally concealing epistome. Adult male chelae different in size and robustness; inner surface of palm of major chela with longitudinal tuberculated ridge, used as stridulatory plectrum.
</paragraph>
</subSubSection>
<subSubSection id="C3106531FFAFFF9F26D62AF104035321" lastPageId="13" lastPageNumber="228" pageId="12" pageNumber="227" type="description">
<paragraph id="8BB536BAFFAFFF9E26D62AF100135720" blockId="12.[151,1437,656,2015]" pageId="12" pageNumber="227">
<emphasis id="B97EEAA8FFAFFF9E26D62AF1002854C9" bold="true" box="[199,345,1412,1438]" pageId="12" pageNumber="227">Description.</emphasis>
Based on adult male with G1 well developed (
<quantity id="4CF29B5FFFAFFF9E254F2AF102B254C8" box="[862,963,1412,1439]" metricMagnitude="-2" metricUnit="m" metricValue="5.43" pageId="12" pageNumber="227" unit="mm" value="54.3">54.3 mm</quantity>
CL, 
<quantity id="4CF29B5FFFAFFF9E25EB2AF0052C54C8" box="[1018,1117,1413,1439]" metricMagnitude="-2" metricUnit="m" metricValue="7.17" pageId="12" pageNumber="227" unit="mm" value="71.7">71.7 mm</quantity>
CW). Carapace transversely oval, wider anteriorly, strongly arched at level of branchial regions, anterolateral margin incipient, posterior portion of external orbital angle slightly marked, armed with minuscule granules. Regions well defined, delimited by shallow grooves. Branchial regions prominent. Fronto-orbital margin about 1/2 of maximum carapace width (Fig. 1A). Front about 1/4 length of carapace, deflexed, surface slightly concave, occasionally straight, inferior margin granulated, raised from surface, straight, generally concealing basal segments of antennules; frontal lobes poorly defined (
<figureCitation id="13312A3FFFAFFF9E26EB292900245720" box="[250,341,1628,1655]" captionStart="FIGURE 2" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[197,1389,181,1904]" captionTargetId="figure-26@2.[197,1389,181,1904]" captionTargetPageId="2" captionText="FIGURE 2. Gecarcinus lateralis from Río Prieto, Veracruz (CNCR 33942): A) frontal view, B) ventral vie, C) rows of spines in the dactyli of the pereiopods, D) major chela. Gecarcinus quadratus from Boca del Cielo, Chiapas (CNCR 34630): E) frontal view, F) ventral view, G) rows of spines in the dactyli of the pereiopods, H) major chela." figureDoi="http://doi.org/10.5281/zenodo.5552007" httpUri="https://zenodo.org/record/5552007/files/figure.png" pageId="12" pageNumber="227">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="8BB536BAFFAFFF9E26D629F501895625" blockId="12.[151,1437,656,2015]" pageId="12" pageNumber="227">
Orbits delimited by well defined, smooth superior border. Infra-orbital margin granulated, interrupted by deep notch, continued as triangular plate forming mesial lobe. Suborbital ridge well marked (stridulating crest of 
<bibRefCitation id="EF9B4B4BFFAFFF9E235F29D1006357B4" author="Guinot, D. &amp; Ng, N. K. &amp; Moreno, P. A. R." pageId="12" pageNumber="227" pagination="547 - 605" refId="ref12756" refString="Guinot, D., Ng, N. K. &amp; Moreno, P. A. R. (2018) Review of grapsoid families for the establishment of a new family for Leptograpsodes Montgomery, 1931, and a new genus of Gecarcinidae H. Milne-Edwards, 1837 (Crustacea, Decapoda, Brachyura, Grapsoidea MacLeay, 1838). Zoosystema, 40, 547 - 605. https: // doi. org / 10.5252 / zoosystema 2018 v 40 a 26" type="journal article" year="2018">
Guinot 
<emphasis id="B97EEAA8FFAFFF9E268629BC01B957B5" box="[151,200,1736,1762]" italics="true" pageId="12" pageNumber="227">et al</emphasis>
. 2018
</bibRefCitation>
), extending from superior border of epistome to subhepatic suture. Surface of anterolateral wall of carapace reticulated (pars strident). Third maxillipeds not closing completely, leaving rhomboidal gap; merus covering epistome anteriorly, anterior border with medial notch, occasionally straight; ischium longer than merus, subequal in width. Palp short, hidden and articulated behind merus, directed laterally (
<figureCitation id="13312A3FFFAFFF9E25F52841054C5618" box="[996,1085,1844,1871]" captionStart="FIGURE 2" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[197,1389,181,1904]" captionTargetId="figure-26@2.[197,1389,181,1904]" captionTargetPageId="2" captionText="FIGURE 2. Gecarcinus lateralis from Río Prieto, Veracruz (CNCR 33942): A) frontal view, B) ventral vie, C) rows of spines in the dactyli of the pereiopods, D) major chela. Gecarcinus quadratus from Boca del Cielo, Chiapas (CNCR 34630): E) frontal view, F) ventral view, G) rows of spines in the dactyli of the pereiopods, H) major chela." figureDoi="http://doi.org/10.5281/zenodo.5552007" httpUri="https://zenodo.org/record/5552007/files/figure.png" pageId="12" pageNumber="227">Fig. 2A</figureCitation>
). Short exognath hidden behind ischium.
</paragraph>
<paragraph id="8BB536BAFFAFFF9F26D6280800B351AE" blockId="12.[151,1437,656,2015]" lastBlockId="13.[151,1436,151,803]" lastPageId="13" lastPageNumber="228" pageId="12" pageNumber="227">
Strong heterochely in males, subequal chelipeds in females; fingers longer than palm, with corneous tips; longitudinal rows of granules throughout surface; cutting margins bearing 6–8 irregularly spaced blunt teeth; dactyl slightly curved inwards; fingers gapping (
<figureCitation id="13312A3FFFAFFF9E247728B103B15688" box="[614,704,1988,2015]" captionStart="FIGURE 2" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[197,1389,181,1904]" captionTargetId="figure-26@2.[197,1389,181,1904]" captionTargetPageId="2" captionText="FIGURE 2. Gecarcinus lateralis from Río Prieto, Veracruz (CNCR 33942): A) frontal view, B) ventral vie, C) rows of spines in the dactyli of the pereiopods, D) major chela. Gecarcinus quadratus from Boca del Cielo, Chiapas (CNCR 34630): E) frontal view, F) ventral view, G) rows of spines in the dactyli of the pereiopods, H) major chela." figureDoi="http://doi.org/10.5281/zenodo.5552007" httpUri="https://zenodo.org/record/5552007/files/figure.png" pageId="12" pageNumber="227">Fig. 2D</figureCitation>
). Internal surface of palm swollen, with distinct longitudinal ridge along midsection, used as stridulatory plectrum, internal depression next to articulation with carpus. Carpus subtriangular, dorso-mesial margin smooth in large organisms, spinulous in small ones. Merus elongated, subtriangular, internal border granulated.
</paragraph>
<paragraph id="8BB536BAFFAEFF9F26D62E74034A50D4" blockId="13.[151,1436,151,803]" pageId="13" pageNumber="228">
Pereiopods (2–4) long, slender, with short setae; merus elongated, surface striated; carpus subtriangular, shorter than propodus, or almost of same length in fifth pair, dorsal margin toothed; propodus shorter than dactyl, dorsal and ventral margins formed by short, slender spines; dactyli armed with 4–6 rows of spines, those on lateral carina poorly developed to absent (
<figureCitation id="13312A3FFFAEFF9F27C42E1C035F50D4" box="[469,558,361,387]" captionStart="FIGURE 2" captionStartId="2.[151,250,1928,1952]" captionTargetBox="[197,1389,181,1904]" captionTargetId="figure-26@2.[197,1389,181,1904]" captionTargetPageId="2" captionText="FIGURE 2. Gecarcinus lateralis from Río Prieto, Veracruz (CNCR 33942): A) frontal view, B) ventral vie, C) rows of spines in the dactyli of the pereiopods, D) major chela. Gecarcinus quadratus from Boca del Cielo, Chiapas (CNCR 34630): E) frontal view, F) ventral view, G) rows of spines in the dactyli of the pereiopods, H) major chela." figureDoi="http://doi.org/10.5281/zenodo.5552007" httpUri="https://zenodo.org/record/5552007/files/figure.png" pageId="13" pageNumber="228">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8BB536BAFFAEFF9F26D62EF9038D50F1" blockId="13.[151,1436,151,803]" box="[199,764,396,423]" pageId="13" pageNumber="228">Male and female abdomen as in species diagnosis.</paragraph>
<paragraph id="8BB536BAFFAEFF9F26D62EDA03EA509E" blockId="13.[151,1436,151,803]" box="[199,667,431,457]" pageId="13" pageNumber="228">
Male G1, as in genus diagnosis (
<figureCitation id="13312A3FFFAEFF9F24252EDA03FF509E" box="[564,654,431,457]" captionStart="FIGURE 3" captionStartId="3.[151,250,1801,1825]" captionTargetBox="[155,1432,181,1777]" captionTargetId="figure-17@3.[155,1432,181,1777]" captionTargetPageId="3" captionText="FIGURE 3. Left male first gonopod (G1): Cephalic view, A) G. lateralis from Río Prieto, Veracruz (CNCR 33942); B) G. quadratus from Boca del Cielo, Chiapas (CNCR 34630). Lateral view, C) G. quadratus from Boca del Cielo, Chiapas (CNCR 34630). Ventral surface, D) G. lateralis female from Barra de Sontecomapan, Veracruz (CNCR 33943); E) G. quadratus female from Playa Higuera Blanca, Nayarit (CNCR 34012); F) G. ruricola male from Cuba (MNHN-IU-2017-8392). Frontal view, G) G. ruricola from Cuba (MNHN-IU-2017-8392) (G. ruricola photos modified from Guinot et al., 2018)." figureDoi="http://doi.org/10.5281/zenodo.5552009" httpUri="https://zenodo.org/record/5552009/files/figure.png" pageId="13" pageNumber="228">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="8BB536BAFFAEFF9F26D62EA4029B5359" blockId="13.[151,1436,151,803]" pageId="13" pageNumber="228">
Female (
<quantity id="4CF29B5FFFAEFF9F27362EA4000750BB" box="[295,374,465,492]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="228" unit="mm" value="39.0">39 mm</quantity>
CL, 
<quantity id="4CF29B5FFFAEFF9F27BD2EA4035250BB" box="[428,547,465,492]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="5.0" metricValueMin="3.3" pageId="13" pageNumber="228" unit="mm" value="41.5" valueMax="50.0" valueMin="33.0">33–50 mm</quantity>
CW), with gonopore on sixth plate of thoracic sternum, protruding. Ring around operculum oval, with sharp edges; operculum a large, oval, protruding lobe.
</paragraph>
<paragraph id="8BB536BAFFAEFF9F26D62D62033C5304" blockId="13.[151,1436,151,803]" pageId="13" pageNumber="228">
<emphasis id="B97EEAA8FFAEFF9F26D62D6200335366" box="[199,322,535,561]" italics="true" pageId="13" pageNumber="228">Coloration</emphasis>
. Dorsal surface of carapace dark, lateral margins orange, posteriorly light cream. Chelipeds and lower portion of front orange (Fig. 1A)
</paragraph>
<paragraph id="8BB536BAFFAEFF9F26D62D2904035321" blockId="13.[151,1436,151,803]" box="[199,1394,604,631]" pageId="13" pageNumber="228">
<emphasis id="B97EEAA8FFAEFF9F26D62D29000F5321" bold="true" box="[199,382,604,630]" pageId="13" pageNumber="228">Measurements.</emphasis>
Males: 
<quantity id="4CF29B5FFFAEFF9F27C92D29030A5320" box="[472,635,604,631]" metricMagnitude="-2" metricUnit="m" metricValue="3.885" metricValueMax="4.91" metricValueMin="2.86" pageId="13" pageNumber="228" unit="mm" value="38.85" valueMax="49.1" valueMin="28.6">28.6–49.1 mm</quantity>
CL; 
<quantity id="4CF29B5FFFAEFF9F24A52D2902265320" box="[692,855,604,631]" metricMagnitude="-2" metricUnit="m" metricValue="4.92" metricValueMax="6.45" metricValueMin="3.3899999999999997" pageId="13" pageNumber="228" unit="mm" value="49.2" valueMax="64.5" valueMin="33.9">33.9–64.5 mm</quantity>
CW. Females 
<quantity id="4CF29B5FFFAEFF9F25EB2D2905F95320" box="[1018,1160,604,631]" metricMagnitude="-2" metricUnit="m" metricValue="3.315" metricValueMax="3.9" metricValueMin="2.73" pageId="13" pageNumber="228" unit="mm" value="33.15" valueMax="39.0" valueMin="27.3">27.3–39 mm</quantity>
CL, 
<quantity id="4CF29B5FFFAEFF9F22D12D2904485320" box="[1216,1337,604,631]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="5.0" metricValueMin="3.3" pageId="13" pageNumber="228" unit="mm" value="41.5" valueMax="50.0" valueMin="33.0">33–50 mm</quantity>
CW.
</paragraph>
</subSubSection>
<subSubSection id="C3106531FFAEFF9F26D62D0A00F75274" pageId="13" pageNumber="228" type="discussion">
<paragraph id="8BB536BAFFAEFF9F26D62D0A00F75274" blockId="13.[151,1436,151,803]" pageId="13" pageNumber="228">
<emphasis id="B97EEAA8FFAEFF9F26D62D0A004A53CE" bold="true" box="[199,315,639,665]" pageId="13" pageNumber="228">Remarks.</emphasis>
The authorship of the species was attributed to 
<bibRefCitation id="EF9B4B4BFFAEFF9F25722D0A054853CE" author="Freminville, M." box="[867,1081,638,665]" pageId="13" pageNumber="228" pagination="213 - 224" refId="ref12257" refString="Freminville, M. (1835) Notice sur les tourlouraux ou crabes de terre des Antilles. Annales des Sciences Naturelles, Zoologie, 2 (2), 213 - 224. [https: // www. biodiversitylibrary. org / item / 111230 # page / 219 / mode / 1 up]" type="journal article" year="1835">Fréminville (1835)</bibRefCitation>
for many years who originally named organisms from the French West Indies: Marie-Galante, La Désirade, 
<collectingCountry id="F31D762AFFAEFF9F25EC2DD405F553EC" box="[1021,1156,673,699]" name="Guadeloupe" pageId="13" pageNumber="228">Guadeloupe</collectingCountry>
and 
<collectingCountry id="F31D762AFFAEFF9F22AD2DD4044A53EC" box="[1212,1339,673,699]" name="Martinique" pageId="13" pageNumber="228">Martinique</collectingCountry>
, as 
<emphasis id="B97EEAA8FFAEFF9F23772DD7004A5389" italics="true" pageId="13" pageNumber="228">
<taxonomicName id="4C0A4D39FFAEFF9F23772DD700465389" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="228" phylum="Arthropoda" rank="species" species="lateralis">Ocypode lateralis</taxonomicName>
.
</emphasis>
However, 
<bibRefCitation id="EF9B4B4BFFAEFF9F27A92DB003F05389" author="Low, M. E. Y. &amp; Ng, P. K. L. &amp; Evenhuis, N. L." box="[440,641,708,735]" pageId="13" pageNumber="228" pagination="101 - 127" refId="ref14196" refString="Low, M. E. Y., Ng, P. K. L. &amp; Evenhuis, N. L. (2013) Names and publication dates of the Brachyura in F. E. Guerin (Guerin-Meneville) (Crustacea: Decapoda). Zootaxa, 3736 (2), 101 - 127. https: // doi. org / 10.11646 / zootaxa. 3736.2.1" type="journal article" year="2013">
Low 
<emphasis id="B97EEAA8FFAEFF9F27E52DB0035F5389" box="[500,558,708,734]" italics="true" pageId="13" pageNumber="228">et al.</emphasis>
(2013)
</bibRefCitation>
reported that Guérin in 1832 had already used the name “ 
<taxonomicName id="4C0A4D39FFAEFF9F230D2DB100145256" authority="Freminv." authorityName="Freminville in Guerin" authorityYear="1832" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="228" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B97EEAA8FFAEFF9F230D2DB101855256" italics="true" pageId="13" pageNumber="228">Gecarcinus lateralis</emphasis>
Freminv.
</taxonomicName>
” in his 
<emphasis id="B97EEAA8FFAEFF9F27AE2D9202515256" box="[447,800,743,769]" italics="true" pageId="13" pageNumber="228">Iconographie du Règne animal</emphasis>
by G. Cuvier and, hence, he must be recognized as the author of the species.
</paragraph>
</subSubSection>
</treatment>
</document>