312 lines
38 KiB
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312 lines
38 KiB
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<document ID-CLB-Dataset="280021" ID-DOI="http://dx.doi.org/10.3897/zookeys.1185.104120" ID-GBIF-Dataset="49dd3650-1e38-4eed-b72c-f38067e15783" ID-Pensoft-Pub="1313-2970-1185-277" ID-Pensoft-UUID="337CDF6D1F90587495D1E8C94581B3B5" ID-ZooBank="912F2D3B2DD24771A895FDD7B1443511" ModsDocID="1313-2970-1185-277" checkinTime="1701378453538" checkinUser="pensoft" docAuthor="Zhang, Jianshuang, Yu, Hao & Lin, Yucheng" docDate="2023" docId="00C66505BCF85CFB825CC345C05BB383" docLanguage="en" docName="ZooKeys 1185: 277-294" docOrigin="ZooKeys 1185" docPubDate="2023-11-30" docSource="http://dx.doi.org/10.3897/zookeys.1185.104120" docTitle="Simonia youyiensis Zhang & Yu & Lin 2023, comb. nov." docType="treatment" docVersion="3" id="337CDF6D1F90587495D1E8C94581B3B5" lastPageNumber="277" masterDocId="337CDF6D1F90587495D1E8C94581B3B5" masterDocTitle="Simonia gen. nov., a new spider genus (Araneae, Theridiosomatidae) from Southeast Asia" masterLastPageNumber="294" masterPageNumber="277" pageNumber="277" updateTime="1701396588983" updateUser="ExternalLinkService">
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<mods:titleInfo id="13C19812F61A47ED9E72D4979D75FD91">
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<mods:title id="4F63B32A69A307687861B237F47043C1">Simonia gen. nov., a new spider genus (Araneae, Theridiosomatidae) from Southeast Asia</mods:title>
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<mods:name id="78E9D5D9C57AF46E36FD8D61F3BA0E45" type="personal">
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<mods:namePart id="20C2B817F15C2A6F09589F69F5AD39C9">Zhang, Jianshuang</mods:namePart>
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<mods:nameIdentifier id="CDD2F698157BB634793B01BCE4DB01D6" type="ORCID">https://orcid.org/0000-0003-4010-3082</mods:nameIdentifier>
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<mods:affiliation id="178B50B20AC020D3FE4C0D9D9E227D3D">School of Life Sciences, Guizhou Normal University, Guiyang, Guizhou 550025, China</mods:affiliation>
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</mods:name>
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<mods:name id="AF85D3CC38729B4E2173F56A6CBEE820" type="personal">
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<mods:roleTerm id="5A791D5326919A9B195B885D81137AAD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7E0B093AD56EF94464CF0C21B0BD5EC9">Yu, Hao</mods:namePart>
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<mods:nameIdentifier id="932E14D7AC992DD581792EE2EC96289F" type="ORCID">https://orcid.org/0000-0002-9113-2425</mods:nameIdentifier>
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<mods:affiliation id="852DB5A4BEC6D2712E6F33B1B3087A0A">School of Life Sciences, Guizhou Normal University, Guiyang, Guizhou 550025, China</mods:affiliation>
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</mods:name>
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<mods:name id="163A4A2E4A0AF5ED341796E7D715032C" type="personal">
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</mods:role>
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<mods:namePart id="8A043FD5EBD957B9B9ABDD2FA15643FE">Lin, Yucheng</mods:namePart>
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<mods:nameIdentifier id="1FEDDBE7606D848C6A6BD2AC3BCEE58F" type="ORCID">https://orcid.org/0000-0002-5054-0633</mods:nameIdentifier>
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<mods:affiliation id="30903543600DFD78F715EFFABC7F4505">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
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<mods:nameIdentifier id="2C17EA5CD0339E7FB10A5D6E3E0D8A7B" type="email">linyucheng@scu.edu.cn</mods:nameIdentifier>
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<mods:typeOfResource id="02748144A052342A75B2CB63E1FA3C74">text</mods:typeOfResource>
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<mods:title id="EC887CAA9A25BE1ED99E02FEE438AE30">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="EFD9E1F664AFE708339E8523636375E7">
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<mods:date id="1A61C3F334791E16DA383B389D373257">2023</mods:date>
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<mods:number id="DA5AD36FEDF3A9A9A9904FE96E91CC02">2023-11-30</mods:number>
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<mods:number id="1463D94B5746084FF12E9712C7EBE523">1185</mods:number>
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<mods:start id="0A29BE6748119C164F68C11FFCE912B0">277</mods:start>
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<mods:end id="DB0C2CE49905D71BBF21FD34EE761EE3">294</mods:end>
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<mods:url id="6A8BC82DAAEC2788FC0EAE77C5B6C14B">http://dx.doi.org/10.3897/zookeys.1185.104120</mods:url>
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</mods:location>
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<mods:classification id="DA7A9C073E891656B66FF8242C332807">journal article</mods:classification>
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<mods:identifier id="CEA7AD8C2AD0A0C4F4D1B8726081F16B" type="DOI">http://dx.doi.org/10.3897/zookeys.1185.104120</mods:identifier>
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<mods:identifier id="E8FFBCEA15A2B21F29A8321E24BEF93D" type="Pensoft-Pub">1313-2970-1185-277</mods:identifier>
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<mods:identifier id="FC08FC2DA53DF94BA4F546E9546E7B1F" type="ZooBank">912F2D3B2DD24771A895FDD7B1443511</mods:identifier>
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<mods:identifier id="ACD552271D5B81147EE45464D704EC3E" type="Pensoft-UUID">337CDF6D1F90587495D1E8C94581B3B5</mods:identifier>
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</mods:mods>
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<treatment id="00C66505BCF85CFB825CC345C05BB383" ID-GBIF-Taxon="219492150" LSID="urn:lsid:plazi:treatment:00C66505BCF85CFB825CC345C05BB383" httpUri="http://treatment.plazi.org/id/00C66505BCF85CFB825CC345C05BB383" lastPageNumber="277" pageId="0" pageNumber="277">
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<subSubSection id="BAE313A8E9B7B955624DBF19094600AF" pageId="0" pageNumber="277" type="nomenclature">
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<paragraph id="7398B45931FCE759F3FBC0F5D43B290F" pageId="0" pageNumber="277">
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<taxonomicName id="8E0295E7F01F531D705C967C8E84E2B0" LSID="00C66505-BCF8-5CFB-825C-C345C05BB383" authority="(Zhao & Li, 2012)" authorityName="Zhang & Yu & Lin" authorityYear="2023" baseAuthorityName="Zhao & Li" baseAuthorityYear="2012" family="Theridiosomatidae" genus="Simonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Simonia youyiensis" order="Araneae" pageId="0" pageNumber="277" phylum="Mollusca" rank="species" species="youyiensis" status="comb. nov.">Simonia youyiensis (Zhao & Li, 2012)</taxonomicName>
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<taxonomicNameLabel id="6C7E36894E7F7A265F10A832F2025EF6" pageId="0" pageNumber="277">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="2748BEA14F258C21F2CC9D70583B1BAA" pageId="0" pageNumber="277" type="description">
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<paragraph id="FDA413B6052F17527E58BFE89184AD2A" pageId="0" pageNumber="277">
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<figureCitation id="4296D41738D105D73753E46AD6B5D225" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male palp of Simonia youyiensis A prolateral B retrolateral C dorsal D ventral. Abbreviations: Co = conductor; CL = cymbial lobe; Cy = cymbium; ED = embolic division; MA = median apophysis; Pc = paracymbium; SCy = cymbial setae; St = subtegulum; Te = tegulum. Scale bars: 0.2 mm (A-D)." figureDoi="10.3897/zookeys.1185.104120.figure1" httpUri="https://binary.pensoft.net/fig/945107" pageId="0" pageNumber="277">Figs 1</figureCitation>
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<figureCitation id="60A4CC7075B2C633804BDDAC0BA19093" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bulb (A) and male palp (B-D) of Simonia youyiensis, cited from Zhang and Lin (2016) A apical B ventral C prolateral D retrolateral. Abbreviations: Co = conductor; CL = cymbial lobe; Cy = cymbium; MA = median apophysis; Pc = paracymbium; SCy = cymbial setae; SD = sperm duct; St = subtegulum; Te = tegulum. Scale bars: 0.2 mm (A-D)." figureDoi="10.3897/zookeys.1185.104120.figure2" httpUri="https://binary.pensoft.net/fig/945108" pageId="0" pageNumber="277">, 2</figureCitation>
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<figureCitation id="C287040A52B4C196D15BF000DFD2DC32" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palpal sclerites of Simonia youyiensis, cited from Zhang and Lin (2016) A bulb, prolateral B embolic division, ventral C conductor, ventral D cymbium, retrolateral. Abbreviations: Co = conductor; CL = cymbial lobe; EA = embolic apophysis; ED = embolic division; EL = embolic lobe; ET = embolic terminal; MA = median apophysis; Pc = paracymbium; SCy = cymbial setae; St = subtegulum; Te = tegulum. Scale bars: 0.1 mm (A-D)." figureDoi="10.3897/zookeys.1185.104120.figure3" httpUri="https://binary.pensoft.net/fig/945109" pageId="0" pageNumber="277">, 3</figureCitation>
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<figureCitation id="61B47CBC8CF212BBAB0CB434E22C1F34" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Simonia youyiensis, male habitus (A, B), female habitus (C, D) and epigyne (E-G) A dorsal B ventral C dorsal D ventral E intact, ventral F cleared, ventral (blue dashed line showing the anterior margin of the epigynal plate) G cleared, dorsal (green dashed line showing schematic course of copulatory duct). Abbreviations: CB = copulatory bursa; CD = copulatory duct; FD = fertilization duct; Sc = scape; SH = spermathecal head; SS = spermathecal stalk; Sp = spermatheca; TF = transverse fold of epigynal plate. Scale bars: 0.5 mm (A-D); 0.2 mm (E-G)." figureDoi="10.3897/zookeys.1185.104120.figure4" httpUri="https://binary.pensoft.net/fig/945110" pageId="0" pageNumber="277">, 4</figureCitation>
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<figureCitation id="ED20426CF51C2ACD475B36DE7C8E8D38" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution records of Simonia gen. nov. species." figureDoi="10.3897/zookeys.1185.104120.figure7" httpUri="https://binary.pensoft.net/fig/945113" pageId="0" pageNumber="277">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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|||
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<subSubSection id="C33F0992BF75CFBC6271D8C620F2E2DC" pageId="0" pageNumber="277" type="reference_group">
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<paragraph id="1A824ECA8942BCFA08301D9EE3D59E66" pageId="0" pageNumber="277">
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<taxonomicName id="6A1339E396CF4820838E282FD71C707B" authorityName="Zhang & Yu & Lin" authorityYear="2023" baseAuthorityName="Zhang & Yu & Lin" baseAuthorityYear="2023" class="Arachnida" family="Theridiosomatidae" genus="Baalzebub" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baalzebub youyiensis" order="Araneae" pageId="0" pageNumber="277" phylum="Arthropoda" rank="species" species="youyiensis" status="comb. nov.">Baalzebub youyiensis</taxonomicName>
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Zhao & Li, 2012: 17, figs 9A-E, 10A, B (♀);
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<bibRefCitation id="21FC8F71B7D1CDE254D742C0E07B3AB2" DOI="https://doi.org/10.11984/j.issn.1000-7083.20150367" author="Zhang, T" journalOrPublisher="Sichuan Journal of Zoology" pageId="0" pageNumber="277" pagination="221 - 226" refId="B14" refString="Zhang, T, Lin, Y, 2016. First description of the male of Baalzebub youyiensis Zhao & Li (Araneae, Theridiosomatidae). Sichuan Journal of Zoology 35 (2): 221 - 226, DOI: https://doi.org/10.11984/j.issn.1000-7083.20150367" title="First description of the male of Baalzebub youyiensis Zhao & Li (Araneae, Theridiosomatidae)." url="https://doi.org/10.11984/j.issn.1000-7083.20150367" volume="35" year="2016">Zhang and Lin 2016</bibRefCitation>
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: 222, figs 1-17 (♂♀).
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</paragraph>
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</subSubSection>
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<subSubSection id="4A932704E588CB7B04AA3375C532BFB2" pageId="0" pageNumber="277" type="type material">
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<paragraph id="FAB70D0E8DD0983B752927ACB81AE93D" pageId="0" pageNumber="277">Type material.</paragraph>
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<paragraph id="4A410307A8B9383B5066D4D97DD395EB" pageId="0" pageNumber="277">
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<emphasis id="347E21D477CBC23DE8FE0DDF74D9BA1C" bold="true" italics="true" pageId="0" pageNumber="277">Holotype</emphasis>
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♀ and
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<emphasis id="8ACC051F8B55146E870B1E3A464A4381" bold="true" italics="true" pageId="0" pageNumber="277">paratypes</emphasis>
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3♀ (IZCAS),
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<emphasis id="FB63FD60F77E9BD061B45F436DEF1EA9" bold="true" pageId="0" pageNumber="277">China</emphasis>
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: Guangxi, Pingxiang City, Youyi Town, Bantou Vill., Niuyan Cave,
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<geoCoordinate id="96127B3AE9B3B7037A288E8D508A3B37" degrees="22" direction="north" minutes="05.666" orientation="latitude" precision="1" value="22.094433">22°05.666'N</geoCoordinate>
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,
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<geoCoordinate id="C4C2C6DBDBF4027E186F3099C19D7894" degrees="106" direction="east" minutes="45.439" orientation="longitude" precision="1" value="106.75732">106°45.439'E</geoCoordinate>
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, 251 m, 18.I.2011, Z. Chen and Z. Zha leg. Examined.
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</paragraph>
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</subSubSection>
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<subSubSection id="C5B96AB705FDFFE2C6B80DFF91C30A24" pageId="0" pageNumber="277" type="materials_examined">
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<paragraph id="91488F32DD5D1A2F05D49B88CD6D09C9" pageId="0" pageNumber="277">Other material examined.</paragraph>
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<paragraph id="5EA9EA81F4399CB5B3552B0900E77D7C" pageId="0" pageNumber="277">
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<materialsCitation id="8807BD9B2E77AB0EBCE3224A35F823EE" collectingDate="2015-09-22" collectorName="J. Wu, Z. Chen" country="China" county="Pingguo Co." latitude="23.8055" location="Liming Vill." longLatPrecision="1" longitude="107.52544" municipality="Baise City" specimenCount="13" specimenCount-female="12" specimenCount-male="1" stateProvince="Guangxi Prov.">
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<emphasis id="F462B30949817BD84B174B9B4EDBD73F" bold="true" pageId="0" pageNumber="277">
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<collectingCountry id="8DAF85F2A05908BD8D7B02D9C9D2E681" name="China">China</collectingCountry>
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</emphasis>
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:
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<specimenCount id="CD7D201275AE1F268D1D9FAF83F2E674" type="male">1♂</specimenCount>
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<specimenCount id="6D79001BC8540EA44298028AB5823338" type="female">12♀</specimenCount>
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(IZCAS),
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<collectingRegion id="1E35C0848A96C852DFCD668E3EB406C0" country="China" name="Guangxi">Guangxi Prov.</collectingRegion>
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,
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<collectingMunicipality id="9113C6B18A4F3EE83AEFF006746D200B">Baise City</collectingMunicipality>
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,
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<collectingCounty id="FCA7E5F2BFF7588651BC9AA76B839B68">Pingguo Co.</collectingCounty>
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,
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<location id="B3032112B12DF71A4D21879F920F863D" LSID="urn:lsid:plazi:treatment:00C66505BCF85CFB825CC345C05BB383:B3032112B12DF71A4D21879F920F863D" country="China" county="Pingguo Co." latitude="23.8055" longLatPrecision="1" longitude="107.52544" municipality="Baise City" name="Liming Vill." stateProvince="Guangxi Prov.">Liming Vill.</location>
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,
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<location id="C1EC393A9AF5EC2ADF6E9E083F2191FB" LSID="urn:lsid:plazi:treatment:00C66505BCF85CFB825CC345C05BB383:C1EC393A9AF5EC2ADF6E9E083F2191FB" country="China" county="Pingguo Co." latitude="23.8055" longLatPrecision="1" longitude="107.52544" municipality="Baise City" name="Ganmoyan Cave" stateProvince="Guangxi Prov.">Ganmoyan Cave</location>
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,
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<geoCoordinate id="395B1B12DD41FB395DB98143C65833BF" degrees="23" direction="north" minutes="48.330" orientation="latitude" precision="1" value="23.8055">23°48.330'N</geoCoordinate>
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,
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<geoCoordinate id="967647C5A177FEDE22CCD17CF83DDAB7" degrees="107" direction="east" minutes="31.526" orientation="longitude" precision="1" value="107.52544">107°31.526'E</geoCoordinate>
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,
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<collectingDate id="F0D0ECCC2D7166A1DC67CCF4598A0180" value="2015-09-22">22.IX.2015</collectingDate>
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,
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<collectorName id="176FFB3F937124B700642DCAFD67B04A">J. Wu</collectorName>
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and
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<collectorName id="6B0DC600EA7B2627F3D6D6839990FCAF">Z. Chen</collectorName>
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leg.
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</materialsCitation>
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;
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<materialsCitation id="1E12F4265671A13659EBE1849A933516" collectingDate="2015-02-09" collectorName="Y. Li, Z. Chen" country="China" county="Donglan Co." elevation="368" latitude="24.355783" location="Sanshi Town" longLatPrecision="1" longitude="107.3865" municipality="Hechi City" specimenCount="8" specimenCount-female="7" specimenCount-male="1" stateProvince="Guangxi Prov.">
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<specimenCount id="6EC66C03887B4F6AD24B775121798A68" type="male">1♂</specimenCount>
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<specimenCount id="A821B4003D0CB1E7BA0C1745D256A722" type="female">7♀</specimenCount>
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(NHMSU),
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|
|
<collectingRegion id="4BD10C3E9A091C059830C9B8D5F42B6D" country="China" name="Guangxi">Guangxi Prov.</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="55AC1EBA68DB9A0BF4B5B5E7F74B08A0">Hechi City</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<collectingCounty id="3864A22EEB295B065E1465F90C080C62">Donglan Co.</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<location id="CDE865D6AEA179D7A9479DB7D3458E93" LSID="urn:lsid:plazi:treatment:00C66505BCF85CFB825CC345C05BB383:CDE865D6AEA179D7A9479DB7D3458E93" country="China" county="Donglan Co." latitude="24.355783" longLatPrecision="1" longitude="107.3865" municipality="Hechi City" name="Sanshi Town" stateProvince="Guangxi Prov.">Sanshi Town</location>
|
|||
|
|
,
|
|||
|
|
<location id="2822E9C2B0222953CD70C5A2278838B9" LSID="urn:lsid:plazi:treatment:00C66505BCF85CFB825CC345C05BB383:2822E9C2B0222953CD70C5A2278838B9" country="China" county="Donglan Co." latitude="24.355783" longLatPrecision="1" longitude="107.3865" municipality="Hechi City" name="Gongping Vill." stateProvince="Guangxi Prov.">Gongping Vill.</location>
|
|||
|
|
, a nameless cave,
|
|||
|
|
<geoCoordinate id="2165932A31EE7E52D0D0DBFC111114A3" degrees="24" direction="north" minutes="21.347" orientation="latitude" precision="1" value="24.355783">24°21.347'N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="9FF40AA829D6B3EFA18D5A7247B3C2F3" degrees="107" direction="east" minutes="23.190" orientation="longitude" precision="1" value="107.3865">107°23.190'E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="C6B279802BA21D4C9B6028AD5B373B1D" metricMagnitude="2" metricUnit="m" metricValue="3.6799999999999997" unit="m" value="368.0">
|
|||
|
|
<elevation id="9512EC473EC3A9BD751967302266FC06" metricMagnitude="2" metricUnit="m" metricValue="3.6799999999999997" unit="m" value="368.0">368 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<collectingDate id="DDAA024DB0A9FD59F6F5B697EAE84BD2" value="2015-02-09">9.II.2015</collectingDate>
|
|||
|
|
,
|
|||
|
|
<collectorName id="0A0FF0848298ACB6C5F4434BD833035E">Y. Li</collectorName>
|
|||
|
|
and
|
|||
|
|
<collectorName id="976DC78FE599B30CC18F78B305C771E9">Z. Chen</collectorName>
|
|||
|
|
leg.
|
|||
|
|
</materialsCitation>
|
|||
|
|
<materialsCitation id="F3F2A835F00C5E249F28294F73425326" collectingDate="2012-08" collectorName="P. Jaeger" country="Laos" county="Lak Sao City" elevation="318" latitude="18.388634" location="Lang Cave" longLatPrecision="1" longitude="104.544586" municipality="Transiten" specimenCount="20" specimenCount-female="17" specimenCount-male="3" stateProvince="Bolikhamsai Prov.">
|
|||
|
|
<emphasis id="46C3B50E6D7518CEF68028344E562DEB" bold="true" pageId="0" pageNumber="277">
|
|||
|
|
<collectingCountry id="98F64BD794ADF0142B238B14C21CBBC7" name="Laos">Laos</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
:
|
|||
|
|
<specimenCount id="4876ADF3466F18A51D677CAE70A6DC9B" type="male">3♂</specimenCount>
|
|||
|
|
<specimenCount id="CD98EFF6395F2FECB3EF0043F48030B9" type="female">17♀</specimenCount>
|
|||
|
|
(NHMSU),
|
|||
|
|
<collectingRegion id="67187855A0C4ECC775D2739783E95E76" country="Laos" name="Bolikhamxai">Bolikhamsai Prov.</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCounty id="09B89573493DF951BB8E3A5FBBC99CF8">Lak Sao City</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="3DC3B043C648AEB5F239E40A7C9A4AF0">Transiten</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<location id="574CC684392A47BE9A06A2E76880A6EC" LSID="urn:lsid:plazi:treatment:00C66505BCF85CFB825CC345C05BB383:574CC684392A47BE9A06A2E76880A6EC" country="Laos" county="Lak Sao City" latitude="18.388634" longLatPrecision="1" longitude="104.544586" municipality="Transiten" name="Lang Cave" stateProvince="Bolikhamsai Prov.">Lang Cave</location>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="84688CE98102949D38C31BBC7BD13264" degrees="18" direction="north" minutes="23.318" orientation="latitude" precision="1" value="18.388634">18°23.318'N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="AA96AC97815E0233F49D072E2C953012" degrees="104" direction="east" minutes="32.675" orientation="longitude" precision="1" value="104.544586">104°32.675'E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="3EF12324F2EC0F46D287EA1FC5724247" metricMagnitude="2" metricUnit="m" metricValue="3.18" unit="m" value="318.0">
|
|||
|
|
<elevation id="E7F4489F485F73E30FF0943953F61D2B" metricMagnitude="2" metricUnit="m" metricValue="3.18" unit="m" value="318.0">318 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<collectingDate id="840462222CAABA1BDC631555ED8D24E3" value="2012-08">VIII.2012</collectingDate>
|
|||
|
|
,
|
|||
|
|
<collectorName id="DDB6C0855A8AC4BBA84781EB99E63A51">
|
|||
|
|
P.
|
|||
|
|
<normalizedToken id="E9000BA49F27CC1467213CD02A765069" originalValue="Jäger">Jaeger</normalizedToken>
|
|||
|
|
</collectorName>
|
|||
|
|
leg.
|
|||
|
|
</materialsCitation>
|
|||
|
|
<materialsCitation id="6E248365B82D82299D6A0A2E0D73EC33" collectingDate="2012-10-26" collectorName="H. Zhao, Z. Chen" country="Vietnam" county="Thanh Son Dist." elevation="375" latitude="21.11" location="Xuan Son National Park" longLatPrecision="1" longitude="104.96" municipality="Xuan Son Commune" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Phu Tho Prov.">
|
|||
|
|
<emphasis id="2B8BAE0A79911EF21DC58FEBD49FDE47" bold="true" pageId="0" pageNumber="277">
|
|||
|
|
<collectingCountry id="5F81F11EBDB50BF1EFBBB8B3CD7C6549" name="Vietnam">Vietnam</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
:
|
|||
|
|
<specimenCount id="CA284A1FA2989ADEF9F8AE628F762D1C" type="male">1♂</specimenCount>
|
|||
|
|
<specimenCount id="B4332E85843B1242A4692BED0C122C66" type="female">3♀</specimenCount>
|
|||
|
|
(IZCAS),
|
|||
|
|
<collectingRegion id="8D65EE1F819C6E2657E5BD1CC7E010A5" country="Vietnam" name="Phu Tho">
|
|||
|
|
<normalizedToken id="2D49ABF3796985AFD7EEFAD1E59927D0" originalValue="Phú">Phu</normalizedToken>
|
|||
|
|
Tho Prov.
|
|||
|
|
</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCounty id="9FBC236227FDD583CE6C51246632A312">Thanh Son Dist.</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="A209A1E18D7E799F38F97E06A1AAD431">Xuan Son Commune</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<location id="2BE48B163CA81971FBF43F03256913B4" LSID="urn:lsid:plazi:treatment:00C66505BCF85CFB825CC345C05BB383:2BE48B163CA81971FBF43F03256913B4" country="Vietnam" county="Thanh Son Dist." latitude="21.11" longLatPrecision="1" longitude="104.96" municipality="Xuan Son Commune" name="Xuan Son National Park" stateProvince="Phu Tho Prov.">Xuan Son National Park</location>
|
|||
|
|
,
|
|||
|
|
<location id="F4FB0FA8DA34D09F4408F175AEB8549F" LSID="urn:lsid:plazi:treatment:00C66505BCF85CFB825CC345C05BB383:F4FB0FA8DA34D09F4408F175AEB8549F" country="Vietnam" county="Thanh Son Dist." latitude="21.11" longLatPrecision="1" longitude="104.96" municipality="Xuan Son Commune" name="Lang Cave" stateProvince="Phu Tho Prov.">Lang Cave</location>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="B1134E632561DAE65F5F3035BCE25FE1" degrees="21" direction="north" minutes="06.600" orientation="latitude" precision="1" value="21.11">21°06.600'N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="89E4D043C8277EB980CC7CDD95E1C2E9" degrees="104" direction="east" minutes="57.600" orientation="longitude" precision="1" value="104.96">104°57.600'E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="0C71A2ADA1FD6CC5132FC75A067490F6" metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">
|
|||
|
|
<elevation id="2ED9E1E0A95CBCC5AEFB07930853CC79" metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">375 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<collectingDate id="6A4C395211B959F775BCBEA4CE85E0B6" value="2012-10-26">26.X.2012</collectingDate>
|
|||
|
|
,
|
|||
|
|
<collectorName id="7DBDBF92F3E3CF95C0A21900FCBF4557">H. Zhao</collectorName>
|
|||
|
|
and
|
|||
|
|
<collectorName id="16B6B11A1711CAACD93D5F3AC891BD5B">Z. Chen</collectorName>
|
|||
|
|
leg.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="AF6ADBB390F38C479605CCAFAE838D9C" pageId="0" pageNumber="277" type="diagnosis">
|
|||
|
|
<paragraph id="921B0982C3D206E1C85F5C73EB85DE45" pageId="0" pageNumber="277">Diagnosis.</paragraph>
|
|||
|
|
<paragraph id="D4DAAA39B546E7BFBC85162FBFDCA35A" pageId="0" pageNumber="277">
|
|||
|
|
Females of
|
|||
|
|
<taxonomicName id="8B723C5C5FDC77075DDC165E46A4D99E" lsidName="S. youyiensis" pageId="0" pageNumber="277" rank="species" species="youyiensis">
|
|||
|
|
<emphasis id="8FF1A763FEBE8079498FFAD154E9F6D6" italics="true" pageId="0" pageNumber="277">S. youyiensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are most similar to those of
|
|||
|
|
<taxonomicName id="F9993D9B8CEA7D34EB11A12467239C84" lsidName="S. sumatra" pageId="0" pageNumber="277" rank="species" species="sumatra">
|
|||
|
|
<emphasis id="2E93033760938AD40AEBA2D6ACE80183" italics="true" pageId="0" pageNumber="277">S. sumatra</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov. by having similar habitus and general shape of the vulva.
|
|||
|
|
<taxonomicName id="4A5AF51713824CA3FCBD89EC32DE317B" authorityName="Zhang & Yu & Lin" authorityYear="2023" baseAuthorityName="Zhao & Li" baseAuthorityYear="2012" family="Theridiosomatidae" genus="Simonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Simonia youyiensis" order="Araneae" pageId="0" pageNumber="277" phylum="Mollusca" rank="species" species="youyiensis">
|
|||
|
|
<emphasis id="41DD1DCC65539A938CA889BF1170B8A6" italics="true" pageId="0" pageNumber="277">Simonia youyiensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="56BECAF207DBC75B25F29EB90F6ADC6B" lsidName="S. sumatra" pageId="0" pageNumber="277" rank="species" species="sumatra">
|
|||
|
|
<emphasis id="81D4406DFF91A45DBF8CA17F0414C7FF" italics="true" pageId="0" pageNumber="277">S. sumatra</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov. in 1) anterior margin of the epigynal plate slightly concaved (c. 140°) (vs concaved c. 120°) (cf. Fig.
|
|||
|
|
<figureCitation id="CA50947AA130CF2C82782CB02639D0E0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Simonia youyiensis, male habitus (A, B), female habitus (C, D) and epigyne (E-G) A dorsal B ventral C dorsal D ventral E intact, ventral F cleared, ventral (blue dashed line showing the anterior margin of the epigynal plate) G cleared, dorsal (green dashed line showing schematic course of copulatory duct). Abbreviations: CB = copulatory bursa; CD = copulatory duct; FD = fertilization duct; Sc = scape; SH = spermathecal head; SS = spermathecal stalk; Sp = spermatheca; TF = transverse fold of epigynal plate. Scale bars: 0.5 mm (A-D); 0.2 mm (E-G)." figureDoi="10.3897/zookeys.1185.104120.figure4" httpUri="https://binary.pensoft.net/fig/945110" pageId="0" pageNumber="277">4E, F</figureCitation>
|
|||
|
|
and Fig.
|
|||
|
|
<figureCitation id="A8311A37201EF30760AF5BA85E54F70A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype female of Simonia sumatra sp. nov., habitus (A, B) and epigyne (C-E) A dorsal B ventral C intact, ventral D cleared, ventral (blue dashed line showing the anterior margin of the epigynal plate) E cleared, dorsal (green dashed line showing schematic course of copulatory duct). Abbreviations: CB = copulatory bursa; CD = copulatory duct; FD = fertilization duct; SB = spermathecal base; Sc = scape; SH = spermathecal head; SS = spermathecal stalk; Sp = spermatheca; TF = transverse fold of epigynal plate. Scale bars: 0.5 mm (A, B); 0.2 mm (C-E)." figureDoi="10.3897/zookeys.1185.104120.figure6" httpUri="https://binary.pensoft.net/fig/945112" pageId="0" pageNumber="277">6C, D</figureCitation>
|
|||
|
|
), 2) spermathecae comma-shaped (vs spermathecae shaped like the whole hind leg of a frog) (cf. Fig.
|
|||
|
|
<figureCitation id="5A5BFEEDEEE8DEFBBB09094B8DB3F113" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Simonia youyiensis, male habitus (A, B), female habitus (C, D) and epigyne (E-G) A dorsal B ventral C dorsal D ventral E intact, ventral F cleared, ventral (blue dashed line showing the anterior margin of the epigynal plate) G cleared, dorsal (green dashed line showing schematic course of copulatory duct). Abbreviations: CB = copulatory bursa; CD = copulatory duct; FD = fertilization duct; Sc = scape; SH = spermathecal head; SS = spermathecal stalk; Sp = spermatheca; TF = transverse fold of epigynal plate. Scale bars: 0.5 mm (A-D); 0.2 mm (E-G)." figureDoi="10.3897/zookeys.1185.104120.figure4" httpUri="https://binary.pensoft.net/fig/945110" pageId="0" pageNumber="277">4G</figureCitation>
|
|||
|
|
and Fig.
|
|||
|
|
<figureCitation id="FB7E839C27B8022FAB60AA26D3AFD122" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype female of Simonia sumatra sp. nov., habitus (A, B) and epigyne (C-E) A dorsal B ventral C intact, ventral D cleared, ventral (blue dashed line showing the anterior margin of the epigynal plate) E cleared, dorsal (green dashed line showing schematic course of copulatory duct). Abbreviations: CB = copulatory bursa; CD = copulatory duct; FD = fertilization duct; SB = spermathecal base; Sc = scape; SH = spermathecal head; SS = spermathecal stalk; Sp = spermatheca; TF = transverse fold of epigynal plate. Scale bars: 0.5 mm (A, B); 0.2 mm (C-E)." figureDoi="10.3897/zookeys.1185.104120.figure6" httpUri="https://binary.pensoft.net/fig/945112" pageId="0" pageNumber="277">6E</figureCitation>
|
|||
|
|
), and 3) bursae spherical (vs egg-shaped) (cf. Fig.
|
|||
|
|
<figureCitation id="D392E592A348BB8612163957FD13AB09" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Simonia youyiensis, male habitus (A, B), female habitus (C, D) and epigyne (E-G) A dorsal B ventral C dorsal D ventral E intact, ventral F cleared, ventral (blue dashed line showing the anterior margin of the epigynal plate) G cleared, dorsal (green dashed line showing schematic course of copulatory duct). Abbreviations: CB = copulatory bursa; CD = copulatory duct; FD = fertilization duct; Sc = scape; SH = spermathecal head; SS = spermathecal stalk; Sp = spermatheca; TF = transverse fold of epigynal plate. Scale bars: 0.5 mm (A-D); 0.2 mm (E-G)." figureDoi="10.3897/zookeys.1185.104120.figure4" httpUri="https://binary.pensoft.net/fig/945110" pageId="0" pageNumber="277">4G</figureCitation>
|
|||
|
|
and Fig.
|
|||
|
|
<figureCitation id="F1514E40768B2D7801D33500FF67B2FB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype female of Simonia sumatra sp. nov., habitus (A, B) and epigyne (C-E) A dorsal B ventral C intact, ventral D cleared, ventral (blue dashed line showing the anterior margin of the epigynal plate) E cleared, dorsal (green dashed line showing schematic course of copulatory duct). Abbreviations: CB = copulatory bursa; CD = copulatory duct; FD = fertilization duct; SB = spermathecal base; Sc = scape; SH = spermathecal head; SS = spermathecal stalk; Sp = spermatheca; TF = transverse fold of epigynal plate. Scale bars: 0.5 mm (A, B); 0.2 mm (C-E)." figureDoi="10.3897/zookeys.1185.104120.figure6" httpUri="https://binary.pensoft.net/fig/945112" pageId="0" pageNumber="277">6E</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="015FA8735555BABCC078A807471EA730" pageId="0" pageNumber="277" type="description">
|
|||
|
|
<paragraph id="69281C8F0B543E6E7265336B7E29C8F0" pageId="0" pageNumber="277">Description.</paragraph>
|
|||
|
|
<paragraph id="9BD2C0BA4706EFB3398C48B1DEAF1A80" pageId="0" pageNumber="277">
|
|||
|
|
<emphasis id="57C1FD21049526168E0FAE391BE8F0E5" bold="true" pageId="0" pageNumber="277">Male</emphasis>
|
|||
|
|
(Fig.
|
|||
|
|
<figureCitation id="FB9C3FBF00A4E110DA41F5876965298A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Simonia youyiensis, male habitus (A, B), female habitus (C, D) and epigyne (E-G) A dorsal B ventral C dorsal D ventral E intact, ventral F cleared, ventral (blue dashed line showing the anterior margin of the epigynal plate) G cleared, dorsal (green dashed line showing schematic course of copulatory duct). Abbreviations: CB = copulatory bursa; CD = copulatory duct; FD = fertilization duct; Sc = scape; SH = spermathecal head; SS = spermathecal stalk; Sp = spermatheca; TF = transverse fold of epigynal plate. Scale bars: 0.5 mm (A-D); 0.2 mm (E-G)." figureDoi="10.3897/zookeys.1185.104120.figure4" httpUri="https://binary.pensoft.net/fig/945110" pageId="0" pageNumber="277">4A, B</figureCitation>
|
|||
|
|
): Carapace brown, slightly darker in ocular region, without distinct pattern; cervical groove and radial grooves indistinct. AER distinctly recurved, PER slightly procurved. Sternum yellowish brown. Mouthparts reddish brown. Legs uniformly yellowish. Abdomen round, dorsum basically light brown, with 3 pairs of black diagonal bands on sides, forming 3 V-shaped stripes; venter centrally with black trapezoidal speckle, marginally with several discontinuous streaks.
|
|||
|
|
<emphasis id="69CDDE5BBD8C0A7FF059C2C22F3D7D23" bold="true" italics="true" pageId="0" pageNumber="277">Measurements</emphasis>
|
|||
|
|
: Total length 1.62. Carapace 0.83 long, 0.81 wide. Clypeus 0.21 high. Sternum 0.52 long, 0.49 wide. Abdomen 1.14 long, 1.08 wide. Length of legs: I 3.32 (1.01, 0.39, 0.88, 0.63, 0.41); II 2.51 (0.81, 0.32, 0.58, 0.51, 0.29); III 1.84 (0.52, 0.21, 0.38, 0.41, 0.32); IV 2.22 (0.49, 0.32, 0.52, 0.58, 0.31).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="0EFA7F14570B1C64EFD5BE09BADBAC8E" pageId="0" pageNumber="277">
|
|||
|
|
Palp (Figs
|
|||
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<figureCitation id="AC6320D3C97ECBC7EDC2C3E584E55279" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male palp of Simonia youyiensis A prolateral B retrolateral C dorsal D ventral. Abbreviations: Co = conductor; CL = cymbial lobe; Cy = cymbium; ED = embolic division; MA = median apophysis; Pc = paracymbium; SCy = cymbial setae; St = subtegulum; Te = tegulum. Scale bars: 0.2 mm (A-D)." figureDoi="10.3897/zookeys.1185.104120.figure1" httpUri="https://binary.pensoft.net/fig/945107" pageId="0" pageNumber="277">1</figureCitation>
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<figureCitation id="863101226D3961EBA01D85112638D52B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palpal sclerites of Simonia youyiensis, cited from Zhang and Lin (2016) A bulb, prolateral B embolic division, ventral C conductor, ventral D cymbium, retrolateral. Abbreviations: Co = conductor; CL = cymbial lobe; EA = embolic apophysis; ED = embolic division; EL = embolic lobe; ET = embolic terminal; MA = median apophysis; Pc = paracymbium; SCy = cymbial setae; St = subtegulum; Te = tegulum. Scale bars: 0.1 mm (A-D)." figureDoi="10.3897/zookeys.1185.104120.figure3" httpUri="https://binary.pensoft.net/fig/945109" pageId="0" pageNumber="277">3</figureCitation>
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): Femur unmodified. Patella round and small, nearly as long as tibia, dorsally bears strong and long macroseta slightly longer than tibia. Tibia cup-shaped, about 1/6-1/5 length of cymbium, with several sparse setae. Cymbium (Cy) broad, navicular, c. 2.1
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<normalizedToken id="ACAF16819BB92B0C2672BC1885FD9B9A" originalValue="×">x</normalizedToken>
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longer than wide, dorsally clothed with dense setae, basolaterally with lobe (CL) and row of longitudinally arranged setae (SCy) along suture between cymbium and lobe. Cymbial lobe represented by small and semi-transparent sheet, just like outline of thumb. Paracymbium (Pc) L-shaped, about 1/5-1/4 length of cymbium, with sharp and spine-like tip. Tegulum (Te) capacious and inflated, c. 1.5
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<normalizedToken id="9A5C4A08D2CE69DBD3C7B8C71F539EE8" originalValue="×">x</normalizedToken>
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longer than wide, surface with countless delicate texture; sperm duct (SD) indistinct and sinuate. Subtegulum (ST) located prolaterally to tegulum, represented by large and surface smooth tubercle, slightly longer than 1/2 length of tegulum. Median apophysis (MA) small, about 2/5 length of tegulum, shaped like tadpole, consisting of relatively wide head (distal part) and narrow tail (proximal part); distal process blunt, apex round and rough, with many tiny and scale-like tooth; proximal process triangular, gradually narrowing toward its apex, apex sharp. Conductor (Co) large and axe-shaped, aligned transversely on anterior part of bulb; almost all of conductor hyaline and with vein-shaped grains, except membranous distal margin. Embolic division nearly as long as conductor, hidden behind conductor, consisting of broad embolic lobe (EL), embolic terminal (ET), and at least 3 apophyses (EA); embolic terminal short, less than 1/2 length of embolic lobe, shaped like a torch, with cylindric stalk and multiramose apex; all embolic apophyses bristle-like and slender, slightly longer than embolic terminal; some apexes of embolic terminal and embolic apophyses overpass distal margin of conductor.
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</paragraph>
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<paragraph id="1070565C34EC51EBB03DDE44DB82DCE8" pageId="0" pageNumber="277">
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<emphasis id="C42182E1FB34E3708E50FF689E477DF7" bold="true" pageId="0" pageNumber="277">Female.</emphasis>
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Somatic features as in Fig.
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<figureCitation id="B34775E27833F2E55CF7C8CEC6B4771D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Simonia youyiensis, male habitus (A, B), female habitus (C, D) and epigyne (E-G) A dorsal B ventral C dorsal D ventral E intact, ventral F cleared, ventral (blue dashed line showing the anterior margin of the epigynal plate) G cleared, dorsal (green dashed line showing schematic course of copulatory duct). Abbreviations: CB = copulatory bursa; CD = copulatory duct; FD = fertilization duct; Sc = scape; SH = spermathecal head; SS = spermathecal stalk; Sp = spermatheca; TF = transverse fold of epigynal plate. Scale bars: 0.5 mm (A-D); 0.2 mm (E-G)." figureDoi="10.3897/zookeys.1185.104120.figure4" httpUri="https://binary.pensoft.net/fig/945110" pageId="0" pageNumber="277">4C, D</figureCitation>
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and coloration distinctly lighter than in male.
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<emphasis id="4D0B64ADAD9BE769E11E78E86F69E51D" bold="true" italics="true" pageId="0" pageNumber="277">Measurements</emphasis>
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: Total length 2.40. Carapace 0.91 long, 0.90 wide. Clypeus 0.20 high. Sternum 0.53 long, 0.51 wide. Abdomen 1.56 long, 1.27 wide. Length of legs: I 3.81 (1.31, 0.42, 0.78, 0.89, 0.41); II 3.17 (0.87, 0.43, 0.81, 0.68, 0.38); III 2.03 (0.63, 0.28, 0.37, 0.44, 0.31); IV 2.32 (0.62, 0.31, 0.49, 0.57, 0.33).
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</paragraph>
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<paragraph id="5AD31F4B24F62A56C54E2FA3E973AF72" pageId="0" pageNumber="277">
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Epigyne (Fig.
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<figureCitation id="B1344BF84D24EA61CAA412C6652366B7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Simonia youyiensis, male habitus (A, B), female habitus (C, D) and epigyne (E-G) A dorsal B ventral C dorsal D ventral E intact, ventral F cleared, ventral (blue dashed line showing the anterior margin of the epigynal plate) G cleared, dorsal (green dashed line showing schematic course of copulatory duct). Abbreviations: CB = copulatory bursa; CD = copulatory duct; FD = fertilization duct; Sc = scape; SH = spermathecal head; SS = spermathecal stalk; Sp = spermatheca; TF = transverse fold of epigynal plate. Scale bars: 0.5 mm (A-D); 0.2 mm (E-G)." figureDoi="10.3897/zookeys.1185.104120.figure4" httpUri="https://binary.pensoft.net/fig/945110" pageId="0" pageNumber="277">4E-G</figureCitation>
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). Epigynal plate large, c. 1.4
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<normalizedToken id="8CCB005B637473CCC05569FAF271FA13" originalValue="×">x</normalizedToken>
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wider than long, anteriorly slightly concaved by c. 140°, arrangement of various parts of vulva indistinctly visible through tegument. Scape (Sc) T-shaped, small, about 1/5 length of epigynal plate. Spermathecae (Sp) comma-shaped, with bean-shaped head (SH) and slightly curved stalk (SS); anterior surface of spermathecal heads touching anterior margin of epigynal plate; spermathecal stalks extending obliquely, separated by about 2-3
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<normalizedToken id="B1FA48444A52B3702B3334B8FB84F941" originalValue="×">x</normalizedToken>
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diameters of stalks. Copulatory bursae (CB) spherical.
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</paragraph>
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</subSubSection>
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<subSubSection id="E854D050E9043FD44FCB830F9EE3BE50" pageId="0" pageNumber="277" type="distribution">
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<paragraph id="CC035CF35E98472C1AEFF2A262EA7726" pageId="0" pageNumber="277">Distribution.</paragraph>
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<paragraph id="7AFBE1730DF389B33E39795D102E68BE" pageId="0" pageNumber="277">
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China (Guangxi), Laos, Vietnam (Fig.
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<figureCitation id="FE7F44A9621EE4DADF8DEB7975421F04" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution records of Simonia gen. nov. species." figureDoi="10.3897/zookeys.1185.104120.figure7" httpUri="https://binary.pensoft.net/fig/945113" pageId="0" pageNumber="277">7</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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