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<document id="B7DD8B49CC410C53279D0EA891366F7A" ID-DOI="10.11646/zootaxa.3888.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10086415" ID-ZooBank="1B437AE1-CF28-4C1B-95B6-C31A295905A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623730057475" checkinUser="felipe" docAuthor="Schwarzhans, Werner" docDate="2014" docId="463A8F36FF80FFC912979958E249FD50" docLanguage="en" docName="zt03888p073.pdf.imf" docOrigin="Zootaxa 3888 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hymenocephalus hachijoensis Okamura 1970" docType="treatment" docVersion="2" lastPageNumber="50" masterDocId="BA03F74EFFB0FFF812009814E16BFFF8" masterDocTitle="Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species" masterLastPageNumber="73" masterPageNumber="1" pageNumber="49" updateTime="1699488604623" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title id="253BC4B0A270722978962630DA363BBF">Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species</mods:title>
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<mods:namePart id="DF4E02F681A7951F78B298AB9D6E3E41">Schwarzhans, Werner</mods:namePart>
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<mods:title id="0A5D61E5AEC526CD137B4710F6C04D0F">Zootaxa</mods:title>
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<mods:date id="8D7B63269BAB4E93D406D944BF9D0458">2014</mods:date>
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<mods:number id="716FE4D22609C0EC68D6CCE89C77FAFF">2014-11-28</mods:number>
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<mods:detail id="AF09C845D74FDD58A1498CD2DC6CD189" type="volume">
<mods:number id="45D281795F17731962945994F9A1AFC2">3888</mods:number>
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<mods:number id="8ACAB6771032BCE34EEBA29C0BC79C2A">1</mods:number>
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<mods:classification id="8AF30CD5A2512C2E534324EA53C51AF9">journal article</mods:classification>
<mods:identifier id="889E541365F980B5D9F28E12B960AEE7" type="DOI">10.11646/zootaxa.3888.1.1</mods:identifier>
<mods:identifier id="F282DEC4DE1A74E6877B9C8F4ADF2B93" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9A08B491B9A5BA08810A2BCA99ED89B4" type="Zenodo-Dep">10086415</mods:identifier>
<mods:identifier id="20B202064E984DCCF4EBE16441501A2B" type="ZooBank">1B437AE1-CF28-4C1B-95B6-C31A295905A0</mods:identifier>
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<treatment id="463A8F36FF80FFC912979958E249FD50" LSID="urn:lsid:plazi:treatment:463A8F36FF80FFC912979958E249FD50" httpUri="http://treatment.plazi.org/id/463A8F36FF80FFC912979958E249FD50" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<subSubSection id="86896DABFF80FFC812979958E3B8FE9E" box="[151,723,332,358]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="CE2C3E20FF80FFC812979958E3B8FE9E" blockId="48.[151,723,332,391]" box="[151,723,332,358]" pageId="48" pageNumber="49">
<heading id="9564894CFF80FFC812979958E3B8FE9E" bold="true" box="[151,723,332,358]" fontSize="11" level="2" pageId="48" pageNumber="49" reason="4">
<taxonomicName id="099345A3FF80FFC812979958E3B8FE9E" ID-CoL="3NJ8X" authority="Okamura, 1970" authorityName="Okamura" authorityYear="1970" box="[151,723,332,358]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="hachijoensis">
<emphasis id="FCE7E232FF80FFC812979958E3B8FE9E" bold="true" box="[151,723,332,358]" pageId="48" pageNumber="49">
<emphasis id="FCE7E232FF80FFC812979958E36CFE9E" bold="true" box="[151,519,332,358]" italics="true" pageId="48" pageNumber="49">Hymenocephalus hachijoensis</emphasis>
<bibRefCitation id="AA0243D1FF80FFC8100F9958E3B8FE9E" author="Okamura, O." box="[527,723,332,358]" pageId="48" pageNumber="49" pagination="1 - 216" refId="ref51620" refString="Okamura, O. (1970) Fauna Japonica. Macrourina (Pisces). Academic Press of Japan, Tokyo, pp. 1 - 216." type="book chapter" year="1970">Okamura, 1970</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="86896DABFF80FFC81297997AE039FE7F" box="[151,338,366,391]" pageId="48" pageNumber="49" type="description">
<paragraph id="CE2C3E20FF80FFC81297997AE039FE7F" blockId="48.[151,723,332,391]" box="[151,338,366,391]" pageId="48" pageNumber="49">
<figureCitation id="56A822A5FF80FFC81297997AE040FE7F" box="[151,299,366,391]" captionStart="FIGURE 23" captionStartId="48.[151,250,1573,1595]" captionTargetBox="[225,1363,838,1540]" captionTargetId="figure-379@48.[220,1367,831,1548]" captionTargetPageId="48" captionText="FIGURE 23. Hymenocephalus hachijoensis, Holotype, BSKU 14171: A—Lateral view of head, B—Dorsal view of head, C—Inner face of otolith, D—Ventral view of otolith." figureDoi="http://doi.org/10.5281/zenodo.10086465" httpUri="https://zenodo.org/record/10086465/files/figure.png" pageId="48" pageNumber="49">Figs. 23A–D</figureCitation>
,
<figureCitation id="56A822A5FF80FFC81336997AE039FE7F" box="[310,338,366,391]" captionStart="FIGURE 27" captionStartId="53.[151,250,1329,1351]" captionTargetBox="[159,1430,656,1300]" captionTargetId="figure-554@53.[153,1434,649,1305]" captionTargetPageId="53" captionText="FIGURE 27. Geographical distribution of Hymenocephalus antraeus, H. hachijoensis, H. heterolepis and H. punt." figureDoi="http://doi.org/10.5281/zenodo.10086473" httpUri="https://zenodo.org/record/10086473/files/figure.png" pageId="48" pageNumber="49">27</figureCitation>
</paragraph>
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<subSubSection id="86896DABFF80FFC8129799A2E28CFE13" pageId="48" pageNumber="49" type="reference_group">
<paragraph id="CE2C3E20FF80FFC8129799A2E5E2FE34" blockId="48.[151,1161,438,491]" box="[151,1161,438,460]" pageId="48" pageNumber="49">
<treatmentCitationGroup id="EE83190EFF80FFC8129799A2E5E2FE34" box="[151,1161,438,460]" pageId="48" pageNumber="49">
<treatmentCitation id="4F321831FF80FFC8129799A2E270FE34" author="Okamura, O." box="[151,795,438,460]" page="50" pageId="48" pageNumber="49" year="1970">
<taxonomicName id="099345A3FF80FFC8129799A2E270FE34" ID-CoL="3NJ8X" authority="Okamura, 1970: 50" authorityName="Okamura" authorityPageNumber="50" authorityYear="1970" box="[151,795,438,460]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="subSpecies" species="striatissimus" subSpecies="hachijoensis">
<emphasis id="FCE7E232FF80FFC8129799A2E33AFE34" box="[151,593,438,460]" italics="true" pageId="48" pageNumber="49">Hymenocephalus striatissimus hachijoensis</emphasis>
<bibRefCitation id="AA0243D1FF80FFC8105799A2E270FE34" author="Okamura, O." box="[599,795,438,460]" pageId="48" pageNumber="49" pagination="1 - 216" refId="ref51620" refString="Okamura, O. (1970) Fauna Japonica. Macrourina (Pisces). Academic Press of Japan, Tokyo, pp. 1 - 216." type="book chapter" year="1970">Okamura, 1970: 50</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
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locality:
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,
<geoCoordinate id="ABA758E7FF80FFC8161999A2E515FE34" box="[1049,1150,438,460]" degrees="140" direction="east" minutes="03" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="140.05">140°03’E</geoCoordinate>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="CE2C3E20FF80FFC8129799C1E28CFE13" blockId="48.[151,1161,438,491]" box="[151,999,469,491]" pageId="48" pageNumber="49">
<treatmentCitationGroup id="EE83190EFF80FFC8129799C1E28CFE13" box="[151,999,469,491]" pageId="48" pageNumber="49">
<treatmentCitation id="4F321831FF80FFC8129799C1E3F5FE13" author="Okamura, O." box="[151,670,469,491]" page="93" pageId="48" pageNumber="49" year="1984">
<taxonomicName id="099345A3FF80FFC8129799C1E3F5FE13" ID-CoL="3NJ8X" authority=": Okamura, 1984: 93" authorityName=": Okamura" authorityPageNumber="93" authorityYear="1984" box="[151,670,469,491]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="hachijoensis">
<emphasis id="FCE7E232FF80FFC8129799C1E0A1FE13" box="[151,458,469,491]" italics="true" pageId="48" pageNumber="49">Hymenocephalus hachijoensis</emphasis>
:
<bibRefCitation id="AA0243D1FF80FFC813D799C1E3F5FE13" author="Okamura, O." box="[471,670,469,491]" pageId="48" pageNumber="49" refId="ref51648" refString="Okamura, O. (1984) The Fishes of the Japanese Archipelago. Vol. 1. &amp; 2. Tokai University Press, 437 pp. &amp; 370 pls. [Masuda, H., Amaoka, K., Araga, C., Uyeno, T. &amp; Yoshino, T.]" type="book" year="1984">Okamura, 1984: 93</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="4F321831FF80FFC810A999C1E288FE13" author="Sazonov, Y. I. &amp; Iwamoto, T." box="[681,995,469,491]" page="62" pageId="48" pageNumber="49" year="1992">
<bibRefCitation id="AA0243D1FF80FFC810A999C1E288FE13" author="Sazonov, Y. I. &amp; Iwamoto, T." box="[681,995,469,491]" pageId="48" pageNumber="49" pagination="27 - 95" refId="ref51836" refString="Sazonov, Y. I. &amp; Iwamoto, T. (1992) Grenadiers (Pisces, Gadiformes) of the Nazca and Sala y Gomez Ridges, southeastern Pacific. Proceedings of the California Academy of Sciences, 48 (2), 27 - 95." type="journal article" year="1992">Sazonov &amp; Iwamoto, 1992: 62</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF80FFC812979A03E095FDAB" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="CE2C3E20FF80FFC812979A03E095FDAB" blockId="48.[151,1437,535,776]" pageId="48" pageNumber="49">
<emphasis id="FCE7E232FF80FFC812979A03E0E8FDC8" bold="true" box="[151,387,535,560]" pageId="48" pageNumber="49">Material examined.</emphasis>
<materialsCitation id="7EFB347DFF80FFC8138C9A03E092FDAB" collectionCode="BSKU" elevation="570" latitude="33.266666" longLatPrecision="1205" longitude="140.05" pageId="48" pageNumber="49" specimenCode="BSKU 14171, BSKU 14172" specimenCount="2" typeStatus="holotype">
<specimenCount id="D895F5A9FF80FFC8138C9A03E373FDC8" box="[396,536,535,560]" pageId="48" pageNumber="49" type="generic">2 specimens</specimenCount>
<specimenCode id="9E35965BFF80FFC810219A03E3D0FDC8" box="[545,699,535,560]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="48" pageNumber="49">BSKU 14171</specimenCode>
(
<typeStatus id="11288082FF80FFC810CC9A03E25FFDC8" box="[716,820,535,560]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
),
<quantity id="096B93C5FF80FFC8114A9A03E2C2FDD7" box="[842,937,535,560]" metricMagnitude="-1" metricUnit="m" metricValue="1.95" pageId="48" pageNumber="49" unit="mm" value="195.0">195 mm</quantity>
TL, and
<specimenCode id="9E35965BFF80FFC816169A03E5DBFDC8" box="[1046,1200,535,560]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="48" pageNumber="49">BSKU 14172</specimenCode>
(
<typeStatus id="11288082FF80FFC816C19A03E44CFDC8" box="[1217,1319,535,560]" pageId="48" pageNumber="49" type="paratype">paratype</typeStatus>
),
<quantity id="096B93C5FF80FFC8173D9A03E4F6FDD7" box="[1341,1437,535,560]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="48" pageNumber="49" unit="mm" value="130.0">130 mm</quantity>
TL,
<geoCoordinate id="ABA758E7FF80FFC812C79A28E041FDAD" box="[199,298,572,597]" degrees="33" direction="north" minutes="16" orientation="latitude" pageId="48" pageNumber="49" precision="925" value="33.266666">33°16’N</geoCoordinate>
,
<geoCoordinate id="ABA758E7FF80FFC813369A28E0CFFDAD" box="[310,420,572,597]" degrees="140" direction="east" minutes="03" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="140.05">140°03’E</geoCoordinate>
,
<quantity id="096B93C5FF80FFC813AF9A28E092FDAB" box="[431,505,572,597]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="48" pageNumber="49" unit="m" value="570.0">
<specimenCount id="D895F5A9FF80FFC813AF9A28E092FDAB" box="[431,505,572,597]" pageId="48" pageNumber="49" type="generic">
<elevation id="45BED913FF80FFC813AF9A28E092FDAB" box="[431,505,572,597]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="48" pageNumber="49" unit="m" value="570.0">570 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF80FFC812C79A4BE41DF88F" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="CE2C3E20FF80FFC812C79A4BE2B4FCF0" blockId="48.[151,1437,535,776]" pageId="48" pageNumber="49">
<emphasis id="FCE7E232FF80FFC812C79A4BE02AFD80" bold="true" box="[199,321,607,632]" pageId="48" pageNumber="49">Diagnosis.</emphasis>
Pelvic fin rays 7–8; pectoral fin rays 14–15; snout barely projecting, short, 15–17% HL; barbel long, 30–35% HL, reaching vertical through center of orbit; orbit diameter large, 35–38% HL; infraorbital width 10–12% HL; preopercular supporter moderately long, with straight rear margin, 6–7% HL; gill rakers 18–20; ventral striae reaching to ½ distance from pelvic fin bases to periproct; otolith with high predorsal lobe, colliculi separated, narrowly placed across collum; OL:OH = 1.05; TCL:PCL = 1.6.
</paragraph>
<caption id="9AEC6EA8FF80FFC812979E31E3DAF9A2" ID-DOI="http://doi.org/10.5281/zenodo.10086465" ID-Zenodo-Dep="10086465" httpUri="https://zenodo.org/record/10086465/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1573,1595]" targetBox="[225,1363,838,1540]" targetPageId="48">
<paragraph id="CE2C3E20FF80FFC812979E31E3DAF9A2" blockId="48.[151,1436,1573,1626]" pageId="48" pageNumber="49">
<emphasis id="FCE7E232FF80FFC812979E31E048F9C2" bold="true" box="[151,291,1573,1595]" pageId="48" pageNumber="49">FIGURE 23.</emphasis>
<taxonomicName id="099345A3FF80FFC8132C9E31E308F9C3" authorityName=": Okamura" authorityYear="1984" box="[300,611,1573,1595]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="hachijoensis">
<emphasis id="FCE7E232FF80FFC8132C9E31E308F9C3" box="[300,611,1573,1595]" italics="true" pageId="48" pageNumber="49">Hymenocephalus hachijoensis</emphasis>
</taxonomicName>
, Holotype, BSKU 14171: A—Lateral view of head, B—Dorsal view of head, C—Inner face of otolith, D—Ventral view of otolith.
</paragraph>
</caption>
<paragraph id="CE2C3E20FF80FFC812C79E92E41DF88F" blockId="48.[151,1436,1670,2019]" pageId="48" pageNumber="49">
<emphasis id="FCE7E232FF80FFC812C79E92E009F967" bold="true" box="[199,354,1670,1695]" pageId="48" pageNumber="49">Comparison.</emphasis>
<taxonomicName id="099345A3FF80FFC813699E92E3D6F967" authorityName=": Okamura" authorityYear="1984" box="[361,701,1670,1695]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="hachijoensis">
<emphasis id="FCE7E232FF80FFC813699E92E3D6F967" box="[361,701,1670,1695]" italics="true" pageId="48" pageNumber="49">Hymenocephalus hachijoensis</emphasis>
</taxonomicName>
is readily recognized by the long barbel, which is shorter than in
<taxonomicName id="099345A3FF80FFC812979EBFE051F93B" box="[151,314,1706,1731]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="longibarbis">
<emphasis id="FCE7E232FF80FFC812979EBFE051F93B" box="[151,314,1706,1731]" italics="true" pageId="48" pageNumber="49">H. longibarbis</emphasis>
</taxonomicName>
and
<taxonomicName id="099345A3FF80FFC8136F9EBFE092F93A" authorityName="Schwarzhans" authorityYear="2014" box="[367,505,1707,1730]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="iwamotoi">
<emphasis id="FCE7E232FF80FFC8136F9EBFE092F93A" box="[367,505,1707,1730]" italics="true" pageId="48" pageNumber="49">H. iwamotoi</emphasis>
</taxonomicName>
, of about the same length as in
<taxonomicName id="099345A3FF80FFC811599EBEE53EF93B" authorityName=": Okamura" authorityYear="1970" box="[857,1109,1706,1731]" class="Actinopterygii" family="Macrouridae" genus="Hymenogadus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="FCE7E232FF80FFC811599EBEE53EF93B" box="[857,1109,1706,1731]" italics="true" pageId="48" pageNumber="49">Hymenogadus gracilis</emphasis>
</taxonomicName>
, and longer than in any other species of the genus
<taxonomicName id="099345A3FF80FFC813849EDAE32EF91F" authorityName="Giglioli" authorityYear="1884" box="[388,581,1742,1767]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="FCE7E232FF80FFC813849EDAE32EF91F" box="[388,581,1742,1767]" italics="true" pageId="48" pageNumber="49">Hymenocephalus</emphasis>
</taxonomicName>
, with those of
<taxonomicName id="099345A3FF80FFC810F19EC4E23EF91F" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[753,853,1744,1767]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="aeger">
<emphasis id="FCE7E232FF80FFC810F19EC4E23EF91F" box="[753,853,1744,1767]" italics="true" pageId="48" pageNumber="49">H. aeger</emphasis>
</taxonomicName>
coming closest at 20–25% HL. The short, rounded snout resembles species in the
<taxonomicName id="099345A3FF80FFC813F59EE7E3E8F8F2" authorityName=": Weber" authorityYear="1913" box="[501,643,1779,1802]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="striatissimus">
<emphasis id="FCE7E232FF80FFC813F59EE7E3E8F8F2" box="[501,643,1779,1802]" italics="true" pageId="48" pageNumber="49">striatissimus</emphasis>
</taxonomicName>
Group, like the low number of pelvic fin rays (7–8 vs
<quantity id="096B93C5FF80FFC816F69EE6E431F8F3" box="[1270,1370,1778,1803]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="48" pageNumber="49" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10–12 in</quantity>
other members of the
<taxonomicName id="099345A3FF80FFC813509F0CE0D9F8D7" authorityName="Gilbert &amp; Cramer" authorityYear="1897" box="[336,434,1816,1839]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="antraeus">
<emphasis id="FCE7E232FF80FFC813509F0CE0D9F8D7" box="[336,434,1816,1839]" italics="true" pageId="48" pageNumber="49">antraeus</emphasis>
</taxonomicName>
Group) and the long infraorbital supporter. The separated colliculi and only moderately enlarged pseudocolliculum easily distinguish
<taxonomicName id="099345A3FF80FFC810A19F2FE239F8AB" authorityName=": Okamura" authorityYear="1984" box="[673,850,1850,1875]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="hachijoensis">
<emphasis id="FCE7E232FF80FFC810A19F2FE239F8AB" box="[673,850,1850,1875]" italics="true" pageId="48" pageNumber="49">H. hachijoensis</emphasis>
</taxonomicName>
from species of the
<taxonomicName id="099345A3FF80FFC816449F2FE59DF8AA" authorityName=": Weber" authorityYear="1913" box="[1092,1270,1851,1874]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="striatissimus">
<emphasis id="FCE7E232FF80FFC816449F2FE59DF8AA" box="[1092,1270,1851,1874]" italics="true" pageId="48" pageNumber="49">H. striatissimus</emphasis>
</taxonomicName>
Group. These observations support the concept first expressed by
<bibRefCitation id="AA0243D1FF80FFC810DA9F4AE2FFF88F" author="Okamura, O." box="[730,916,1886,1911]" pageId="48" pageNumber="49" refId="ref51648" refString="Okamura, O. (1984) The Fishes of the Japanese Archipelago. Vol. 1. &amp; 2. Tokai University Press, 437 pp. &amp; 370 pls. [Masuda, H., Amaoka, K., Araga, C., Uyeno, T. &amp; Yoshino, T.]" type="book" year="1984">Okamura (1984)</bibRefCitation>
to elevate
<taxonomicName id="099345A3FF80FFC8160E9F74E5D7F88F" authorityName=": Okamura" authorityYear="1984" box="[1038,1212,1886,1911]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="hachijoensis">
<emphasis id="FCE7E232FF80FFC8160E9F74E5D7F88F" box="[1038,1212,1886,1911]" italics="true" pageId="48" pageNumber="49">H. hachijoensis</emphasis>
</taxonomicName>
to species level.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF80FFC912C79F96E23BFDE0" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="description">
<paragraph id="CE2C3E20FF80FFC912C79F96E198FEE5" blockId="48.[151,1436,1670,2019]" lastBlockId="49.[151,1436,151,680]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<emphasis id="FCE7E232FF80FFC812C79F96E033F863" bold="true" box="[199,344,1922,1947]" pageId="48" pageNumber="49">Description.</emphasis>
Head morphology (n = 2) (
<figureCitation id="56A822A5FF80FFC810999F96E24DF863" box="[665,806,1922,1947]" captionStart="FIGURE 23" captionStartId="48.[151,250,1573,1595]" captionTargetBox="[225,1363,838,1540]" captionTargetId="figure-379@48.[220,1367,831,1548]" captionTargetPageId="48" captionText="FIGURE 23. Hymenocephalus hachijoensis, Holotype, BSKU 14171: A—Lateral view of head, B—Dorsal view of head, C—Inner face of otolith, D—Ventral view of otolith." figureDoi="http://doi.org/10.5281/zenodo.10086465" httpUri="https://zenodo.org/record/10086465/files/figure.png" pageId="48" pageNumber="49">Fig. 23A–B</figureCitation>
): Snout short, rounded, barely projecting, 15–17% HL, orbit diameter large, 35–38% HL, interorbital width 50–60% HW. Barbel long, 30–35% HL, reaching vertical through center of orbit. Head canals well developed, infraorbital width 10–12% HL, supraorbital canal with 5 segments, width 9–10% HL, supratemporal canal not identified, preopercular canal width 9–10% HL, postorbitalpreopercular interspace 8–10% HL. Infranasal supporter very large, touching upper lip; infraorbital supporter long, expanded beyond entire orbit, 90–100% OD; preopercular supporter moderately long, 6–7% HL, rear margin straight.
</paragraph>
<paragraph id="CE2C3E20FF81FFC912C79933E23BFDE0" blockId="49.[151,1436,151,680]" pageId="49" pageNumber="50">
Otolith morphology (n = 1) (
<figureCitation id="56A822A5FF81FFC9101A9933E3C3FEB8" box="[538,680,295,320]" captionStart="FIGURE 23" captionStartId="48.[151,250,1573,1595]" captionTargetBox="[225,1363,838,1540]" captionTargetId="figure-379@48.[220,1367,831,1548]" captionTargetPageId="48" captionText="FIGURE 23. Hymenocephalus hachijoensis, Holotype, BSKU 14171: A—Lateral view of head, B—Dorsal view of head, C—Inner face of otolith, D—Ventral view of otolith." figureDoi="http://doi.org/10.5281/zenodo.10086465" httpUri="https://zenodo.org/record/10086465/files/figure.png" pageId="49" pageNumber="50">Fig. 23C–D</figureCitation>
): Otolith large; OL:OH = 1.05; OH:OT = 3.2. Dorsal rim with a distinct, moderately broad predorsal lobe with rounded tip, distally marked by small incision; posterior tip at about level of sulcus termination; ventral rim deep, regularly curved, smooth, deepest anterior of the middle; anterior rim high, nearly vertical. Inner face slightly convex, with slightly supramedian sulcus. Ostial and caudal colliculi moderately small, narrowly placed across collum, terminating moderately close to anterior rim and at some distance from posterior tip of otolith; pseudocolliculum long. CCL:OCL = 0.75; TCL:PCL = 1.6. Dorsal depression small; ventral furrow distinct, moderately close to ventral rim.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF81FFC912C79A30E249FD50" pageId="49" pageNumber="50" type="distribution">
<paragraph id="CE2C3E20FF81FFC912C79A30E249FD50" blockId="49.[151,1436,151,680]" pageId="49" pageNumber="50">
<emphasis id="FCE7E232FF81FFC912C79A30E032FDC5" bold="true" box="[199,345,548,573]" pageId="49" pageNumber="50">Distribution</emphasis>
(
<figureCitation id="56A822A5FF81FFC913699A30E0D5FDC5" box="[361,446,548,573]" captionStart="FIGURE 27" captionStartId="53.[151,250,1329,1351]" captionTargetBox="[159,1430,656,1300]" captionTargetId="figure-554@53.[153,1434,649,1305]" captionTargetPageId="53" captionText="FIGURE 27. Geographical distribution of Hymenocephalus antraeus, H. hachijoensis, H. heterolepis and H. punt." figureDoi="http://doi.org/10.5281/zenodo.10086473" httpUri="https://zenodo.org/record/10086473/files/figure.png" pageId="49" pageNumber="50">Fig. 27</figureCitation>
).
<taxonomicName id="099345A3FF81FFC913D29A30E24DFDC5" authorityName=": Okamura" authorityYear="1984" box="[466,806,548,573]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="hachijoensis">
<emphasis id="FCE7E232FF81FFC913D29A30E24DFDC5" box="[466,806,548,573]" italics="true" pageId="49" pageNumber="50">Hymenocephalus hachijoensis</emphasis>
</taxonomicName>
was originally described only from the
<typeStatus id="11288082FF81FFC916E99A30E472FDC4" box="[1257,1305,548,572]" pageId="49" pageNumber="50">type</typeStatus>
locality off the small Hachijo Island on the Iwo Jima Ridge off southern central
<collectingCountry id="B6847EB0FF81FFC911B19A5CE29FFDA7" box="[945,1012,584,607]" name="Japan" pageId="49" pageNumber="50">Japan</collectingCountry>
.
<bibRefCitation id="AA0243D1FF81FFC916019A53E40EFD98" author="Sazonov, Y. I. &amp; Iwamoto, T." box="[1025,1381,583,608]" pageId="49" pageNumber="50" pagination="27 - 95" refId="ref51836" refString="Sazonov, Y. I. &amp; Iwamoto, T. (1992) Grenadiers (Pisces, Gadiformes) of the Nazca and Sala y Gomez Ridges, southeastern Pacific. Proceedings of the California Academy of Sciences, 48 (2), 27 - 95." type="journal article" year="1992">Sazonov &amp; Iwamoto (1992:63)</bibRefCitation>
also recorded specimens caught over seamounts of the Emperor Ridge (
<geoCoordinate id="ABA758E7FF81FFC911949A78E2B8FD7D" box="[916,979,620,645]" degrees="32" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="55555" value="32.0">32°N</geoCoordinate>
,
<geoCoordinate id="ABA758E7FF81FFC911E09A78E54DFD7D" box="[992,1062,620,645]" degrees="173" direction="east" orientation="longitude" pageId="49" pageNumber="50" precision="55555" value="173.0">173°E</geoCoordinate>
and 41°
<geoCoordinate id="ABA758E7FF81FFC916869A78E5AEFD7D" box="[1158,1221,620,645]" degrees="04" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="55555" value="4.0">04°N</geoCoordinate>
,
<geoCoordinate id="ABA758E7FF81FFC916D29A78E455FD7D" box="[1234,1342,620,645]" degrees="170" direction="east" minutes="32" orientation="longitude" pageId="49" pageNumber="50" precision="925" value="170.53334">170°32’E</geoCoordinate>
) and on the Kyushu-Palau Ridge (no geographic details recorded).
</paragraph>
</subSubSection>
</treatment>
</document>
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