treatments-xml/data/03/A3/87/03A38781FFE8FF9EFF70FB6AFE55C564.xml

<document id="72CD634806A07983B99978849B99B595" ID="10.11646/zootaxa.4267.1.1SLASH11021" ID-CLB-Dataset="33061" ID-DOI="10.5281/zenodo.575641" ID-GBIF-Dataset="810b55e4-b8ba-4368-bf9c-8517ee33e7d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="575641" ID-ZooBank="8920893E-E5F8-482A-A60D-7A248D2B0DCB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494577389872" checkinUser="plazi" docAuthor="Skelley, Paul, Xu, Guang, Tang, William, Lindström, Anders J., Marler, Thomas, Khuraijam, Jibankumar Singh, Singh, Rita &amp; Rich, Stephen" docDate="2017" docId="03A38781FFE8FF9EFF70FB6AFE55C564" docLanguage="en" docName="zootaxa.4267.1.1SLASH11021.pdf" docOrigin="Zootaxa 4267 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cycadophila (Cycadophila) collina Skelley, Xu &amp; Tang, new species" docType="treatment" docVersion="8" lastPageNumber="18" masterDocId="FF9AFFF9FFE6FF8FFFE7FFEBFFD5C639" masterDocTitle="Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species" masterLastPageNumber="63" masterPageNumber="1" pageNumber="15" updateTime="1698439510239" updateUser="ExternalLinkService">
<mods:mods id="8C2D0D94F392BC2488CFED20CBA15F03" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B5B16851840AAE72E1CDB11F5F541BFC">
<mods:title id="19EE3FAA3E4283DA6B0DD080637B5A34">Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species</mods:title>
</mods:titleInfo>
<mods:name id="1505828C7C767AE30E014456567638C3" type="personal">
<mods:role id="2BBE2F303B632883F2DEAC050BE09CD9">
<mods:roleTerm id="527377D06757C0C2DAFEB929D3536512">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F8422B59FE7FBA078D1AF9CD3C136999">Skelley, Paul</mods:namePart>
</mods:name>
<mods:name id="FBEEBF95D8408CC81E166D183F704A65" type="personal">
<mods:role id="2C17C19A57932A57E112E9E6360D3A09">
<mods:roleTerm id="B8CF808B4CA686FF91DE21E49E6188CC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08A97DF8D04F3CB5DD2EF7316525B105">Xu, Guang</mods:namePart>
</mods:name>
<mods:name id="B7C88E5DF203DD0EC208ED9012800AF5" type="personal">
<mods:role id="238E9B90A766023F6851BBB9F91E1B9E">
<mods:roleTerm id="C9E5AA2D8AA49052F08A64504691A745">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EF601A449B1BE0DB70FEC9829E9A4DA3">Tang, William</mods:namePart>
</mods:name>
<mods:name id="73871AC96220C4D2CD0B616EBD447B02" type="personal">
<mods:role id="7724EEAE93D9616C53438CC075ECE6B2">
<mods:roleTerm id="7C69F18A826BEE0217881426479BAC9C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F52C413A80064269CE667C93A9534CEA">Lindström, Anders J.</mods:namePart>
</mods:name>
<mods:name id="C7A990EE9A8FF65C874A3570233481FC" type="personal">
<mods:role id="2E7EF179F1E68BB9035B1F2A4396EA27">
<mods:roleTerm id="7CD049E6480CBECE8F188BC5817CF7F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BBC9D2A750AFA0ED34F8A660280578D6">Marler, Thomas</mods:namePart>
</mods:name>
<mods:name id="CAE724D5B3D892FBD7B221D278BC47AF" type="personal">
<mods:role id="A8CF5666ADC08EC842F129C94CE06D5F">
<mods:roleTerm id="CE862B45B40A6F4C898BD51FA017075C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="135F9AAD8EF211221894C338991EC921">Khuraijam, Jibankumar Singh</mods:namePart>
</mods:name>
<mods:name id="63EDC74CE31D35DEBECB21B2D760D2BD" type="personal">
<mods:role id="B2CD0FE83F1C0E18884E12B4069AE3FD">
<mods:roleTerm id="EE1F0DAD5DE32F3DB6914F9BFBDCA8E7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70F02C5175AD83924CF3EEE4E9A5F980">Singh, Rita</mods:namePart>
</mods:name>
<mods:name id="5C0307A8191673F1A282677E8AE16C17" type="personal">
<mods:role id="7DA5E1776670242229C346F27EFC8696">
<mods:roleTerm id="F4885179FF8C1BEA278C93087363308A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="85D0C9008A7C202D623F7AD6211EC4DA">Rich, Stephen</mods:namePart>
</mods:name>
<mods:typeOfResource id="B2C8A2974A1550B3EB00CE5E975A4779">text</mods:typeOfResource>
<mods:relatedItem id="F6EA3D99B334D87302B6AF26C60ED072" type="host">
<mods:titleInfo id="3B39149CC1F69F7FC4E10F7FBE153322">
<mods:title id="780FD51E388276E2B2A1E14DA66C4585">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B2DC3DE7DF0C280F707D4899F867F0E3">
<mods:date id="018AF0F25324AAC309B8C13A2F056242">2017</mods:date>
<mods:detail id="016B74EEF2908562D44CEA53B7C7466F" type="volume">
<mods:number id="3BF1598FF428053AA0D695859DB1FA7D">4267</mods:number>
</mods:detail>
<mods:detail id="4798D14BCF868D3E3A0ECB071944FF11" type="issue">
<mods:number id="0FBC1A47AA85B3145796AF05EAA9C9DC">1</mods:number>
</mods:detail>
<mods:extent id="9244BF75A1C7438F103297F82564460A" unit="page">
<mods:start id="8E60123D67DD90F387BF30DDDF31F19F">1</mods:start>
<mods:end id="13BE3C2EE4B49E970D7A357405ED8AF9">63</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9DF5EF8B3562D6C61F6931F87AB3AEF4">journal article</mods:classification>
<mods:identifier id="6C0B0CAFAA8BF8F2CFEFB2EF72879D63" type="CLB-Dataset">33061</mods:identifier>
<mods:identifier id="105989A629DBD798DAF27D02073496E5" type="DOI">10.5281/zenodo.575641</mods:identifier>
<mods:identifier id="C92A8877A5644E6F500BB80ED547EC42" type="GBIF-Dataset">810b55e4-b8ba-4368-bf9c-8517ee33e7d0</mods:identifier>
<mods:identifier id="0EDACFEB31D492B517AA73C87E34E4D6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E85EC9DFA2CEA0BFC12F8FABE82216B7" type="Zenodo-Dep">575641</mods:identifier>
<mods:identifier id="0364C1677AE9131EF84FDB191EEAA89C" type="ZooBank">8920893E-E5F8-482A-A60D-7A248D2B0DCB</mods:identifier>
</mods:mods>
<treatment id="03A38781FFE8FF9EFF70FB6AFE55C564" ID-DOI="http://doi.org/10.5281/zenodo.5999177" ID-GBIF-Taxon="130809497" ID-Zenodo-Dep="5999177" LSID="urn:lsid:plazi:treatment:03A38781FFE8FF9EFF70FB6AFE55C564" httpUri="http://treatment.plazi.org/id/03A38781FFE8FF9EFF70FB6AFE55C564" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection id="C310651CFFE8FF81FF70FB6AFEA3C285" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BB53697FFE8FF81FF70FB6AFC07C2A2" blockId="14.[151,978,1153,1213]" box="[151,978,1153,1179]" pageId="14" pageNumber="15">
<heading id="D0FD81FBFFE8FF81FF70FB6AFC07C2A2" bold="true" box="[151,978,1153,1179]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B97EEA85FFE8FF81FF70FB6AFC07C2A2" bold="true" box="[151,978,1153,1179]" pageId="14" pageNumber="15">
<taxonomicName id="4C0A4D14FFE8FF81FF70FB6AFCE0C2A2" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[151,821,1153,1179]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="collina" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFE8FF81FF70FB6AFEFAC2A2" bold="true" box="[151,303,1153,1179]" italics="true" pageId="14" pageNumber="15">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFE8FF81FED8FB6AFE03C2A2" bold="true" box="[319,470,1153,1179]" italics="true" pageId="14" pageNumber="15">Cycadophila</emphasis>
) 
<emphasis id="B97EEA85FFE8FF81FE0FFB6AFDEEC2A2" bold="true" box="[488,571,1153,1179]" italics="true" pageId="14" pageNumber="15">collina</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
, 
<taxonomicNameLabel id="A24D57FEFFE8FF81FCA5FB6AFC07C2A2" box="[834,978,1153,1179]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB53697FFE8FF81FF70FB4FFEA3C285" blockId="14.[151,978,1153,1213]" box="[151,374,1188,1213]" pageId="14" pageNumber="15">
<figureCitation id="13312A12FFE8FF81FF70FB4FFF36C285" box="[151,227,1188,1213]" captionStart="FIGURE 2" captionStartId="12.[151,250,1846,1868]" captionTargetBox="[170,1433,205,1803]" captionTargetId="figure@12.[162,1447,195,1811]" captionTargetPageId="12" captionText="FIGURE 2. Key characters of adults: photographs of lateral head and pronotal margin: A) Cycadophila (C.) fupingensis; B) C. (Strobilophila) tansachai; C) C. (C.) yunnanensis; SEM of protibia, ventral view: D) C. (C.) fupingensis; E) C. (C.) debaonica; F) C. (S.) tansachai; G) C. (S.) yangi; photographs of mesotibia, ventral view: H) C. (C.) debaonica female; I) C. (C.) debaonica male; J) C. (C.) lata male; photographs of left half of pronotum: K) C. (C.) collina; L) C. (C.) debaonica; M) C. (S.) kwaiensis; N) C. (S.) tansachai; photographs of mesofemur of major male: O) C. (C.) collina ex Vietnam; P) C. (C.) debaonica." httpUri="https://zenodo.org/record/575643/files/figure.png" pageId="14" pageNumber="15">Figs. 2</figureCitation>
K, 2O, 4A–E
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFE8FF81FF70FB07FEB6C0EE" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BB53697FFE8FF81FF70FB07FE9AC3E4" blockId="14.[151,1437,1260,2005]" pageId="14" pageNumber="15">
<emphasis id="B97EEA85FFE8FF81FF70FB07FE8DC33C" bold="true" box="[151,344,1260,1285]" pageId="14" pageNumber="15">Adult diagnosis.</emphasis>
Morphologically distinguishable from other 
<emphasis id="B97EEA85FFE8FF81FCA9FB06FCBDC33D" box="[846,872,1261,1284]" italics="true" pageId="14" pageNumber="15">C.</emphasis>
(
<taxonomicName id="4C0A4D14FFE8FF81FC9FFB07FBD2C33C" box="[888,1031,1260,1285]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFE8FF81FC9FFB07FBD2C33C" box="[888,1031,1260,1285]" italics="true" pageId="14" pageNumber="15">Cycadophila</emphasis>
</taxonomicName>
) spp. by the long supraocular striae, head width/ventral interocular distance ratio &lt;3.50; width of elytra greatest at middle, without fine acute projection at posterior pronotal hind angle, and strong distal dilation of mesotibia and mesofemora in major males. Morphologically most similar to 
<taxonomicName id="4C0A4D14FFE8FF81FDF6FAB3FD0BC349" box="[529,734,1367,1392]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="debaonica" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE8FF81FDF6FAB3FDFEC356" box="[529,555,1368,1391]" italics="true" pageId="14" pageNumber="15">C.</emphasis>
(
<emphasis id="B97EEA85FFE8FF81FDDAFAB3FD82C356" box="[573,599,1368,1391]" italics="true" pageId="14" pageNumber="15">C.</emphasis>
) 
<emphasis id="B97EEA85FFE8FF81FD8EFABCFD0BC349" box="[617,734,1367,1392]" italics="true" pageId="14" pageNumber="15">debaonica</emphasis>
</taxonomicName>
which has a fine obtuse projection at posterior pronotal hind angle and major males with mesofemora narrowing from middle to apex. Most confidently distinguished from 
<taxonomicName id="4C0A4D14FFE8FF81FA65FA96FEEAC381" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="debaonica" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE8FF81FA65FA96FA49C3AD" box="[1410,1436,1405,1428]" italics="true" pageId="14" pageNumber="15">C.</emphasis>
(
<emphasis id="B97EEA85FFE8FF81FF47FA4BFF6FC38E" box="[160,186,1440,1463]" italics="true" pageId="14" pageNumber="15">C.</emphasis>
) 
<emphasis id="B97EEA85FFE8FF81FF2DFA74FEEAC381" box="[202,319,1439,1464]" italics="true" pageId="14" pageNumber="15">debaonica</emphasis>
</taxonomicName>
by analysis of the 16S rRNA gene (GenBank Accession numbers 
<accessionNumber id="9459AB74FFE8FF81FBC6FA4BFB70C381" box="[1057,1189,1439,1464]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY365227" pageId="14" pageNumber="15">KY 365227</accessionNumber>
and 
<accessionNumber id="9459AB74FFE8FF81FB3DFA74FA8DC381" box="[1242,1368,1439,1464]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY365247" pageId="14" pageNumber="15">KY365247</accessionNumber>
), host and distribution.
</paragraph>
<paragraph id="8BB53697FFE8FF81FF20FA0CFEB6C0EE" blockId="14.[151,1437,1260,2005]" pageId="14" pageNumber="15">
<emphasis id="B97EEA85FFE8FF81FF20FA0CFE32C039" bold="true" box="[199,487,1511,1536]" pageId="14" pageNumber="15">Adult male description.</emphasis>
Length 
<quantity id="4CF29B72FFE8FF81FDABFA0CFD21C3C6" box="[588,756,1511,1536]" metricMagnitude="-3" metricUnit="m" metricValue="5.02" metricValueMax="5.85" metricValueMin="4.19" pageId="14" pageNumber="15" unit="mm" value="5.02" valueMax="5.85" valueMin="4.19">4.19–5.85 mm</quantity>
, width 
<quantity id="4CF29B72FFE8FF81FCA9FA03FC2DC3C6" box="[846,1016,1512,1536]" metricMagnitude="-3" metricUnit="m" metricValue="2.1399999999999997" metricValueMax="2.4" metricValueMin="1.88" pageId="14" pageNumber="15" unit="mm" value="2.1399999999999997" valueMax="2.4" valueMin="1.88">1.88–2.40 mm</quantity>
. Body elongate-oval in dorsal view, greatest width at middle of elytra; in lateral view convex dorsally. General body color reddish-brown (
<figureCitation id="13312A12FFE8FF81FAEEF9E7FA80C01D" box="[1289,1365,1548,1573]" captionStart="FIGURE 4" captionStartId="16.[151,250,1671,1693]" captionTargetBox="[151,1436,373,1650]" captionTargetId="figure@16.[151,1436,373,1650]" captionTargetPageId="16" captionText="FIGURE 4. Photographs of Cycadophila (Cycadophila) collina major male from Vietnam, paratype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia." httpUri="https://zenodo.org/record/575645/files/figure.png" pageId="14" pageNumber="15">Figs. 4</figureCitation>
A–D), head darker than pronotum and lateral halves of elytra; elytra with large central black maculation occupying roughly half of elytral length from near the scutellum to 2/3 length of elytra and roughly 2/5 of the elytral width, dorsal surface punctate, shining and dorsally appearing glabrous, short procumbent hairs associated with punctation behind eyes and along lateral sides and declivity of elytra, ventrally mostly covered with short procumbent setae.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFE8FF9FFF20F90FFB5DC706" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph id="8BB53697FFE8FF80FF20F90FFADAC491" blockId="14.[151,1437,1260,2005]" lastBlockId="15.[151,1437,151,2013]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B97EEA85FFE8FF81FF20F90FFED0C0C4" bold="true" box="[199,261,1764,1789]" italics="true" pageId="14" pageNumber="15">Head</emphasis>
in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, sparsely punctured; width 
<quantity id="4CF29B72FFE8FF81FEBAF8E3FE10C126" box="[349,453,1800,1824]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="14" pageNumber="15" unit="mm" value="1.12">1.12 mm</quantity>
; dorsal interocular distance 
<quantity id="4CF29B72FFE8FF81FCE5F8ECFCBDC126" box="[770,872,1799,1824]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="14" pageNumber="15" unit="mm" value="0.68">0.68 mm</quantity>
, head width/dorsal interocular distance ratio 1.65, ventral interocular distance 
<quantity id="4CF29B72FFE8FF81FE30F8C7FD95C17D" box="[471,576,1836,1860]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="14" pageNumber="15" unit="mm" value="0.39">0.39 mm</quantity>
, head width/ventral interocular distance ratio 3.05. Eye globular; bordered dorsally with long supraocular stria reaching anterior angle of eye; projected laterally, with large black facets and short, inconspicuous interfacetal setae. Frons with short, dark, subcuticular suture on each side, approximately 1/4 width of frons in length, extending obliquely to lateral margin anterior to antennal insertion. Antennal length slightly shorter than pronotal width, slightly longer than head width, antennomere relative lengths from base to end approximately 14:10:12:10:10:10:9:9:10:10:10, antennomere widths of II–VII subequal, I and VIII subequal and about 1.30× width of II–VII, width equals length; IX and X subequal and about 2.00× width of II–VII, XI about 1.50× width of II–VII, globular. Clypeus truncate anteriorly, anterior margin with long setae; densely punctate; somewhat emarginate, with narrow margin. Mandible with 3 apical teeth; prostheca extending from mola halfway to apical teeth, length about ½ width, edge nearly straight, with fringe of hair along dorsal base that extends to the proximal apical tooth; mola with carinulation and small spines. Labrum transverse, about 2.00× broader than long; anterior margin fringed with long bristles, especially dense medially. Hypopharynx with thin setae directed medially. Maxilla with cardo expanded apically; stipes narrowly triangular, apically acute, partially overlapping with palpiger; galea and lacinia with long, densely aligned setae; setae on the lacinia thick and strong, but distinguishable hooks absent from surrounding dense setae; galea wider than lacinia. Maxillary palpomeres II, III and IV longer than width; IV elongate oval, apex densely papillate; relative lengths of II–IV approximately 5:3:7. Labium with mentum pentagonal with carina forming triangular plate projecting medially, lacking distinct lateral pockets; labial palpomere III elongate-oval. 
<collectingRegion id="49CEF875FFE9FF80FD6EFDA3FD3DC466" box="[649,744,584,607]" country="Myanmar" name="Chin" pageId="15" pageNumber="16">Mentum</collectingRegion>
and submentum with moderate punctation and setation, setae short. Gular area smooth, without setae or punctation, except near suture with submentum, where shallow transversely elongate foveae are present bearing dense setose punctures (
<figureCitation id="13312A12FFE9FF80FC20FD64FBDFC491" box="[967,1034,655,680]" captionStart="FIGURE 4" captionStartId="16.[151,250,1671,1693]" captionTargetBox="[151,1436,373,1650]" captionTargetId="figure@16.[151,1436,373,1650]" captionTargetPageId="16" captionText="FIGURE 4. Photographs of Cycadophila (Cycadophila) collina major male from Vietnam, paratype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia." httpUri="https://zenodo.org/record/575645/files/figure.png" pageId="15" pageNumber="16">Fig. 4</figureCitation>
D), foveae dark brown.
</paragraph>
<paragraph id="8BB53697FFE9FF80FF20FD5FFB04C299" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="16">
<emphasis id="B97EEA85FFE9FF80FF20FD5FFEC8C4F4" bold="true" box="[199,285,692,717]" italics="true" pageId="15" pageNumber="16">Thorax</emphasis>
with pronotum transverse in dorsal view with marginal beads on all sides; length/width ratio 0.69, convex, wider at base; anterior angles weakly developed; posterior angles exteriorly angulate; posterior margin projecting medially; with a narrow longitudinal furrow posterolaterally, beginning at pore on posterior margin extending forward and ending abruptly, length of furrows relative to pronotal length 0.22. Prosternum with anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae; prosternal process round and convex apically. Hypomeron laterally with few punctures, medially without longitudinal striations. Scutellar shield transverse, posterior margin clearly projected, pentagonal. Elytra in dorsal view elongate-oval, convex; length/width 3.30, greatest width near midlength; with marginal line basally, scutellary striole extending ¼ elytral length, with 10–13 punctures; intervals of striae with fine, shallow punctures. All punctures of elytral striae bearing a single short seta; seta only visible in profile, extending slightly out of puncture. Punctation on meso- and metaventrites shallow, small. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over half metaventrite length. Legs stout, relatively similar in length and shape. Femora compressed laterally; tibiae shorter than femora, gradually dilated to narrow apex; protibia distinctly dilated apically, with apical fringe of short, stout spinules ventrally, inner margin with fine setae.
</paragraph>
<paragraph id="8BB53697FFE9FF80FF20FB47FC7CC341" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="16">
<emphasis id="B97EEA85FFE9FF80FF20FB47FEE0C2FC" bold="true" box="[199,309,1196,1221]" italics="true" pageId="15" pageNumber="16">Abdomen</emphasis>
with 5 ventrites bearing fine punctures; anterior margin with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I lacking subcoxal line, slightly longer medially than II; I–IV each with pair of long erect sensory hairs located adjacent to midline (often abraded); II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex bearing row of short, appressed, densely arranged setae.
</paragraph>
<paragraph id="8BB53697FFE9FF80FF20FA6FFDDCC069" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="16">
<emphasis id="B97EEA85FFE9FF80FF20FA6FFEA5C3A4" bold="true" box="[199,368,1412,1437]" italics="true" pageId="15" pageNumber="16">Male genitalia</emphasis>
(tegmen and median lobe of aedeagus) twisted approximately 90˚ toward the left side of the animal and resting on side. Tegmen sclerotized, triangular, but twisted medially; with anterior region ring-like, posterior region sheath-like, posterior margin slightly arcuate; lateral margins gradually converging posteriorly; dorsally with 2 large, elongate, somewhat coniform parameres. Parameres apically with long setae; length/width ratio 2.60. Aedeagus with median lobe slender, highly sclerotized, apex long and acuminate. Ratio of penile strut length to median lobe length 3:1.
</paragraph>
<paragraph id="8BB53697FFE9FF80FF20F9B7FC2DC13C" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="16">
<emphasis id="B97EEA85FFE9FF80FF20F9B7FECAC04C" bold="true" box="[199,287,1628,1653]" italics="true" pageId="15" pageNumber="16">Female</emphasis>
similar to male except with pro- and mesotibiae and femora less robust and not dilated apically, pygidium truncate, apical edge fairly straight with 7 long setae and a fringe of setae about 1/2 as long. Genitalia with gonocoxites triangular, gradually narrowing posteriorly; apices of gonocoxites laterally with concave impression and setae. Gonostylus inserted at the concave impression of the gonocoxite, with several short setae and 1 long seta. Valvifers expanded posteriorly. Spermatheca elongate, C-shaped.
</paragraph>
<paragraph id="8BB53697FFE9FF9FFF20F8E4FB5DC706" blockId="15.[151,1437,151,2013]" lastBlockId="16.[151,1436,151,320]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<materialsCitation id="3B623CCAFFE9FF9FFF20F8E4FBEEC724" ID-GBIF-Occurrence="1502685035" collectorName="E. Range. Known" country="Laos" county="Houphan Province" lastPageId="16" lastPageNumber="17" location="Vieng Xai village" municipality="Viengxay District" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Son La Province" typeStatus="holotype">
<emphasis id="B97EEA85FFE9FF80FF20F8E4FEECC111" bold="true" box="[199,313,1807,1832]" pageId="15" pageNumber="16">
<emphasis id="B97EEA85FFE9FF80FF20F8E4FEE7C111" bold="true" box="[199,306,1807,1832]" italics="true" pageId="15" pageNumber="16">Variation</emphasis>
.
</emphasis>
In some individuals the dark elytral mark is slightly larger or smaller, and varies from a lighter to darker brown. In some this mark is pale enough to be indistinct. Variation in pro- and mesotibial dilation appears to be sexually dimorphic. Males usually have these tibiae more strongly dilated. However, there is enough variation that distinguishing weakly developed males from strongly developed females can be problematic. Weakly developed females have the mesotibia almost parallel-sided to the apex. Strongly developed males have the outer margin of the mesotibia slightly swollen to a widened apex. Similarly, mesofemoral development varies in males from strongly to weakly dilated toward apex. Major males of 
<taxonomicName id="4C0A4D14FFF6FF9FFCA0FF73FC60C689" box="[839,949,151,176]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="collina">
<emphasis id="B97EEA85FFF6FF9FFCA0FF73FC60C689" box="[839,949,151,176]" italics="true" pageId="16" pageNumber="17">C. collina</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A4D14FFF6FF9FFC0CFF73FB54C689" box="[1003,1153,151,176]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="debaonica">
<emphasis id="B97EEA85FFF6FF9FFC0CFF73FB54C689" box="[1003,1153,151,176]" italics="true" pageId="16" pageNumber="17">C. debaonica</emphasis>
</taxonomicName>
are distinctly different in this character. Minor males of either species are difficult to distinguish. Figures used in the key are major males. 
<emphasis id="B97EEA85FFF6FF9FFF20FF34FEB0C6C1" bold="true" box="[199,357,223,248]" pageId="16" pageNumber="17">
<typeStatus id="54B18835FFF6FF9FFF20FF34FED7C6C1" box="[199,258,223,248]" pageId="16" pageNumber="17">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31D7607FFF6FF9FFE8BFF0BFE70C6CE" box="[364,421,224,247]" name="Laos" pageId="16" pageNumber="17">Laos</collectingCountry>
, 
<collectingCounty id="62D44E1BFFF6FF9FFE57FF34FD50C6C1" box="[432,645,223,248]" pageId="16" pageNumber="17">Houphan Province</collectingCounty>
, 
<collectingMunicipality id="6BD1ACEDFFF6FF9FFD68FF34FC8CC6C1" box="[655,857,223,248]" pageId="16" pageNumber="17">Viengxay District</collectingMunicipality>
, 
<location id="8ED5604CFFF6FF9FFC83FF34FBF9C6C1" LSID="urn:lsid:plazi:treatment:03A38781FFE8FF9EFF70FB6AFE55C564:8ED5604CFFF6FF9FFC83FF34FBF9C6C1" box="[868,1068,223,248]" country="Laos" county="Houphan Province" municipality="Viengxay District" name="Vieng Xai village" pageId="16" pageNumber="17" stateProvince="Son La Province">Vieng Xai village</location>
, 20˚23’41”N 104˚13’44” 
<collectorName id="26FF5341FFF6FF9FFAA0FF34FEA1C725" pageId="16" pageNumber="17">
E. 
<emphasis id="B97EEA85FFF6FF9FFF20FEEFFECFC724" bold="true" box="[199,282,260,285]" pageId="16" pageNumber="17">Range.</emphasis>
Known
</collectorName>
from 
<location id="8ED5604CFFF6FF9FFE50FEEFFD5CC724" LSID="urn:lsid:plazi:treatment:03A38781FFE8FF9EFF70FB6AFE55C564:8ED5604CFFF6FF9FFE50FEEFFD5CC724" box="[439,649,260,285]" country="Laos" county="Houphan Province" municipality="Viengxay District" name="Houphan Province" pageId="16" pageNumber="17" stateProvince="Son La Province">Houphan Province</location>
in 
<location id="8ED5604CFFF6FF9FFD4CFEEFFC92C725" LSID="urn:lsid:plazi:treatment:03A38781FFE8FF9EFF70FB6AFE55C564:8ED5604CFFF6FF9FFD4CFEEFFC92C725" box="[683,839,260,285]" country="Laos" county="Houphan Province" municipality="Viengxay District" name="northern Laos" pageId="16" pageNumber="17" stateProvince="Son La Province">northern Laos</location>
and 
<collectingRegion id="49CEF875FFF6FF9FFC9AFEEFFBEEC724" box="[893,1083,260,285]" country="Vietnam" name="Son La" pageId="16" pageNumber="17">
Son 
<collectingRegion id="49CEF875FFF6FF9FFC48FEEEFBEEC724" box="[943,1083,260,285]" inferredBySuffix="true" pageId="16" pageNumber="17">La Province</collectingRegion>
</collectingRegion>
</materialsCitation>
, 
<materialsCitation id="3B623CCAFFF6FF9FFBA2FEEFFDB4C779" ID-GBIF-Occurrence="1502685039" country="Vietnam" location="Yunnan Province" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Yunnan Province" typeStatus="holotype">
<collectingCountry id="F31D7607FFF6FF9FFBA2FEEFFB71C724" box="[1093,1188,260,285]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
<collectedFrom id="3816756AFFF6FF9FFB4DFEEEFAB3C724" box="[1194,1382,260,285]" pageId="16" pageNumber="17">
on 
<taxonomicName id="4C0A4D14FFF6FF9FFB2AFEEEFAB3C724" box="[1229,1382,260,285]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="collina">
<emphasis id="B97EEA85FFF6FF9FFB2AFEEEFAB3C724" box="[1229,1382,260,285]" italics="true" pageId="16" pageNumber="17">Cycas collina</emphasis>
</taxonomicName>
</collectedFrom>
, and the southern portion of 
<collectingRegion id="49CEF875FFF6FF9FFE79FEC3FDB4C779" box="[414,609,295,320]" country="China" name="Yunnan" pageId="16" pageNumber="17">Yunnan Province</collectingRegion>
</materialsCitation>
, 
<materialsCitation id="3B623CCAFFF6FF9FFD8BFECCFB5DC706" ID-GBIF-Occurrence="1502685050" box="[620,1160,295,320]" country="China" location="China" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F31D7607FFF6FF9FFD8BFECCFD7AC779" box="[620,687,295,320]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
<collectedFrom id="3816756AFFF6FF9FFD51FEC3FCC8C706" box="[694,797,296,319]" pageId="16" pageNumber="17">
on 
<taxonomicName id="4C0A4D14FFF6FF9FFD3EFEC3FCC8C706" box="[729,797,296,319]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFF6FF9FFD3EFEC3FCC8C706" box="[729,797,296,319]" italics="true" pageId="16" pageNumber="17">Cycas</emphasis>
</taxonomicName>
</collectedFrom>
sp. 
<taxonomicName id="4C0A4D14FFF6FF9FFCACFECCFC10C779" box="[843,965,295,320]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="collina">
aff. 
<emphasis id="B97EEA85FFF6FF9FFC90FECCFC10C779" box="[887,965,295,320]" italics="true" pageId="16" pageNumber="17">collina</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A4D14FFF6FF9FFC1BFEC3FB54C779" box="[1020,1153,296,320]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="tanqingii">
<emphasis id="B97EEA85FFF6FF9FFC1BFEC3FB54C779" box="[1020,1153,296,320]" italics="true" pageId="16" pageNumber="17">C. tanqingii</emphasis>
</taxonomicName>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF75661FFFF6FF9FFF70F96CFC2AC0F9" httpUri="https://zenodo.org/record/575645/files/figure.png" pageId="16" pageNumber="17" targetBox="[151,1436,373,1650]" targetPageId="16">
<paragraph id="8BB53697FFF6FF9FFF70F96CFC2AC0F9" blockId="16.[151,1436,1671,1728]" pageId="16" pageNumber="17">
<emphasis id="B97EEA85FFF6FF9FFF70F96CFEC0C0A4" bold="true" box="[151,277,1671,1693]" pageId="16" pageNumber="17">FIGURE 4.</emphasis>
Photographs of 
<taxonomicName id="4C0A4D14FFF6FF9FFE23F96CFCE5C0A4" box="[452,816,1671,1693]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="collina" subGenus="Cycadophila">
<emphasis id="B97EEA85FFF6FF9FFE23F96CFD93C0A4" box="[452,582,1671,1693]" italics="true" pageId="16" pageNumber="17">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFF6FF9FFDB0F96CFD0DC0A4" box="[599,728,1671,1693]" italics="true" pageId="16" pageNumber="17">Cycadophila</emphasis>
) 
<emphasis id="B97EEA85FFF6FF9FFD0DF96CFCE5C0A4" box="[746,816,1671,1693]" italics="true" pageId="16" pageNumber="17">collina</emphasis>
</taxonomicName>
major male from Vietnam, paratype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia.
</paragraph>
</caption>
<subSubSection id="C310651CFFF6FF9EFF20F91BFDEFC724" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BB53697FFF6FF9EFF20F91BFDEFC724" blockId="16.[151,1436,1776,2017]" lastBlockId="17.[151,1437,151,861]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<materialsCitation id="3B623CCAFFF6FF9FFF20F91BFE7BC1A0" ID-GBIF-Occurrence="1502685049" collectingDate="2015-04-07" collectionCode="FSCA" collectorName="N. T. Hiep &amp; L. Averyanov &amp; N. S. Khang &amp; N. Q. Hieu &amp; T. Maisak &amp; Somneux &amp; Deposited" country="Laos" latitude="20.394722" location="Vieng Xai" longLatPrecision="21" longitude="104.22889" municipality="Material" pageId="16" pageNumber="17" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B97EEA85FFF6FF9FFF20F91BFDF6C130" bold="true" box="[199,547,1776,1801]" pageId="16" pageNumber="17">
<collectingMunicipality id="6BD1ACEDFFF6FF9FFF20F91BFEFAC130" box="[199,303,1776,1801]" pageId="16" pageNumber="17">Material</collectingMunicipality>
examined. 
<typeStatus id="54B18835FFF6FF9FFE50F91BFDF6C130" box="[439,547,1776,1801]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
</emphasis>
(by designation) male with the following labels: 1) [rectangular; white; printed in black ink] “ 
<collectingCountry id="F31D7607FFF6FF9FFED1F8FEFE57C114" box="[310,386,1813,1837]" name="Laos" pageId="16" pageNumber="17">LAOS</collectingCountry>
, Houphan Pr., Viengxay Ds., 
<location id="8ED5604CFFF6FF9FFD2CF8FFFCE8C114" LSID="urn:lsid:plazi:treatment:03A38781FFE8FF9EFF70FB6AFE55C564:8ED5604CFFF6FF9FFD2CF8FFFCE8C114" box="[715,829,1812,1837]" country="Laos" latitude="20.394722" longLatPrecision="21" longitude="104.22889" municipality="Material" name="Vieng Xai" pageId="16" pageNumber="17">Vieng Xai</location>
vill., 
<geoCoordinate id="EE3E5050FFF6FF9FFC9AF8FFFBD4C114" box="[893,1025,1812,1837]" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="15" value="20.394722">20°23'41″N</geoCoordinate>
, 
<geoCoordinate id="EE3E5050FFF6FF9FFBEBF8FFFB49C114" box="[1036,1180,1812,1837]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="15" value="104.22889">104°13'44″E</geoCoordinate>
, 
<date id="FFB41057FFF6FF9FFB41F8FFFAC9C114" box="[1190,1308,1812,1837]" pageId="16" pageNumber="17" value="2015-04-07">
<collectingDate id="EFF0E9BFFFF6FF9FFB41F8FFFAC9C114" box="[1190,1308,1812,1837]" pageId="16" pageNumber="17" value="2015-04-07">7-IV-2015</collectingDate>
</date>
, 
<collectorName id="26FF5341FFF6FF9FFAC0F8FEFA4DC114" box="[1319,1432,1812,1837]" pageId="16" pageNumber="17">N.T. Hiep</collectorName>
, 
<collectorName id="26FF5341FFF6FF9FFF70F8D2FEE1C168" box="[151,308,1849,1873]" pageId="16" pageNumber="17">L. Averyanov</collectorName>
, 
<collectorName id="26FF5341FFF6FF9FFEA7F8D2FE1DC168" box="[320,456,1848,1873]" pageId="16" pageNumber="17">N.S. Khang</collectorName>
, 
<collectorName id="26FF5341FFF6FF9FFE31F8D2FD9BC168" box="[470,590,1848,1873]" pageId="16" pageNumber="17">N.Q. Hieu</collectorName>
, 
<collectorName id="26FF5341FFF6FF9FFDBCF8D2FD05C168" box="[603,720,1848,1873]" pageId="16" pageNumber="17">T. Maisak</collectorName>
, 
<collectorName id="26FF5341FFF6FF9FFD3AF8D2FC99C168" box="[733,844,1849,1873]" pageId="16" pageNumber="17">Somneux</collectorName>
”; 2) “ex 
<specimenCount id="9D0CFD1EFFF6FF9FFC54F8D2FC1DC169" box="[947,968,1849,1872]" pageId="16" pageNumber="17" type="male">♂</specimenCount>
cones 
<taxonomicName id="4C0A4D14FFF6FF9FFBFDF8D2FB60C168" box="[1050,1205,1848,1873]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="collina">
<emphasis id="B97EEA85FFF6FF9FFBFDF8D2FB60C168" box="[1050,1205,1848,1873]" italics="true" pageId="16" pageNumber="17">Cycas collina</emphasis>
</taxonomicName>
(LA-VN 1124)”; 3) [rectangular; red, printed in black ink] “ 
<typeStatus id="54B18835FFF6FF9FFDBAF8B6FD24C14C" box="[605,753,1885,1909]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D0CFD1EFFF6FF9FFD1EF8B6FCDBC14D" box="[761,782,1885,1908]" pageId="16" pageNumber="17" type="male">♂</specimenCount>
<taxonomicName id="4C0A4D14FFF6FF9FFCF1F8B7FA58C14C" authority="P. Skelley, G. Xu &amp; W. Tang 2017" authorityName="P. Skelley, G. Xu &amp; W. Tang" authorityYear="2017" box="[790,1421,1884,1909]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="collina">
<emphasis id="B97EEA85FFF6FF9FFCF1F8B7FC29C14C" box="[790,1020,1884,1909]" italics="true" pageId="16" pageNumber="17">Cycadophila collina</emphasis>
P. Skelley, G. Xu &amp; W. Tang 2017
</taxonomicName>
”. 
<collectorName id="26FF5341FFF6FF9FFF70F86BFEDCC1A0" box="[151,265,1920,1945]" pageId="16" pageNumber="17">Deposited</collectorName>
in the 
<collectionCode id="ED1BAE52FFF6FF9FFE83F86AFE7BC1A0" box="[356,430,1921,1945]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="17">FSCA</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B623CCAFFF6FF9EFE5BF86BFDC2C689" ID-GBIF-Occurrence="1502685045" collectingDate="1996-05-10" collectionCode="FSCA" collectorName="W. Tang" country="China" lastPageId="17" lastPageNumber="18" latitude="22.166666" location="Guanping Reserve" longLatPrecision="1261" longitude="100.833336" pageId="16" pageNumber="17" specimenCount="48" specimenCount-female="1" specimenCount-male="2" stateProvince="Yunnan Province" typeStatus="allotype">
<emphasis id="B97EEA85FFF6FF9FFE5BF86BFDF4C1A0" bold="true" box="[444,545,1920,1945]" pageId="16" pageNumber="17">
<typeStatus id="54B18835FFF6FF9FFE5BF86BFDF4C1A0" box="[444,545,1920,1945]" pageId="16" pageNumber="17" type="allotype">Allotype</typeStatus>
</emphasis>
female and 
<specimenCount id="9D0CFD1EFFF6FF9FFD5EF86BFC82C1A0" box="[697,855,1920,1945]" pageId="16" pageNumber="17" type="generic" typeStatus="paratypes">
45 
<emphasis id="B97EEA85FFF6FF9FFD06F86BFC82C1A0" bold="true" box="[737,855,1920,1945]" pageId="16" pageNumber="17">paratypes</emphasis>
</specimenCount>
: 
<emphasis id="B97EEA85FFF6FF9FFC8CF86BFB78C1A0" bold="true" box="[875,1197,1920,1945]" pageId="16" pageNumber="17">
<collectingCountry id="F31D7607FFF6FF9FFC8CF86BFC1FC1A0" box="[875,970,1920,1945]" name="China" pageId="16" pageNumber="17">CHINA</collectingCountry>
, 
<collectingRegion id="49CEF875FFF6FF9FFC3CF86BFB78C1A0" box="[987,1197,1920,1945]" country="China" name="Yunnan" pageId="16" pageNumber="17">Yunnan Province</collectingRegion>
</emphasis>
, 
<location id="8ED5604CFFF6FF9FFB26F86BFA4DC1A1" LSID="urn:lsid:plazi:treatment:03A38781FFE8FF9EFF70FB6AFE55C564:8ED5604CFFF6FF9FFB26F86BFA4DC1A1" box="[1217,1432,1920,1945]" country="China" latitude="22.166666" longLatPrecision="1261" longitude="100.833336" name="Guanping Reserve" pageId="16" pageNumber="17" stateProvince="Yunnan Province">Guanping Reserve</location>
, 
<geoCoordinate id="EE3E5050FFF6FF9FFF70F84EFF24C184" box="[151,241,1957,1981]" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="22.166666">22°10'N</geoCoordinate>
<geoCoordinate id="EE3E5050FFF6FF9FFF1CF84FFEB7C184" box="[251,354,1956,1981]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="100.833336">100°50'E</geoCoordinate>
, 
<date id="FFB41057FFF6FF9FFE96F84EFDC5C184" box="[369,528,1956,1981]" pageId="16" pageNumber="17" value="1996-05-10">
<collectingDate id="EFF0E9BFFFF6FF9FFE96F84EFDC5C184" box="[369,528,1956,1981]" pageId="16" pageNumber="17" value="1996-05-10">10 May 1996</collectingDate>
</date>
, 
<collectorName id="26FF5341FFF6FF9FFDF8F84EFD57C184" box="[543,642,1957,1981]" pageId="16" pageNumber="17">W. Tang</collectorName>
, ex spent 
<specimenCount id="9D0CFD1EFFF6FF9FFD1BF84EFCC4C185" box="[764,785,1957,1980]" pageId="16" pageNumber="17" type="male">♂</specimenCount>
strobilus of 
<taxonomicName id="4C0A4D14FFF6FF9FFC40F84EFC3EC185" box="[935,1003,1957,1980]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFF6FF9FFC40F84EFC3EC185" box="[935,1003,1957,1980]" italics="true" pageId="16" pageNumber="17">Cycas</emphasis>
</taxonomicName>
sp. 
<taxonomicName id="4C0A4D14FFF6FF9FFBC6F84FFB4BC184" box="[1057,1182,1956,1981]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="collina">
aff. 
<emphasis id="B97EEA85FFF6FF9FFBB7F84FFB4BC184" box="[1104,1182,1956,1981]" italics="true" pageId="16" pageNumber="17">collina</emphasis>
</taxonomicName>
, WT 44 (2); ex early pollen-shedding 
<specimenCount id="9D0CFD1EFFF6FF9FFEB2F822FEBFC1D9" box="[341,362,1993,2016]" pageId="16" pageNumber="17" type="male">♂</specimenCount>
strobilus of 
<taxonomicName id="4C0A4D14FFF6FF9FFDE7F822FD90C1D9" box="[512,581,1993,2016]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFF6FF9FFDE7F822FD90C1D9" box="[512,581,1993,2016]" italics="true" pageId="16" pageNumber="17">Cycas</emphasis>
</taxonomicName>
sp. 
<taxonomicName id="4C0A4D14FFF6FF9FFD9FF823FD23C1D8" box="[632,758,1992,2017]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="collina">
aff. 
<emphasis id="B97EEA85FFF6FF9FFD4FF823FD23C1D8" box="[680,758,1992,2017]" italics="true" pageId="16" pageNumber="17">collina</emphasis>
</taxonomicName>
, WT 45 (
<typeStatus id="54B18835FFF6FF9FFC8DF823FC1CC1D8" box="[874,969,1992,2017]" pageId="16" pageNumber="17" type="allotype">allotype</typeStatus>
, 
<collectionCode id="ED1BAE52FFF6FF9FFC30F822FBF4C1D8" box="[983,1057,1993,2017]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="17">FSCA</collectionCode>
; 20); ex receptive 
<specimenCount id="9D0CFD1EFFF6FF9FFB1CF822FAC5C1D9" box="[1275,1296,1993,2016]" pageId="16" pageNumber="17" type="female">♀</specimenCount>
strobilus of 
<taxonomicName id="4C0A4D14FFF7FF9EFF70FF73FF09C696" box="[151,220,152,175]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFF7FF9EFF70FF73FF09C696" box="[151,220,152,175]" italics="true" pageId="17" pageNumber="18">Cycas</emphasis>
</taxonomicName>
sp. 
<taxonomicName id="4C0A4D14FFF7FF9EFEE9FF7CFE5FC689" box="[270,394,151,176]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="collina">
aff. 
<emphasis id="B97EEA85FFF7FF9EFEDBFF7CFE5FC689" box="[316,394,151,176]" italics="true" pageId="17" pageNumber="18">collina</emphasis>
</taxonomicName>
, WT 46 (2)
</materialsCitation>
; 
<materialsCitation id="3B623CCAFFF7FF9EFDC1FF7CFB8CC6ED" ID-GBIF-Occurrence="1502685042" collectingDate="1996-04-04" collectionCode="ANIC, FSCA" collectorName="S. L. Yang" country="Vietnam" location="Mai Son" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Son La Province" typeStatus="paratype">
<emphasis id="B97EEA85FFF7FF9EFDC1FF7CFC52C689" bold="true" box="[550,903,151,176]" pageId="17" pageNumber="18">
<collectingCountry id="F31D7607FFF7FF9EFDC1FF7CFD7AC689" box="[550,687,151,176]" name="Vietnam" pageId="17" pageNumber="18">VIETNAM</collectingCountry>
, 
<collectingRegion id="49CEF875FFF7FF9EFD59FF7CFC52C689" box="[702,903,151,176]" country="Vietnam" name="Son La" pageId="17" pageNumber="18">
Son 
<collectingRegion id="49CEF875FFF7FF9EFD13FF7CFC52C689" box="[756,903,151,176]" inferredBySuffix="true" pageId="17" pageNumber="18">La Province</collectingRegion>
</collectingRegion>
</emphasis>
, nr. 
<location id="8ED5604CFFF7FF9EFC59FF7CFBF7C689" LSID="urn:lsid:plazi:treatment:03A38781FFE8FF9EFF70FB6AFE55C564:8ED5604CFFF7FF9EFC59FF7CFBF7C689" box="[958,1058,151,176]" country="Vietnam" name="Mai Son" pageId="17" pageNumber="18" stateProvince="Son La Province">Mai Son</location>
, 21°8'630″N 104°1'731″E, 
<taxonomicName id="4C0A4D14FFF7FF9EFAB0FF73FEF5C6ED" authority="Apr" authorityName="Apr" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="collina">
<emphasis id="B97EEA85FFF7FF9EFAB0FF73FF30C6EC" italics="true" pageId="17" pageNumber="18">Cycas collina</emphasis>
, 
<collectingDate id="EFF0E9BFFFF7FF9EFF13FF57FEF5C6ED" box="[244,288,188,212]" pageId="17" pageNumber="18" value="1996-04-04">Apr</collectingDate>
</taxonomicName>
4, 1996, 
<collectorName id="26FF5341FFF7FF9EFE69FF57FE2AC6EC" box="[398,511,188,213]" pageId="17" pageNumber="18">S.L. Yang</collectorName>
SLY777 (21). 
<typeStatus id="54B18835FFF7FF9EFD53FF56FCF4C6EC" box="[692,801,189,213]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
deposited in 
<collectionCode id="ED1BAE52FFF7FF9EFC5BFF57FBD6C6ED" box="[956,1027,188,212]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="17" pageNumber="18">ANIC</collectionCode>
, 
<collectionCode id="ED1BAE52FFF7FF9EFBF7FF57FB8CC6ED" box="[1040,1113,188,212]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="17" pageNumber="18">FSCA</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B623CCAFFF7FF9EFB82FF57FDEFC724" ID-GBIF-Occurrence="1502685046" collectingDate="2008-03-27" collectionCode="DNA" collectorName="He Jianghai" country="China" location="Luchun Co" municipality="Others Examined" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="paratype">
<emphasis id="B97EEA85FFF7FF9EFB82FF57FEB0C6C1" bold="true" pageId="17" pageNumber="18">
<collectingMunicipality id="6BD1ACEDFFF7FF9EFB82FF57FAEBC6EC" box="[1125,1342,188,213]" pageId="17" pageNumber="18">Others Examined</collectingMunicipality>
: 
<collectingCountry id="F31D7607FFF7FF9EFAABFF57FA42C6EC" box="[1356,1431,188,213]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
, 
<collectingRegion id="49CEF875FFF7FF9EFF70FF34FEB0C6C1" box="[151,357,223,248]" country="China" name="Yunnan" pageId="17" pageNumber="18">Yunnan Province</collectingRegion>
</emphasis>
, 
<location id="8ED5604CFFF7FF9EFE93FF34FE20C6C1" LSID="urn:lsid:plazi:treatment:03A38781FFE8FF9EFF70FB6AFE55C564:8ED5604CFFF7FF9EFE93FF34FE20C6C1" box="[372,501,223,248]" country="China" municipality="Others Examined" name="Luchun Co" pageId="17" pageNumber="18" stateProvince="Yunnan Province">Luchun Co</location>
, 
<taxonomicName id="4C0A4D14FFF7FF9EFDE6FF0BFD67C6C1" box="[513,690,224,248]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="tanqingii">
<emphasis id="B97EEA85FFF7FF9EFDE6FF0BFD67C6C1" box="[513,690,224,248]" italics="true" pageId="17" pageNumber="18">Cycas tanqingii</emphasis>
</taxonomicName>
<specimenCount id="9D0CFD1EFFF7FF9EFD5DFF0BFD1AC6CE" box="[698,719,224,247]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
cone, 
<date id="FFB41057FFF7FF9EFCFCFF0BFC1FC6C1" box="[795,970,223,248]" pageId="17" pageNumber="18" value="2008-03-27">
<collectingDate id="EFF0E9BFFFF7FF9EFCFCFF0BFC1FC6C1" box="[795,970,223,248]" pageId="17" pageNumber="18" value="2008-03-27">27 March 2008</collectingDate>
</date>
, 
<collectorName id="26FF5341FFF7FF9EFC31FF0BFB8EC6C1" box="[982,1115,223,248]" pageId="17" pageNumber="18">He Jianghai</collectorName>
[specimen destroyed during 
<collectionCode id="ED1BAE52FFF7FF9EFF70FEEFFF01C725" box="[151,212,260,284]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="17" pageNumber="18">DNA</collectionCode>
analysis; pin and label remain].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFF7FF9EFF20FECCFBFDC706" box="[199,1064,295,320]" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BB53697FFF7FF9EFF20FECCFBFDC706" blockId="17.[151,1437,151,861]" box="[199,1064,295,320]" pageId="17" pageNumber="18">
<emphasis id="B97EEA85FFF7FF9EFF20FECCFE9BC779" bold="true" box="[199,334,295,320]" pageId="17" pageNumber="18">Etymology.</emphasis>
The species is named after one of its host species, 
<taxonomicName id="4C0A4D14FFF7FF9EFC6FFEC3FBF7C779" box="[904,1058,295,320]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="collina">
<emphasis id="B97EEA85FFF7FF9EFC6FFEC3FBF7C779" box="[904,1058,295,320]" italics="true" pageId="17" pageNumber="18">Cycas collina</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFF7FF9EFF20FEA7FE55C564" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BB53697FFF7FF9EFF20FEA7FC4EC405" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="18">
<emphasis id="B97EEA85FFF7FF9EFF20FEA7FEE1C75C" bold="true" box="[199,308,332,357]" pageId="17" pageNumber="18">Remarks</emphasis>
. DNA analysis of the 16S rRNA gene indicates that 
<taxonomicName id="4C0A4D14FFF7FF9EFC43FEA7FB5EC75C" box="[932,1163,332,357]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="collina">
<emphasis id="B97EEA85FFF7FF9EFC43FEA7FB5EC75C" box="[932,1163,332,357]" italics="true" pageId="17" pageNumber="18">Cycadophila collina</emphasis>
</taxonomicName>
forms part of a cryptic species complex with 
<taxonomicName id="4C0A4D14FFF7FF9EFE76FE9BFDCCC7B1" box="[401,537,367,392]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="debonica">
<emphasis id="B97EEA85FFF7FF9EFE76FE9BFDCCC7B1" box="[401,537,367,392]" italics="true" pageId="17" pageNumber="18">C. debonica</emphasis>
</taxonomicName>
and that this beetle is associated with two hosts, 
<taxonomicName id="4C0A4D14FFF7FF9EFBA2FE9BFB23C7B1" box="[1093,1270,368,392]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="tanqingii">
<emphasis id="B97EEA85FFF7FF9EFBA2FE9BFB23C7B1" box="[1093,1270,368,392]" italics="true" pageId="17" pageNumber="18">Cycas tanqingii</emphasis>
</taxonomicName>
and the 
<taxonomicName id="4C0A4D14FFF7FF9EFAB0FE9BFF30C794" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="collina">
<emphasis id="B97EEA85FFF7FF9EFAB0FE9BFF30C794" italics="true" pageId="17" pageNumber="18">Cycas collina</emphasis>
</taxonomicName>
species complex. These two host 
<taxonomicName id="4C0A4D14FFF7FF9EFD85FE7EFD73C795" box="[610,678,405,428]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFF7FF9EFD85FE7EFD73C795" box="[610,678,405,428]" italics="true" pageId="17" pageNumber="18">Cycas</emphasis>
</taxonomicName>
species inhabit rugged terrain south of the Red River while the two known hosts of its sister taxon, 
<taxonomicName id="4C0A4D14FFF7FF9EFE10FE5CFCD7C7E9" box="[503,770,439,464]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="debaonica">
<emphasis id="B97EEA85FFF7FF9EFE10FE5CFCD7C7E9" box="[503,770,439,464]" italics="true" pageId="17" pageNumber="18">Cycadophila debaonica</emphasis>
</taxonomicName>
, occur either far north of the Red River (
<taxonomicName id="4C0A4D14FFF7FF9EFB2CFE53FA47C7E9" box="[1227,1426,439,464]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="debaoensis">
<emphasis id="B97EEA85FFF7FF9EFB2CFE53FA47C7E9" box="[1227,1426,439,464]" italics="true" pageId="17" pageNumber="18">Cycas debaoensis</emphasis>
</taxonomicName>
) or farther east and south of the Red River on limestone towers (
<taxonomicName id="4C0A4D14FFF7FF9EFC90FE36FB8CC7CC" box="[887,1113,476,501]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="hoabinhensis">
<emphasis id="B97EEA85FFF7FF9EFC90FE36FB8CC7CC" box="[887,1113,476,501]" italics="true" pageId="17" pageNumber="18">Cycas hoabinhensis</emphasis>
</taxonomicName>
). Other possible host 
<taxonomicName id="4C0A4D14FFF7FF9EFAB0FE36FA49C7CD" box="[1367,1436,477,500]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFF7FF9EFAB0FE36FA49C7CD" box="[1367,1436,477,500]" italics="true" pageId="17" pageNumber="18">Cycas</emphasis>
</taxonomicName>
species for 
<taxonomicName id="4C0A4D14FFF7FF9EFEFEFE14FE2AC421" box="[281,511,511,536]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="collina">
<emphasis id="B97EEA85FFF7FF9EFEFEFE14FE2AC421" box="[281,511,511,536]" italics="true" pageId="17" pageNumber="18">Cycadophila collina</emphasis>
</taxonomicName>
include 
<taxonomicName id="4C0A4D14FFF7FF9EFD84FDEBFC9CC421" box="[611,841,511,536]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="dolichophylla">
<emphasis id="B97EEA85FFF7FF9EFD84FDEBFC9CC421" box="[611,841,511,536]" italics="true" pageId="17" pageNumber="18">Cycas dolichophylla</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0A4D14FFF7FF9EFCBEFDEBFC68C421" box="[857,957,511,536]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="bifida">
<emphasis id="B97EEA85FFF7FF9EFCBEFDEBFC68C421" box="[857,957,511,536]" italics="true" pageId="17" pageNumber="18">C. bifida</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A4D14FFF7FF9EFC11FDEBFB83C421" box="[1014,1110,512,536]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="fugax">
<emphasis id="B97EEA85FFF7FF9EFC11FDEBFB83C421" box="[1014,1110,512,536]" italics="true" pageId="17" pageNumber="18">C. fugax</emphasis>
</taxonomicName>
, which occur in regions just north or east of 
<taxonomicName id="4C0A4D14FFF7FF9EFEAFFDCEFE37C404" box="[328,482,548,573]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="collina">
<emphasis id="B97EEA85FFF7FF9EFEAFFDCEFE37C404" box="[328,482,548,573]" italics="true" pageId="17" pageNumber="18">Cycas collina</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A4D14FFF7FF9EFDFFFDCEFC42C405" authority="Osborne et al. 2007" authorityName="Osborne et al." authorityYear="2007" box="[536,919,548,573]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="tanqingii">
<emphasis id="B97EEA85FFF7FF9EFDFFFDCEFD48C405" box="[536,669,549,572]" italics="true" pageId="17" pageNumber="18">C. tanqingii</emphasis>
(
<bibRefCitation id="EF9B4B66FFF7FF9EFD4BFDCFFC45C405" author="Osborne" box="[684,912,548,573]" pageId="17" pageNumber="18" refString="Osborne, R., Hill, K. D., Nguyen, H. T. &amp; Ke, L. P. (2007) Cycads of Fietnam. Tien Wah Press (Pte) Ltd, Singapore, 116 pp." type="book" year="2007">
Osborne 
<emphasis id="B97EEA85FFF7FF9EFCF3FDCEFC99C404" box="[788,844,548,573]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2007
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BB53697FFF7FF9EFF20FDACFC30C4F4" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="18">
Morphologically, 
<taxonomicName id="4C0A4D14FFF7FF9EFE74FDA3FDD0C459" box="[403,517,583,608]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="collina">
<emphasis id="B97EEA85FFF7FF9EFE74FDA3FDD0C459" box="[403,517,583,608]" italics="true" pageId="17" pageNumber="18">C. collina</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C0A4D14FFF7FF9EFD0FFDA3FCEDC459" box="[744,824,583,608]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B97EEA85FFF7FF9EFD0FFDA3FCEDC459" box="[744,824,583,608]" italics="true" pageId="17" pageNumber="18">C. lata</emphasis>
</taxonomicName>
in development of male sexual dimorphisms. These species also appear to occupy distinct, but neighboring distributions (see Remarks under 
<taxonomicName id="4C0A4D14FFF7FF9EFB7DFD86FB3FC4BC" box="[1178,1258,620,645]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B97EEA85FFF7FF9EFB7DFD86FB3FC4BC" box="[1178,1258,620,645]" italics="true" pageId="17" pageNumber="18">C. lata</emphasis>
</taxonomicName>
). Even without molecular analyses, the striking difference in ventral interocular distance and different distributions allow us to maintain 
<taxonomicName id="4C0A4D14FFF7FF9EFEE7FD5EFE99C4F4" box="[256,332,692,717]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B97EEA85FFF7FF9EFEE7FD5EFE99C4F4" box="[256,332,692,717]" italics="true" pageId="17" pageNumber="18">C. lata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A4D14FFF7FF9EFE65FD5EFE24C4F4" box="[386,497,692,717]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="collina">
<emphasis id="B97EEA85FFF7FF9EFE65FD5EFE24C4F4" box="[386,497,692,717]" italics="true" pageId="17" pageNumber="18">C. collina</emphasis>
</taxonomicName>
as valid species within the species complex.
</paragraph>
<paragraph id="8BB53697FFF7FF9EFF20FD3CFE55C564" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="18">
Considering the morphological similarity in species of the 
<taxonomicName id="4C0A4D14FFF7FF9EFC83FD3CFC59C4C9" box="[868,908,727,752]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lata">lata</taxonomicName>
complex, the 
<collectingCountry id="F31D7607FFF7FF9EFBC8FD33FBB0C4D6" box="[1071,1125,728,751]" name="Laos" pageId="17" pageNumber="18">Laos</collectingCountry>
<typeStatus id="54B18835FFF7FF9EFB89FD3CFB05C4C9" box="[1134,1232,727,752]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C0A4D14FFF7FF9EFB1FFD33FABDC4C9" box="[1272,1384,727,752]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="collina">
<emphasis id="B97EEA85FFF7FF9EFB1FFD33FABDC4C9" box="[1272,1384,727,752]" italics="true" pageId="17" pageNumber="18">C. collina</emphasis>
</taxonomicName>
was chosen because it was distinguished with both molecular and morphological evidence. Although the series of Chinese and Vietnamese 
<typeStatus id="54B18835FFF7FF9EFE54FCCBFDCAC501" box="[435,543,800,824]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
match the 
<collectingCountry id="F31D7607FFF7FF9EFD7AFCCBFD06C50E" box="[669,723,800,823]" name="Laos" pageId="17" pageNumber="18">Laos</collectingCountry>
specimen morphologically, they have not been confirmed with 16S rRNA evidence.
</paragraph>
</subSubSection>
</treatment>
</document>