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<document id="0052470D463EAE06B1E2B621B56DFAFE" ID-CLB-Dataset="299703" ID-DOI="10.11646/zootaxa.5476.1.23" ID-GBIF-Dataset="f19cec98-eb1f-4305-8027-69ce085404c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681748" ID-ZooBank="4250D709-E06C-4CFF-84D6-7A9E62427FE2" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1720375291131" checkinUser="plazi" docAuthor="Mazancourt, Valentin De, Mazel, Virgile, Marquet, Gérard, Poupin, Joseph &amp; Keith, Philippe" docDate="2024" docId="03FF87F3E318FFEA1CBEFE78FAB9FE25" docLanguage="en" docName="zootaxa.5476.1.23.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.23" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptychognathus similis Mazancourt, Mazel, Marquet, Poupin &amp; Keith, 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="292" masterDocId="FFC6FF8BE30EFFF31C29FFF2FFA1FFD1" masterDocTitle="Integrative taxonomy study of brackish water crabs of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from Polynesia, with description of two new species" masterLastPageNumber="297" masterPageNumber="267" pageNumber="289" updateTime="1720727319533" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="73A398ECEDE11C044AEE3EF4084AC84C">Integrative taxonomy study of brackish water crabs of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from Polynesia, with description of two new species</mods:title>
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<mods:namePart id="DAC5A6C3491B7467F1EF40F359848FFA">Mazancourt, Valentin De</mods:namePart>
<mods:affiliation id="C9CF7418B395BFF819ED62389C438FAE">Muséum national d’Histoire naturelle, UMR BOREA, CP 026, 57, rue Cuvier, F- 75231 Paris, Cedex 05, France.</mods:affiliation>
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<mods:namePart id="3DAD9D68280D2C66445B9C12F4BD64FC">Mazel, Virgile</mods:namePart>
<mods:affiliation id="CFD95918A3D6D0ED2D79DBD204C12099">Ichtyo-Pacific, Virgile Mazel BP 381816 Punavai, 98718 Punaauia, Tahiti, Polynésie française.</mods:affiliation>
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<mods:namePart id="29F61E3E62C3D156D323EA89FCBD12B0">Marquet, Gérard</mods:namePart>
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<mods:namePart id="4E3AE14472B9C4CEEDEA9EDB0421A548">Keith, Philippe</mods:namePart>
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<mods:affiliation id="F7AEC60CBC0947FDE13761DFCCB0A421">Muséum national d’Histoire naturelle, UMR BOREA, CP 026, 57, rue Cuvier, F- 75231 Paris, Cedex 05, France.</mods:affiliation>
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<mods:date id="CA2AB14B5F2A89D795B657A776BD7557">2024</mods:date>
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<treatment id="03FF87F3E318FFEA1CBEFE78FAB9FE25" ID-DOI="http://doi.org/10.5281/zenodo.12727237" ID-GBIF-Taxon="235040518" ID-Zenodo-Dep="12727237" LSID="urn:lsid:plazi:treatment:03FF87F3E318FFEA1CBEFE78FAB9FE25" httpUri="http://treatment.plazi.org/id/03FF87F3E318FFEA1CBEFE78FAB9FE25" lastPageId="25" lastPageNumber="292" pageId="22" pageNumber="289">
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<taxonomicName id="4C564D66E318FFE51CBEFE78FE00FE74" authorityName="Mazancourt &amp; Mazel &amp; Marquet &amp; Poupin &amp; Keith" authorityYear="2024" box="[151,417,394,421]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="289" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B922EAF7E318FFE51CBEFE78FE00FE74" bold="true" box="[151,417,394,421]" italics="true" pageId="22" pageNumber="289">Ptychognathus similis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211578CE318FFE51D80FE79FDA6FE74" box="[425,519,395,421]" pageId="22" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BE936E5E318FFE51CBEFE5DFE24FE18" blockId="22.[151,519,394,457]" box="[151,389,430,457]" pageId="22" pageNumber="289">
(
<figureCitation id="136D2A60E318FFE51CB6FE5DFEBEFE18" box="[159,287,431,457]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="22" pageNumber="289">Fig. 14A–J</figureCitation>
,
<figureCitation id="136D2A60E318FFE51D02FE5CFEDCFE18" box="[299,381,430,457]" captionStart="FIGURE 15" captionStartId="24.[151,250,1075,1099]" captionTargetBox="[152,1417,181,1052]" captionTargetId="figure-26@24.[151,1436,181,1052]" captionTargetPageId="24" captionText="FIGURE 15. Ptychognathus similis sp. nov.:A, B, dorsal view of specimens from Futuna; C, typical habitat of species, Vainifao River, Futuna. Photo credits: J. Poupin (A), P. Keith (B) and A. Dutartre (C)." figureDoi="http://doi.org/10.5281/zenodo.12681781" httpUri="https://zenodo.org/record/12681781/files/figure.png" pageId="22" pageNumber="289">15A–B</figureCitation>
)
</paragraph>
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<subSubSection id="C34C656EE318FFE51CBEFE1DFE44FDF7" pageId="22" pageNumber="289" type="reference_group">
<paragraph id="8BE936E5E318FFE51CBEFE1DFE44FDF7" blockId="22.[151,1437,495,550]" pageId="22" pageNumber="289">
<treatmentCitationGroup id="AB4611CBE318FFE51CBEFE1DFE44FDF7" pageId="22" pageNumber="289">
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<taxonomicName id="4C564D66E318FFE51CBEFE1DFE15FDD6" ID-CoL="6WNLQ" authorityName="Sakai" authorityYear="1955" box="[151,436,495,519]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="289" phylum="Arthropoda" rank="species" species="hachijoensis">Ptychognathus hachijoensis</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0AF710F4E318FFE51DFBFE1DFD5BFDD6" author="Poupin, J. &amp; Juncker, M." box="[466,762,495,519]" page="36" pageId="22" pageNumber="289" year="2008">
<bibRefCitation id="EFC74B14E318FFE51DFBFE1DFD5BFDD6" author="Poupin, J. &amp; Juncker, M." box="[466,762,495,519]" pageId="22" pageNumber="289" refId="ref19175" refString="Poupin, J. &amp; Juncker, M. (2008) Crustaces des iles Wallis &amp; Futuna: Inventaire illustre, especes commercialisables et capture des formes larvaires. Rapport Technique du CRISP, janvier 2008, 43 pp. [http: // poupin. joseph. free. fr / pdf / poupin-juncker- 2008 - crustacea-wallis. pdf]" type="book" year="2008">Poupin &amp; Juncker, 2008: 36</bibRefCitation>
</treatmentCitation>
, fig. 4c–d; 2010: 75, fig. C.—
<treatmentCitation id="0AF710F4E318FFE5181EFE1DFADFFDD6" author="Keith, P. &amp; Marquet, G." box="[1079,1406,495,519]" page="64" pageId="22" pageNumber="289" year="2011">
<bibRefCitation id="EFC74B14E318FFE5181EFE1DFADFFDD6" author="Keith, P. &amp; Marquet, G." box="[1079,1406,495,519]" pageId="22" pageNumber="289" refId="ref17968" refString="Keith, P. &amp; Marquet, G. (2011) Poissons et crustaces d'eau douce de Wallis et Futuna. Societe Francaise d'Ichtyologie, 119 pp." type="book" year="2011">Keith &amp; Marquet, 2011: 64–65</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="0AF710F4E318FFE51CEEFDFCFE40FDF7" author="Keith, P. &amp; Marquet, G. &amp; Gerbeaux, P. &amp; Vigneux, E. &amp; Lord, C." box="[199,481,526,550]" page="124" pageId="22" pageNumber="289" year="2013">
<bibRefCitation id="EFC74B14E318FFE51CEEFDFCFE40FDF7" author="Keith, P. &amp; Marquet, G. &amp; Gerbeaux, P. &amp; Vigneux, E. &amp; Lord, C." box="[199,481,526,550]" pageId="22" pageNumber="289" refId="ref17997" refString="Keith, P., Marquet, G., Gerbeaux, P., Vigneux, E. &amp; Lord, C. (2013) Poissons et crustaces d'eau douce de Polynesie. Taxonomie, ecologie, biologie et gestion. Societe francaise d'Ichtyologie, Paris, 282 pp." type="book" year="2013">
Keith
<emphasis id="B922EAF7E318FFE51D2FFDFDFE98FDF7" box="[262,313,526,550]" italics="true" pageId="22" pageNumber="289">et al.</emphasis>
, 2013: 124–125
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<emphasis id="B922EAF7E318FFE51CBEFDA2FE9EFDB8" bold="true" box="[151,319,591,618]" pageId="22" pageNumber="289">Type material</emphasis>
.
<materialsCitation id="3B3E3CB8E318FFE51D67FDBDFE97FD61" ID-GBIF-Occurrence="4905160343" collectingDate="2007-10-14" collectionCode="CA" collectorName="J. Poupin &amp; M. Juncker" country="Wallis and Futuna" latitude="-14.307683" location="Vainifao River" longLatPrecision="1" longitude="-178.14273" municipality="Station" pageId="22" pageNumber="289" specimenCode="CA2659" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B922EAF7E318FFE51D67FDBDFE14FDB8" box="[334,437,591,617]" italics="true" pageId="22" pageNumber="289">
<typeStatus id="54ED8847E318FFE51D67FDBDFE14FDB8" box="[334,437,591,617]" pageId="22" pageNumber="289" type="holotype">Holotype</typeStatus>
</emphasis>
: MNHN-IU-2022-4125 (DNA voucher
<specimenCode id="DBF09E9EE318FFE51F53FDBDFC7FFDBB" box="[890,990,591,618]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="22" pageNumber="289" type="Museum">CA2659</specimenCode>
),
<specimenCount id="9D50FD6CE318FFE51FDBFDA2FBA8FDBB" box="[1010,1033,592,618]" count="1" pageId="22" pageNumber="289" type="male">♂</specimenCount>
, 9.3 mm × 7.6 mm,
<location id="8E89603EE318FFE518D8FDBDFA39FDB8" LSID="urn:lsid:plazi:treatment:03FF87F3E318FFEA1CBEFE78FAB9FE25:8E89603EE318FFE518D8FDBDFA39FDB8" box="[1265,1432,591,617]" country="Wallis and Futuna" latitude="-14.307683" longLatPrecision="1" longitude="-178.14273" municipality="Station" name="Vainifao River" pageId="22" pageNumber="289">Vainifao River</location>
,
<collectingCountry id="F3417675E318FFE51CBEFD81FF49FD5C" box="[151,232,627,653]" name="Wallis and Futuna" pageId="22" pageNumber="289">Futuna</collectingCountry>
, Wallis and
<taxonomicName id="4C564D66E318FFE51D57FD81FE6DFD5C" authority="Territory" authorityName="Territory" box="[382,460,627,653]" family="Gobiidae" genus="Akihito" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="289" phylum="Chordata" rank="species" species="futuna">Futuna</taxonomicName>
Territory,
<collectingMunicipality id="6B8DAC9FE318FFE51E62FD81FD3BFD5C" box="[587,666,627,653]" pageId="22" pageNumber="289">Station</collectingMunicipality>
13,
<geoCoordinate id="EE625022E318FFE51EFCFD81FCC0FD5C" box="[725,865,627,653]" degrees="14" direction="south" minutes="18.461" orientation="latitude" pageId="22" pageNumber="289" precision="1" value="-14.307683">14°18.461'S</geoCoordinate>
,
<geoCoordinate id="EE625022E318FFE51F58FD81FBA4FD5F" box="[881,1029,627,654]" degrees="178" direction="west" minutes="8.564" orientation="longitude" pageId="22" pageNumber="289" precision="1" value="-178.14273">178°8.564'W</geoCoordinate>
, coll.
<collectorName id="26A35333E318FFE51878FD81FB1FFD5C" box="[1105,1214,627,653]" pageId="22" pageNumber="289">J. Poupin</collectorName>
&amp;
<collectorName id="26A35333E318FFE518C5FD81FAD1FD5C" box="[1260,1392,627,653]" pageId="22" pageNumber="289">M. Juncker</collectorName>
,
<date id="FFE81025E318FFE519A9FD81FE97FD61" pageId="22" pageNumber="289" value="2007-10-14">
<collectingDate id="EFACE9CDE318FFE519A9FD81FE97FD61" pageId="22" pageNumber="289" value="2007-10-14">14 October 2007</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B3E3CB8E318FFE51D6BFD65FEEBFD26" ID-GBIF-Occurrence="4905160318" collectingDate="2007-10-14" collectionCode="CA" collectorName="J. Poupin &amp; M. Juncker" country="Wallis and Futuna" latitude="-14.307683" location="Vainifao River" longLatPrecision="1" longitude="-178.14273" municipality="Station" pageId="22" pageNumber="289" specimenCode="CA2658" specimenCount="10" specimenCount-female="5" specimenCount-male="5" typeStatus="paratype">
<emphasis id="B922EAF7E318FFE51D6BFD65FE15FD61" box="[322,436,663,688]" italics="true" pageId="22" pageNumber="289">
<typeStatus id="54ED8847E318FFE51D6BFD65FE15FD61" box="[322,436,663,688]" pageId="22" pageNumber="289" type="paratype">Paratypes</typeStatus>
</emphasis>
: MNHN-IU-2022-4126 (DNA voucher
<specimenCode id="DBF09E9EE318FFE51F57FD64FC41FD60" box="[894,992,662,689]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="22" pageNumber="289" type="Museum">CA2658</specimenCode>
),
<specimenCount id="9D50FD6CE318FFE51FDFFD64FB83FD60" box="[1014,1058,662,689]" count="1" pageId="22" pageNumber="289" type="female">1 ♀</specimenCount>
, 7.3 mm × 6.1 mm,
<specimenCount id="9D50FD6CE318FFE5193AFD64FA9EFD60" box="[1299,1343,662,689]" count="5" pageId="22" pageNumber="289" type="male">5 ♂</specimenCount>
6.4–7.9 mm × 5.6–6.5 mm,
<specimenCount id="9D50FD6CE318FFE51DAAFD48FE10FD05" box="[387,433,698,724]" count="2" pageId="22" pageNumber="289" type="female">2 ♀</specimenCount>
7.0–7.2 mm × 5.9–6.3 mm,
<specimenCount id="9D50FD6CE318FFE51F2CFD48FC92FD05" box="[773,819,698,724]" count="2" pageId="22" pageNumber="289" type="female">2 ♀</specimenCount>
ovig. 6.6–7.9 mm × 5.2–7.6 mm, same locality data as for holotype
</materialsCitation>
;
<materialsCitation id="3B3E3CB8E318FFE51D71FD2FFCA7FCCA" ID-GBIF-Occurrence="4905160368" collectingDate="2023-08-06" collectionCode="V" collectorName="N. Charpin &amp; Mazel" country="Wallis and Futuna" latitude="-14.253817" location="Vai River" longLatPrecision="1" longitude="-178.1532" municipality="Station" pageId="22" pageNumber="289" specimenCount="4" specimenCount-male="4" typeStatus="paratype">
MNHN-IU-2022-4127,
<specimenCount id="9D50FD6CE318FFE51E41FD2FFD39FD29" box="[616,664,733,760]" count="4" pageId="22" pageNumber="289" type="male">4 ♂</specimenCount>
, 6.5–15.2 × 5.8–13.2 mm,
<location id="8E89603EE318FFE51FFFFD2FFBE7FD26" LSID="urn:lsid:plazi:treatment:03FF87F3E318FFEA1CBEFE78FAB9FE25:8E89603EE318FFE51FFFFD2FFBE7FD26" box="[982,1094,733,759]" country="Wallis and Futuna" latitude="-14.253817" longLatPrecision="1" longitude="-178.1532" municipality="Station" name="Vai River" pageId="22" pageNumber="289">Vai River</location>
,
<collectingMunicipality id="6B8DAC9FE318FFE5187DFD2FFB02FD26" box="[1108,1187,733,759]" pageId="22" pageNumber="289">Station</collectingMunicipality>
3,
<geoCoordinate id="EE625022E318FFE518E4FD2FFAF8FD29" box="[1229,1369,733,760]" degrees="14" direction="south" minutes="15.229" orientation="latitude" pageId="22" pageNumber="289" precision="1" value="-14.253817">14°15.229'S</geoCoordinate>
,
<geoCoordinate id="EE625022E318FFE5194EFD2FFF57FCCA" degrees="178" direction="west" minutes="9.192" orientation="longitude" pageId="22" pageNumber="289" precision="1" value="-178.1532">178° 9.192'W</geoCoordinate>
, coll.
<collectorName id="26A35333E318FFE51D17FCF3FE1DFCCA" box="[318,444,769,795]" pageId="22" pageNumber="289">N. Charpin</collectorName>
&amp;
<collectionCode id="ED47AE20E318FFE51DCDFCF3FE59FCCA" box="[484,504,769,795]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="289" type="Museum">V</collectionCode>
.
<collectorName id="26A35333E318FFE51E2CFCF3FDEFFCCA" box="[517,590,769,795]" pageId="22" pageNumber="289">Mazel</collectorName>
,
<date id="FFE81025E318FFE51E72FCF3FCA7FCCA" box="[603,774,769,795]" pageId="22" pageNumber="289" value="2023-08-06">
<collectingDate id="EFACE9CDE318FFE51E72FCF3FCA7FCCA" box="[603,774,769,795]" pageId="22" pageNumber="289" value="2023-08-06">6 August 2023</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3CB8E318FFE51F3DFCF3FE8BFCB3" ID-GBIF-Occurrence="4905160332" collectingDate="2023-08-06" collectionCode="V" collectorName="N. Charpin &amp; Mazel" country="Wallis and Futuna" latitude="-14.259566" location="Galoli River" longLatPrecision="1" longitude="-178.13841" municipality="Station" pageId="22" pageNumber="289" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
MNHN-IU-2022-4128,
<specimenCount id="9D50FD6CE318FFE5180DFCF3FBF3FCCA" box="[1060,1106,769,795]" count="2" pageId="22" pageNumber="289" type="male">2 ♂</specimenCount>
, 6.4–6.6 × 5.5–5.6 mm,
<specimenCount id="9D50FD6CE318FFE51959FCF3FA3DFCCA" box="[1392,1436,769,795]" count="2" pageId="22" pageNumber="289" type="female">2 ♀</specimenCount>
ovig., 10.9–11.4 × 9.6–9.9 mm,
<location id="8E89603EE318FFE51E2AFCD6FD31FCEF" LSID="urn:lsid:plazi:treatment:03FF87F3E318FFEA1CBEFE78FAB9FE25:8E89603EE318FFE51E2AFCD6FD31FCEF" box="[515,656,804,830]" country="Wallis and Futuna" latitude="-14.259566" longLatPrecision="1" longitude="-178.13841" municipality="Station" name="Galoli River" pageId="22" pageNumber="289">Galoli River</location>
,
<collectingMunicipality id="6B8DAC9FE318FFE51EB5FCD6FD4AFCEF" box="[668,747,804,830]" pageId="22" pageNumber="289">Station</collectingMunicipality>
5
<geoCoordinate id="EE625022E318FFE51F20FCD6FC35FCEE" box="[777,916,804,831]" degrees="14" direction="south" minutes="15.574" orientation="latitude" pageId="22" pageNumber="289" precision="1" value="-14.259566">14°15.574'S</geoCoordinate>
,
<geoCoordinate id="EE625022E318FFE51F89FCD6FB95FCEE" box="[928,1076,804,831]" degrees="178" direction="west" minutes="8.305" orientation="longitude" pageId="22" pageNumber="289" precision="1" value="-178.13841">178°8.305'W</geoCoordinate>
, coll.
<collectorName id="26A35333E318FFE51850FCD7FB57FCEF" box="[1145,1270,804,831]" pageId="22" pageNumber="289">N. Charpin</collectorName>
&amp;
<collectionCode id="ED47AE20E318FFE51933FCD7FA8FFCEE" box="[1306,1326,805,831]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="289" type="Museum">V</collectionCode>
.
<collectorName id="26A35333E318FFE51913FCD6FA22FCEF" box="[1338,1411,804,830]" pageId="22" pageNumber="289">Mazel</collectorName>
,
<date id="FFE81025E318FFE519A7FCD6FE8BFCB3" pageId="22" pageNumber="289" value="2023-08-06">
<collectingDate id="EFACE9CDE318FFE519A7FCD6FE8BFCB3" pageId="22" pageNumber="289" value="2023-08-06">6 August 2023</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3CB8E318FFE51D1EFCBAFC49FC54" ID-GBIF-Occurrence="4905160371" collectingDate="2023-08-08" collectionCode="V" collectorName="N. Charpin &amp; Mazel" country="Wallis and Futuna" latitude="-14.2933" location="Leava River" longLatPrecision="1" longitude="-178.15906" municipality="Station" pageId="22" pageNumber="289" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
MNHN-IU-2022-4129,
<specimenCount id="9D50FD6CE318FFE51E6DFCBAFDD0FCB3" box="[580,625,840,866]" count="2" pageId="22" pageNumber="289" type="male">2 ♂</specimenCount>
, 13.7–15.3 × 12.5–11.6 mm,
<specimenCount id="9D50FD6CE318FFE51F95FCBAFC47FCB3" box="[956,998,840,866]" count="1" pageId="22" pageNumber="289" type="female">1 ♀</specimenCount>
, 13.7 × 11.4 mm,
<location id="8E89603EE318FFE5189CFCBAFAE0FCB3" LSID="urn:lsid:plazi:treatment:03FF87F3E318FFEA1CBEFE78FAB9FE25:8E89603EE318FFE5189CFCBAFAE0FCB3" box="[1205,1345,840,866]" country="Wallis and Futuna" latitude="-14.2933" longLatPrecision="1" longitude="-178.15906" municipality="Station" name="Leava River" pageId="22" pageNumber="289">Leava River</location>
,
<collectingMunicipality id="6B8DAC9FE318FFE51964FCBAFA3DFCB3" box="[1357,1436,840,866]" pageId="22" pageNumber="289">Station</collectingMunicipality>
10,
<geoCoordinate id="EE625022E318FFE51CE8FC99FEECFC57" box="[193,333,875,902]" degrees="14" direction="south" minutes="17.598" orientation="latitude" pageId="22" pageNumber="289" precision="1" value="-14.2933">14°17.598'S</geoCoordinate>
,
<geoCoordinate id="EE625022E318FFE51D71FC99FE4DFC57" box="[344,492,875,902]" degrees="178" direction="west" minutes="9.543" orientation="longitude" pageId="22" pageNumber="289" precision="1" value="-178.15906">178°9.543'W</geoCoordinate>
, coll.
<collectorName id="26A35333E318FFE51E06FC9EFD0AFC54" box="[559,683,875,902]" pageId="22" pageNumber="289">N. Charpin</collectorName>
&amp;
<collectionCode id="ED47AE20E318FFE51EE6FC9EFD42FC57" box="[719,739,876,902]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="289" type="Museum">V</collectionCode>
.
<collectorName id="26A35333E318FFE51EC4FC99FC97FC54" box="[749,822,875,901]" pageId="22" pageNumber="289">Mazel</collectorName>
,
<date id="FFE81025E318FFE51F68FC99FC49FC54" box="[833,1000,875,901]" pageId="22" pageNumber="289" value="2023-08-08">
<collectingDate id="EFACE9CDE318FFE51F68FC99FC49FC54" box="[833,1000,875,901]" pageId="22" pageNumber="289" value="2023-08-08">8 August 2023</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3CB8E318FFE51FDAFC99FA91FC78" ID-GBIF-Occurrence="4905160357" collectingDate="2023-08-09" collectionCode="V" collectorName="N. Charpin &amp; Mazel" country="Wallis and Futuna" latitude="-14.28475" location="Vainifao River" longLatPrecision="1" longitude="-178.13634" municipality="Station" pageId="22" pageNumber="289" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
MNHN-IU-2022-4130,
<specimenCount id="9D50FD6CE318FFE51929FC99FA8DFC57" box="[1280,1324,875,902]" count="1" pageId="22" pageNumber="289" type="male">1 ♂</specimenCount>
, 9.2 × 7.3 mm,
<location id="8E89603EE318FFE51CF8FC7DFED9FC78" LSID="urn:lsid:plazi:treatment:03FF87F3E318FFEA1CBEFE78FAB9FE25:8E89603EE318FFE51CF8FC7DFED9FC78" box="[209,376,911,937]" country="Wallis and Futuna" latitude="-14.28475" longLatPrecision="1" longitude="-178.13634" municipality="Station" name="Vainifao River" pageId="22" pageNumber="289">Vainifao River</location>
,
<collectingMunicipality id="6B8DAC9FE318FFE51DADFC7DFE72FC78" box="[388,467,911,937]" pageId="22" pageNumber="289">Station</collectingMunicipality>
11,
<geoCoordinate id="EE625022E318FFE51E2FFC7CFD2FFC78" box="[518,654,910,937]" degrees="14" direction="south" minutes="17.085" orientation="latitude" pageId="22" pageNumber="289" precision="1" value="-14.28475">14°17.085'S</geoCoordinate>
<geoCoordinate id="EE625022E318FFE51EBEFC7DFC8AFC78" box="[663,811,911,937]" degrees="178" direction="west" minutes="8.180" orientation="longitude" pageId="22" pageNumber="289" precision="1" value="-178.13634">178°8.180'W</geoCoordinate>
, coll.
<collectorName id="26A35333E318FFE51F59FC7DFC4CFC78" box="[880,1005,911,937]" pageId="22" pageNumber="289">N. Charpin</collectorName>
&amp;
<collectionCode id="ED47AE20E318FFE5183AFC7DFB86FC78" box="[1043,1063,911,937]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="289" type="Museum">V</collectionCode>
.
<collectorName id="26A35333E318FFE5181AFC7DFBDDFC78" box="[1075,1148,911,937]" pageId="22" pageNumber="289">Mazel</collectorName>
,
<date id="FFE81025E318FFE518A1FC7DFA91FC78" box="[1160,1328,911,937]" pageId="22" pageNumber="289" value="2023-08-09">
<collectingDate id="EFACE9CDE318FFE518A1FC7DFA91FC78" box="[1160,1328,911,937]" pageId="22" pageNumber="289" value="2023-08-09">9 August 2023</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3CB8E318FFE51914FC7DFE9FFC1D" ID-GBIF-Occurrence="4905160319" collectingDate="2023-08-14" collectionCode="MNHN-, V" collectorName="N. Charpin &amp; Mazel" country="Wallis and Futuna" latitude="-14.291634" location="Leava River" longLatPrecision="1" longitude="-178.15509" municipality="Station" pageId="22" pageNumber="289" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectionCode id="ED47AE20E318FFE51914FC7DFA3DFC78" box="[1341,1436,911,937]" pageId="22" pageNumber="289">MNHNIU-</collectionCode>
2022-4131,
</materialsCitation>
<materialsCitation id="3B3E3CB8E318FFE51D6EFC40FE01FC21" ID-GBIF-Occurrence="4905160337" collectingDate="2023-08-14" collectionCode="MNHN-, V" collectorName="N. Charpin &amp; Mazel" country="Wallis and Futuna" latitude="-14.291634" location="Leava River" longLatPrecision="1" longitude="-178.15509" municipality="Station" pageId="22" pageNumber="289" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D50FD6CE318FFE51D6EFC40FED4FC1C" box="[327,373,946,973]" count="1" pageId="22" pageNumber="289" type="male">1 ♂</specimenCount>
, 14.5 × 12.5 mm,
<location id="8E89603EE318FFE51E65FC41FD78FC1D" LSID="urn:lsid:plazi:treatment:03FF87F3E318FFEA1CBEFE78FAB9FE25:8E89603EE318FFE51E65FC41FD78FC1D" box="[588,729,946,973]" country="Wallis and Futuna" latitude="-14.291634" longLatPrecision="1" longitude="-178.15509" municipality="Station" name="Leava River" pageId="22" pageNumber="289">Leava River</location>
,
<collectingMunicipality id="6B8DAC9FE318FFE51ECFFC40FC94FC1D" box="[742,821,946,972]" pageId="22" pageNumber="289">Station</collectingMunicipality>
20,
<geoCoordinate id="EE625022E318FFE51F42FC40FC57FC1D" box="[875,1014,946,972]" degrees="14" direction="south" minutes="17.498" orientation="latitude" pageId="22" pageNumber="289" precision="1" value="-14.291634">14°17.498'S</geoCoordinate>
,
<geoCoordinate id="EE625022E318FFE5182DFC40FB39FC1C" box="[1028,1176,946,973]" degrees="178" direction="west" minutes="9.305" orientation="longitude" pageId="22" pageNumber="289" precision="1" value="-178.15509">178°9.305'W</geoCoordinate>
, coll.
<collectorName id="26A35333E318FFE518F6FC41FAFCFC1D" box="[1247,1373,946,973]" pageId="22" pageNumber="289">N. Charpin</collectorName>
&amp;
<collectionCode id="ED47AE20E318FFE519ACFC41FA38FC1C" box="[1413,1433,947,973]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="289" type="Museum">V</collectionCode>
.
<collectorName id="26A35333E318FFE51CBEFC27FF41FC3E" box="[151,224,981,1007]" pageId="22" pageNumber="289">Mazel</collectorName>
,
<date id="FFE81025E318FFE51CC2FC24FE01FC21" box="[235,416,982,1008]" pageId="22" pageNumber="289" value="2023-08-14">
<collectingDate id="EFACE9CDE318FFE51CC2FC24FE01FC21" box="[235,416,982,1008]" pageId="22" pageNumber="289" value="2023-08-14">14 August 2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE936E5E318FFE51CEEFC0BFB6FFB8B" blockId="22.[151,1437,591,2037]" pageId="22" pageNumber="289">
<emphasis id="B922EAF7E318FFE51CEEFC0BFE75FBC2" bold="true" box="[199,468,1017,1043]" pageId="22" pageNumber="289">Comparative material</emphasis>
.
<taxonomicName id="4C564D66E318FFE51DCCFC0BFC12FBC5" authority="Sakai, 1955" authorityName="Sakai" authorityYear="1955" box="[485,947,1017,1044]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="289" phylum="Arthropoda" rank="species" species="hachijoensis">
<emphasis id="B922EAF7E318FFE51DCCFC0BFC83FBC2" box="[485,802,1017,1043]" italics="true" pageId="22" pageNumber="289">Ptychognathus hachijoensis</emphasis>
<materialsCitation id="3B3E3CB8E318FFE51F05FC0BFC12FBC5" ID-GBIF-Occurrence="4905160302" box="[812,947,1017,1044]" collectionCode="TAIWAN" collectorName="J. - W. Hsu" location="Gengfang River" pageId="22" pageNumber="289" specimenCode="CA2866, CA2865" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Yilan">
<bibRefCitation id="EFC74B14E318FFE51F05FC0BFC12FBC5" author="Sakai, T." box="[812,947,1017,1044]" pageId="22" pageNumber="289" pagination="193 - 201" refId="ref19642" refString="Sakai, T. (1955) Further Notes on the Brachyuran Crustacea of the Hachijo Island. Records of Oceanographic. Works in Japan, 2 (1), 193 - 201." type="journal article" year="1955">Sakai, 1955</bibRefCitation>
</materialsCitation>
</taxonomicName>
(
<collectionCode id="ED47AE20E318FFE51FECFC0BFB9BFBC2" box="[965,1082,1017,1043]" pageId="22" pageNumber="289">TAIWAN</collectionCode>
): MNHN-IU-2022-4001 (DNA voucher
<specimenCode id="DBF09E9EE318FFE51CDEFBEEFEFAFBE7" box="[247,347,1052,1078]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="22" pageNumber="289" type="Museum">CA2866</specimenCode>
),
<specimenCount id="9D50FD6CE318FFE51D44FBEFFE39FBE6" box="[365,408,1053,1079]" count="1" pageId="22" pageNumber="289" type="male">1 ♂</specimenCount>
,
<quantity id="4CAE9B00E318FFE51D8AFBEEFE59FBE6" box="[419,504,1052,1079]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="22" pageNumber="289" unit="mm" value="9.6">9.6 mm</quantity>
×
<quantity id="4CAE9B00E318FFE51E3AFBEFFDCAFBE6" box="[531,619,1053,1079]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="22" pageNumber="289" unit="mm" value="7.9">7.9 mm</quantity>
,
<specimenCount id="9D50FD6CE318FFE51E5CFBEFFD01FBE6" box="[629,672,1053,1079]" count="1" pageId="22" pageNumber="289" type="male">1 ♂</specimenCount>
,
<quantity id="4CAE9B00E318FFE51E82FBEFFCA1FBE6" box="[683,768,1053,1079]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="22" pageNumber="289" unit="mm" value="8.8">8.8 mm</quantity>
×
<quantity id="4CAE9B00E318FFE51F35FBEFFCD5FBE6" box="[796,884,1053,1079]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="22" pageNumber="289" unit="mm" value="7.3">7.3 mm</quantity>
; DNA voucher
<specimenCode id="DBF09E9EE318FFE51809FBEEFB20FBE6" box="[1056,1153,1052,1079]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="22" pageNumber="289" type="Museum">CA2865</specimenCode>
,
<specimenCount id="9D50FD6CE318FFE518A5FBEFFB14FBE6" box="[1164,1205,1053,1079]" count="1" pageId="22" pageNumber="289" type="female">1 ♀</specimenCount>
,
<quantity id="4CAE9B00E318FFE518EBFBEFFA84FBE6" box="[1218,1317,1053,1079]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="22" pageNumber="289" unit="mm" value="10.2">10.2 mm</quantity>
× 9.0 mm,
<specimenCount id="9D50FD6CE318FFE51CBEFBB2FF60FB8A" box="[151,193,1088,1115]" count="1" pageId="22" pageNumber="289" type="female">1 ♀</specimenCount>
,
<quantity id="4CAE9B00E318FFE51CE6FBB2FE92FB8A" box="[207,307,1088,1115]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="22" pageNumber="289" unit="mm" value="10.8">10.8 mm</quantity>
×
<quantity id="4CAE9B00E318FFE51D78FBB2FE08FB8A" box="[337,425,1088,1115]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="22" pageNumber="289" unit="mm" value="9.2">9.2 mm</quantity>
,
<location id="8E89603EE318FFE51D9DFBB2FDC9FB8B" LSID="urn:lsid:plazi:treatment:03FF87F3E318FFEA1CBEFE78FAB9FE25:8E89603EE318FFE51D9DFBB2FDC9FB8B" box="[436,616,1088,1114]" name="Gengfang River" pageId="22" pageNumber="289" stateProvince="Yilan">Gengfang River</location>
estuary, Toucheng,
<collectingRegion id="4992F807E318FFE51F6FFBB2FC7FFB8B" box="[838,990,1088,1114]" country="Taiwan" name="Yilan" pageId="22" pageNumber="289">Yilan County</collectingRegion>
, coll.
<collectorName id="26A35333E318FFE51808FBB3FB2EFB8A" box="[1057,1167,1089,1115]" pageId="22" pageNumber="289">J.-W. Hsu</collectorName>
<emphasis id="B922EAF7E318FFE518BFFBB3FB6FFB8B" box="[1174,1230,1088,1114]" italics="true" pageId="22" pageNumber="289">et al.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34C656EE318FFE51CEEFB91FBC0FAFE" pageId="22" pageNumber="289" type="diagnosis">
<paragraph id="8BE936E5E318FFE51CEEFB91FBC0FAFE" blockId="22.[151,1437,591,2037]" pageId="22" pageNumber="289">
<emphasis id="B922EAF7E318FFE51CEEFB91FEE3FBAC" bold="true" box="[199,322,1123,1149]" pageId="22" pageNumber="289">Diagnosis.</emphasis>
Carapace subquadrate, dorsal surface smooth, flat; front broad, anterior margin slightly concave, frontal width 0.45–0.47 times CW; anterolateral margin with single orbital tooth followed by faint lobe. Third maxillipeds with merus anterolateral angle inflated; exopod broad, inflated, greatest width 1.0–1.3 times median width of ischium. Chelipeds stout, symmetrical in males; fingers with tuft of long setae on outer surface restricted at base of fingers; pulvinus at base of fingers. Ambulatory legs slender, somewhat compressed laterally; propodus and dactylus with dense long setae on posterior margins, glabrous on anterior margins.
</paragraph>
</subSubSection>
<subSubSection id="C34C656EE318FFEA1CEEFACAFA99FF28" lastPageId="25" lastPageNumber="292" pageId="22" pageNumber="289" type="description">
<paragraph id="8BE936E5E318FFE51CEEFACAFD95F9D2" blockId="22.[151,1437,591,2037]" pageId="22" pageNumber="289">
<emphasis id="B922EAF7E318FFE51CEEFACAFEF8FA83" bold="true" box="[199,345,1336,1362]" pageId="22" pageNumber="289">Description.</emphasis>
Carapace (
<figureCitation id="136D2A60E318FFE51DF7FACAFDE6FA83" box="[478,583,1336,1362]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="22" pageNumber="289">Fig. 14A</figureCitation>
) subquadrate, slightly broader than long, 1.1–1.2 times as broad as long, flat, dorsal surface smooth, glabrous. Regions poorly defined, central H-shaped grooved distinct; epigastric cristae strong. Front broad, anterior margin slightly concave medially frontal width 0.45–0.47 times CW, fronto-orbital width 0.78–0.84 times CW. Anterolateral margins with single orbital tooth, followed by faint lobe. Posterolateral margins divergent posteriorly, moderately sloping outwards. Infraorbital ridge consisting of several small rounded granules, decreasing in size laterally.
</paragraph>
<paragraph id="8BE936E5E318FFE51CEEF9FFFB8EF9F6" blockId="22.[151,1437,591,2037]" box="[199,1071,1549,1576]" pageId="22" pageNumber="289">
Epistome broad (
<figureCitation id="136D2A60E318FFE51DAEF9FFFE4CF9F6" box="[391,493,1549,1575]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="22" pageNumber="289">Fig. 14B</figureCitation>
), median part triangular, margin with tiny granules.
</paragraph>
<paragraph id="8BE936E5E318FFE51CEEF9C2FEFCF964" blockId="22.[151,1437,591,2037]" pageId="22" pageNumber="289">
Third maxillipeds (
<figureCitation id="136D2A60E318FFE51D8CF9C2FDADF99A" box="[421,524,1584,1611]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="22" pageNumber="289">Fig. 14C</figureCitation>
) with ischium subequal in proximal and distal widths, lateral margin concave, ventral surface with longitudinal median ridge. Merus broad, with anterolateral angle expanded; ventral surface with oblique ridge on mesial part. Exopod broad, inflated, greatest width 1.2–1.3 times (male), 1.0 (female) of median width of ischium.
</paragraph>
<paragraph id="8BE936E5E318FFE51CEEF94CFAE9F85B" blockId="22.[151,1437,591,2037]" pageId="22" pageNumber="289">
Chelipeds (
<figureCitation id="136D2A60E318FFE51D61F94CFE0EF909" box="[328,431,1726,1752]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="22" pageNumber="289">Fig. 14D</figureCitation>
) symmetrical in male, not granular at all. Merus longer than broad (length to width ratio 1.6–1.7), anterior and posterior margins glabrous, posterior distal margins bluntly carinate. Carpus, unarmed, strongly inflated, slightly longer than wide (length to width ratio 1.1–1.2). Palm much inflated, smooth, without distinct granules; inner surface glabrous. Fingers with tuft of long setae on outer surface restricted at the base of fingers. Dactylus as long as palm, terminating in small, hoof-shaped corneous claw, with 9 or 10 small blunt teeth. Pollex with only faint indication of longitudinal groove, with 4 or 5 blunt teeth. Pulvinus at base of fingers.
</paragraph>
<paragraph id="8BE936E5E318FFE51CEEF861FBBCF824" blockId="22.[151,1437,591,2037]" pageId="22" pageNumber="289">
Female chelipeds (
<figureCitation id="136D2A60E318FFE51DB2F861FE5EF87C" box="[411,511,1939,1965]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="22" pageNumber="289">Fig. 14E</figureCitation>
) distinctly smaller than those of male, chela much narrower than that of male. No tuft of setae on outer surface of fingers. Carpus unarmed. Palm not strongly inflated, dactylus distinctly shorter than palm; narrow hiatus between fingers. Two distinct rows of granules on the palm.
</paragraph>
<caption id="DF29666DE319FFE41CBEF865FBB8F826" ID-DOI="http://doi.org/10.5281/zenodo.12681779" ID-Zenodo-Dep="12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="23" pageNumber="290" startId="23.[151,250,1943,1967]" targetBox="[158,1431,186,1914]" targetPageId="23" targetType="figure">
<paragraph id="8BE936E5E319FFE41CBEF865FBB8F826" blockId="23.[151,1436,1943,2040]" pageId="23" pageNumber="290">
<emphasis id="B922EAF7E319FFE41CBEF865FE80F87E" bold="true" box="[151,289,1943,1967]" pageId="23" pageNumber="290">FIGURE 14.</emphasis>
<taxonomicName id="4C564D66E319FFE41D00F86AFDA8F861" authorityName="Mazancourt &amp; Mazel &amp; Marquet &amp; Poupin &amp; Keith" authorityYear="2024" box="[297,521,1944,1968]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="290" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B922EAF7E319FFE41D00F86AFDA8F861" box="[297,521,1944,1968]" italics="true" pageId="23" pageNumber="290">Ptychognathus similis</emphasis>
</taxonomicName>
<emphasis id="B922EAF7E319FFE41E38F86AFDC2F861" bold="true" box="[529,611,1944,1968]" pageId="23" pageNumber="290">
<taxonomicNameLabel id="A211578CE319FFE41E38F86AFDC2F861" box="[529,611,1944,1968]" pageId="23" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna.
</paragraph>
</caption>
<caption id="DF29666DE316FFEB1CBEFBC1FC3EFBA1" ID-DOI="http://doi.org/10.5281/zenodo.12681781" ID-Zenodo-Dep="12681781" httpUri="https://zenodo.org/record/12681781/files/figure.png" pageId="24" pageNumber="291" startId="24.[151,250,1075,1099]" targetBox="[152,1417,181,1052]" targetPageId="24" targetType="figure">
<paragraph id="8BE936E5E316FFEB1CBEFBC1FC3EFBA1" blockId="24.[151,1436,1075,1136]" pageId="24" pageNumber="291">
<emphasis id="B922EAF7E316FFEB1CBEFBC1FEBEFB9D" bold="true" box="[151,287,1075,1100]" pageId="24" pageNumber="291">FIGURE 15.</emphasis>
<taxonomicName id="4C564D66E316FFEB1D0DFBC6FDA0FB9D" authorityName="Mazancourt &amp; Mazel &amp; Marquet &amp; Poupin &amp; Keith" authorityYear="2024" box="[292,513,1076,1100]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="291" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B922EAF7E316FFEB1D0DFBC6FDA0FB9D" box="[292,513,1076,1100]" italics="true" pageId="24" pageNumber="291">Ptychognathus similis</emphasis>
</taxonomicName>
<emphasis id="B922EAF7E316FFEB1E2EFBC6FDF7FB9D" bold="true" box="[519,598,1076,1100]" pageId="24" pageNumber="291">
<taxonomicNameLabel id="A211578CE316FFEB1E2EFBC6FDF7FB9D" box="[519,598,1076,1100]" pageId="24" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, B, dorsal view of specimens from Futuna; C, typical habitat of species, Vainifao River, Futuna. Photo credits: J. Poupin (A), P. Keith (B) and A. Dutartre (C).
</paragraph>
</caption>
<paragraph id="8BE936E5E316FFEB1CEEFB6CFB42FA2D" blockId="24.[151,1437,1182,2001]" pageId="24" pageNumber="291">
Ambulatory legs (P2–5), slender, somewhat compressed laterally; P4 longest (male: length 1.4–1.6 times carapace width, female: length 1.5 times carapace width); P5 shortest. Merus elongated, subrectangular in lateral view, unarmed; anterior margins of merus of P2–4 with long soft setae; posterior margins with sparsely long setae. Anterior and posterior margins of carpus glabrous. Propodus with dense long setae distally on posterior margins and glabrous on anterior margins. Dactylus terminating in subacute corneous claw, with dense long setae on posterior margins and anterior margins glabrous; ventral surface with rows of short setae. P3 merus (
<figureCitation id="136D2A60E316FFEB18AEFAA0FB46FABD" box="[1159,1255,1362,1388]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="24" pageNumber="291">Fig. 14F</figureCitation>
)
<quantity id="4CAE9B00E316FFEB18DCFAA0FA95FABD" box="[1269,1332,1362,1388]" metricMagnitude="-2" metricUnit="m" metricValue="7.366" pageId="24" pageNumber="291" unit="in" value="2.9">2.9 in</quantity>
<typeStatus id="54ED8847E316FFEB1913FAA0FA3DFABD" box="[1338,1436,1362,1388]" pageId="24" pageNumber="291" type="holotype">holotype</typeStatus>
(
<quantity id="4CAE9B00E316FFEB1CB6FA84FEB0FA41" box="[159,273,1398,1424]" metricMagnitude="-2" metricUnit="m" metricValue="7.366" metricValueMax="8.128" metricValueMin="6.604" pageId="24" pageNumber="291" unit="in" value="2.9" valueMax="3.2" valueMin="2.6">2.6–3.2 in</quantity>
<typeStatus id="54ED8847E316FFEB1D31FA84FE24FA41" box="[280,389,1398,1424]" pageId="24" pageNumber="291" type="paratype">paratypes</typeStatus>
) as long as wide, carpus 2.4 (1.7–2.4) as long as wide, propodus 1.9 (2.0–2.7) as long as wide dactylus 6.3 (4.4–5.4) as long as wide, dactylus 0.8 (0.8–0.9) times as long as propodus. P5 merus (
<figureCitation id="136D2A60E316FFEB18C0FA68FAEFFA65" box="[1257,1358,1434,1460]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="24" pageNumber="291">Fig. 14G</figureCitation>
)
<quantity id="4CAE9B00E316FFEB1974FA68FA3DFA65" box="[1373,1436,1434,1460]" metricMagnitude="-2" metricUnit="m" metricValue="5.588" pageId="24" pageNumber="291" unit="in" value="2.2">2.2 in</quantity>
<typeStatus id="54ED8847E316FFEB1CBEFA4CFF58FA09" box="[151,249,1470,1496]" pageId="24" pageNumber="291" type="holotype">holotype</typeStatus>
(
<quantity id="4CAE9B00E316FFEB1D2EFA4CFED9FA09" box="[263,376,1470,1496]" metricMagnitude="-2" metricUnit="m" metricValue="5.969" metricValueMax="6.604" metricValueMin="5.334" pageId="24" pageNumber="291" unit="in" value="2.35" valueMax="2.6" valueMin="2.1">2.1–2.6 in</quantity>
<typeStatus id="54ED8847E316FFEB1D57FA4CFE4AFA09" box="[382,491,1470,1496]" pageId="24" pageNumber="291" type="paratype">paratypes</typeStatus>
) as long as wide, carpus 2.0 (1.9–2.3) as long as wide; propodus 1.3 (1.2–1.5) as long as wide dactylus 3.4 (2.2–2.7) 4 as long as wide, dactylus 1.4 (0.9–1.1) times as long as propodus.
</paragraph>
<paragraph id="8BE936E5E316FFEB1CEEF9F4FE80F95D" blockId="24.[151,1437,1182,2001]" pageId="24" pageNumber="291">
Male pleon (
<figureCitation id="136D2A60E316FFEB1D7DF9F4FE18F9F1" box="[340,441,1542,1568]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="24" pageNumber="291">Fig. 14H</figureCitation>
) narrow, broadly triangular, nearly smooth, distinctly narrowed at base of telson; margins without short setae. Telson linguiform, margins without setae, terminal margin broadly rounded; length 1.2 (
<typeStatus id="54ED8847E316FFEB1906F9D8FA35F995" box="[1327,1428,1578,1604]" pageId="24" pageNumber="291" type="holotype">holotype</typeStatus>
) (
<quantity id="4CAE9B00E316FFEB1C89F9BCFF43F9B9" box="[160,226,1614,1640]" metricMagnitude="-2" metricUnit="m" metricValue="3.302" pageId="24" pageNumber="291" unit="in" value="1.3">1.3 in</quantity>
<typeStatus id="54ED8847E316FFEB1CC3F9BCFEF6F9B9" box="[234,343,1614,1640]" pageId="24" pageNumber="291" type="paratype">paratypes</typeStatus>
) times basal width, 1.3 (1.4) times long as sixth somite, sixth somite 1.5 (1.3) times longer than fifth somite.
</paragraph>
<paragraph id="8BE936E5E316FFEB1CEEF964FF5EF8CD" blockId="24.[151,1437,1182,2001]" pageId="24" pageNumber="291">
Female pleon (
<figureCitation id="136D2A60E316FFEB1D47F964FE6BF961" box="[366,458,1686,1712]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="24" pageNumber="291">Fig. 14J</figureCitation>
) very broad, covering most of thoracic sternum, fourth somite broadest. Telson broadly subtriangular, terminal margin broadly rounded, length 0.42 times basal width, 1.0 times longer than sixth somite. Sixth somite 1.2–1.3 times longer than fifth somite, measured on midline. Small round eggs
<quantity id="4CAE9B00E316FFEB18FFF92CFADDF929" box="[1238,1404,1758,1784]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="24" pageNumber="291" unit="mm" value="0.27" valueMax="0.28" valueMin="0.26">0.26–0.28 mm</quantity>
in diameter.
</paragraph>
<paragraph id="8BE936E5E316FFEB1CEEF8D4FEABF859" blockId="24.[151,1437,1182,2001]" pageId="24" pageNumber="291">
Male G1 (
<figureCitation id="136D2A60E316FFEB1D10F8D4FE35F891" box="[313,404,1830,1856]" captionStart="FIGURE 14" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[158,1431,186,1914]" captionTargetId="figure-19@23.[152,1436,181,1919]" captionTargetPageId="23" captionText="FIGURE 14. Ptychognathus similis sp. nov.: A, carapace; B, epistome, C, left third maxilliped; D, male cheliped; E, female cheliped; F, third pereiopod; G, fifth pereiopod; H, male pleon; I, left first gonopod; J, female pleon. Holotype MNHN-IU-2022- 4125 (A–D, F–I) male; paratype MNHN-IU-2022-4126 (E, J) female, both from Futuna." figureDoi="http://doi.org/10.5281/zenodo.12681779" httpUri="https://zenodo.org/record/12681779/files/figure.png" pageId="24" pageNumber="291">Fig. 14I</figureCitation>
) slender, 5 times as long as wide, slightly curving dorsally reaching suture between thoracic sternite 4 and 5; tip bluntly rounded, forming short chitinous structure; subdistal lobe obtuse, partially obscured by stiff setae.
</paragraph>
<paragraph id="8BE936E5E316FFEB1CEEF860FF77F800" blockId="24.[151,1437,1182,2001]" pageId="24" pageNumber="291">
<emphasis id="B922EAF7E316FFEB1CEEF860FE08F87D" bold="true" box="[199,425,1938,1964]" pageId="24" pageNumber="291">Colouration in life.</emphasis>
Body and chelipeds (
<figureCitation id="136D2A60E316FFEB1EB5F860FCA0F87C" box="[668,769,1938,1965]" captionStart="FIGURE 15" captionStartId="24.[151,250,1075,1099]" captionTargetBox="[152,1417,181,1052]" captionTargetId="figure-26@24.[151,1436,181,1052]" captionTargetPageId="24" captionText="FIGURE 15. Ptychognathus similis sp. nov.:A, B, dorsal view of specimens from Futuna; C, typical habitat of species, Vainifao River, Futuna. Photo credits: J. Poupin (A), P. Keith (B) and A. Dutartre (C)." figureDoi="http://doi.org/10.5281/zenodo.12681781" httpUri="https://zenodo.org/record/12681781/files/figure.png" pageId="24" pageNumber="291">Fig. 15A</figureCitation>
) yellowish to brownish or greyish, sometimes reddish (
<figureCitation id="136D2A60E316FFEB1946F860FF68F800" captionStart="FIGURE 15" captionStartId="24.[151,250,1075,1099]" captionTargetBox="[152,1417,181,1052]" captionTargetId="figure-26@24.[151,1436,181,1052]" captionTargetPageId="24" captionText="FIGURE 15. Ptychognathus similis sp. nov.:A, B, dorsal view of specimens from Futuna; C, typical habitat of species, Vainifao River, Futuna. Photo credits: J. Poupin (A), P. Keith (B) and A. Dutartre (C)." figureDoi="http://doi.org/10.5281/zenodo.12681781" httpUri="https://zenodo.org/record/12681781/files/figure.png" pageId="24" pageNumber="291">Fig. 15B</figureCitation>
).
</paragraph>
<paragraph id="8BE936E5E317FFEA1CEEFF64FEC9FF04" blockId="25.[151,1436,150,501]" pageId="25" pageNumber="292">
<emphasis id="B922EAF7E317FFEA1CEEFF64FF5FFF61" bold="true" box="[199,254,150,176]" pageId="25" pageNumber="292">Size.</emphasis>
Small-sized. The largest male specimen examined is
<quantity id="4CAE9B00E317FFEA1F72FF64FC13FF60" box="[859,946,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="25" pageNumber="292" unit="mm" value="9.3">9.3 mm</quantity>
×
<quantity id="4CAE9B00E317FFEA1FF8FF64FB86FF60" box="[977,1063,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="25" pageNumber="292" unit="mm" value="7.6">7.6 mm</quantity>
and the largest ovigerous female
<quantity id="4CAE9B00E317FFEA1CBEFF48FF4CFF04" box="[151,237,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="25" pageNumber="292" unit="mm" value="7.9">7.9 mm</quantity>
×
<quantity id="4CAE9B00E317FFEA1D22FF48FEC5FF04" box="[267,356,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="25" pageNumber="292" unit="mm" value="7.6">7.6 mm</quantity>
.
</paragraph>
<paragraph id="8BE936E5E317FFEA1CEEFF2CFA99FF28" blockId="25.[151,1436,150,501]" box="[199,1336,222,249]" pageId="25" pageNumber="292">
<emphasis id="B922EAF7E317FFEA1CEEFF2CFE8BFF29" bold="true" box="[199,298,222,248]" pageId="25" pageNumber="292">Habitat.</emphasis>
On
<collectingCountry id="F3417675E317FFEA1D73FF2DFE57FF29" box="[346,502,222,249]" name="Futuna Island" pageId="25" pageNumber="292">Futuna Island</collectingCountry>
, this species is found in rivers in brackish water, near estuaries (
<figureCitation id="136D2A60E317FFEA18EDFF2CFA8AFF28" box="[1220,1323,222,249]" captionStart="FIGURE 15" captionStartId="24.[151,250,1075,1099]" captionTargetBox="[152,1417,181,1052]" captionTargetId="figure-26@24.[151,1436,181,1052]" captionTargetPageId="24" captionText="FIGURE 15. Ptychognathus similis sp. nov.:A, B, dorsal view of specimens from Futuna; C, typical habitat of species, Vainifao River, Futuna. Photo credits: J. Poupin (A), P. Keith (B) and A. Dutartre (C)." figureDoi="http://doi.org/10.5281/zenodo.12681781" httpUri="https://zenodo.org/record/12681781/files/figure.png" pageId="25" pageNumber="292">Fig. 15C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C656EE317FFEA1CEEFEF0FF53FE91" pageId="25" pageNumber="292" type="distribution">
<paragraph id="8BE936E5E317FFEA1CEEFEF0FF53FE91" blockId="25.[151,1436,150,501]" pageId="25" pageNumber="292">
<emphasis id="B922EAF7E317FFEA1CEEFEF0FEC0FECD" bold="true" box="[199,353,258,284]" pageId="25" pageNumber="292">Distribution.</emphasis>
<taxonomicName id="4C564D66E317FFEA1D4FFEF0FDFBFECD" authorityName="Mazancourt &amp; Mazel &amp; Marquet &amp; Poupin &amp; Keith" authorityYear="2024" box="[358,602,258,284]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="292" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B922EAF7E317FFEA1D4FFEF0FDFBFECD" box="[358,602,258,284]" italics="true" pageId="25" pageNumber="292">Ptychognathus similis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211578CE317FFEA1E49FEF1FD14FECC" box="[608,693,259,285]" pageId="25" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
seems to be endemic to
<collectingCountry id="F3417675E317FFEA1F94FEF1FBF5FECD" box="[957,1108,258,285]" name="Futuna Island" pageId="25" pageNumber="292">Futuna Island</collectingCountry>
(
<collectingCountry id="F3417675E317FFEA1848FEF0FA8AFECC" box="[1121,1323,258,285]" name="Wallis and Futuna" pageId="25" pageNumber="292">
Wallis and
<taxonomicName id="4C564D66E317FFEA18F4FEF1FA8AFECC" authority="Territory" authorityName="Territory" box="[1245,1323,259,285]" family="Gobiidae" genus="Akihito" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="292" phylum="Chordata" rank="species" species="futuna">Futuna</taxonomicName>
</collectingCountry>
Territory) (
<figureCitation id="136D2A60E317FFEA1CB6FED4FF45FE91" box="[159,228,294,320]" captionStart="FIGURE 6" captionStartId="9.[151,250,1462,1486]" captionTargetBox="[154,1434,183,1438]" captionTargetId="figure-19@9.[151,1436,181,1441]" captionTargetPageId="9" captionText="FIGURE 6. Distribution map of the species studied. Stars indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.12681762" httpUri="https://zenodo.org/record/12681762/files/figure.png" pageId="25" pageNumber="292">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C656EE317FFEA1CEEFEB8FBE1FEB4" box="[199,1088,330,357]" pageId="25" pageNumber="292" type="etymology">
<paragraph id="8BE936E5E317FFEA1CEEFEB8FBE1FEB4" blockId="25.[151,1436,150,501]" box="[199,1088,330,357]" pageId="25" pageNumber="292">
<emphasis id="B922EAF7E317FFEA1CEEFEB8FEEFFEB5" bold="true" box="[199,334,330,356]" pageId="25" pageNumber="292">Etymology.</emphasis>
This new species is named after its resemblance to
<taxonomicName id="4C564D66E317FFEA1FBBFEB9FB98FEB5" authorityName="Sakai" authorityYear="1955" box="[914,1081,330,356]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="292" phylum="Arthropoda" rank="species" species="hachijoensis">
<emphasis id="B922EAF7E317FFEA1FBBFEB9FB98FEB5" box="[914,1081,330,356]" italics="true" pageId="25" pageNumber="292">P. hachijoensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C656EE317FFEA1CEEFE9CFAB9FE25" pageId="25" pageNumber="292" type="discussion">
<paragraph id="8BE936E5E317FFEA1CEEFE9CFAB9FE25" blockId="25.[151,1436,150,501]" pageId="25" pageNumber="292">
<emphasis id="B922EAF7E317FFEA1CEEFE9CFE9AFE59" bold="true" box="[199,315,366,392]" pageId="25" pageNumber="292">Remarks.</emphasis>
By the anterolateral margins of its carapace having a single orbital tooth followed by a faint lobe, and by the propodus and dactylus of its ambulatory legs having dense, long setae on the anterior margins,
<taxonomicName id="4C564D66E317FFEA18DFFE60FF41FE01" authorityName="Mazancourt &amp; Mazel &amp; Marquet &amp; Poupin &amp; Keith" authorityYear="2024" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="292" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B922EAF7E317FFEA18DFFE60FF41FE01" italics="true" pageId="25" pageNumber="292">Ptychognathus similis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211578CE317FFEA1CC1FE45FE9EFE00" box="[232,319,439,465]" pageId="25" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C564D66E317FFEA1E24FE45FCE5FE00" authority="Sakai, 1955" authorityName="Sakai" authorityYear="1955" box="[525,836,438,465]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="292" phylum="Arthropoda" rank="species" species="hachijoensis">
<emphasis id="B922EAF7E317FFEA1E24FE45FD14FE01" box="[525,693,438,464]" italics="true" pageId="25" pageNumber="292">P. hachijoensis</emphasis>
<bibRefCitation id="EFC74B14E317FFEA1E94FE44FCE5FE00" author="Sakai, T." box="[701,836,438,465]" pageId="25" pageNumber="292" pagination="193 - 201" refId="ref19642" refString="Sakai, T. (1955) Further Notes on the Brachyuran Crustacea of the Hachijo Island. Records of Oceanographic. Works in Japan, 2 (1), 193 - 201." type="journal article" year="1955">Sakai, 1955</bibRefCitation>
</taxonomicName>
, but it is possible to distinguish them by the width of front (0.45–0.47 times CW vs 0.55–0.56) and by its P5 dactylus (0.8 times as long as propodus vs 1.2).
</paragraph>
</subSubSection>
</treatment>
</document>
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