238 lines
36 KiB
XML
238 lines
36 KiB
XML
|
<document id="649D97AAE0EB78FCF348F29E1098743E" ID-ISSN="0024-4082" ID-ZooBank="74C07292-2BD6-4E3E-B68D-B144B81BBD83" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629585783401" checkinUser="felipe" docAuthor="Bonifácio, Paulo & Menot, Lénaïck" docDate="2019" docId="0B1B8791FFD10605F967EBDE7F385D50" docLanguage="en" docName="ZoolJLinnSoc.185.555–635.pdf" docOrigin="Zoological Journal of the Linnean Society 185" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Hodor hodor hodor 2019, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="600" masterDocId="F722FFE9FFFA0628F820E81A7D025E58" masterDocTitle="New genera and species from the Equatorial Pacific provide phylogenetic insights into deep-sea Polynoidae (Annelida)" masterLastPageNumber="635" masterPageNumber="555" pageNumber="598" updateTime="1699081480826" updateUser="plazi">
|
|||
|
|
<mods:mods id="B1E7383E680C794701E8218C8A994F6F" xmlns:mods="http://www.loc.gov/mods/v3">
|
|||
|
|
<mods:titleInfo id="436A6BAC376269CDC69A76F0B255A860">
|
|||
|
|
<mods:title id="16A3501536CE1F3BE6CD953E581274D8">New genera and species from the Equatorial Pacific provide phylogenetic insights into deep-sea Polynoidae (Annelida)</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:name id="FFDB21F15D21CC4C572B32FD9209369E" type="personal">
|
|||
|
|
<mods:role id="AA1FE3809A0637F2F217410C38C63601">
|
|||
|
|
<mods:roleTerm id="1ED564981745557F4A18CAD45714D832">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="066DBE9F86FAED70385EDE61C292D713">Bonifácio, Paulo</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name id="BAF44EA88D9185AFC2CDE477BF826489" type="personal">
|
|||
|
|
<mods:role id="D49CB5F597AE004E3830F306A77BCF64">
|
|||
|
|
<mods:roleTerm id="0F7D6F6A954F7823CC89CEA70807145F">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="491C16AC24CBBABF92A581F89357E1F3">Menot, Lénaïck</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:typeOfResource id="DB7F93E7B1845FA76B54AC088A95F9D5">text</mods:typeOfResource>
|
|||
|
|
<mods:relatedItem id="CADDFD8DEB5FE4D365FBE927FE858769" type="host">
|
|||
|
|
<mods:titleInfo id="122B2784C57368CD196803BD6076F693">
|
|||
|
|
<mods:title id="DA1E8588DB6892FEAF0FED1D62B71295">Zoological Journal of the Linnean Society</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:part id="8D5B3CC8B67DFAFCB616D855150CB8EE">
|
|||
|
|
<mods:date id="8BF1FEF3EC3EE7EA56E333827EDC3874">2019</mods:date>
|
|||
|
|
<mods:detail id="B7548F477A4C913D2B13A628B619890B" type="volume">
|
|||
|
|
<mods:number id="2F76A05C13CA227FD101AD8A970FE4BB">185</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:extent id="60B70FA84E55A3CE4D45EA13CEA1EF0E" unit="page">
|
|||
|
|
<mods:start id="F1C3620CBD9191426FE2B33EDDA383E6">555</mods:start>
|
|||
|
|
<mods:end id="37A4CADB15B25B77BDFA119619597B48">635</mods:end>
|
|||
|
|
</mods:extent>
|
|||
|
|
</mods:part>
|
|||
|
|
</mods:relatedItem>
|
|||
|
|
<mods:classification id="9588F52A5E9843E2D442D0B672422BC5">journal article</mods:classification>
|
|||
|
|
<mods:identifier id="951A3D691A4D07A7D198F562F7B7396D" type="ISSN">0024-4082</mods:identifier>
|
|||
|
|
<mods:identifier id="9361077C0317941F1C68862CA41B4B95" type="ZooBank">74C07292-2BD6-4E3E-B68D-B144B81BBD83</mods:identifier>
|
|||
|
|
</mods:mods>
|
|||
|
|
<treatment id="0B1B8791FFD10605F967EBDE7F385D50" ID-DOI="http://doi.org/10.5281/zenodo.5240831" ID-Zenodo-Dep="5240831" LSID="urn:lsid:plazi:treatment:0B1B8791FFD10605F967EBDE7F385D50" httpUri="http://treatment.plazi.org/id/0B1B8791FFD10605F967EBDE7F385D50" lastPageId="45" lastPageNumber="600" pageId="43" pageNumber="598">
|
|||
|
|
<subSubSection id="CBA8650CFFD10603F967EBDE7F645D84" box="[327,614,964,988]" pageId="43" pageNumber="598" type="nomenclature">
|
|||
|
|
<paragraph id="830D3687FFD10603F967EBDE7F645D84" blockId="43.[327,614,964,988]" box="[327,614,964,988]" pageId="43" pageNumber="598">
|
|||
|
|
<heading id="D84581EBFFD10603F967EBDE7F645D84" box="[327,614,964,988]" centered="true" fontSize="9" level="2" pageId="43" pageNumber="598" reason="2">
|
|||
|
|
<taxonomicName id="44B24D04FFD10603F967EBDE7CF95D82" ID-CoL="3M62S" authority="Bonifácio & Menot, 2019" authorityName="hodor" authorityYear="2019" box="[327,507,964,987]" class="Polychaeta" family="Polynoidae" genus="Hodor" kingdom="Animalia" order="Phyllodocida" pageId="43" pageNumber="601" phylum="Annelida" rank="species" species="hodor" status="gen. nov., sp. nov.">HODOR HODOR</taxonomicName>
|
|||
|
|
<emphasis id="B1C6EA95FFD10603FA23EBDD7F645D84" bold="true" box="[515,614,964,988]" pageId="43" pageNumber="598">
|
|||
|
|
<taxonomicNameLabel id="AAF557EEFFD10603FA23EBDD7F645D84" box="[515,614,964,988]" pageId="43" pageNumber="598" rank="species">SP. NOV.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="CBA8650CFFD10605F961EBF17F385D50" lastPageId="45" lastPageNumber="600" pageId="43" pageNumber="598" type="description">
|
|||
|
|
<paragraph id="830D3687FFD10603F961EBF17F6F5A5C" blockId="43.[321,621,1003,1028]" box="[321,621,1003,1028]" pageId="43" pageNumber="598">
|
|||
|
|
(
|
|||
|
|
<figureCitation id="1B892A02FFD10603F969EBF17F365A5B" box="[329,564,1003,1028]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">FIG. 12A–H; TABLES</figureCitation>
|
|||
|
|
1, 2)
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD10603F883EC2F7F765B7D" blockId="43.[163,779,1076,1317]" pageId="43" pageNumber="598">
|
|||
|
|
<emphasis id="B1C6EA95FFD10603F883EC2F7C5B5A11" box="[163,345,1076,1098]" italics="true" pageId="43" pageNumber="598">Type material:</emphasis>
|
|||
|
|
<materialsCitation id="33DA3CDAFFD10603F951EC2E7F765B7D" collectingDate="2015-04-21" collectionCode="IFR, APEI" county="Clarion-Clipperton Fracture Zone" elevation="0" latitude="18.755634" location="Equatorial Eastern Pacific Ocean" longLatPrecision="1" longitude="-128.3403" municipality="Eastern Pacific Ocean" pageId="43" pageNumber="598" specimenCode="IFR655-2, 1, APEI#3" specimenCount="1" typeStatus="holotype">
|
|||
|
|
<typeStatus id="5C098825FFD10603F951EC2E7CE15A12" box="[369,483,1076,1098]" pageId="43" pageNumber="598" type="holotype">Holotype</typeStatus>
|
|||
|
|
, MNHN-IA-TYPE 1825 (
|
|||
|
|
<specimenCode id="D3149EFCFFD10603F88CEC497C295A31" box="[172,299,1107,1129]" collectionCode="IFR" pageId="43" pageNumber="598">IFR655-2</specimenCode>
|
|||
|
|
-
|
|||
|
|
<specimenCode id="D3149EFCFFD10603F913EC497C3F5A31" box="[307,317,1107,1129]" collectionCode="IFR" pageId="43" pageNumber="598">1</specimenCode>
|
|||
|
|
), complete, length
|
|||
|
|
<quantity id="444A9B62FFD10603FA14EC497FB15A31" box="[564,691,1107,1129]" metricMagnitude="-2" metricUnit="m" metricValue="1.135" pageId="43" pageNumber="598" unit="mm" value="11.35">11.35 mm</quantity>
|
|||
|
|
, width
|
|||
|
|
<quantity id="444A9B62FFD10603F883EC697C055AD0" box="[163,263,1139,1161]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="43" pageNumber="598" unit="mm" value="1.68">1.68 mm</quantity>
|
|||
|
|
, 24 segments (including tentacular segment),
|
|||
|
|
<location id="866D605CFFD10603F883EC887F245AFF" LSID="urn:lsid:plazi:treatment:0B1B8791FFD10605F967EBDE7F385D50:866D605CFFD10603F883EC887F245AFF" box="[163,550,1170,1192]" county="Clarion-Clipperton Fracture Zone" latitude="18.755634" longLatPrecision="1" longitude="-128.3403" municipality="Eastern Pacific Ocean" name="Equatorial Eastern Pacific Ocean" pageId="43" pageNumber="598">
|
|||
|
|
Equatorial
|
|||
|
|
<collectingMunicipality id="6369ACFDFFD10603F906EC897F245AFF" box="[294,550,1170,1192]" pageId="43" pageNumber="598">Eastern Pacific Ocean</collectingMunicipality>
|
|||
|
|
</location>
|
|||
|
|
,
|
|||
|
|
<collectingCounty id="6A6C4E0BFFD10603FA11EC887C445A9F" pageId="43" pageNumber="598">Clarion-Clipperton Fracture Zone</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<specimenCode id="D3149EFCFFD10603F971ECAB7CAF5A9E" box="[337,429,1201,1222]" collectionCode="APEI" country="Canada" httpUri="http://grbio.org/cool/pg9t-riai" name="Agriculture Canada Research Station" pageId="43" pageNumber="598">APEI#3</specimenCode>
|
|||
|
|
, station 192, collected
|
|||
|
|
<date id="F70C1047FFD10603FA8EECAB7DDD5ABF" pageId="43" pageNumber="598" value="2015-04-21">
|
|||
|
|
<collectingDate id="E748E9AFFFD10603FA8EECAB7DDD5ABF" pageId="43" pageNumber="598" value="2015-04-21">21 April 2015</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
, epibenthic sledge epi-net, start
|
|||
|
|
<geoCoordinate id="E6865040FFD10603FA52ECCB7E055ABE" box="[626,775,1231,1254]" degrees="18" direction="north" minutes="44.807" orientation="latitude" pageId="43" pageNumber="598" precision="1" value="18.746784">18°44.807′N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="E6865040FFD10603F883ECEA7C405B5D" box="[163,322,1263,1286]" degrees="128" direction="west" minutes="21.874" orientation="longitude" pageId="43" pageNumber="598" precision="1" value="-128.36456">128°21.874′W</geoCoordinate>
|
|||
|
|
, end
|
|||
|
|
<geoCoordinate id="E6865040FFD10603F9A0ECEA7F0C5B5D" box="[384,526,1263,1286]" degrees="18" direction="north" minutes="45.338" orientation="latitude" pageId="43" pageNumber="598" precision="1" value="18.755634">18°45.338′N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="E6865040FFD10603FA3AECEA7FB85B5D" box="[538,698,1263,1286]" degrees="128" direction="west" minutes="20.418" orientation="longitude" pageId="43" pageNumber="598" precision="1" value="-128.3403">128°20.418′W</geoCoordinate>
|
|||
|
|
, 4821–
|
|||
|
|
<elevation id="089FD1B4FFD10603F883ED157C9D5B7D" box="[163,415,1295,1317]" isEnumeration="true" pageId="43" pageNumber="598">
|
|||
|
|
<quantity id="444A9B62FFD10603F883ED157DF45B7D" box="[163,246,1295,1317]" metricMagnitude="3" metricUnit="m" metricValue="4.82" pageId="43" pageNumber="598" unit="m" value="4820.0">4820 m</quantity>
|
|||
|
|
depth,
|
|||
|
|
<quantity id="444A9B62FFD10603F96BED157C9D5B7D" box="[331,415,1295,1317]" metricMagnitude="3" metricUnit="m" metricValue="2.799" pageId="43" pageNumber="598" unit="m" value="2799.0">2799 m</quantity>
|
|||
|
|
</elevation>
|
|||
|
|
trawling distance.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD10603F883ED557C1B5859" blockId="43.[163,779,1359,1881]" pageId="43" pageNumber="598">
|
|||
|
|
<emphasis id="B1C6EA95FFD10603F883ED557F1A5B3C" box="[163,536,1359,1380]" italics="true" pageId="43" pageNumber="598">
|
|||
|
|
Description (based on
|
|||
|
|
<typeStatus id="5C098825FFD10603F987ED557F0B5B3C" box="[423,521,1359,1380]" pageId="43" pageNumber="598" type="holotype">holotype</typeStatus>
|
|||
|
|
):
|
|||
|
|
</emphasis>
|
|||
|
|
<typeStatus id="5C098825FFD10603FA0CED557F975B3D" box="[556,661,1359,1381]" pageId="43" pageNumber="598" type="holotype">Holotype</typeStatus>
|
|||
|
|
complete,
|
|||
|
|
<quantity id="444A9B62FFD10603F883ED747C185BDB" box="[163,282,1390,1412]" metricMagnitude="-2" metricUnit="m" metricValue="1.135" pageId="43" pageNumber="598" unit="mm" value="11.35">11.35 mm</quantity>
|
|||
|
|
long and
|
|||
|
|
<quantity id="444A9B62FFD10603F9B6ED747CFD5BDB" box="[406,511,1390,1412]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="43" pageNumber="598" unit="mm" value="1.68">1.68 mm</quantity>
|
|||
|
|
wide for 24 segments (including tentacular segment), dorsoventrally flattened, posteriorly tapering; colour of live animal not known; ethanol-preserved specimen pale white (
|
|||
|
|
<figureCitation id="1B892A02FFD10603F88BEDF17C0B5858" box="[171,265,1515,1537]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12A</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD10603F89BEE107DF45901" blockId="43.[163,779,1359,1881]" pageId="43" pageNumber="598">
|
|||
|
|
Prostomium bilobed, about as long as wide, lobes subtriangular, poorly developed, anteriorly tapering into blunt peaks, extending until superior lip; frontal filaments absent; median notch between prostomial lobes narrow and shallow (
|
|||
|
|
<figureCitation id="1B892A02FFD10603F9C7EE927F4858C5" box="[487,586,1672,1694]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12C</figureCitation>
|
|||
|
|
); eyes absent; a pair of internal white ganglia visible through translucent epidermis, dorsolaterally located on prostomium. Median and lateral antennae absent. Palps smooth, tapering, very long (reaching segment 9), inserted on large, rounded palpophores (
|
|||
|
|
<figureCitation id="1B892A02FFD10603F9CBEF3E7F495961" box="[491,587,1828,1850]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12C</figureCitation>
|
|||
|
|
). Facial tubercle absent.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD10603FB73E8DF7ECE5F8A" blockId="43.[827,1444,197,1881]" pageId="43" pageNumber="598">
|
|||
|
|
Tentacular segment fused to prostomium, well developed, with a pair of short lobes, inserted laterally and slightly below prostomium; without acicula or chaetae; tentaculophores large, bulbous, equal sized; dorsal tentacular style smooth, tapering, short (reaching segment 4); ventral tentacular style missing (
|
|||
|
|
<figureCitation id="1B892A02FFD10603FB63E9647EA05FCB" box="[835,930,382,404]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12C</figureCitation>
|
|||
|
|
). Pharynx not everted. Second segment with elytrophores, subbiramous parapodia, chaetae and ventral cirri.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD10603FB73E9C07EAF5C14" blockId="43.[827,1444,197,1881]" pageId="43" pageNumber="598">
|
|||
|
|
Nine pairs of large, globular elytrophores (
|
|||
|
|
<figureCitation id="1B892A02FFD10603FD1EE9C07E4C5C57" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12B, D</figureCitation>
|
|||
|
|
) present on segments 2, 4, 5, 7, 9, 11, 13, 15 and 17 (all elytra missing); with dorsal cirrophores on last segments.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD10603FB73EA4C7EE15CF1" blockId="43.[827,1444,197,1881]" pageId="43" pageNumber="598">
|
|||
|
|
Cirrigerous segments with large, bulbous dorsal cirrophores (
|
|||
|
|
<figureCitation id="1B892A02FFD10603FB8DEA6E790E5CD2" box="[941,1036,628,650]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12B</figureCitation>
|
|||
|
|
), inserted subdistally on notopodia; styles missing.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD10603FB73EAA879D25D7C" blockId="43.[827,1444,197,1881]" pageId="43" pageNumber="598">
|
|||
|
|
Segments 6 and 8 with large, swollen dorsal structure (
|
|||
|
|
<figureCitation id="1B892A02FFD10603FB5DEACB79025CBE" box="[893,1024,721,743]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12A, B</figureCitation>
|
|||
|
|
), located basally to cirrophores, interiorly whitish; smaller on segment 6, bigger on segment 8. Dorsal tubercles absent.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD10603FB73EB37791E5D85" blockId="43.[827,1444,197,1881]" pageId="43" pageNumber="598">
|
|||
|
|
Ventral cirri smooth, tapering, present from segment 2 to last segment; inserted basally on neuropodia of segment 2, style long (longer than tip of neuroacicular lobe); in subsequent segments inserted medially on neuropodia (
|
|||
|
|
<figureCitation id="1B892A02FFD10603FBECEBB2792F5DE5" box="[972,1069,936,958]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12D</figureCitation>
|
|||
|
|
), style short (shorter than tip of neuroacicular lobe).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD10603FB73EBFC7ED458E7" blockId="43.[827,1444,197,1881]" pageId="43" pageNumber="598">
|
|||
|
|
Parapodia subbiramous; notopodia reduced, much short than neuropodia (
|
|||
|
|
<figureCitation id="1B892A02FFD10603FC74EC1F79BA5A42" box="[1108,1208,1029,1051]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12D</figureCitation>
|
|||
|
|
). Notopodia arising from the dorsum as two thickened ridges; narrow, subtriangular, tapering into long acicular lobe, tip of notoacicula not penetrating epidermis. Neuropodia large, rectangular to subtriangular, tapering into long acicular lobe, tip of neuroacicula not penetrating epidermis. Notochaetae variable in number (one to 16 observed), long, slender, slightly curved with distinct, faint spinous rows on convex side, with blunt tips preceded by subdistally smooth margin (
|
|||
|
|
<figureCitation id="1B892A02FFD10603FB64ED227EA95B16" box="[836,939,1336,1358]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12E</figureCitation>
|
|||
|
|
); notochaetae more slender than neurochaetae. Neurochaetae of
|
|||
|
|
<specimenCount id="95B4FD0EFFD10603FC5EED4279F55B35" box="[1150,1271,1368,1389]" pageId="43" pageNumber="598" type="generic" typeStatus="types">two types</specimenCount>
|
|||
|
|
: (1) moderate in number (12–28 observed), long to very long, distally flattened to concave, serrated along both margins, with pointed tips (
|
|||
|
|
<figureCitation id="1B892A02FFD10603FC70EDAE79B25B91" box="[1104,1200,1460,1482]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12F</figureCitation>
|
|||
|
|
); and (2) middle and lower group on segments 3–7 modified, moderate in number (16 observed), stouter, short to long, distally flattened to concave, with coarse spines along both margins, spines concentrated basally and well spaced later, middle part with smooth margins (most of or less of their length), subdistally smooth or with one or two spines, blunt tip, tip margin slightly lighter (
|
|||
|
|
<figureCitation id="1B892A02FFD10603FB63EEB07EC558E7" box="[835,967,1706,1728]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12G, H</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD10603FB73EED37E745901" blockId="43.[827,1444,197,1881]" pageId="43" pageNumber="598">
|
|||
|
|
Nephridial papillae present on segments 11, 12 and 13, small, bulbous. Last segment very reduced. Pygidium rounded, not enclosed by last segment; with terminal anus (
|
|||
|
|
<figureCitation id="1B892A02FFD10603FBDAEF3F79625962" box="[1018,1120,1829,1851]" captionStart="Figure 12" captionStartId="44.[145,224,1602,1624]" captionTargetBox="[144,1425,195,1541]" captionTargetId="figure-195@44.[693,968,758,946]" captionTargetPageId="44" captionText="Figure 12. Hodor hodor gen. nov., sp. nov., holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5–10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure." pageId="43" pageNumber="598">Fig. 12A</figureCitation>
|
|||
|
|
). Anal cirri lost, scars not seen.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="D7CD660FFFD60604F8B1EE5879045896" pageId="44" pageNumber="599" startId="44.[145,224,1602,1624]" targetBox="[144,1425,195,1541]" targetPageId="44">
|
|||
|
|
<paragraph id="830D3687FFD60604F8B1EE5879045896" blockId="44.[145,1425,1602,1742]" pageId="44" pageNumber="599">
|
|||
|
|
<emphasis id="B1C6EA95FFD60604F8B1EE587CA55800" bold="true" box="[145,423,1602,1624]" pageId="44" pageNumber="599">
|
|||
|
|
Figure 12.
|
|||
|
|
<taxonomicName id="44B24D04FFD60604F932EE587CA55800" authority="Bonifácio & Menot, 2019" authorityName="hodor" authorityYear="2019" box="[274,423,1602,1624]" class="Polychaeta" family="Polynoidae" genus="Hodor" kingdom="Animalia" order="Phyllodocida" pageId="44" pageNumber="599" phylum="Annelida" rank="species" species="hodor" status="gen. nov., sp. nov.">
|
|||
|
|
<emphasis id="B1C6EA95FFD60604F932EE587CA55800" bold="true" box="[274,423,1602,1624]" italics="true" pageId="44" pageNumber="599">Hodor hodor</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
<taxonomicNameLabel id="AAF557EEFFD60604F98EEE587F785800" box="[430,634,1602,1624]" pageId="44" pageNumber="599" rank="species">
|
|||
|
|
<emphasis id="B1C6EA95FFD60604F98EEE587F77580F" bold="true" box="[430,629,1602,1623]" pageId="44" pageNumber="599">gen. nov., sp. nov</emphasis>
|
|||
|
|
.
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
, holotype MNHN-IA-TYPE 1825 (A–H). A, dorsal view of a preserved complete specimen. B, dorsal view of segments 6 and 8 with swollen dorsal structure, segments 5
|
|||
|
|
<emphasis id="B1C6EA95FFD60604FC52EE7A797F582D" box="[1138,1149,1632,1653]" italics="true" pageId="44" pageNumber="599">–</emphasis>
|
|||
|
|
10. C, anterior end, dorsolateral view, chaetae omitted. D, left parapodia, posterior view, segment 9. E, notochaeta with very faint spinous rows, segment 12. F, neurochaeta, segment 11. G, stouter neurochaeta, lower group, segment 7. H, stout neurochaeta, lower group, segment 4. Abbreviations: ci, cirrophore; el, elytrophore; sw, swollen dorsal structure.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="830D3687FFD60605F8B1EEE07CAD5FE9" blockId="44.[145,761,1786,1900]" lastBlockId="45.[163,779,197,434]" lastPageId="45" lastPageNumber="600" pageId="44" pageNumber="599">
|
|||
|
|
<emphasis id="B1C6EA95FFD60604F8B1EEE07CAB5957" box="[145,425,1786,1807]" italics="true" pageId="44" pageNumber="599">
|
|||
|
|
Remarks:
|
|||
|
|
<taxonomicName id="44B24D04FFD60604F92FEEE07CAB5957" authority="Bonifácio & Menot, 2019" authorityName="hodor" authorityYear="2019" box="[271,425,1786,1807]" class="Polychaeta" family="Polynoidae" genus="Hodor" kingdom="Animalia" order="Phyllodocida" pageId="44" pageNumber="599" phylum="Annelida" rank="species" species="hodor" status="gen. nov., sp. nov.">Hodor hodor</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
<taxonomicNameLabel id="AAF557EEFFD60604F993EEE07F8A5957" box="[435,648,1786,1807]" pageId="44" pageNumber="599" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
|
|||
|
|
presents many similarities with
|
|||
|
|
<taxonomicName id="44B24D04FFD60604F9B3EF037F345976" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[403,566,1817,1838]" class="Polychaeta" family="Polynoidae" genus="Hodor" kingdom="Animalia" order="Phyllodocida" pageId="44" pageNumber="599" phylum="Annelida" rank="species" species="anduril" status="gen. nov., sp. nov.">
|
|||
|
|
<emphasis id="B1C6EA95FFD60604F9B3EF037F345976" box="[403,566,1817,1838]" italics="true" pageId="44" pageNumber="599">Hodor anduril</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="AAF557EEFFD60604FA1AEF037FF75976" box="[570,757,1817,1838]" pageId="44" pageNumber="599" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
|
|||
|
|
, including the number of segments, prostomium and parapodial shape, chaetae and number of nephridial papillae pairs. However, the palps in
|
|||
|
|
<taxonomicName id="44B24D04FFD60604FCECEEE0785E5957" authority="Bonifácio & Menot, 2019" authorityName="hodor" authorityYear="2019" box="[1228,1372,1786,1807]" class="Polychaeta" family="Polynoidae" genus="Hodor" kingdom="Animalia" order="Phyllodocida" pageId="44" pageNumber="599" phylum="Annelida" rank="species" species="hodor" status="gen. nov., sp. nov.">
|
|||
|
|
<emphasis id="B1C6EA95FFD60604FCECEEE0785E5957" box="[1228,1372,1786,1807]" italics="true" pageId="44" pageNumber="599">Hodor hodor</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="AAF557EEFFD60604FD42EEE07EB75976" pageId="44" pageNumber="599" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
|
|||
|
|
are longer (reaching segment 9) than the palps in
|
|||
|
|
<taxonomicName id="44B24D04FFD60604FBB2EF2D793D5914" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[914,1087,1847,1868]" class="Polychaeta" family="Polynoidae" genus="Hodor" kingdom="Animalia" order="Phyllodocida" pageId="44" pageNumber="599" phylum="Annelida" rank="species" species="anduril" status="gen. nov., sp. nov.">
|
|||
|
|
<emphasis id="B1C6EA95FFD60604FBB2EF2D793D5914" box="[914,1087,1847,1868]" italics="true" pageId="44" pageNumber="599">Hodor anduril</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="AAF557EEFFD60604FC69EF22781A5915" box="[1097,1304,1848,1869]" pageId="44" pageNumber="599" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
|
|||
|
|
(reaching segment 2). Furthermore, the average K2P distance between these species was 14.2% for
|
|||
|
|
<emphasis id="B1C6EA95FFD70605FA44E8DC7F965E83" box="[612,660,198,219]" italics="true" pageId="45" pageNumber="600">COI</emphasis>
|
|||
|
|
and 6.1% for 16S. Interestingly, the morphological variation observed in
|
|||
|
|
<taxonomicName id="44B24D04FFD70605F909E9197CCF5F40" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[297,461,259,280]" class="Polychaeta" family="Polynoidae" genus="Hodor" kingdom="Animalia" order="Phyllodocida" pageId="45" pageNumber="600" phylum="Annelida" rank="species" species="anduril" status="gen. nov., sp. nov.">
|
|||
|
|
<emphasis id="B1C6EA95FFD70605F909E9197CCF5F40" box="[297,461,259,280]" italics="true" pageId="45" pageNumber="600">Hodor anduril</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="AAF557EEFFD70605F9F3E9197F925F40" box="[467,656,259,280]" pageId="45" pageNumber="600" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
|
|||
|
|
specimens could also indicate some dissimilarity within
|
|||
|
|
<taxonomicName id="44B24D04FFD70605FAE2E93B7DEE5F0D" authority="Bonifácio & Menot, 2019" authorityName="hodor" authorityYear="2019" class="Polychaeta" family="Polynoidae" genus="Hodor" kingdom="Animalia" order="Phyllodocida" pageId="45" pageNumber="600" phylum="Annelida" rank="species" species="hodor" status="gen. nov., sp. nov.">
|
|||
|
|
<emphasis id="B1C6EA95FFD70605FAE2E93B7DEE5F0D" italics="true" pageId="45" pageNumber="600">Hodor hodor</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="AAF557EEFFD70605F8D8E95A7CE25F0D" box="[248,480,320,341]" pageId="45" pageNumber="600" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
|
|||
|
|
specimens, which will necessitate care in the identification of any further specimens. In addition, both species have overlapping distribution in APEI#3.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD70605F883E9C17F145C76" blockId="45.[163,779,475,558]" pageId="45" pageNumber="600">
|
|||
|
|
<emphasis id="B1C6EA95FFD70605F883E9C17C205FA8" box="[163,290,475,496]" italics="true" pageId="45" pageNumber="600">Etymology:</emphasis>
|
|||
|
|
This species is dedicated to
|
|||
|
|
<taxonomicName id="44B24D04FFD70605FA55E9C17FBE5FA9" authorityName="Bonifácio & Menot" authorityYear="2019" box="[629,700,475,497]" class="Polychaeta" family="Polynoidae" genus="Hodor" kingdom="Animalia" order="Phyllodocida" pageId="45" pageNumber="600" phylum="Annelida" rank="genus">Hodor</taxonomicName>
|
|||
|
|
, one of P.B.’s favourite characters in the novel ‘
|
|||
|
|
<emphasis id="B1C6EA95FFD70605FA58E9E07DFE5C75" italics="true" pageId="45" pageNumber="600">A song of ice and fire</emphasis>
|
|||
|
|
’ by George R. R. Martin.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD70605F883EA4D7F115CD3" blockId="45.[163,779,599,652]" pageId="45" pageNumber="600">
|
|||
|
|
<emphasis id="B1C6EA95FFD70605F883EA4D7C405C34" box="[163,322,599,620]" italics="true" pageId="45" pageNumber="600">Genetic data:</emphasis>
|
|||
|
|
DNA sequencing for this species was successful for
|
|||
|
|
<emphasis id="B1C6EA95FFD70605F963EA6C7C705CD3" box="[323,370,630,651]" italics="true" pageId="45" pageNumber="600">COI</emphasis>
|
|||
|
|
, 16S and 18S.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="830D3687FFD70605F883EAAE7F385D50" blockId="45.[163,779,692,776]" pageId="45" pageNumber="600">
|
|||
|
|
<emphasis id="B1C6EA95FFD70605F883EAAE7C3E5C91" box="[163,316,692,713]" italics="true" pageId="45" pageNumber="600">Distribution:</emphasis>
|
|||
|
|
Only
|
|||
|
|
<specimenCount id="95B4FD0EFFD70605F9B2EAAF7F305C92" box="[402,562,692,714]" pageId="45" pageNumber="600" type="generic">one specimen</specimenCount>
|
|||
|
|
was sampled at a single station within the Clarion-Clipperton Fracture Zone in APEI#3 area (
|
|||
|
|
<typeStatus id="5C098825FFD70605F9BFEAE87CD35D5F" box="[415,465,754,775]" pageId="45" pageNumber="600">type</typeStatus>
|
|||
|
|
locality).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|