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<document id="8993E469FB1FC754DE3AFCAA27E32F9F" ID-DOI="10.5281/zenodo.5299545" ID-ISSN="0374-1036" ID-Zenodo-Dep="5299545" ID-ZooBank="D94600BD-1221-47B6-9C70-BA8C82B74CEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630093724692" checkinUser="marcus" docAuthor="Janák, JiĜí" docDate="2014" docId="03CD1A4EFFDDFE0ABBCCFDF8A061FD69" docLanguage="en" docName="ActaEntMusNatPra.2014.54.1.195-231.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Octavius muellerae Janák 2014, sp. nov." docType="treatment" docVersion="4" lastPageNumber="200" masterDocId="FFF46236FFD9FE0FBA2BFF83A27AFF9A" masterDocTitle="New species and subspecies of Octavius from South Africa, with a key and additional distribution records (Coleoptera: Staphylinidae: Euaesthetinae)" masterLastPageNumber="231" masterPageNumber="195" pageNumber="199" updateTime="1699142974711" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="0E130A1072CD3906E040C4983A8CF2E9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9778D3FE795BA2CF3D37F4AEF4EABF3F">
<mods:title id="03CBCFEA6E30586AE57362FD8CE5C1C2">New species and subspecies of Octavius from South Africa, with a key and additional distribution records (Coleoptera: Staphylinidae: Euaesthetinae)</mods:title>
</mods:titleInfo>
<mods:name id="467473DFAC6A90D146633D4A55E06E42" type="personal">
<mods:role id="5A261F36071BFEF22ECC4D48AC95CFCF">
<mods:roleTerm id="1D28A173C742B0FD39FD7F79C3140EEC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5DF61533499F63711E9ADD2DF29ED99F">Janák, JiĜí</mods:namePart>
</mods:name>
<mods:typeOfResource id="5599A3B143B0451F68F21427764FEDBE">text</mods:typeOfResource>
<mods:relatedItem id="6C2513B2887EC1D37C0B98E316DBE238" type="host">
<mods:titleInfo id="6B45DCC6DF02495C2FC16A8E5CD27048">
<mods:title id="31C3D62E1D57311205B282165426E9C3">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="926A7EE744F7BE2CB9D6FFFAC8EC3DF5">
<mods:date id="47C34A92CDAB1F6EE6F53EB72A97EABE">2014</mods:date>
<mods:detail id="CD7B554AA60F9D13925C09EEF0207BE0" type="pubDate">
<mods:number id="7F735E9DB1AD8DECB5805CB66CD8C6B2">2014-04-30</mods:number>
</mods:detail>
<mods:detail id="95D5F235B626B120477E33A982E2E4AA" type="volume">
<mods:number id="74A232EBB9EA6411DB97319B85870B6D">54</mods:number>
</mods:detail>
<mods:detail id="4C9DD343D06ACC30A51E1017EAE6C559" type="issue">
<mods:number id="3A7E64A630CE74ED9473170621B4B9BD">1</mods:number>
</mods:detail>
<mods:extent id="E8D94CBF3DA011F34C052120FD062831" unit="page">
<mods:start id="38225ED8D4EB0E5FDF7C0510AD6A76B5">195</mods:start>
<mods:end id="97B74D28D9BF51B6AA7A95602BB5B6A0">231</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="89DDE45D69242BA60F8DCB984F55D072">journal article</mods:classification>
<mods:identifier id="441049DC95E0A7F69B5D1D3A87101B1E" type="DOI">10.5281/zenodo.5299545</mods:identifier>
<mods:identifier id="7C274381D7A3A42BDB186B6650767F60" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="3A15E5E0937DE0AF30AB785D177BEC1F" type="Zenodo-Dep">5299545</mods:identifier>
<mods:identifier id="F552E43D44A8EF9C124F503F1CEECD0F" type="ZooBank">D94600BD-1221-47B6-9C70-BA8C82B74CEC</mods:identifier>
</mods:mods>
<treatment id="03CD1A4EFFDDFE0ABBCCFDF8A061FD69" ID-DOI="http://doi.org/10.5281/zenodo.5306997" ID-Zenodo-Dep="5306997" LSID="urn:lsid:plazi:treatment:03CD1A4EFFDDFE0ABBCCFDF8A061FD69" httpUri="http://treatment.plazi.org/id/03CD1A4EFFDDFE0ABBCCFDF8A061FD69" lastPageId="5" lastPageNumber="200" pageId="4" pageNumber="199">
<subSubSection id="C37EF8D3FFDDFE0BBBCCFDF8A16DFD09" box="[487,791,635,659]" pageId="4" pageNumber="199" type="nomenclature">
<paragraph id="8BDBAB58FFDDFE0BBBCCFDF8A16DFD09" blockId="4.[487,791,635,659]" box="[487,791,635,659]" pageId="4" pageNumber="199">
<heading id="D0931C34FFDDFE0BBBCCFDF8A16DFD09" bold="true" box="[487,791,635,659]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="199" reason="3">
<emphasis id="B910774AFFDDFE0BBBCCFDF8A16DFD09" bold="true" box="[487,791,635,659]" pageId="4" pageNumber="199">
<taxonomicName id="4C64D0DBFFDDFE0BBBCCFDF8A0C6FD09" ID-CoL="48JLK" authority="Janák, 2014" authorityName="Janák" authorityYear="2014" box="[487,700,635,659]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B910774AFFDDFE0BBBCCFDF8A0C6FD09" bold="true" box="[487,700,635,659]" italics="true" pageId="4" pageNumber="199">Octavius muellerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223CA31FFDDFE0BB8E9FDFFA16DFD09" box="[706,791,636,659]" pageId="4" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37EF8D3FFDDFE0BB878FD19A0D1FD37" box="[595,683,666,685]" pageId="4" pageNumber="199" type="description">
<paragraph id="8BDBAB58FFDDFE0BB878FD19A0D1FD37" blockId="4.[595,683,666,685]" box="[595,683,666,685]" pageId="4" pageNumber="199">
(
<figureCitation id="135FB7DDFFDDFE0BB872FD19A0DEFD37" box="[601,676,666,685]" captionStart="Figs" captionStartId="3.[113,152,1565,1584]" captionTargetBox="[140,1068,235,1462]" captionTargetId="figure-13@3.[128,1074,219,1466]" captionTargetPageId="3" captionText="Figs. 4–9. 4–6 – Octavius mostovskii sp. nov., HT male. 7–8 – O. muellerae sp. nov., HT male. 4, 7 – aedeagus ventral; 5, 8 – sternite 8; 6, 9 – sternite 9. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5299549" httpUri="https://zenodo.org/record/5299549/files/figure.png" pageId="4" pageNumber="199">Figs 7–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37EF8D3FFDDFE0BBABCFD47A160FCC6" pageId="4" pageNumber="199" type="materials_examined">
<paragraph id="8BDBAB58FFDDFE0BBABCFD47A1EAFD4D" blockId="4.[151,1127,708,861]" box="[151,912,708,727]" pageId="4" pageNumber="199">
<emphasis id="B910774AFFDDFE0BBABCFD47A36AFD4D" bold="true" box="[151,272,708,727]" pageId="4" pageNumber="199">
<typeStatus id="54DF15FAFFDDFE0BBABCFD47A2BEFD4D" box="[151,196,708,727]" pageId="4" pageNumber="199">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F373EBC8FFDDFE0BBB3EFD47A3FEFD4D" box="[277,388,708,727]" name="South Africa" pageId="4" pageNumber="199">South Africa</collectingCountry>
,
<collectingRegion id="49A065BAFFDDFE0BBBA7FD47A01FFD4D" box="[396,613,708,727]" country="South Africa" name="KwaZulu-Natal" pageId="4" pageNumber="199">KwaZulu-Natal Province</collectingRegion>
, Ongoye Forest, 28°50ƍS, 31°44ƍE.
</paragraph>
<paragraph id="8BDBAB58FFDDFE0BBABCFD5CA1A2FCBD" blockId="4.[151,1127,708,861]" pageId="4" pageNumber="199">
<emphasis id="B910774AFFDDFE0BBABCFD5CA366FD68" bold="true" box="[151,284,735,754]" pageId="4" pageNumber="199">Type material.</emphasis>
<materialsCitation id="3B0CA105FFDDFE0BBB0AFD5CA344FCBD" collectingDate="2010-12-04" collectingDateMax="2010-12-05" collectingDateMin="2010-12-04" collectionCode="TMSA" collectorName="Ruth Muller" country="South Africa" determinerName="J. Janak" elevation="294" latitude="-28.5" location="Ongoye forest" longLatPrecision="739" longitude="31.44" municipality="Natal" pageId="4" pageNumber="199" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<typeStatus id="54DF15FAFFDDFE0BBB0AFD5CA30FFD6A" box="[289,373,735,754]" pageId="4" pageNumber="199" type="holotype">HOLOTYPE</typeStatus>
: 3: ‘
<collectingCountry id="F373EBC8FFDDFE0BBBB5FD5CA044FD68" box="[414,574,735,754]" name="South Africa" pageId="4" pageNumber="199">SOUTH AFRICA</collectingCountry>
, [
<collectingRegion id="49A065BAFFDDFE0BB860FD5CA150FD68" box="[587,810,735,754]" country="South Africa" name="KwaZulu-Natal" pageId="4" pageNumber="199">KwaZulu-Natal Province</collectingRegion>
]: KWZ
<collectingMunicipality id="6BBF3122FFDDFE0BB945FD5CA1E4FD68" box="[878,926,735,754]" pageId="4" pageNumber="199">Natal</collectingMunicipality>
,
<location id="8EBBFD83FFDDFE0BB98CFD5CA659FD68" LSID="urn:lsid:plazi:treatment:03CD1A4EFFDDFE0ABBCCFDF8A061FD69:8EBBFD83FFDDFE0BB98CFD5CA659FD68" box="[935,1059,735,754]" country="South Africa" latitude="-28.5" longLatPrecision="739" longitude="31.44" municipality="Natal" name="Ongoye forest" pageId="4" pageNumber="199" stateProvince="KwaZulu-Natal">Ongoye forest</location>
,
<quantity id="4C9C06BDFFDDFE0BBE00FD5CA61EFD68" box="[1067,1124,735,754]" metricMagnitude="2" metricUnit="m" metricValue="2.94" pageId="4" pageNumber="199" unit="m" value="294.0">
<elevation id="00494C6BFFDDFE0BBE00FD5CA61EFD68" box="[1067,1124,735,754]" metricMagnitude="2" metricUnit="m" metricValue="2.94" pageId="4" pageNumber="199" unit="m" value="294.0">294 m</elevation>
</quantity>
,
<geoCoordinate id="EE50CD9FFFDDFE0BBABCFD7AA2ADFC96" box="[151,215,761,780]" degrees="28.50" direction="south" orientation="latitude" pageId="4" pageNumber="199" precision="555" value="-28.5">28.50 S</geoCoordinate>
–
<geoCoordinate id="EE50CD9FFFDDFE0BBAC1FD7AA356FC96" box="[234,300,761,780]" degrees="31.44" direction="east" orientation="longitude" pageId="4" pageNumber="199" precision="555" value="31.44">31.44 E</geoCoordinate>
, sifting forest litter,
<date id="FFDA8D98FFDDFE0BBBF8FD7AA03CFC96" box="[467,582,761,780]" pageId="4" pageNumber="199" value="2010-12-04" valueMax="2010-12-05" valueMin="2010-12-04">
<collectingDate id="EF9E7470FFDDFE0BBBF8FD7AA03CFC96" box="[467,582,761,780]" pageId="4" pageNumber="199" value="2010-12-04" valueMax="2010-12-05" valueMin="2010-12-04">4.–5.xii.2010</collectingDate>
</date>
, E-Y 3890, leg.
<collectorName id="2691CE8EFFDDFE0BB8E0FD7AA149FC96" box="[715,819,761,780]" pageId="4" pageNumber="199">Ruth Müller</collectorName>
//
<taxonomicName id="4C64D0DBFFDDFE0BB96DFD7AA191FC96" authority="Janák, 2014" authorityName="Janák" authorityYear="2014" box="[838,1003,761,780]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B910774AFFDDFE0BB96DFD7AA191FC96" box="[838,1003,761,780]" italics="true" pageId="4" pageNumber="199">Octavius muellerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223CA31FFDDFE0BB9C4FD7AA667FC96" box="[1007,1053,761,780]" pageId="4" pageNumber="199" rank="species">sp. n.</taxonomicNameLabel>
,
<determinerName id="7E07479CFFDDFE0BBE0FFD7AA61CFC96" box="[1060,1126,761,780]" pageId="4" pageNumber="199">J. Janák</determinerName>
det. 2013’ (
<collectionCode id="ED75339DFFDDFE0BBAD3FC97A342FCBD" box="[248,312,788,807]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="199">TMSA</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54DF15FAFFDDFE0BBB6DFC97A3E6FCBF" box="[326,412,788,807]" pageId="4" pageNumber="199" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D6260D1FFDDFE0BBB83FC97A3AEFCB2" box="[424,468,785,808]" pageId="4" pageNumber="199" type="female">2 ♀♀</specimenCount>
: same data as
<typeStatus id="54DF15FAFFDDFE0BB87FFC97A0E5FCBD" box="[596,671,788,807]" pageId="4" pageNumber="199" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D6260D1FFDDFE0BB887FC97A092FCBD" box="[684,744,788,807]" pageId="4" pageNumber="199" type="generic">1 spec.</specimenCount>
in
<collectionCode id="ED75339DFFDDFE0BB92AFC97A144FCBD" box="[769,830,788,807]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="199">TMSA</collectionCode>
,
<specimenCount id="9D6260D1FFDDFE0BB96CFC97A1FEFCBD" box="[839,900,788,807]" pageId="4" pageNumber="199" type="generic">1 spec.</specimenCount>
in
<collectionCode id="ED75339DFFDDFE0BB9B5FC97A1B5FCBD" box="[926,975,788,807]" pageId="4" pageNumber="199">JJRC</collectionCode>
).
</paragraph>
<paragraph id="8BDBAB58FFDDFE0BBABCFCACA160FCC6" blockId="4.[151,1127,708,861]" pageId="4" pageNumber="199">
<emphasis id="B910774AFFDDFE0BBABCFCACA087FCD8" bold="true" box="[151,765,815,834]" pageId="4" pageNumber="199">
Additional material examined.
<collectingCountry id="F373EBC8FFDDFE0BBB98FCACA022FCD8" box="[435,600,815,834]" name="South Africa" pageId="4" pageNumber="199">SOUTH AFRICA</collectingCountry>
: KWAZULU- NATAL:
</emphasis>
Ongoye forest,
<geoCoordinate id="EE50CD9FFFDDFE0BB9ACFCACA1B2FCD8" box="[903,968,815,834]" degrees="28.50" direction="south" orientation="latitude" pageId="4" pageNumber="199" precision="555" value="-28.5">28.50 S</geoCoordinate>
–
<geoCoordinate id="EE50CD9FFFDDFE0BB9F5FCACA658FCD8" box="[990,1058,815,834]" degrees="31.44" direction="east" orientation="longitude" pageId="4" pageNumber="199" precision="555" value="31.44">31.44 E</geoCoordinate>
,
<quantity id="4C9C06BDFFDDFE0BBE00FCACA619FCD8" box="[1067,1123,815,834]" metricMagnitude="2" metricUnit="m" metricValue="2.94" pageId="4" pageNumber="199" unit="m" value="294.0">294 m</quantity>
, sifting forest litter,
<date id="FFDA8D98FFDDFE0BBB10FCCAA3D4FCC6" box="[315,430,841,860]" pageId="4" pageNumber="199" value="2010-12-04" valueMax="2010-12-05" valueMin="2010-12-04">4.–5.xii.2010</date>
, E-Y 3890, leg. Ruth Müller,
<specimenCount id="9D6260D1FFDDFE0BB887FCCAA0B0FCC7" box="[684,714,838,861]" pageId="4" pageNumber="199" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED75339DFFDDFE0BB8FCFCCAA16FFCC6" box="[727,789,841,860]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="199">TMSA</collectionCode>
)
</paragraph>
</subSubSection>
<subSubSection id="C37EF8D3FFDDFE0BBABCFCF6A3B0F9D6" pageId="4" pageNumber="199" type="description">
<paragraph id="8BDBAB58FFDDFE0BBABCFCF6A3C4FC56" blockId="4.[151,1127,885,1612]" pageId="4" pageNumber="199">
<emphasis id="B910774AFFDDFE0BBABCFCF6A361FC16" bold="true" box="[151,283,885,908]" pageId="4" pageNumber="199">Description</emphasis>
(n = 3). Body length
<quantity id="4C9C06BDFFDDFE0BBBD6FCF6A006FC16" box="[509,636,885,908]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="4" pageNumber="199" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 mm</quantity>
(M
<quantity id="4C9C06BDFFDDFE0BB882FCF6A087FC16" box="[681,765,885,908]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="199" unit="mm" value="1.3">1.3 mm</quantity>
, HT
<quantity id="4C9C06BDFFDDFE0BB91BFCF6A1FEFC16" box="[816,900,885,908]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="199" unit="mm" value="1.3">1.3 mm</quantity>
), forebody length
<quantity id="4C9C06BDFFDDFE0BBE6EFCF6A2BAFC36" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="199" unit="mm" value="0.6">0.6 mm</quantity>
(HT, PT). Microphthalmous, apterous, light rusty, head and pronotum dull, elytra and abdomen moderately shiny.
</paragraph>
<paragraph id="8BDBAB58FFDDFE0BBA9EFC56A10FFBD6" blockId="4.[151,1127,885,1612]" pageId="4" pageNumber="199">Head slightly narrower than pronotum (R 0.89–0.92, M 0.90, HT 0.89), eyes very small, temples about three times as long as eyes (R 3.00–3.27, M 3.12, HT 3.27), subparallel, posterior angles moderately angular, median impression on disc present, lateral parts of head very ¿nely granulose, median part very densely and ¿nely reticulate.</paragraph>
<paragraph id="8BDBAB58FFDDFE0BBA9EFBD6A3E4FB76" blockId="4.[151,1127,885,1612]" pageId="4" pageNumber="199">Pronotum moderately broader than long (R 1.12–1.17, M 1.09, HT 1.14), strongly narrowed posteriorly; anterior angles slightly angular, dorsal impressions moderately deep, transverse impression deep, lateral impressions deep, but not delimited by a sharp longitudinal ridge laterally; lateral parts beside lateral impressions densely granulose, remainder of surface very densely ¿nely reticulate.</paragraph>
<paragraph id="8BDBAB58FFDDFE0BBA9EFB76A0A2FAD6" blockId="4.[151,1127,885,1612]" pageId="4" pageNumber="199">Elytra subquadrate, much broader than long (R 1.42–1.55, M 1.47, HT 1.45), with a sharp longitudinal ridge laterally; between the latter and suture with a moderate longitudinal undulation, irregularly sculptured, and moderately reticulate.</paragraph>
<paragraph id="8BDBAB58FFDDFE0BBA9EFAD6A3E8FA76" blockId="4.[151,1127,885,1612]" pageId="4" pageNumber="199">
Male. Sternite 8 moderately emarginate in posterior one-¿fteenth (
<figureCitation id="135FB7DDFFDDFE0BB9A1FAD6A1B6FAF6" box="[906,972,1365,1388]" captionStart="Figs" captionStartId="3.[113,152,1565,1584]" captionTargetBox="[140,1068,235,1462]" captionTargetId="figure-13@3.[128,1074,219,1466]" captionTargetPageId="3" captionText="Figs. 4–9. 4–6 – Octavius mostovskii sp. nov., HT male. 7–8 – O. muellerae sp. nov., HT male. 4, 7 – aedeagus ventral; 5, 8 – sternite 8; 6, 9 – sternite 9. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5299549" httpUri="https://zenodo.org/record/5299549/files/figure.png" pageId="4" pageNumber="199">Fig. 8</figureCitation>
), sternite 9 as in
<figureCitation id="135FB7DDFFDDFE0BBA98FAF6A28FFA16" box="[179,245,1397,1420]" captionStart="Figs" captionStartId="3.[113,152,1565,1584]" captionTargetBox="[140,1068,235,1462]" captionTargetId="figure-13@3.[128,1074,219,1466]" captionTargetPageId="3" captionText="Figs. 4–9. 4–6 – Octavius mostovskii sp. nov., HT male. 7–8 – O. muellerae sp. nov., HT male. 4, 7 – aedeagus ventral; 5, 8 – sternite 8; 6, 9 – sternite 9. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5299549" httpUri="https://zenodo.org/record/5299549/files/figure.png" pageId="4" pageNumber="199">Fig. 9</figureCitation>
. Aedeagus elongate and asymmetrical (length
<quantity id="4C9C06BDFFDDFE0BB8CCFAF6A130FA16" box="[743,842,1397,1420]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="4" pageNumber="199" unit="mm" value="0.41">0.41 mm</quantity>
), pointed apically, internal structure with long, apically pointed plate with several spines and long narrow tube reaching nearly to the apex of median lobe; parameres slightly shorter than median lobe, with about 10 apical setae (
<figureCitation id="135FB7DDFFDDFE0BBB68FA56A3FFFA76" box="[323,389,1493,1516]" captionStart="Figs" captionStartId="3.[113,152,1565,1584]" captionTargetBox="[140,1068,235,1462]" captionTargetId="figure-13@3.[128,1074,219,1466]" captionTargetPageId="3" captionText="Figs. 4–9. 4–6 – Octavius mostovskii sp. nov., HT male. 7–8 – O. muellerae sp. nov., HT male. 4, 7 – aedeagus ventral; 5, 8 – sternite 8; 6, 9 – sternite 9. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5299549" httpUri="https://zenodo.org/record/5299549/files/figure.png" pageId="4" pageNumber="199">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BDBAB58FFDDFE0BBABCFA76A3B0F9D6" blockId="4.[151,1127,885,1612]" pageId="4" pageNumber="199">
<emphasis id="B910774AFFDDFE0BBABCFA76A36CF996" bold="true" box="[151,278,1525,1548]" pageId="4" pageNumber="199">Variability.</emphasis>
The female listed in additional material examined has most characters in variability range of the
<typeStatus id="54DF15FAFFDDFE0BBB71F996A3F2F9B6" box="[346,392,1557,1580]" pageId="4" pageNumber="199">type</typeStatus>
series, but differs from the
<typeStatus id="54DF15FAFFDDFE0BB887F996A09EF9B6" box="[684,740,1557,1580]" pageId="4" pageNumber="199">types</typeStatus>
by larger eyes (R = 2.30). It was not included into the
<typeStatus id="54DF15FAFFDDFE0BBB7AF9B6A305F9D6" box="[337,383,1589,1612]" pageId="4" pageNumber="199">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C37EF8D3FFDCFE0ABA5AFF5FA3A4FDA9" pageId="5" pageNumber="200" type="diagnosis">
<paragraph id="8BDBAB58FFDCFE0ABA5AFF5FA3A4FDA9" blockId="5.[113,1089,220,755]" pageId="5" pageNumber="200">
<emphasis id="B910774AFFDCFE0ABA5AFF5FA311FF69" bold="true" box="[113,363,220,243]" pageId="5" pageNumber="200">Differential diagnosis.</emphasis>
<taxonomicName id="4C64D0DBFFDCFE0ABB5AFF5FA03BFF69" authority="Janák, 2014" authorityName="Janák" authorityYear="2014" box="[369,577,220,243]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B910774AFFDCFE0ABB5AFF5FA03BFF69" box="[369,577,220,243]" italics="true" pageId="5" pageNumber="200">Octavius muellerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223CA31FFDCFE0AB86CFF5FA0E0FF69" box="[583,666,220,243]" pageId="5" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
belongs among species with very small eyes, with temples more than three times as long as eyes and with the head not or at most slightly widened posteriorly. From already described species, it is externally most similar to
<taxonomicName id="4C64D0DBFFDCFE0ABA5AFEBFA30EFEC9" authority="Puthz, 2006" authorityName="Puthz" authorityYear="2006" box="[113,372,316,339]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="ocellifer">
<emphasis id="B910774AFFDCFE0ABA5AFEBFA291FEC9" box="[113,235,316,339]" italics="true" pageId="5" pageNumber="200">O. ocellifer</emphasis>
<bibRefCitation id="EFF5D6A9FFDCFE0ABAD9FEBFA30EFEC9" author="PUTHZ V." box="[242,372,316,339]" pageId="5" pageNumber="200" pagination="1 - 28" refId="ref19005" refString="PUTHZ V. 2006: New southern African species and distribution records of the genus Octavius Fauvel (Coleoptera: Staphylinidae). Annals of the Transvaal Museum 43: 1 - 28." type="journal article" year="2006">Puthz, 2006</bibRefCitation>
</taxonomicName>
, but it differs by slightly larger eyes and more transverse pronotum with deeper dorsal impressions and by the different sexual characters of male.
<taxonomicName id="4C64D0DBFFDCFE0AB99DFEDFA63BFEE9" authority="Janák, 2014" authorityName="Janák" authorityYear="2014" box="[950,1089,348,371]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B910774AFFDCFE0AB99DFEDFA63BFEE9" box="[950,1089,348,371]" italics="true" pageId="5" pageNumber="200">O. muellerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223CA31FFDCFE0ABA5AFEFFA2BFFE09" box="[113,197,380,403]" pageId="5" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
is externally very simillar to
<taxonomicName id="4C64D0DBFFDCFE0AB82FFEFFA0DBFE09" authority="Janák, 2014" authorityName="Janák" authorityYear="2014" box="[516,673,380,403]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="acutipennis" status="sp. nov.">
<emphasis id="B910774AFFDCFE0AB82FFEFFA0DBFE09" box="[516,673,380,403]" italics="true" pageId="5" pageNumber="200">O. acutipennis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223CA31FFDCFE0AB882FEFFA087FE09" box="[681,765,380,403]" pageId="5" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
occuring in the same locality, from which it can be distinguished (sometimes with dif¿culties) by deeper dorsal impressions on pronotum, coarsely reticulate head and pronotum, deeper median impression on head and smaller eyes. The new species differs from that species by the completely different shape of aedeagus with an apically pointed plate with several spines and a long narrow tube reaching nearly to the apex of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C37EF8D3FFDCFE0ABA5AFDBFA655FD29" pageId="5" pageNumber="200" type="etymology">
<paragraph id="8BDBAB58FFDCFE0ABA5AFDBFA07BFD09" blockId="5.[113,1089,220,755]" pageId="5" pageNumber="200">
<emphasis id="B910774AFFDCFE0ABA5AFDBFA288FDC9" bold="true" box="[113,242,572,595]" pageId="5" pageNumber="200">Etymology.</emphasis>
This species is dedicated to Ruth Müller (Ditsong Museum, Pretoria,
<collectingCountry id="F373EBC8FFDCFE0ABE29FDBFA2C0FDE9" name="South Africa" pageId="5" pageNumber="200">South Africa</collectingCountry>
), who helped me signi¿cantly during my trips to the
<collectingCountry id="F373EBC8FFDCFE0AB92AFDDFA668FDE9" box="[769,1042,604,627]" name="South Africa" pageId="5" pageNumber="200">Republic of South Africa</collectingCountry>
and supported my study of
<taxonomicName id="4C64D0DBFFDCFE0ABB4CFDFFA387FD09" box="[359,509,636,659]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="200" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
.
</paragraph>
<paragraph id="8BDBAB58FFDCFE0ABA5AFD1FA655FD29" blockId="5.[113,1089,220,755]" box="[113,1071,668,691]" pageId="5" pageNumber="200">
<emphasis id="B910774AFFDCFE0ABA5AFD1FA295FD29" bold="true" box="[113,239,668,691]" pageId="5" pageNumber="200">Bionomics.</emphasis>
All specimens have been found in siftings of forest litter in indigenous forest.
</paragraph>
</subSubSection>
<subSubSection id="C37EF8D3FFDCFE0ABA5AFD3FA061FD69" pageId="5" pageNumber="200" type="distribution">
<paragraph id="8BDBAB58FFDCFE0ABA5AFD3FA061FD69" blockId="5.[113,1089,220,755]" pageId="5" pageNumber="200">
<emphasis id="B910774AFFDCFE0ABA5AFD3FA37EFD49" bold="true" box="[113,260,700,723]" pageId="5" pageNumber="200">Distribution.</emphasis>
<taxonomicName id="4C64D0DBFFDCFE0ABB22FD3FA3A3FD49" authority="Janák, 2014" authorityName="Janák" authorityYear="2014" box="[265,473,700,723]" class="Insecta" family="Staphylinidae" genus="Octavius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="muellerae" status="sp. nov.">
<emphasis id="B910774AFFDCFE0ABB22FD3FA3A3FD49" box="[265,473,700,723]" italics="true" pageId="5" pageNumber="200">Octavius muellerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223CA31FFDCFE0ABBF4FD3FA04BFD49" box="[479,561,700,723]" pageId="5" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
is currently recorded only from Ongoye Forest in
<collectingRegion id="49A065BAFFDCFE0ABA5AFD5FA3FBFD69" box="[113,385,732,755]" country="South Africa" name="KwaZulu-Natal" pageId="5" pageNumber="200">KwaZulu-Natal Province</collectingRegion>
,
<collectingCountry id="F373EBC8FFDCFE0ABBA7FD5FA06DFD69" box="[396,535,732,755]" name="South Africa" pageId="5" pageNumber="200">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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