157 lines
18 KiB
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157 lines
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<document id="42289483EECE51F47604D45E9D637390" ID-DOI="10.5281/zenodo.178251" ID-GBIF-Dataset="905dea16-dd69-45f9-9fe2-5b1aa6af93b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="178251" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459886785444" checkinUser="plazi" docAuthor="Hippa, Heikki & Vilkamaa, Pekka" docDate="2007" docId="03DB45439C78FFC443DDFF77FB61FBC5" docLanguage="en" docName="zt01556p050.pdf" docOrigin="Zootaxa 1556" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Keilbachia bicuspis Hippa & Vilkamaa, 2007, sp. n." docType="treatment" docVersion="9" lastPageNumber="39" masterDocId="FFE23D3B9C70FFCC434AFFE0FFC7FFA9" masterDocTitle="The flagria group of Keilbachia Mohrig (Diptera, Sciaridae) in a biodiversity hot spot: nine new sympatric species from Kambaiti, Myanmar" masterLastPageNumber="50" masterPageNumber="31" pageNumber="39" updateTime="1698227854892" updateUser="plazi">
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<mods:title id="2CC290AECBD972138F762641BAEF4885">The flagria group of Keilbachia Mohrig (Diptera, Sciaridae) in a biodiversity hot spot: nine new sympatric species from Kambaiti, Myanmar</mods:title>
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<mods:name id="BA7CD820D698D69C06630FAE8ED8BE51" type="personal">
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<mods:namePart id="40960F0D8A103E3AEA59915FD2EF0E6C">Hippa, Heikki</mods:namePart>
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<mods:name id="88CB5EBF70E091ECD522CE31A7E173C9" type="personal">
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</mods:role>
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<mods:namePart id="5F06B1D253A1FBB2E140C8DB739E4D8C">Vilkamaa, Pekka</mods:namePart>
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<mods:typeOfResource id="E18D8D9D7972C54BB409B5B4CD399976">text</mods:typeOfResource>
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<mods:title id="C846E866B2E9D7FC607B5CB64EE82CB9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6AA49EF4EA316ADA887AF5C86AC8BC8E">
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<mods:date id="1E2A175BE8AD625651C9826E0AFD4D34">2007</mods:date>
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<mods:detail id="1C16A3E48F42308BF022C3721ECA5042" type="volume">
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<mods:number id="FBB005012548BCE4A5E68C5E89C395A6">1556</mods:number>
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<mods:classification id="6720C2028B0C33234BE58F1F00B85FAC">journal article</mods:classification>
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<mods:identifier id="B712B757E45A910271DC05C7C7BA0F85" type="DOI">10.5281/zenodo.178251</mods:identifier>
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<mods:identifier id="BA12653547D7DA95006DF0306C1838BC" type="GBIF-Dataset">905dea16-dd69-45f9-9fe2-5b1aa6af93b6</mods:identifier>
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<mods:identifier id="DDD019DDF9FA49EA9DBE4B8D309CBA52" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="CB4ED83ED45B3E04595DFD9C9973038A" type="Zenodo-Dep">178251</mods:identifier>
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<treatment id="03DB45439C78FFC443DDFF77FB61FBC5" ID-DOI="http://doi.org/10.5281/zenodo.6241292" ID-GBIF-Taxon="119371648" ID-Zenodo-Dep="6241292" LSID="urn:lsid:plazi:treatment:03DB45439C78FFC443DDFF77FB61FBC5" httpUri="http://treatment.plazi.org/id/03DB45439C78FFC443DDFF77FB61FBC5" lastPageNumber="39" pageId="8" pageNumber="39">
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<subSubSection id="C368A7DE9C78FFC443DDFF77FE7FFF7D" pageId="8" pageNumber="39" type="nomenclature">
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<paragraph id="8BCDF4559C78FFC443DDFF77FE0CFF18" blockId="8.[151,459,151,212]" box="[151,459,151,177]" pageId="8" pageNumber="39">
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<heading id="D08543399C78FFC443DDFF77FE0CFF18" bold="true" box="[151,459,151,177]" fontSize="11" level="1" pageId="8" pageNumber="39" reason="1">
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<emphasis id="B90628479C78FFC443DDFF77FE0CFF18" bold="true" box="[151,459,151,177]" pageId="8" pageNumber="39">
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<taxonomicName id="4C728FD69C78FFC443DDFF77FE45FF18" ID-CoL="3R4L5" box="[151,386,151,177]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="bicuspis" status="sp. nov.">
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<emphasis id="B90628479C78FFC443DDFF77FE45FF18" bold="true" box="[151,386,151,177]" italics="true" pageId="8" pageNumber="39">Keilbachia bicuspis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A235953C9C78FFC442C3FF77FE0CFF18" box="[393,459,151,177]" pageId="8" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCDF4559C78FFC443DDFF5AFE7FFF7D" blockId="8.[151,459,151,212]" box="[151,440,186,212]" pageId="8" pageNumber="39">
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(
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<figureCitation id="1349E8D09C78FFC443D5FF5AFF22FF7D" box="[159,229,186,212]" captionStart="FIGURE 1" captionStartId="3.[151,255,1939,1963]" captionTargetBox="[232,1367,193,1884]" captionTargetId="figure@3.[198,1385,154,1913]" captionTargetPageId="3" captionText="FIGURE 1. Maxillary palpus. A, Keilbachia acamptochaeta sp. n. (paratype). B, K. flabellata sp. n. (holotype). C, K. gigas sp. n. (holotype). D, K. fasciata sp. n. (holotype). E, K. foveolata sp. n. (holotype). F, K. bicuspis sp. n. (holotype). G, K. rima sp. n. (holotype). H, K. filigera sp. n. (holotype). I, K. oligonema sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178252/files/figure.png" pageId="8" pageNumber="39">Figs 1</figureCitation>
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F, 2A, 3H, 5D, E)
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</paragraph>
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</subSubSection>
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<subSubSection id="C368A7DE9C78FFC443DDFEEAFE5EFCFD" pageId="8" pageNumber="39" type="materials_examined">
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<paragraph id="8BCDF4559C78FFC443DDFEEAFF3BFEE5" blockId="8.[151,1436,266,1132]" pageId="8" pageNumber="39">
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<emphasis id="B90628479C78FFC443DDFEEAFEB6FE8D" bold="true" box="[151,369,266,292]" pageId="8" pageNumber="39">Material studied.</emphasis>
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<emphasis id="B90628479C78FFC44237FEEAFDEAFE8A" box="[381,557,266,291]" italics="true" pageId="8" pageNumber="39">
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<typeStatus id="54C94AF79C78FFC44237FEEAFE2FFE8A" box="[381,488,266,291]" pageId="8" pageNumber="39" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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(in
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<collectionCode id="ED636C909C78FFC44129FEEAFD7CFE8D" box="[611,699,266,292]" pageId="8" pageNumber="39">NRMS</collectionCode>
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). N.E.
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<collectingCountry id="F365B4C59C78FFC4405EFEEAFCA3FE8D" box="[788,868,266,292]" name="Myanmar" pageId="8" pageNumber="39">Burma</collectingCountry>
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(
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<collectingCountry id="F365B4C59C78FFC4403DFEEAFC28FE8D" box="[887,1007,266,292]" name="Myanmar" pageId="8" pageNumber="39">Myanmar</collectingCountry>
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), Kambaiti,
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<quantity id="4C8A59B09C78FFC447CDFEEAFB22FE8D" box="[1159,1253,266,292]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="8" pageNumber="39" unit="ft" value="7000.0">7000 ft</quantity>
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.,
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<date id="FFCCD2959C78FFC447B3FEEAFAB6FE8D" box="[1273,1393,266,292]" pageId="8" pageNumber="39" value="1934-04-15">15.4.1934</date>
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, R. Malaise.
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</paragraph>
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<paragraph id="8BCDF4559C78FFC4438CFEBAFAB4FC85" blockId="8.[151,1436,266,1132]" pageId="8" pageNumber="39">
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Male.
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<emphasis id="B90628479C78FFC4425FFEBAFEACFEDD" bold="true" box="[277,363,346,372]" pageId="8" pageNumber="39">Colour</emphasis>
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. The specimen is strongly faded, almost unicolorous pale yellowish-brown.
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<emphasis id="B90628479C78FFC44652FEBAFA9CFEDD" bold="true" box="[1304,1371,346,372]" pageId="8" pageNumber="39">Head</emphasis>
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. Eye bridge 3 facets wide. Face (prefrons) with 5 setae in a transverse row. Clypeus with 1 seta. Maxillary palpus (
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<figureCitation id="1349E8D09C78FFC443D5FE4AFF25FE6D" box="[159,226,426,452]" captionStart="FIGURE 1" captionStartId="3.[151,255,1939,1963]" captionTargetBox="[232,1367,193,1884]" captionTargetId="figure@3.[198,1385,154,1913]" captionTargetPageId="3" captionText="FIGURE 1. Maxillary palpus. A, Keilbachia acamptochaeta sp. n. (paratype). B, K. flabellata sp. n. (holotype). C, K. gigas sp. n. (holotype). D, K. fasciata sp. n. (holotype). E, K. foveolata sp. n. (holotype). F, K. bicuspis sp. n. (holotype). G, K. rima sp. n. (holotype). H, K. filigera sp. n. (holotype). I, K. oligonema sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178252/files/figure.png" pageId="8" pageNumber="39">Fig. 1</figureCitation>
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F) with three palpomeres. Palpomere 1 with one seta, with numerous hyaline sensilla widely scattered on dorsal surface, no sensory pit; palpomere 2 with ca. 7 setae; palpomere 3 with ca. 7 setae. Antennal flagellomere 4,
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<figureCitation id="1349E8D09C78FFC44211FE1AFE62FDBD" box="[347,421,506,532]" captionStart="FIGURE 2" captionStartId="4.[151,255,1425,1449]" captionTargetBox="[152,1424,202,1424]" captionTargetId="figure@4.[151,1436,194,1467]" captionTargetPageId="4" captionText="FIGURE 2. Antennal flagellomeres 3 and 4 (A) and flagellomere 4 (B – E). A, Keilbachia bicuspis sp. n. (holotype). B, K. oligonema sp. n. (holotype). C, K. fasciata sp. n. (holotype). D, K. filigera sp. n. (holotype). E, K. rima sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178253/files/figure.png" pageId="8" pageNumber="39">Fig. 2</figureCitation>
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A.
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<emphasis id="B90628479C78FFC44284FE1AFDECFDBD" bold="true" box="[462,555,506,532]" pageId="8" pageNumber="39">Thorax</emphasis>
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. Number of setae on anterior pronotum and episternum 1 not counted because the sclerites are badly damaged in the single specimen.
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<emphasis id="B90628479C78FFC440D5FDC2FC25FD95" bold="true" box="[927,994,546,572]" pageId="8" pageNumber="39">Wing</emphasis>
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. Length
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<quantity id="4C8A59B09C78FFC4471AFDC2FB77FD95" box="[1104,1200,546,572]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="39" unit="mm" value="2.7">2.7 mm</quantity>
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. Width/length 0.42. R1/R 1.21. c/
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<geoCoordinate id="EE4692929C78FFC44279FDAAFE41FDCD" box="[307,390,586,612]" direction="west" orientation="longitude" pageId="8" pageNumber="39" precision="555" value="-0.79">w 0.79</geoCoordinate>
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. r-m and bM equal in length, r-m/bM 1.00, both non-setose.
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<emphasis id="B90628479C78FFC4470CFDAAFB46FDCD" bold="true" box="[1094,1153,586,612]" pageId="8" pageNumber="39">Legs</emphasis>
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. Apex of fore tibia,
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<figureCitation id="1349E8D09C78FFC44627FDAAFF62FD25" captionStart="FIGURE 3" captionStartId="5.[151,255,1905,1929]" captionTargetBox="[188,1393,195,1864]" captionTargetId="figure@5.[163,1428,158,1890]" captionTargetPageId="5" captionText="FIGURE 3. Apical part of fore tibia, prolateral view. A, Keilbachia oligonema sp. n. (Burma). B, K. fasciata sp. n. (paratype). C, K. rima sp. n. (holotype). D, K. acamptochaeta sp. n. (paratype). E, K. flabellata sp. n. (holotype). F, K. foveolata sp. n. (holotype). G, K. filigera sp. n. (holotype). H, K. bicuspis sp. n. (holotype). I, K. gigas sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178254/files/figure.png" pageId="8" pageNumber="39">Fig. 3</figureCitation>
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H. Length of fore basitarsomere/length of fore tibia 0.79. Length of fore tibia/length of fore femur 1.48. Length of hind tibia/length of scutum+scutellum not counted because the thorax is distorted.
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<emphasis id="B90628479C78FFC44791FD7AFA92FD1D" bold="true" box="[1243,1365,666,692]" pageId="8" pageNumber="39">Abdomen</emphasis>
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. Sternite 8 with ca. 20 setae, widely spread.
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<emphasis id="B90628479C78FFC44139FD22FCDBFD75" bold="true" box="[627,796,706,732]" pageId="8" pageNumber="39">Hypopygium.</emphasis>
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<figureCitation id="1349E8D09C78FFC4406CFD22FCABFD75" box="[806,876,706,732]" captionStart="FIGURE 5" captionStartId="7.[151,255,1812,1836]" captionTargetBox="[175,1403,213,1759]" captionTargetId="figure@7.[151,1436,194,1788]" captionTargetPageId="7" captionText="FIGURE 5. Eye bridge, frontal view (A), and part of hypopygium (B, D) and gonostylus (C, E), ventral view. A, B, C, Keilbachia acamptochaeta sp. n. (holotype). D, E, K. bicuspis sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178256/files/figure.png" pageId="8" pageNumber="39">Fig. 5</figureCitation>
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D, E: Gonostylus with two rather short arcuate mesial megasetae arising from a common basal body and one megaseta at the base of the apical tooth. Tegmen laterally compressed so that the sides partly cover each other, and many of the details difficult to interpret.
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</paragraph>
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<paragraph id="8BCDF4559C78FFC4438CFCDAFE5EFCFD" blockId="8.[151,1436,266,1132]" box="[198,409,826,852]" pageId="8" pageNumber="39">Female unknown.</paragraph>
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</subSubSection>
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<subSubSection id="C368A7DE9C78FFC4438CFC82FB61FBC5" pageId="8" pageNumber="39" type="discussion">
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<paragraph id="8BCDF4559C78FFC4438CFC82FB61FBC5" blockId="8.[151,1436,266,1132]" pageId="8" pageNumber="39">
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<emphasis id="B90628479C78FFC4438CFC82FE95FCD5" bold="true" box="[198,338,866,892]" pageId="8" pageNumber="39">Discussion.</emphasis>
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<taxonomicName id="4C728FD69C78FFC44213FC82FD87FCD2" box="[345,576,866,891]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="bicuspis">
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<emphasis id="B90628479C78FFC44213FC82FD87FCD2" box="[345,576,866,891]" italics="true" pageId="8" pageNumber="39">Keilbachia bicuspis</emphasis>
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</taxonomicName>
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is not especially similar to any other
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<taxonomicName id="4C728FD69C78FFC440BDFC82FBB0FCD2" box="[1015,1143,866,891]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="genus">
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<emphasis id="B90628479C78FFC440BDFC82FBB0FCD2" box="[1015,1143,866,891]" italics="true" pageId="8" pageNumber="39">Keilbachia</emphasis>
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</taxonomicName>
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. It is one of the five species with only two curved megasetae on the gonostylus, the other four being
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<emphasis id="B90628479C78FFC44701FC6AFF3EFC62" italics="true" pageId="8" pageNumber="39">
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<taxonomicName id="4C728FD69C78FFC44701FC6AFAE0FC0A" box="[1099,1319,906,931]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="acamptochaeta">K. acamptochaeta</taxonomicName>
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,
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<taxonomicName id="4C728FD69C78FFC4467CFC6AFF3EFC62" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="biflagrispina">K. biflagrispina</taxonomicName>
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</emphasis>
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,
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<taxonomicName id="4C728FD69C78FFC44240FC52FE6BFC62" box="[266,428,946,971]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="oligonema">
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<emphasis id="B90628479C78FFC44240FC52FE6BFC62" box="[266,428,946,971]" italics="true" pageId="8" pageNumber="39">K. oligonema</emphasis>
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</taxonomicName>
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and
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<emphasis id="B90628479C78FFC442A1FC52FDC1FC62" box="[491,518,946,971]" italics="true" pageId="8" pageNumber="39">K.</emphasis>
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sp. 2.
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<taxonomicName id="4C728FD69C78FFC44116FC52FD25FC62" box="[604,738,946,971]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="bicuspis">
|
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<emphasis id="B90628479C78FFC44116FC52FD25FC62" box="[604,738,946,971]" italics="true" pageId="8" pageNumber="39">K. bicuspis</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
differs from all of these except for
|
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|
|
<taxonomicName id="4C728FD69C78FFC447D6FC52FAB3FC62" box="[1180,1396,946,971]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="acamptochaeta">
|
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|
|
<emphasis id="B90628479C78FFC447D6FC52FAB3FC62" box="[1180,1396,946,971]" italics="true" pageId="8" pageNumber="39">K. acamptochaeta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having the curved megasetae short and stout, only about twice the length of their basal body instead of being several times longer than the basal body.
|
|||
|
|
<taxonomicName id="4C728FD69C78FFC441D3FBE2FCD8FBB2" box="[665,799,1026,1051]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="bicuspis">
|
|||
|
|
<emphasis id="B90628479C78FFC441D3FBE2FCD8FBB2" box="[665,799,1026,1051]" italics="true" pageId="8" pageNumber="39">K. bicuspis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="4C728FD69C78FFC44082FBE2FB65FBB2" box="[968,1186,1026,1051]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="acamptochaeta">
|
|||
|
|
<emphasis id="B90628479C78FFC44082FBE2FB65FBB2" box="[968,1186,1026,1051]" italics="true" pageId="8" pageNumber="39">K. acamptochaeta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having only one megaseta in addition to the curved ones on its gonostylus, whilst in
|
|||
|
|
<taxonomicName id="4C728FD69C78FFC44096FBCAFB72FBEA" box="[988,1205,1066,1091]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="acamptochaeta">
|
|||
|
|
<emphasis id="B90628479C78FFC44096FBCAFB72FBEA" box="[988,1205,1066,1091]" italics="true" pageId="8" pageNumber="39">K. acamptochaeta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
there are three.
|
|||
|
|
<taxonomicName id="4C728FD69C78FFC446CBFBCAFF30FBC5" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="bicuspis">
|
|||
|
|
<emphasis id="B90628479C78FFC446CBFBCAFF30FBC5" italics="true" pageId="8" pageNumber="39">K. bicuspis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
differs further by having maxillary palpus three-segmented, not one-segmented.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|