<documentid="33CB8B678AB47A69A85B7D535094583A"ID-CLB-Dataset="38930"ID-DOI="10.11646/zootaxa.4118.1.1"ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619"ID-ISSN="1175-5326"ID-Zenodo-Dep="264597"ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="guilherme"checkinTime="1465242077321"checkinUser="plazi"docAuthor="C, Lees David"docDate="2016"docId="038747324C78C6471EB729E2FC722384"docLanguage="en"docName="zootaxa.4118.pdf.imf"docOrigin="Zootaxa 4118 (1)"docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph"docStyleId="5EBBA59367AD13919D70D935FA04F6A3"docStyleName="Zootaxa.2013-.monograph"docStyleVersion="19"docTitle="Gallienia alaokola"docType="treatment"docVersion="12"lastPageNumber="15"masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174"masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision"masterLastPageNumber="97"masterPageNumber="1"pageNumber="15"updateTime="1735917000942"updateUser="ExternalLinkService">
<mods:titleid="A289CB6842787D8F32B42C0588805FCC">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
<figureCitationid="1315EAA14C78C6591F9E283BFE2D245B"box="[446,510,1302,1327]"captionStart="FIGURE 3"captionStartId="15.[151,250,1650,1672]"captionTargetBox="[303,1292,195,1608]"captionTargetId="figure@15.[275,1307,177,1626]"captionTargetPageId="15"captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)."figureDoi="http://doi.org/10.5281/zenodo.264600"httpUri="https://zenodo.org/record/264600/files/figure.png"pageId="14"pageNumber="15">Fig. 3</figureCitation>
<bibRefCitationid="EFBF8BD54C78C6591FC42817FD402427"author="Oberthur"box="[484,659,1338,1363]"pageId="14"pageNumber="15"refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244."type="journal article"year="1916">Oberthür, 1916</bibRefCitation>
</taxonomicName>
, according to a label under the specimen (but it is certain that Viette never published this). That on the right (#3044) represents the specimen that
<bibRefCitationid="EFBF8BD54C78C6591DD02873FB732403"author="d'Abrera"box="[1008,1184,1374,1399]"pageId="14"pageNumber="15"refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp."type="book"year="1980">d’Abrera (1980</bibRefCitation>
,
<bibRefCitationid="EFBF8BD54C78C6591A8A2872FB302403"author="d'Abrera"box="[1194,1251,1375,1399]"pageId="14"pageNumber="15"refString="d'Abrera, B. L. (1997) Butterflies of the Afrotropical Region. Part 1. Papilionidae, Pieridae, Acraeidae, Danaidae, Satyridae. Hill House Publishers, Melbourne & London, 263 pp."type="book"year="1997">1997</bibRefCitation>
’. I contend, considering dorsal colouration, ocellus expression in space-M3 and relatively more pointed HW tails, that both specimens in fact represent
<bibRefCitationid="EFBF8BD54C78C6591EDC28E7FE582497"author="Butler"box="[252,395,1482,1507]"pageId="14"pageNumber="15"refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820"type="journal article"year="1879">Butler, 1879</bibRefCitation>
<figureCitationid="1315EAA14C78C6591D0028E7FCB72497"box="[800,868,1482,1507]"captionStart="FIGURE 3"captionStartId="15.[151,250,1650,1672]"captionTargetBox="[303,1292,195,1608]"captionTargetId="figure@15.[275,1307,177,1626]"captionTargetPageId="15"captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)."figureDoi="http://doi.org/10.5281/zenodo.264600"httpUri="https://zenodo.org/record/264600/files/figure.png"pageId="14"pageNumber="15">Fig. 3</figureCitation>
<bibRefCitationid="EFBF8BD54C78C6591A5828E7FAF82497"author="d'Abrera"box="[1144,1323,1482,1507]"pageId="14"pageNumber="15"refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp."type="book"year="1980">d’Abrera (1980</bibRefCitation>
: 190) had placed a red label next to photograph of “
<bibRefCitationid="EFBF8BD54C78C6591ED92B3FFE66275F"box="[249,437,1554,1579]"pageId="14"pageNumber="15"refString="d'Abrera, B. L. (1997) Butterflies of the Afrotropical Region. Part 1. Papilionidae, Pieridae, Acraeidae, Danaidae, Satyridae. Hill House Publishers, Melbourne & London, 263 pp."type="book">d’Abrera (1997)</bibRefCitation>
had removed the red label. Without DNA data, choice of a ♀
would be extremely unsatisfactory in this difficult-to-distinguish species group, and thus unlikely to promote stability. It appears that d’Abrera’s (1980) action does not, however, represent a valid lectotypification, since it is not an unambiguous selection of the name-bearing
<figureCitationid="1315EAA14C78C6591D682BC7FC592677"box="[840,906,1770,1795]"captionStart="FIGURE 3"captionStartId="15.[151,250,1650,1672]"captionTargetBox="[303,1292,195,1608]"captionTargetId="figure@15.[275,1307,177,1626]"captionTargetPageId="15"captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)."figureDoi="http://doi.org/10.5281/zenodo.264600"httpUri="https://zenodo.org/record/264600/files/figure.png"pageId="14"pageNumber="15">Fig. 3</figureCitation>
C) and a much darker upperside. They lack the ocellus expression in both space-M3 and CuA1 of the ♂ HW upperside, with sometimes neither ocellus expressed (this variation in his mixed series is alluded to in Oberthür’s 1916: 205–206 description). Among Oberthür’s 75
<figureCitationid="1315EAA14C78C6591FC22A50FDF526E2"box="[482,550,1917,1942]"captionStart="FIGURE 3"captionStartId="15.[151,250,1650,1672]"captionTargetBox="[303,1292,195,1608]"captionTargetId="figure@15.[275,1307,177,1626]"captionTargetPageId="15"captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)."figureDoi="http://doi.org/10.5281/zenodo.264600"httpUri="https://zenodo.org/record/264600/files/figure.png"pageId="14"pageNumber="15">Fig. 3</figureCitation>
<bibRefCitationid="EFBF8BD54C78C6591CE32AE8FCA726AA"author="d'Abrera"box="[707,884,1989,2014]"pageId="14"pageNumber="15"refString="d'Abrera, B. L. (1997) Butterflies of the Afrotropical Region. Part 1. Papilionidae, Pieridae, Acraeidae, Danaidae, Satyridae. Hill House Publishers, Melbourne & London, 263 pp."type="book"year="1997">d’Abrera (1997</bibRefCitation>
: 229) in DV/VV, following preference for a previously illustrated specimen (Recommendation 74B of the code), although in this case, not the one that the original author intended. This action is also in the spirit of the Code as it promotes stability and obviates the need to describe a new species within Oberthür’s syntypic series. The majority of the series of
<bibRefCitationid="EFBF8BD54C79C6581A8A2ABDFAB926DD"author="Oberthur"box="[1194,1386,1936,1961]"pageId="15"pageNumber="16"refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244."type="journal article"year="1916">Oberthür (1916)</bibRefCitation>
, and are then automatically among the PLTs of Oberthür’s nominal species. Although not illustrated by
<bibRefCitationid="EFBF8BD54C79C6581C872AF5FCB72685"author="Oberthur"box="[679,868,2008,2033]"pageId="15"pageNumber="16"refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244."type="journal article"year="1916">Oberthür (1916)</bibRefCitation>
, the chosen LT represents the more widespread/ common and generally darker-upperside species (formerly known as sp. 56;
<bibRefCitationid="EFBF8BD54C66C6471DD82DB5FBA221C4"author="Lees"box="[1016,1137,152,176]"pageId="16"pageNumber="17"refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp."type="book"year="1997">Lees 1997</bibRefCitation>
: 65), which tends strongly either to lack space-M3 ocellus expression, or to lack expression in both space-M3 and CuA1 on HWD. This species also lacks a ventral abdominal black androconial patch (usually bilateral of midline, sometimes obscured by overlying ochreous scales) that is characteristic of
<bibRefCitationid="EFBF8BD54C66C6471DE32C28FB902068"author="Lees"box="[963,1091,260,284]"pageId="16"pageNumber="17"refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp."type="book"year="1997">Lees, 1997</bibRefCitation>
<bibRefCitationid="EFBF8BD54C66C6471FF22C05FD862034"author="Lees"box="[466,597,296,320]"pageId="16"pageNumber="17"refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp."type="book"year="1997">Lees, 1997</bibRefCitation>
: 105) also has multiply toothed long valves, which do not have such a projected sharply inbent tip, as does the genitalia of Oberthür’s ♂ ‘iconotype’ of
<figureCitationid="1315EAA14C66C6471B4E2C61FF7620FC"captionStart="FIGURE 3"captionStartId="15.[151,250,1650,1672]"captionTargetBox="[303,1292,195,1608]"captionTargetId="figure@15.[275,1307,177,1626]"captionTargetPageId="15"captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)."figureDoi="http://doi.org/10.5281/zenodo.264600"httpUri="https://zenodo.org/record/264600/files/figure.png"pageId="16"pageNumber="17">Fig. 3</figureCitation>
, although the abdomen is worn and may even not be the original., The strong expression of the space-M3 ocellus in ♂♂ (in contrast to its strong reduction or absence) confirms the identity of specimens currently referred to
, as here fixed, also has a darker dorsal colouration). Note the androconial scales on abdominal sternum third segment and the inwards recurved valve tips characteristic of
<figureCitationid="1315EAA14C66C6471B2A2CD2FA9E236C"box="[1290,1357,511,536]"captionStart="FIGURE 3"captionStartId="15.[151,250,1650,1672]"captionTargetBox="[303,1292,195,1608]"captionTargetId="figure@15.[275,1307,177,1626]"captionTargetPageId="15"captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)."figureDoi="http://doi.org/10.5281/zenodo.264600"httpUri="https://zenodo.org/record/264600/files/figure.png"pageId="16"pageNumber="17">Fig. 3</figureCitation>
<figureCitationid="1315EAA14C66C6471EF42F09FEC62348"box="[212,277,548,573]"captionStart="FIGURE 3"captionStartId="15.[151,250,1650,1672]"captionTargetBox="[303,1292,195,1608]"captionTargetId="figure@15.[275,1307,177,1626]"captionTargetPageId="15"captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)."figureDoi="http://doi.org/10.5281/zenodo.264600"httpUri="https://zenodo.org/record/264600/files/figure.png"pageId="16"pageNumber="17">Fig. 3</figureCitation>
has relatively long tails especially at M3, a lighter brown dorsal colouration, and three or four white stripes on the FWV tending to be more strongly represented as marks on the FWD costa, than the species/specimen here selected to represent
<figureCitationid="1315EAA14C66C6471D742F41FC4423F0"box="[852,919,620,645]"captionStart="FIGURE 3"captionStartId="15.[151,250,1650,1672]"captionTargetBox="[303,1292,195,1608]"captionTargetId="figure@15.[275,1307,177,1626]"captionTargetPageId="15"captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)."figureDoi="http://doi.org/10.5281/zenodo.264600"httpUri="https://zenodo.org/record/264600/files/figure.png"pageId="16"pageNumber="17">Fig. 3</figureCitation>
