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<document id="220436E4BD35A131BF863BEF857EE5FA" ID-ISSN="0024-4082" ID-ZooBank="3A70B56-000D-4974-AAC6-F1B198C86BC4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738885802293" checkinUser="plazi" docAuthor="Golestani, Haleh, Crocetta, Fabio, Padula, Vinicius, Camacho-García, Yolanda, Langeneck, Joachim, Poursanidis, Dimitris, Pola, Marta, Yokeş, M. Baki, Cervera, Juan Lucas, Jung, Dae-Wui, Gosliner, Terrence M., Araya, Juan Francisco, Hooker, Yuri, Schrödl, Michael &amp; Valdés, Ángel" docDate="2019" docId="038987ACF57BFFCDB19DF998AEF8F960" docLanguage="en" docName="zlz028.pdf" docOrigin="Zoological Journal of the Linnean Society 187" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Aplysia nigrocincta von Martens 1880" docType="treatment" docVersion="1" lastPageNumber="314" masterDocId="FFB0FFD4F55BFFEEB153FFC9AF15FFA9" masterDocTitle="The little Aplysia coming of age: from one species to a complex of species complexes in Aplysia parvula (Mollusca: Gastropoda: Heterobranchia)" masterLastPageNumber="330" masterPageNumber="279" pageNumber="311" updateTime="1738889083971" updateUser="GgImagineBatch">
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<mods:title id="D3C141AB8D0C7D725E1C90A6623F6752">The little Aplysia coming of age: from one species to a complex of species complexes in Aplysia parvula (Mollusca: Gastropoda: Heterobranchia)</mods:title>
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<mods:namePart id="53CCA7A1F39A3DDF7C6645DA6EB10D02">Padula, Vinicius</mods:namePart>
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<mods:namePart id="D094CCEF9AACB107B6C74BD2ADDC3C43">Pola, Marta</mods:namePart>
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<mods:namePart id="4F0010E268460B2A44060F2C66AF84A7">Gosliner, Terrence M.</mods:namePart>
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<mods:namePart id="D743C6DD52A91251EEA2D5C51C97F2AD">Araya, Juan Francisco</mods:namePart>
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<mods:namePart id="9DD43AC705CB8BC000BD70816F9B8F94">Schrödl, Michael</mods:namePart>
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<mods:title id="09D3197D3BBFDB224C18CB29DDEE9662">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="5F1BCD818F05FCD730385F77B115FD70">2019</mods:date>
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<mods:number id="4C1ECC50A46F311ACADECD1AFA31C8F7">187</mods:number>
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<treatment id="038987ACF57BFFCDB19DF998AEF8F960" LSID="urn:lsid:plazi:treatment:038987ACF57BFFCDB19DF998AEF8F960" httpUri="http://treatment.plazi.org/id/038987ACF57BFFCDB19DF998AEF8F960" lastPageId="35" lastPageNumber="314" pageId="32" pageNumber="311">
<subSubSection id="C33A6531F57BFFCEB19DF998ADAEF9C0" box="[206,699,1616,1641]" pageId="32" pageNumber="311" type="nomenclature">
<paragraph id="8B9F36BAF57BFFCEB19DF998ADAEF9C0" blockId="32.[206,699,1616,1641]" box="[206,699,1616,1641]" pageId="32" pageNumber="311">
<heading id="D0D781D6F57BFFCEB19DF998ADAEF9C0" box="[206,699,1616,1641]" centered="true" fontSize="9" level="2" pageId="32" pageNumber="311" reason="2">
<taxonomicName id="4C204D39F57BFFCEB19DF998ADAEF9C0" ID-CoL="7Q6DX" authority="VON MARTENS, 1880" authorityName="von Martens" authorityYear="1880" box="[206,699,1616,1641]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="32" pageNumber="311" phylum="Mollusca" rank="species" species="nigrocincta">
<emphasis id="B954EAA8F57BFFCEB19DF998AEDEF9C1" box="[206,459,1617,1640]" italics="true" pageId="32" pageNumber="311">
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<smallCapsWord id="8D79A066F57BFFCEB061F99DAEDEF9C1" baselines="1635" box="[306,459,1620,1640]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nigrocincta" pageId="32" pageNumber="311">NIGROCINCTA</smallCapsWord>
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<bibRefCitation id="EFB14B4BF57BFFCEB081F99DADAEF9C0" author="von Martens E" box="[466,699,1616,1641]" pageId="32" pageNumber="311" pagination="19 - 22" refId="ref32340" refString="von Martens E. 1880. Mollusken. In: Mobius K, Richters F, von Martens E, eds. Beitrage zur Meeresfauna der Insel Mauritius und der Seychellen. Berlin: Gutmann'schen Buchhandlung, 181 - 352, pls. 19-22." type="book chapter" year="1880">
<smallCapsWord id="8D79A066F57BFFCEB081F99DAD14F9C1" baselines="1635" box="[466,513,1620,1640]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="von" pageId="32" pageNumber="311">VON</smallCapsWord>
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, 1880
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<subSubSection id="C33A6531F57BFFCEB197F9BEADD0F939" box="[196,709,1655,1680]" pageId="32" pageNumber="311" type="description">
<paragraph id="8B9F36BAF57BFFCEB197F9BEADD0F939" blockId="32.[196,709,1655,1680]" box="[196,709,1655,1680]" pageId="32" pageNumber="311">
(
<figureCitation id="131B2A3FF57BFFCEB19FF9BEAE44F939" box="[204,337,1655,1680]" captionStart="Figure 4" captionStartId="15.[162,241,1223,1245]" captionTargetBox="[163,1443,195,1183]" captionTargetId="figure-194@15.[163,1443,195,1183]" captionTargetPageId="15" captionText="Figure 4. Photographs of type specimens of Aplysia parvula s.l. A–B, shell of syntype of A. parvula Pease, 1868 from the ‘West Indies’ (NHMD 288589), ventral view (A), dorsal view (B), photos: Kathe Jensen NHMD. C–D, shell of syntype of Syphonota punctata Pease, 1868 from Tahiti (MCZM 297868), ventral view (C), dorsal view (D), photos: Jennifer Trimble MCZM. E–F, shell of syntype of S. elongata Pease, 1860 from the Hawaiian Is. (MCZM 298486), ventral view (E), dorsal view (F), photos: Jennifer Trimble MCZM. G–H, shell of syntype of S. elongata Pease, 1860 from the Hawaiian Is. (ANSP 20686), ventral view (G), dorsal view (H), photos: ANSP. I–J, shell of syntype of A. japonica Clessin, 1899 (ZMB Moll 38136), ventral view (I), dorsal view (J), photos: ZMB. K–L, shell of syntype of A. nigrocincta von Martens, 1880 (ZMB Moll 32165), ventral view (K), dorsal view (L), photos: Christine Zorn ZMB. M, dorsal view of shell with tissue of syntype of A.nigrocincta von Martens, 1880 (ZMB Moll 51472), photo: ZMB. N–O, shell of syntype of A. orientalis G. D. Sowerby II, 1869 (NHMUK 1958.1.9.270-1), ventral view (N), dorsal view (O), photos: Harry Taylor NHMUK. P–Q, shell of syntype of A. japonica G. D. Sowerby II, 1869 (NHMUK 1958.1.9.267-9), ventral view (P), dorsal view (Q), photos: Harry Taylor NHMUK." pageId="32" pageNumber="311">
<smallCapsWord id="8D79A066F57BFFCEB19FF9BEAE17F926" baselines="1674,1674" box="[204,258,1655,1680]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="32" pageNumber="311">FIGS</smallCapsWord>
4K–M
</figureCitation>
,
<figureCitation id="131B2A3FF57BFFCEB00DF9BEAEFFF939" box="[350,490,1655,1680]" captionStart="Figure 14" captionStartId="28.[146,224,1540,1562]" captionTargetBox="[304,1264,197,1499]" captionTargetId="figure-94@28.[303,1266,195,1500]" captionTargetPageId="28" captionText="Figure 14. Photographs of living animals in the Aplysia nigrocincta–elongata species complex. A, A. nigrocincta, specimen from Mindoro, Philippines (CASIZ 208581, isolate HG99). B, A. nigrocincta, specimen from Madang, Papua New Guinea (CASIZ 191092, isolate HG103). C, A. nigrocincta, specimen from Savai’i Is., Samoa (ZSM Mol 20060223). D, A. hooveri, specimen from Tumbes, Peru (ZSM Mol 20090834). E, A. elongata, specimen from Napili Bay, Maui, Hawaiian Is. (CPIC 00300, isolate HG5). F, A.elongata, specimen from Maliko Bay, Maui, Hawaiian Is. (CPIC 00363, isolate HG6). G, A.elongata, specimen from Maliko Bay, Maui, Hawaiian Is. (CPIC 00333, isolate HG9). H, A. nigrocincta, specimen from Madang, Papua New Guinea (CASIZ 191335, isolate HG102)." pageId="32" pageNumber="311">14A–C, 14H</figureCitation>
,
<figureCitation id="131B2A3FF57BFFCEB0ABF9BEAD30F939" box="[504,549,1655,1680]" captionStart="Figure 15" captionStartId="29.[163,242,857,879]" captionTargetBox="[163,1443,195,818]" captionTargetId="figure-440@29.[163,1443,195,818]" captionTargetPageId="29" captionText="Figure 15. Scanning electron micrographs of shells of specimens in the Aplysia nigrocincta–elongata species complex. A, A. elongata, specimen from Maui, Hawaiian Is. (CPIC 00325, isolate HG18) B, A. hooveri, specimen from Punta Belches, Mexico (CPIC 01851, isolate HG94). C, A. nigrocincta, specimen from Madang, Papua New Guinea (CASIZ 191335, isolate HG102)." pageId="32" pageNumber="311">15C</figureCitation>
,
<figureCitation id="131B2A3FF57BFFCEB361F9BEAD95F939" box="[562,640,1655,1680]" captionStart="Figure 16" captionStartId="30.[145,223,1363,1385]" captionTargetBox="[147,1426,197,1321]" captionTargetId="figure-162@30.[145,1428,195,1323]" captionTargetPageId="30" captionText="Figure 16. Scanning electron micrographs of the radular teeth and jaws of specimens in the Aplysia nigrocincta–elongata species complex. A–C, A. nigrocincta, specimen from Mindoro, Philippines (CASIZ 208581, isolate HG105): A, rachidian and inner radular teeth; B, outer radular teeth; C, jaw elements. D–F, A. elongata, specimen from Napili Bay, Maui, Hawaiian Isalnds (CPIC 00333, isolate HG9): D, rachidian and inner radular teeth; E, outer radular teeth; F, jaw elements. G–I, A. hooveri, specimen from the Galapagos Islands (LACM 72–189): G, rachidian and inner radular teeth; H, outer radular teeth; I, jaw elements." pageId="32" pageNumber="311">16A–C</figureCitation>
,
<figureCitation id="131B2A3FF57BFFCEB3DEF9BEADA8F939" box="[653,701,1655,1680]" captionStart="Figure 17" captionStartId="31.[163,243,1079,1101]" captionTargetBox="[227,1379,195,1039]" captionTargetId="figure-67@31.[227,1379,195,1039]" captionTargetPageId="31" captionText="Figure 17. Photographs of the penis of specimens in the Aplysia nigrocincta–elongata species complex. A, A. elongata, specimen from Napili Bay, Maui, Hawaiian Is. (CPIC 00333, isolate HG9). B, A.hooveri, specimen from Bahia de Los Angeles, Baja California, Mexico (CASIZ 071868). C, A. hooveri, specimen from the Galapagos Is. (LACM 72–189). D, A. nigrocincta, specimen from Madang, Papua New Guinea (CASIZ 191335, isolate HG102). Abbreviations: pp, penis; pr, prostate; rm, retractor muscle." pageId="32" pageNumber="311">17D</figureCitation>
)
</paragraph>
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<paragraph id="8B9F36BAF57BFFCEB1C2F956AD51F899" blockId="32.[145,761,1695,1901]" pageId="32" pageNumber="311">
<treatmentCitationGroup id="AB301194F57BFFCEB1C2F956AD51F899" pageId="32" pageNumber="311">
<treatmentCitation id="0A8110ABF57BFFCEB1C2F956ADB7F91C" author="von Martens E" box="[145,674,1695,1717]" page="307" pageId="32" pageNumber="311" year="1880">
<taxonomicName id="4C204D39F57BFFCEB1C2F956ADB7F91C" ID-CoL="7Q6DX" authority="von Martens, 1880: 307" authorityName="von Martens" authorityPageNumber="307" authorityYear="1880" box="[145,674,1695,1717]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="32" pageNumber="311" phylum="Mollusca" rank="species" species="nigrocincta">
<emphasis id="B954EAA8F57BFFCEB1C2F956AE6EF91C" box="[145,379,1695,1717]" italics="true" pageId="32" pageNumber="311">Aplysia nigrocincta</emphasis>
<materialsCitation id="3B483CE7F57BFFCEB0D7F969ADB7F91C" box="[388,674,1696,1717]" collectorName="Mobius" country="Mauritius" location="Mauritius" pageId="32" pageNumber="311" specimenCount="1" typeStatus="syntype">
<bibRefCitation id="EFB14B4BF57BFFCEB0D7F969ADB7F91C" author="von Martens E" box="[388,674,1696,1717]" pageId="32" pageNumber="311" pagination="19 - 22" refId="ref32340" refString="von Martens E. 1880. Mollusken. In: Mobius K, Richters F, von Martens E, eds. Beitrage zur Meeresfauna der Insel Mauritius und der Seychellen. Berlin: Gutmann'schen Buchhandlung, 181 - 352, pls. 19-22." type="book chapter" year="1880">von Martens, 1880: 307</bibRefCitation>
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, pl. 21, fig. 3.
<emphasis id="B954EAA8F57BFFCEB1A5F976AE82F97A" box="[246,407,1726,1748]" italics="true" pageId="32" pageNumber="311">Type locality:</emphasis>
Île aux Fouquets,
<collectingCountry id="F337762AF57BFFCEB329F977ADE1F97D" box="[634,756,1726,1748]" name="Mauritius" pageId="32" pageNumber="311">Mauritius</collectingCountry>
.
<emphasis id="B954EAA8F57BFFCEB1F9F914AE0FF95B" box="[170,282,1757,1778]" italics="true" pageId="32" pageNumber="311">
<typeStatus id="549B8818F57BFFCEB1F9F914AE06F95B" box="[170,275,1757,1778]" pageId="32" pageNumber="311" type="syntype">Syntypes</typeStatus>
:
</emphasis>
<collectingCountry id="F337762AF57BFFCEB070F914AE8EF95A" box="[291,411,1757,1779]" name="Mauritius" pageId="32" pageNumber="311">Mauritius</collectingCountry>
, dry shell, 14.2-mm long, leg.
<collectorName id="26D5536CF57BFFCEB1F9F932AFEBF8B8" box="[170,254,1787,1809]" pageId="32" pageNumber="311">Möbius</collectorName>
(ZMB Moll 32165), shell with tissue 10-mm long, leg.
<collectorName id="26D5536CF57BFFCEB047F8D3AE7DF899" box="[276,360,1818,1840]" pageId="32" pageNumber="311">Möbius</collectorName>
(ZMB Moll 51472).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33A6531F57BFFCDB1C2F8F0AEF8F960" lastPageId="35" lastPageNumber="314" pageId="32" pageNumber="311" type="materials_examined">
<paragraph id="8B9F36BAF57BFFCEB1C2F8F0ADE2F8E6" blockId="32.[145,761,1695,1901]" box="[145,759,1849,1871]" pageId="32" pageNumber="311">
<emphasis id="B954EAA8F57BFFCEB1C2F8F0AD37F8E7" box="[145,546,1849,1870]" italics="true" pageId="32" pageNumber="311">Additional material examined:</emphasis>
Savai’i Island,
</paragraph>
<paragraph id="8B9F36BAF57BFFCEB1C2F891ADECF8C4" blockId="32.[145,761,1695,1901]" box="[145,761,1879,1901]" pageId="32" pageNumber="311">
<collectingCountry id="F337762AF57BFFCEB1C2F891AFF5F8C4" box="[145,224,1880,1901]" name="Samoa" pageId="32" pageNumber="311">Samoa</collectingCountry>
, leg. M. Schrödl (
<collectionCode id="ED31AE7FF57BFFCEB0F2F891AECDF8C4" box="[417,472,1880,1901]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="32" pageNumber="311" type="Museum">ZSM</collectionCode>
Mol 20060223). Mindoro,
</paragraph>
<paragraph id="8B9F36BAF57BFFCEB27AFCB3ACD8FBCE" blockId="32.[809,1425,890,1127]" pageId="32" pageNumber="311">
<materialsCitation id="3B483CE7F57BFFCEB27AFCB3AA98FC06" collectingDate="2015-04-28" collectionCode="CASIZ" collectorName="P. J Aristorenas" country="Philippines" location="Madang" pageId="32" pageNumber="311" specimenCode="CASIZ 208581" specimenCount="1" stateProvince="Madang">
<collectingCountry id="F337762AF57BFFCEB27AFCB3ACA0FC39" box="[809,949,890,912]" name="Philippines" pageId="32" pageNumber="311">Philippines</collectingCountry>
,
<date id="FF9E107AF57BFFCEB297FCB3AB78FC39" box="[964,1133,890,912]" pageId="32" pageNumber="311" value="2015-04-28">
<collectingDate id="EFDAE992F57BFFCEB297FCB3AB78FC39" box="[964,1133,890,912]" pageId="32" pageNumber="311" value="2015-04-28">28 April 2015</collectingDate>
</date>
,
<specimenCount id="9D26FD33F57BFFCEB52FFCB2AA36FC39" box="[1148,1315,890,912]" count="1" pageId="32" pageNumber="311" type="generic">one specimen</specimenCount>
, leg.
<collectorName id="26D5536CF57BFFCEB437FCB2ACA5FC06" pageId="32" pageNumber="311">P. J Aristorenas</collectorName>
(
<specimenCode id="DB869EC1F57BFFCEB2EDFC50AB7DFC06" box="[958,1128,921,943]" collectionCode="CASIZ" pageId="32" pageNumber="311">CASIZ 208581</specimenCode>
, isolate HG105).
<collectingRegion id="49E4F858F57BFFCEB47DFC50AA98FC06" box="[1326,1421,921,943]" country="Papua New Guinea" name="Madang" pageId="32" pageNumber="311">Madang</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B483CE7F57BFFCEB27AFC71AA61FC42" collectingDate="2012-11-19" collectionCode="CASIZ" collectorName="V. Knutson" country="Papua New Guinea" location="Papua New Guinea" pageId="32" pageNumber="311" specimenCode="CASIZ 191335" specimenCount="1">
<collectingCountry id="F337762AF57BFFCEB27AFC71AB12FC64" box="[809,1031,951,973]" name="Papua New Guinea" pageId="32" pageNumber="311">Papua New Guinea</collectingCountry>
,
<date id="FF9E107AF57BFFCEB542FC71ABFDFC64" box="[1041,1256,951,973]" pageId="32" pageNumber="311" value="2012-11-19">
<collectingDate id="EFDAE992F57BFFCEB542FC71ABFDFC64" box="[1041,1256,951,973]" pageId="32" pageNumber="311" value="2012-11-19">19 November 2012</collectingDate>
</date>
,
<specimenCount id="9D26FD33F57BFFCEB5A0FC71AA98FC64" box="[1267,1421,951,973]" count="1" pageId="32" pageNumber="311" type="generic">one specimen</specimenCount>
, leg.
<collectorName id="26D5536CF57BFFCEB20FFC1EACF1FC45" box="[860,996,983,1004]" pageId="32" pageNumber="311">V. Knutson</collectorName>
(
<specimenCode id="DB869EC1F57BFFCEB2A6FC1FABBEFC45" box="[1013,1195,982,1004]" collectionCode="CASIZ" pageId="32" pageNumber="311">CASIZ 191335</specimenCode>
, isolate HG102)
</materialsCitation>
;
<materialsCitation id="3B483CE7F57BFFCEB4D0FC1FAB54FB80" collectingDate="2012-11-09" collectionCode="CASIZ" collectorName="V. Knutson" country="Papua New Guinea" pageId="32" pageNumber="311" specimenCode="CASIZ 191092" specimenCount="1">
<date id="FF9E107AF57BFFCEB4D0FC1FACC8FBA3" pageId="32" pageNumber="311" value="2012-11-09">
<collectingDate id="EFDAE992F57BFFCEB4D0FC1FACC8FBA3" pageId="32" pageNumber="311" value="2012-11-09">9 November 2012</collectingDate>
</date>
,
<specimenCount id="9D26FD33F57BFFCEB2BBFC3CAB97FBA2" box="[1000,1154,1013,1035]" count="1" pageId="32" pageNumber="311" type="generic">one specimen</specimenCount>
, leg.
<collectorName id="26D5536CF57BFFCEB5EAFC3CAA23FBA3" box="[1209,1334,1013,1034]" pageId="32" pageNumber="311">V. Knutson</collectorName>
(
<specimenCode id="DB869EC1F57BFFCEB417FC3CAC95FB80" collectionCode="CASIZ" pageId="32" pageNumber="311">CASIZ 191092</specimenCode>
, isolate HG103)
</materialsCitation>
.
<materialsCitation id="3B483CE7F57BFFCEB51EFBDAACDCFBCE" collectionCode="CPIC" collectorName="M. Pola" country="Mozambique" location="Mozambique" pageId="32" pageNumber="311" specimenCode="CPIC 01386" specimenCount="1">
<collectingCountry id="F337762AF57BFFCEB51EFBDAABF0FB80" box="[1101,1253,1043,1065]" name="Mozambique" pageId="32" pageNumber="311">Mozambique</collectingCountry>
,
<specimenCount id="9D26FD33F57BFFCEB5A1FBDDAA85FB80" box="[1266,1424,1043,1065]" count="1" pageId="32" pageNumber="311" type="generic">one specimen</specimenCount>
<quantity id="4CD89B5FF57BFFCEB27AFBFBAC78FBE1" box="[809,877,1074,1096]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="32" pageNumber="311" unit="mm" value="9.0">9 mm</quantity>
preserved length, leg.
<collectorName id="26D5536CF57BFFCEB5DAFBFAABF0FBE1" box="[1161,1253,1074,1096]" pageId="32" pageNumber="311">M. Pola</collectorName>
(
<specimenCode id="DB869EC1F57BFFCEB5A5FBFBAA99FBE1" box="[1270,1420,1074,1096]" collectionCode="CPIC" pageId="32" pageNumber="311">CPIC 01386</specimenCode>
, isolate HG85)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9F36BAF57BFFCEB27AFB59AC9EFA33" blockId="32.[809,1425,1167,1434]" pageId="32" pageNumber="311">
<emphasis id="B954EAA8F57BFFCEB27AFB59ACA0FB0C" box="[809,949,1168,1189]" italics="true" pageId="32" pageNumber="311">Description:</emphasis>
Morphological characteristics as in the description of species complex 3. Radular formula 22 × 3.6.1.6.
<quantity id="4CD89B5FF57BFFCEB2E8FB04ACFDFB4A" box="[955,1000,1229,1251]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="32" pageNumber="311" unit="in" value="3.0">3 in</quantity>
a
<quantity id="4CD89B5FF57BFFCEB554FB04AB5CFB4B" box="[1031,1097,1229,1250]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="32" pageNumber="311" unit="mm" value="9.0">9 mm</quantity>
preserved length specimen from
<collectingCountry id="F337762AF57BFFCEB23AFB22AB14FAA8" box="[873,1025,1259,1281]" name="Mozambique" pageId="32" pageNumber="311">Mozambique</collectingCountry>
(
<collectionCode id="ED31AE7FF57BFFCEB541FB25AB46FAA8" box="[1042,1107,1260,1281]" pageId="32" pageNumber="311">CPIC</collectionCode>
01386, isolate HG85), n × 2.6.1.6.
<quantity id="4CD89B5FF57BFFCEB22FFAC3ACBCFA89" box="[892,937,1290,1312]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="32" pageNumber="311" unit="in" value="2.0">2 in</quantity>
a
<quantity id="4CD89B5FF57BFFCEB295FAC3AB00FA89" box="[966,1045,1290,1312]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="311" unit="mm" value="15.0">15 mm</quantity>
preserved length specimen from the
<collectingCountry id="F337762AF57BFFCEB204FAE0ACCBFA96" box="[855,990,1321,1343]" name="Philippines" pageId="32" pageNumber="311">Philippines</collectingCountry>
(
<collectionCode id="ED31AE7FF57BFFCEB2BDFAE0AB2BFA97" box="[1006,1086,1321,1342]" pageId="32" pageNumber="311">CASIZ</collectionCode>
208581, isolate HG105) and n × 3.7.1.7.
<quantity id="4CD89B5FF57BFFCEB2FDFA81ACCEFAF4" box="[942,987,1351,1373]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="32" pageNumber="311" unit="in" value="3.0">3 in</quantity>
a
<quantity id="4CD89B5FF57BFFCEB2A9FA81AB5FFAF4" box="[1018,1098,1352,1373]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="311" unit="mm" value="20.0">20 mm</quantity>
preserved length specimen from
<collectingCountry id="F337762AF57BFFCEB23EFAAEAB77FAD5" box="[877,1122,1382,1404]" name="Papua New Guinea" pageId="32" pageNumber="311">Papua New Guinea</collectingCountry>
(
<collectionCode id="ED31AE7FF57BFFCEB527FAAFABD2FAD2" box="[1140,1223,1382,1403]" pageId="32" pageNumber="311">CASIZ</collectionCode>
191335, isolate HG102).
</paragraph>
<paragraph id="8B9F36BAF57BFFCEB27AFA0DAC87F9FA" blockId="32.[809,1425,1475,1619]" pageId="32" pageNumber="311">
<emphasis id="B954EAA8F57BFFCEB27AFA0DAC6FFA70" box="[809,890,1476,1497]" italics="true" pageId="32" pageNumber="311">Range:</emphasis>
The range of
<taxonomicName id="4C204D39F57BFFCEB560FA0AAA08FA70" authorityName="von Martens" authorityYear="1880" box="[1075,1309,1475,1497]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="32" pageNumber="311" phylum="Mollusca" rank="species" species="nigrocincta">
<emphasis id="B954EAA8F57BFFCEB560FA0AAA08FA70" box="[1075,1309,1475,1497]" italics="true" pageId="32" pageNumber="311">Aplysia nigrocincta</emphasis>
</taxonomicName>
has been confirmed to include the Indian and eestern Pacific Oceans, from East Africa (
<collectingCountry id="F337762AF57BFFCEB505F9C8ABFEF9BE" box="[1110,1259,1537,1559]" name="Mozambique" pageId="32" pageNumber="311">Mozambique</collectingCountry>
) to
<collectingCountry id="F337762AF57BFFCEB44BF9C8AA98F9BE" box="[1304,1421,1537,1559]" name="Mauritius" pageId="32" pageNumber="311">Mauritius</collectingCountry>
, the
<collectingCountry id="F337762AF57BFFCEB20BF9D6ACF7F99C" box="[856,994,1567,1589]" name="Philippines" pageId="32" pageNumber="311">Philippines</collectingCountry>
,
<collectingCountry id="F337762AF57BFFCEB2BCF9E9ABCCF99C" box="[1007,1241,1567,1589]" name="Papua New Guinea" pageId="32" pageNumber="311">Papua New Guinea</collectingCountry>
,
<collectingCountry id="F337762AF57BFFCEB5B4F9D6AA49F99C" box="[1255,1372,1567,1589]" name="Indonesia" pageId="32" pageNumber="311">Indonesia</collectingCountry>
and
<collectingCountry id="F337762AF57BFFCEB27AF9F7AC9BF9FA" box="[809,910,1598,1619]" name="Vanuatu" pageId="32" pageNumber="311">Vanuatu</collectingCountry>
.
</paragraph>
<paragraph id="8B9F36BAF57BFFCEB27AF95EAC83F906" blockId="32.[809,918,1687,1711]" box="[809,918,1687,1711]" pageId="32" pageNumber="311">
<emphasis id="B954EAA8F57BFFCEB27AF95EAC83F906" box="[809,918,1687,1711]" italics="true" pageId="32" pageNumber="311">Remarks</emphasis>
</paragraph>
<paragraph id="8B9F36BAF57BFFCFB27AF976AD7BF9FE" blockId="32.[809,1425,1727,1903]" lastBlockId="33.[163,779,1540,1899]" lastPageId="33" lastPageNumber="312" pageId="32" pageNumber="311">
Several specimens from the tropical Indo-Pacific of
<emphasis id="B954EAA8F57BFFCEB27AF917ACC6F95A" box="[809,979,1758,1779]" italics="true" pageId="32" pageNumber="311">
<taxonomicName id="4C204D39F57BFFCEB27AF917ACBDF95A" authorityName="Morch" authorityYear="1863" box="[809,936,1758,1779]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="32" pageNumber="311" phylum="Mollusca" rank="species" species="parvula">A. parvula</taxonomicName>
<taxonomicNameLabel id="A26757D3F57BFFCEB2E2F917ACC6F95A" box="[945,979,1758,1779]" pageId="32" pageNumber="311" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
sequenced here constitute a distinct species, as confirmed by the ABGD analysis. These specimens are morphologically and genetically members of species complex 3, but are distinct from other species in this complex from the eastern Pacific and the Hawaiian Islands. A review of the literature reveals that the name
<taxonomicName id="4C204D39F57AFFCFB0EAF9EAAFCFF9FF" authority="von Martens, 1880" authorityName="von Martens" authorityYear="1880" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="33" pageNumber="312" phylum="Mollusca" rank="species" species="nigrocincta">
<emphasis id="B954EAA8F57AFFCFB0EAF9EAAD77F991" box="[441,610,1571,1592]" italics="true" pageId="33" pageNumber="312">A. nigrocincta</emphasis>
<bibRefCitation id="EFB14B4BF57AFFCFB33FF9EAAFCFF9FF" author="von Martens E" pageId="33" pageNumber="312" pagination="19 - 22" refId="ref32340" refString="von Martens E. 1880. Mollusken. In: Mobius K, Richters F, von Martens E, eds. Beitrage zur Meeresfauna der Insel Mauritius und der Seychellen. Berlin: Gutmann'schen Buchhandlung, 181 - 352, pls. 19-22." type="book chapter" year="1880">von Martens, 1880</bibRefCitation>
</taxonomicName>
most likely represents this species.
</paragraph>
<caption id="DF5F6632F57AFFCFB1F7FA9BACD2FA77" pageId="33" pageNumber="312" startId="33.[164,243,1362,1384]" targetBox="[163,1442,197,1320]" targetPageId="33" targetType="figure">
<paragraph id="8B9F36BAF57AFFCFB1F7FA9BACD2FA77" blockId="33.[163,1444,1362,1502]" pageId="33" pageNumber="312">
<emphasis id="B954EAA8F57AFFCFB1F7FA9BAE0FFACE" bold="true" box="[164,282,1362,1384]" pageId="33" pageNumber="312">Figure 20.</emphasis>
Scanning electron micrographs of the radular teeth and jaws of specimens of
<taxonomicName id="4C204D39F57AFFCFB51CFA9BABE8FAC1" authorityName="G. D. Sowerby II" authorityYear="1869" box="[1103,1277,1362,1384]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="33" pageNumber="312" phylum="Mollusca" rank="species" species="japonica">
<emphasis id="B954EAA8F57AFFCFB51CFA9BABE8FAC1" box="[1103,1277,1362,1384]" italics="true" pageId="33" pageNumber="312">Aplysia japonica</emphasis>
</taxonomicName>
. A–C, specimen from southern Hokkaido, Japan (CPIC 01367, isolate HG61): A, rachidian and inner radular teeth; B, outer radular teeth; C, jaw elements. D–F, specimen from Jeju-do, South Korea (CPIC 02176, isolate HG130): D, rachidian and inner radular teeth; E, outer radular teeth; F, jaw elements. G–I, specimen from Gyeongsangbuk-do, South Korea (CPIC 02180, isolate HG134): G, rachidian and inner radular teeth; H, outer radular teeth; I, jaw elements.
</paragraph>
</caption>
<paragraph id="8B9F36BAF57AFFCCB1E8F9A9AB9EF8C0" blockId="33.[163,779,1540,1899]" lastBlockId="34.[809,1425,1813,1897]" lastPageId="34" lastPageNumber="313" pageId="33" pageNumber="312">
<bibRefCitation id="EFB14B4BF57AFFCFB1E8F9A9AEC6F9DC" author="von Martens E" box="[187,467,1632,1653]" pageId="33" pageNumber="312" pagination="19 - 22" refId="ref32340" refString="von Martens E. 1880. Mollusken. In: Mobius K, Richters F, von Martens E, eds. Beitrage zur Meeresfauna der Insel Mauritius und der Seychellen. Berlin: Gutmann'schen Buchhandlung, 181 - 352, pls. 19-22." type="book chapter" year="1880">Von Martens (1880)</bibRefCitation>
introduced the name
<taxonomicName id="4C204D39F57AFFCFB1F0F9B6AE47F93D" authorityName="von Martens" authorityYear="1880" box="[163,338,1663,1684]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="33" pageNumber="312" phylum="Mollusca" rank="species" species="nigrocincta">
<emphasis id="B954EAA8F57AFFCFB1F0F9B6AE47F93D" box="[163,338,1663,1684]" italics="true" pageId="33" pageNumber="312">A. nigrocincta</emphasis>
</taxonomicName>
based on a 17-mm long preserved specimen collected from
<collectingCountry id="F337762AF57AFFCFB0E4F954AD3CF91A" box="[439,553,1693,1715]" name="Mauritius" pageId="33" pageNumber="312">Mauritius</collectingCountry>
and described it as: ‘Wide, smooth, light brown, the free edges of the foot flaps, the edge of the sole of the foot and the edge of the rather large hole in the coat (shields) black, mostly continuous, sometimes slightly stained. Shell strongly domed, comparatively large, vivacious amber-yellow, reddish translucent by the shield, spoon-shaped, with a shallow neck and strongly inflected point, occupying more than half the length of the contracted animal.’ This description matches the characteristics of animals in species complex 3, including the presence of a large mantle foramen edged in black and a very large visceral hump, occupying a large portion of the animal. We also examined the type material of
<emphasis id="B954EAA8F57AFFCFB5FAF975AA41F978" box="[1193,1364,1724,1745]" italics="true" pageId="33" pageNumber="312">
<taxonomicName id="4C204D39F57AFFCFB5FAF975AA45F978" authorityName="von Martens" authorityYear="1880" box="[1193,1360,1724,1745]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="33" pageNumber="312" phylum="Mollusca" rank="species" species="nigrocincta">A. nigrocincta</taxonomicName>
,
</emphasis>
which consists of
<specimenCount id="9D26FD33F57AFFCFB2E2F912AB54F959" box="[945,1089,1755,1776]" count="2" pageId="33" pageNumber="312" type="generic" typeStatus="syntype">two syntypes</specimenCount>
remaining, a dry shell specimen and a wet specimen with a shell and some soft parts remaining (ZMB Moll 32165, ZMB Moll 51472) (
<figureCitation id="131B2A3FF57AFFCFB424F8D1AC91F8E2" captionStart="Figure 4" captionStartId="15.[162,241,1223,1245]" captionTargetBox="[163,1443,195,1183]" captionTargetId="figure-194@15.[163,1443,195,1183]" captionTargetPageId="15" captionText="Figure 4. Photographs of type specimens of Aplysia parvula s.l. A–B, shell of syntype of A. parvula Pease, 1868 from the ‘West Indies’ (NHMD 288589), ventral view (A), dorsal view (B), photos: Kathe Jensen NHMD. C–D, shell of syntype of Syphonota punctata Pease, 1868 from Tahiti (MCZM 297868), ventral view (C), dorsal view (D), photos: Jennifer Trimble MCZM. E–F, shell of syntype of S. elongata Pease, 1860 from the Hawaiian Is. (MCZM 298486), ventral view (E), dorsal view (F), photos: Jennifer Trimble MCZM. G–H, shell of syntype of S. elongata Pease, 1860 from the Hawaiian Is. (ANSP 20686), ventral view (G), dorsal view (H), photos: ANSP. I–J, shell of syntype of A. japonica Clessin, 1899 (ZMB Moll 38136), ventral view (I), dorsal view (J), photos: ZMB. K–L, shell of syntype of A. nigrocincta von Martens, 1880 (ZMB Moll 32165), ventral view (K), dorsal view (L), photos: Christine Zorn ZMB. M, dorsal view of shell with tissue of syntype of A.nigrocincta von Martens, 1880 (ZMB Moll 51472), photo: ZMB. N–O, shell of syntype of A. orientalis G. D. Sowerby II, 1869 (NHMUK 1958.1.9.270-1), ventral view (N), dorsal view (O), photos: Harry Taylor NHMUK. P–Q, shell of syntype of A. japonica G. D. Sowerby II, 1869 (NHMUK 1958.1.9.267-9), ventral view (P), dorsal view (Q), photos: Harry Taylor NHMUK." pageId="33" pageNumber="312">Fig. 4K–M</figureCitation>
). These specimens are consistent with the animals here examined and assigned to this species, and there are no major differences between them. Based on other material of
<emphasis id="B954EAA8F579FFCCB025F8FDAD04F8E0" box="[374,529,1844,1865]" italics="true" pageId="34" pageNumber="313">
<taxonomicName id="4C204D39F579FFCCB025F8FDAEF9F8E0" authorityName="Morch" authorityYear="1863" box="[374,492,1844,1865]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="34" pageNumber="313" phylum="Mollusca" rank="species" species="parvula">A. parvula</taxonomicName>
<taxonomicNameLabel id="A26757D3F579FFCCB0A1F8FDAD04F8E0" box="[498,529,1844,1865]" pageId="34" pageNumber="313" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
here examined from the Indo-Pacific, the descriptions associated with the name
<taxonomicName id="4C204D39F579FFCCB23FF8DFAB11F882" authorityName="von Martens" authorityYear="1880" box="[876,1028,1814,1835]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="34" pageNumber="313" phylum="Mollusca" rank="species" species="nigrocincta">
<emphasis id="B954EAA8F579FFCCB23FF8DFAB11F882" box="[876,1028,1814,1835]" italics="true" pageId="34" pageNumber="313">A. nigrocincta</emphasis>
</taxonomicName>
most closely matches the specimens included in this section and, therefore, we are confidently using this name for this species.
</paragraph>
<caption id="DF5F6632F579FFCCB1C2FD78AFF5FCB6" pageId="34" pageNumber="313" startId="34.[145,224,689,711]" targetBox="[145,1425,195,649]" targetPageId="34" targetType="figure">
<paragraph id="8B9F36BAF579FFCCB1C2FD78AFF5FCB6" blockId="34.[145,1425,689,799]" pageId="34" pageNumber="313">
<emphasis id="B954EAA8F579FFCCB1C2FD78AE12FD6F" bold="true" box="[145,263,689,711]" pageId="34" pageNumber="313">Figure 21.</emphasis>
Photographs of the penis of specimens of
<taxonomicName id="4C204D39F579FFCCB397FD78AC67FD6E" authorityName="G. D. Sowerby II" authorityYear="1869" box="[708,882,689,711]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="34" pageNumber="313" phylum="Mollusca" rank="species" species="japonica">
<emphasis id="B954EAA8F579FFCCB397FD78AC67FD6E" box="[708,882,689,711]" italics="true" pageId="34" pageNumber="313">Aplysia japonica</emphasis>
</taxonomicName>
. A, specimen from Gyeongsangbuk-do, South Korea (CPIC 02180, isolate HG134). B, specimen from Gyeongsangbuk-do, South Korea (CPIC 02175, isolate HG129). C, specimen from Gyeongsangbuk-do, South Korea (CPIC 02179, isolate HG133). Abbreviations: pp, penis; pr, prostate; rm, retractor muscle.
</paragraph>
</caption>
<caption id="DF5F6632F579FFCCB1C2F957AE11F946" pageId="34" pageNumber="313" startId="34.[145,223,1694,1716]" targetBox="[147,1426,840,1653]" targetPageId="34" targetType="figure">
<paragraph id="8B9F36BAF579FFCCB1C2F957AE11F946" blockId="34.[145,1425,1694,1775]" pageId="34" pageNumber="313">
<emphasis id="B954EAA8F579FFCCB1C2F957AE10F91A" bold="true" box="[145,261,1694,1716]" pageId="34" pageNumber="313">Figure 22.</emphasis>
Photographs of living animals of
<taxonomicName id="4C204D39F579FFCCB330F957AC06F91D" box="[611,787,1694,1716]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="34" pageNumber="313" phylum="Mollusca" rank="species" species="punctata">
<emphasis id="B954EAA8F579FFCCB330F957AC06F91D" box="[611,787,1694,1716]" italics="true" pageId="34" pageNumber="313">Aplysia punctata</emphasis>
</taxonomicName>
. A, specimen from Antalya, Turkey. B, specimen from Livorno, Italy (CPIC 01379, isolate HG48). C, specimen from Rhodes, Greece. D, specimen from Cape Kamenjak, Croatia (ZSM Mol 20090003).
</paragraph>
</caption>
<caption id="DF5F6632F578FFCDB1F0FA4FAB8BFA5D" pageId="35" pageNumber="314" startId="35.[163,243,1414,1436]" targetBox="[163,1443,195,1374]" targetPageId="35" targetType="figure">
<paragraph id="8B9F36BAF578FFCDB1F0FA4FAB8BFA5D" blockId="35.[163,1443,1414,1524]" pageId="35" pageNumber="314">
<emphasis id="B954EAA8F578FFCDB1F0FA4FAE0EFA32" bold="true" box="[163,283,1414,1436]" pageId="35" pageNumber="314">Figure 23.</emphasis>
Scanning electron micrographs of shells of specimens of
<taxonomicName id="4C204D39F578FFCDB2D7FA4FAB2CFA35" box="[900,1081,1414,1436]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="35" pageNumber="314" phylum="Mollusca" rank="species" species="punctata">
<emphasis id="B954EAA8F578FFCDB2D7FA4FAB2CFA35" box="[900,1081,1414,1436]" italics="true" pageId="35" pageNumber="314">Aplysia punctata</emphasis>
</taxonomicName>
. A, specimen from Gallipoli, Italy (CPIC 01355, isolate HG14). B, specimen from Livorno, Italy (CPIC 01379, isolate HG48). C, specimen from Santander, Spain (CPIC 01424, isolate HG78). D, specimen from Crete, Greece (CPIC 01387, isolate HG87). E, specimen from Crete, Greece (NHMC 52.36, isolate HG33). F, specimen from Crete, Greece (CPIC 01388, isolate HG86).
</paragraph>
</caption>
<paragraph id="8B9F36BAF578FFCDB1E8F9D3AEF8F960" blockId="35.[163,779,1562,1737]" pageId="35" pageNumber="314">
<bibRefCitation id="EFB14B4BF578FFCDB1E8F9D3AE41F999" author="Engel H" box="[187,340,1562,1584]" pageId="35" pageNumber="314" pagination="1 - 76" refId="ref30851" refString="Engel H. 1936. Uber westindische Aplysiidae und Verwandten anderer Gebiete. Capita Zoologica 8: 1-76." type="journal article" year="1936">Engel (1936)</bibRefCitation>
considered
<taxonomicName id="4C204D39F578FFCDB0B6F9D3AD99F999" authorityName="von Martens" authorityYear="1880" box="[485,652,1562,1584]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="35" pageNumber="314" phylum="Mollusca" rank="species" species="nigrocincta">
<emphasis id="B954EAA8F578FFCDB0B6F9D3AD99F999" box="[485,652,1562,1584]" italics="true" pageId="35" pageNumber="314">A. nigrocincta</emphasis>
</taxonomicName>
as a valid species endemic to East Africa, including specimens identified by
<bibRefCitation id="EFB14B4BF578FFCDB01EF991AEE4F9C4" author="Burne RH" box="[333,497,1623,1645]" pageId="35" pageNumber="314" pagination="51 - 58" refId="ref30003" refString="Burne RH. 1906. Notes on the anatomy of South African Aplysiidae, with descriptions of two new species. Journal of Molluscan Studies 7: 51-58." type="journal article" year="1906">Burne (1906)</bibRefCitation>
as
<taxonomicName id="4C204D39F578FFCDB371F991ADC7F9C4" authorityName="von Martens" authorityYear="1880" box="[546,722,1624,1645]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="35" pageNumber="314" phylum="Mollusca" rank="species" species="nigrocincta">
<emphasis id="B954EAA8F578FFCDB371F991ADC7F9C4" box="[546,722,1624,1645]" italics="true" pageId="35" pageNumber="314">A. nigrocincta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C204D39F578FFCDB1F0F9BFAE25F922" baseAuthorityName="Pease" baseAuthorityYear="1860" box="[163,304,1654,1675]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="35" pageNumber="314" phylum="Mollusca" rank="species" species="elongata">
<emphasis id="B954EAA8F578FFCDB1F0F9BFAE25F922" box="[163,304,1654,1675]" italics="true" pageId="35" pageNumber="314">A. elongata</emphasis>
</taxonomicName>
. In this study we found the range of
<taxonomicName id="4C204D39F578FFCDB1F0F95CAE45F903" authorityName="von Martens" authorityYear="1880" box="[163,336,1685,1706]" class="Gastropoda" family="Aplysiidae" genus="Aplysia" kingdom="Animalia" order="Aplysiida" pageId="35" pageNumber="314" phylum="Mollusca" rank="species" species="nigrocincta">
<emphasis id="B954EAA8F578FFCDB1F0F95CAE45F903" box="[163,336,1685,1706]" italics="true" pageId="35" pageNumber="314">A. nigrocincta</emphasis>
</taxonomicName>
to extend from the western Indian Ocean to the western Pacific.
</paragraph>
</subSubSection>
</treatment>
</document>
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