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<document id="AE74EC17ED0188C6DA4E08B9D78C8E1E" ID-CLB-Dataset="7382" ID-DOI="10.11646/zootaxa.4951.3.8" ID-GBIF-Dataset="a2120d24-68fa-4774-acda-400fbcb0e9e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="4668485" ID-ZooBank="84F55133-748D-4EC8-9EC5-9EC9E5088ECC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617782581144" checkinUser="plazi" docAuthor="Makarchenko, Eugenyi A., Semenchenko, Alexander A. &amp; Palatov, Dmitry M." docDate="2021" docId="261DBB48FF86FFA1758FFF2CFB23F82F" docLanguage="en" docName="zootaxa.4951.3.8.pdf" docOrigin="Zootaxa 4951 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pagastia (Pagastia) hanseni Oliver 1959, sp. nov." docType="treatment" docUuid="13933287-2A4E-4A5B-B3DD-DF64400F8B2F" docUuidSource="ZooBank" docVersion="10" lastPageNumber="565" masterDocId="DA24C330FF80FFA77518FFBBFFEBFFDF" masterDocTitle="New species and findings of Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from Central Asia, with DNA barcoding of known species of the genus" masterLastPageNumber="570" masterPageNumber="559" pageNumber="565" updateTime="1698936128681" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="10C87BE6B0C64C5C1395D2440DE94C70" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E95A9AD2A842E27F6B23FB9A5C5DD915">
<mods:title id="E058B0D8AA3FA2352823D5C337B44BE9">New species and findings of Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from Central Asia, with DNA barcoding of known species of the genus</mods:title>
</mods:titleInfo>
<mods:name id="CCE2F6C68665636981A93F61331A9C81" type="personal">
<mods:role id="85BD00AE740949500B457DCCEB76DD5E">
<mods:roleTerm id="2D2017E8DB439E42964DF5D342D8C82C">Author</mods:roleTerm>
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<mods:namePart id="8B86DA0CF40D55C9107A677C8AFC1544">Makarchenko, Eugenyi A.</mods:namePart>
<mods:nameIdentifier id="DCE5DF83D8C010136DB1D57277CA9CE1" type="ORCID">0000-0003-2765-8729</mods:nameIdentifier>
<mods:affiliation id="D2C23A633A8AD27CBE38F2ED181ADCC5">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, 100 let Vladivostoku 159, 690022 Vladivostok, Russia. makarchenko @ biosoil. ru; http: // orcid. org / 0000 - 0003 - 2765 - 8729</mods:affiliation>
<mods:nameIdentifier id="94F934915FBAC8DFCDB3AD47E028612F" type="email">makarchenko@biosoil.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="3A3BD28174F34BE70A98289A0686BC63" type="personal">
<mods:role id="889045FD83AF188800FD510C116D2C2D">
<mods:roleTerm id="CD66AAC4919D1D5666FDF4A277C3C916">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="43F61D0585BB827993FF33C8A6331F3A">Semenchenko, Alexander A.</mods:namePart>
<mods:nameIdentifier id="D00A77EB7148994E3FAEF8EA8E285CF3" type="ORCID">0000-0001-7207-9529</mods:nameIdentifier>
<mods:affiliation id="CA6ED0C7C3E33117906611C0219CD7C7">Laboratory of Ecology and Evolutionary Biology of Aquatic Organisms, Far Eastern Federal University, Suhanova St. 8, 690950 Vladivostok, Russia. semenchenko _ alexander @ mail. ru; https: // orcid. org / 0000 - 0001 - 7207 - 9529</mods:affiliation>
<mods:nameIdentifier id="3B951196C374663BD462772EC3F08453" type="email">semenchenko_alexander@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="2841EBE759B961F1BBFF4ADCC554221F" type="personal">
<mods:role id="832BBB5FEA8EA8688EC313EB73684886">
<mods:roleTerm id="EE08D5636F5383B9A019CE622723363B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B36431A084F5D2383BE27338FD2B40FF">Palatov, Dmitry M.</mods:namePart>
<mods:nameIdentifier id="4A254D857397FEFB5E4EB92093190793" type="ORCID">0000-0002-8826-9316</mods:nameIdentifier>
<mods:affiliation id="42C1AAD877CA339FCDA0F0773974140C">A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Leninskij prosp. 33, 119071 Moscow, Russia. triops @ yandex. ru; https: // orcid. org / 0000 - 0002 - 8826 - 9316</mods:affiliation>
<mods:nameIdentifier id="B56F4FFEBED8E2A197C906F1874270F7" type="email">triops@yandex.ru</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="27D73EB1989AA1E5890BF1F22184D00F">text</mods:typeOfResource>
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<mods:titleInfo id="0A2763B752DCD0AAC6B3995EDE1C463D">
<mods:title id="0BC4E437ECC7ABCED3D40F733BDE8EF6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A2B0749FE6433530A9EBD238A80BCFDE">
<mods:date id="5FCB2521FF6FA817DE43BE6E762384B0">2021</mods:date>
<mods:detail id="A1366E481AE08CE57402FF0F27CF2713" type="pubDate">
<mods:number id="C4C266DF288EDD9F936C33DD26C2007A">2021-04-07</mods:number>
</mods:detail>
<mods:detail id="AB35574C881A60355877C6B0414B3C1F" type="volume">
<mods:number id="E428876FC867F04A831CDA98130A7E05">4951</mods:number>
</mods:detail>
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<mods:number id="E94C70B95F46DC6243C774F089D5EBB7">3</mods:number>
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<mods:start id="00679AA0B66F197366FBF1F6FE4FD6A6">559</mods:start>
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<mods:classification id="10EF7A3E595768E8A8425E1F2455BA88">journal article</mods:classification>
<mods:identifier id="19B4B9A22C5F469765735DA7F3582221" type="CLB-Dataset">7382</mods:identifier>
<mods:identifier id="E8146F401C3DC96DDB7690581B627EB5" type="DOI">10.11646/zootaxa.4951.3.8</mods:identifier>
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<mods:identifier id="48770063AD0A48175F90746F75136921" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="14F02755381C58621E1495A816E4C37C" type="Zenodo-Dep">4668485</mods:identifier>
<mods:identifier id="941778186B95A84E1AE713EF24411065" type="ZooBank">84F55133-748D-4EC8-9EC5-9EC9E5088ECC</mods:identifier>
</mods:mods>
<treatment id="261DBB48FF86FFA1758FFF2CFB23F82F" ID-DOI="http://doi.org/10.5281/zenodo.4685089" ID-GBIF-Taxon="181003563" ID-Zenodo-Dep="4685089" LSID="urn:lsid:zoobank.org:act:13933287-2A4E-4A5B-B3DD-DF64400F8B2F" httpUri="http://treatment.plazi.org/id/261DBB48FF86FFA1758FFF2CFB23F82F" lastPageNumber="565" pageId="6" pageNumber="565">
<subSubSection id="E6AE59D5FF86FFA1758FFF2CFC17FF6E" box="[151,1020,151,178]" pageId="6" pageNumber="565" type="nomenclature">
<paragraph id="AE0B0A5EFF86FFA1758FFF2CFC17FF6E" blockId="6.[151,1020,151,250]" box="[151,1020,151,178]" pageId="6" pageNumber="565">
<heading id="F543BD32FF86FFA1758FFF2CFC17FF6E" bold="true" box="[151,1020,151,178]" fontSize="11" level="1" pageId="6" pageNumber="565" reason="1">
<emphasis id="9CC0D64CFF86FFA1758FFF2CFC17FF6E" bold="true" box="[151,1020,151,178]" pageId="6" pageNumber="565">
<taxonomicName id="69B471DDFF86FFA1758FFF2CFC79FF6E" ID-CoL="6C9D" ID-ENA="994279" authority="Makarchenko, Semenchenko et Palatov" authorityName="Oliver" authorityYear="1959" box="[151,914,151,178]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="9CC0D64CFF86FFA1758FFF2CFF14FF6E" bold="true" box="[151,255,151,177]" italics="true" pageId="6" pageNumber="565">Pagastia</emphasis>
(
<emphasis id="9CC0D64CFF86FFA17409FF2CFEC8FF6E" bold="true" box="[273,291,151,177]" italics="true" pageId="6" pageNumber="565">P</emphasis>
.)
<emphasis id="9CC0D64CFF86FFA17423FF2CFE70FF6D" bold="true" box="[315,411,151,178]" italics="true" pageId="6" pageNumber="565">hanseni</emphasis>
Makarchenko, Semenchenko et Palatov
</taxonomicName>
,
<taxonomicNameLabel id="87F36B37FF86FFA17686FF2CFC17FF6E" box="[926,1020,151,177]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E6AE59D5FF86FFA1758FFF00FC3AFF25" pageId="6" pageNumber="565" type="description">
<paragraph id="AE0B0A5EFF86FFA1758FFF00FE8EFF0A" blockId="6.[151,1020,151,250]" box="[151,357,187,214]" pageId="6" pageNumber="565">
(
<figureCitation id="368F16DBFF86FFA17587FF00FEE0FF09" box="[159,267,187,214]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 1–7. Pagastia (P.) caelestomontana sp. nov. (1–4) and P. (P.) hanseni sp. nov. (5–7), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 – 200 µm, for figures 2, 4, 7 – 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="6" pageNumber="565">Figs. 5–7</figureCitation>
,
<figureCitation id="368F16DBFF86FFA1740FFF00FEB7FF0A" box="[279,348,187,213]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1704,1728]" captionTargetBox="[151,1435,429,1666]" captionTargetId="figure-134@5.[151,1435,417,1680]" captionTargetPageId="5" captionText="FIGURES 8–11. Pagastia (P.) caelestomontana sp. nov. (8–9) and P. (P.) hanseni sp. nov. (10–11), males. 8, 10, hypopygium in dorsal view; 9, gonostylus; 11, anal point." figureDoi="http://doi.org/10.5281/zenodo.4668491" httpUri="https://zenodo.org/record/4668491/files/figure.png" pageId="6" pageNumber="565">10–11</figureCitation>
)
</paragraph>
<paragraph id="AE0B0A5EFF86FFA1758FFF64FC3AFF25" blockId="6.[151,1020,151,250]" box="[151,977,223,250]" pageId="6" pageNumber="565">
<uri id="DA25065CFF86FFA1758FFF64FC3AFF25" box="[151,977,223,250]" pageId="6" pageNumber="565">
urn:lsid:zoobank.org:act:
<uuid id="DA12308BFF86FFA174AEFF64FC3AFF25" box="[438,977,223,250]" pageId="6" pageNumber="565">13933287-2A4E-4A5B-B3DD-DF64400F8B2F</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="E6AE59D5FF86FFA1758FFE9CFEAFFE2A" pageId="6" pageNumber="565" type="materials_examined">
<paragraph id="AE0B0A5EFF86FFA1758FFE9CFEAFFE2A" blockId="6.[151,1437,295,1186]" pageId="6" pageNumber="565">
<emphasis id="9CC0D64CFF86FFA1758FFE9CFEEDFE9E" bold="true" box="[151,262,295,321]" pageId="6" pageNumber="565">Material.</emphasis>
<materialsCitation id="1EDC0003FF86FFA17415FE9CFD9AFE56" ID-GBIF-Occurrence="3071287301" collectingDate="2016-07-02" collectorName="D. Palatov." country="Tajikistan" county="Central Asia" elevation="3366" latitude="37.378616" location="Roshtqal'a District" longLatPrecision="1" longitude="72.34935" municipality="Autonomous Region" pageId="6" pageNumber="565" specimenCount="1" stateProvince="Kuhistoni Badakhshon" typeStatus="holotype">
<typeStatus id="710FB4FCFF86FFA17415FE9CFE91FE9E" box="[269,378,295,321]" pageId="6" pageNumber="565" type="holotype">Holotype</typeStatus>
: adult male,
<collectingCounty id="476A72D2FF86FFA1771DFE9CFD78FE9E" box="[517,659,295,321]" pageId="6" pageNumber="565">Central Asia</collectingCounty>
,
<collectingCountry id="D6A34ACEFF86FFA17785FE9CFCE5FE9E" box="[669,782,295,321]" name="Tajikistan" pageId="6" pageNumber="565">Tajikistan</collectingCountry>
,
<collectingRegion id="6C70C4BCFF86FFA17600FE9CFC19FE9E" box="[792,1010,295,321]" country="Tajikistan" name="Kuhistoni Badakhshon" pageId="6" pageNumber="565">Gorno-Badakhshan</collectingRegion>
<collectingMunicipality id="4E6F9024FF86FFA176EFFE9CFB09FE9E" box="[1015,1250,295,321]" pageId="6" pageNumber="565">Autonomous Region</collectingMunicipality>
,
<location id="AB6B5C85FF86FFA171F4FE9CFF2CFEBA" LSID="urn:lsid:plazi:treatment:261DBB48FF86FFA1758FFF2CFB23F82F:AB6B5C85FF86FFA171F4FE9CFF2CFEBA" country="Tajikistan" county="Central Asia" latitude="37.378616" longLatPrecision="1" longitude="72.34935" municipality="Autonomous Region" name="Roshtqal'a District" pageId="6" pageNumber="565" stateProvince="Kuhistoni Badakhshon">Roshtqal’a District</location>
,
<location id="AB6B5C85FF86FFA175CDFEF0FE93FEBA" LSID="urn:lsid:plazi:treatment:261DBB48FF86FFA1758FFF2CFB23F82F:AB6B5C85FF86FFA175CDFEF0FE93FEBA" box="[213,376,331,357]" country="Tajikistan" county="Central Asia" latitude="37.378616" longLatPrecision="1" longitude="72.34935" municipality="Autonomous Region" name="Soktosh River" pageId="6" pageNumber="565" stateProvince="Kuhistoni Badakhshon">Soktosh River</location>
,
<quantity id="694CA7BBFF86FFA1749DFEF0FE3DFEBA" box="[389,470,331,357]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="565" unit="km" value="2.3">2.3 km</quantity>
west of the ruins of the
<location id="AB6B5C85FF86FFA177EDFEF0FCA8FEBA" LSID="urn:lsid:plazi:treatment:261DBB48FF86FFA1758FFF2CFB23F82F:AB6B5C85FF86FFA177EDFEF0FCA8FEBA" box="[757,835,331,357]" country="Tajikistan" county="Central Asia" latitude="37.378616" longLatPrecision="1" longitude="72.34935" municipality="Autonomous Region" name="Derouj" pageId="6" pageNumber="565" stateProvince="Kuhistoni Badakhshon">Derouj</location>
fortress,
<date id="DA0A2C9EFF86FFA176A8FEF0FBC5FEBA" box="[944,1070,331,357]" pageId="6" pageNumber="565" value="2016-07-02">
<collectingDate id="CA4ED576FF86FFA176A8FEF0FBC5FEBA" box="[944,1070,331,357]" pageId="6" pageNumber="565" value="2016-07-02">2.VII.2016</collectingDate>
</date>
, alt.
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<quantity id="694CA7BBFF86FFA17170FEF0FB54FEB9" box="[1128,1215,331,358]" metricMagnitude="3" metricUnit="m" metricValue="3.366" pageId="6" pageNumber="565" unit="m" value="3366.0">3366 m</quantity>
above sea level
</elevation>
,
<geoCoordinate id="CB806C99FF86FFA17090FEF7FEFCFE56" degrees="37" direction="north" minutes="22.717" orientation="latitude" pageId="6" pageNumber="565" precision="1" value="37.378616">N 37°22.717’</geoCoordinate>
,
<geoCoordinate id="CB806C99FF86FFA1743AFECBFE51FE56" box="[290,442,367,394]" degrees="72" direction="east" minutes="20.961" orientation="longitude" pageId="6" pageNumber="565" precision="1" value="72.34935">E 72°20.961’</geoCoordinate>
, leg.
<collectorName id="03416F88FF86FFA174EDFECBFD9AFE56" box="[501,625,367,394]" pageId="6" pageNumber="565">D. Palatov.</collectorName>
</materialsCitation>
<typeStatus id="710FB4FCFF86FFA1776FFECBFD03FE55" box="[631,744,368,394]" pageId="6" pageNumber="565" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="B8B2C1D7FF86FFA177EBFED4FCD4FE56" box="[755,831,367,393]" pageId="6" pageNumber="565" type="adult">1 adult</specimenCount>
male,
<materialsCitation id="1EDC0003FF86FFA17692FED4FB12FE56" ID-GBIF-Occurrence="3071287308" box="[906,1273,367,394]" collectingDate="2016-07-02" collectorName="D. Palatov." country="Tajikistan" county="Central Asia" elevation="3366" latitude="37.378616" location="Roshtqal'a District" longLatPrecision="1" longitude="72.34935" municipality="Autonomous Region" pageId="6" pageNumber="565" specimenCount="12" specimenCount-larva="12" stateProvince="Kuhistoni Badakhshon" typeStatus="paratype">
<specimenCount id="B8B2C1D7FF86FFA17692FED4FC1FFE56" box="[906,1012,367,393]" pageId="6" pageNumber="565" type="larva">12 larvae</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="1EDC0003FF86FFA1701BFED4FEAFFE2A" ID-GBIF-Occurrence="3071287310" collectingDate="2016-01-07" collectorName="D. Palatov." country="Tajikistan" county="Autonomous Region" elevation="3717" latitude="37.5137" location="Toguzbulok River" longLatPrecision="1" longitude="72.67575" municipality="Shughnon District" pageId="6" pageNumber="565" specimenCount="10" specimenCount-larva="10" stateProvince="Kuhistoni Badakhshon" typeStatus="paratype">
<specimenCount id="B8B2C1D7FF86FFA1701BFED4FA86FE56" box="[1283,1389,367,393]" pageId="6" pageNumber="565" type="larva">10 larvae</specimenCount>
,
<collectingCountry id="D6A34ACEFF86FFA1706FFECBFF07FE72" name="Tajikistan" pageId="6" pageNumber="565">Tajikistan</collectingCountry>
,
<collectingRegion id="6C70C4BCFF86FFA175EFFE28FE3AFE72" box="[247,465,403,429]" country="Tajikistan" name="Kuhistoni Badakhshon" pageId="6" pageNumber="565">Gorno-Badakhshan</collectingRegion>
<collectingCounty id="476A72D2FF86FFA174CEFE28FD2AFE72" box="[470,705,403,429]" pageId="6" pageNumber="565">Autonomous Region</collectingCounty>
,
<collectingMunicipality id="4E6F9024FF86FFA177D3FE28FC71FE72" box="[715,922,403,429]" pageId="6" pageNumber="565">Shughnon District</collectingMunicipality>
, aryk in the valley of the
<location id="AB6B5C85FF86FFA171AAFE28FA95FE72" LSID="urn:lsid:plazi:treatment:261DBB48FF86FFA1758FFF2CFB23F82F:AB6B5C85FF86FFA171AAFE28FA95FE72" box="[1202,1406,403,429]" country="Tajikistan" county="Autonomous Region" latitude="37.5137" longLatPrecision="1" longitude="72.67575" municipality="Shughnon District" name="Toguzbulok River" pageId="6" pageNumber="565" stateProvince="Kuhistoni Badakhshon">Toguzbulok River</location>
, at the beginning of the road to the
<location id="AB6B5C85FF86FFA174EDFE03FD64FE0D" LSID="urn:lsid:plazi:treatment:261DBB48FF86FFA1758FFF2CFB23F82F:AB6B5C85FF86FFA174EDFE03FD64FE0D" box="[501,655,440,466]" country="Tajikistan" county="Autonomous Region" latitude="37.5137" longLatPrecision="1" longitude="72.67575" municipality="Shughnon District" name="Maysara Pass" pageId="6" pageNumber="565" stateProvince="Kuhistoni Badakhshon">Maysara Pass</location>
,
<date id="DA0A2C9EFF86FFA17782FE0CFCF0FE0E" box="[666,795,439,465]" pageId="6" pageNumber="565" value="2016-01-07">
<collectingDate id="CA4ED576FF86FFA17782FE0CFCF0FE0E" box="[666,795,439,465]" pageId="6" pageNumber="565" value="2016-01-07">01.07.2016</collectingDate>
</date>
, alt.
<elevation id="2599ED6DFF86FFA17656FE0CFBB9FE0E" box="[846,1106,439,466]" metricMagnitude="3" metricUnit="m" metricValue="3.717" pageId="6" pageNumber="565" unit="m" value="3717.0">
<quantity id="694CA7BBFF86FFA17656FE0CFC4AFE0D" box="[846,929,439,466]" metricMagnitude="3" metricUnit="m" metricValue="3.717" pageId="6" pageNumber="565" unit="m" value="3717.0">3717 m</quantity>
above sea level
</elevation>
,
<geoCoordinate id="CB806C99FF86FFA17144FE03FB1DFE0E" box="[1116,1270,439,466]" degrees="37" direction="north" minutes="30.822" orientation="latitude" pageId="6" pageNumber="565" precision="1" value="37.5137">N 37°30.822’</geoCoordinate>
,
<geoCoordinate id="CB806C99FF86FFA17019FE03FA7CFE0D" box="[1281,1431,439,466]" degrees="72" direction="east" minutes="40.545" orientation="longitude" pageId="6" pageNumber="565" precision="1" value="72.67575">E 72°40.545’</geoCoordinate>
, leg.
<collectorName id="03416F88FF86FFA175DFFE67FEAFFE2A" box="[199,324,475,502]" pageId="6" pageNumber="565">D. Palatov.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E6AE59D5FF86FFA175DFFE44FF26F8E4" pageId="6" pageNumber="565" type="etymology">
<paragraph id="AE0B0A5EFF86FFA175DFFE44FE94FDE1" blockId="6.[151,1437,295,1186]" pageId="6" pageNumber="565">
<emphasis id="9CC0D64CFF86FFA175DFFE44FE4DFDC6" bold="true" box="[199,422,511,537]" pageId="6" pageNumber="565">Derivatio nominis.</emphasis>
The species is named in honour of the American chironomid taxonomist of
<taxonomicName id="69B471DDFF86FFA17000FE44FE94FDE1" authority="Dean Cyrus Hansen." authorityName="Dean Cyrus Hansen." class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="subFamily" subFamily="Diamesinae">Diamesinae Dean Cyrus Hansen.</taxonomicName>
</paragraph>
<paragraph id="AE0B0A5EFF86FFA175DFFDFCFE4BFDBE" blockId="6.[151,1437,295,1186]" box="[199,416,583,610]" pageId="6" pageNumber="565">
<emphasis id="9CC0D64CFF86FFA175DFFDFCFEADFDBD" bold="true" box="[199,326,583,610]" italics="true" pageId="6" pageNumber="565">Adult male</emphasis>
(n = 1).
</paragraph>
<paragraph id="AE0B0A5EFF86FFA175DFFDD0FD10FD5A" blockId="6.[151,1437,295,1186]" box="[199,763,619,646]" pageId="6" pageNumber="565">Total length 5.0 mm. Total length/wing length 1.0.</paragraph>
<paragraph id="AE0B0A5EFF86FFA175DFFD34FED0FD12" blockId="6.[151,1437,295,1186]" pageId="6" pageNumber="565">Coloration. Head, thorax, legs, and abdomen brown to dark brown; antennae light brown or yellowish-brown; wings greyish.</paragraph>
<paragraph id="AE0B0A5EFF86FFA175DFFD6CFE3AFC5E" blockId="6.[151,1437,295,1186]" pageId="6" pageNumber="565">
Head. Eyes bare and extended dorsomedially. Temporal setae consisting of 4 coronals, 30–33 orbitals, 8 verticals, and 15–18 postorbitals. Clypeus with
<emphasis id="9CC0D64CFF86FFA1776EFD47FD7BFCCA" box="[630,656,764,789]" italics="true" pageId="6" pageNumber="565">ca</emphasis>
90 setae. Antenna with 13 flagellomeres and a well developed plume; pedicel with 2–3 setae; terminal flagellomere with 1 subapical setae 48 μm long. AR 3.80. Palpomere lengths (in μm): 72; 136; 249; 304; 356. Distal part of palpomere 3 with sensilla capitata of 16–20 μm diameter. Palpomeres 1–5 length/head width 1.09.
</paragraph>
<paragraph id="AE0B0A5EFF86FFA175DFFC30FD2BFC32" blockId="6.[151,1437,295,1186]" pageId="6" pageNumber="565">
Thorax.Antepronotum completely covered with 51 setae, 100–268 μm long (
<figureCitation id="368F16DBFF86FFA17107FC30FBB0FC79" box="[1055,1115,907,934]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 1–7. Pagastia (P.) caelestomontana sp. nov. (1–4) and P. (P.) hanseni sp. nov. (5–7), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 – 200 µm, for figures 2, 4, 7 – 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="6" pageNumber="565">Fig 6</figureCitation>
). Acrostichals 14 (80–88 μm long); dorsocentrals 53–54 (168–260 μm long), in posterior part in 4 rows, in middle and anterior parts in 2–3 rows; prealars 47–50 (152–260 μm long); scutellars 73.
</paragraph>
<paragraph id="AE0B0A5EFF86FFA175DFFC4CFB1DFB86" blockId="6.[151,1437,295,1186]" pageId="6" pageNumber="565">
Wing. Length
<quantity id="694CA7BBFF86FFA17471FC4CFE3BFBCD" box="[361,464,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="4.96" pageId="6" pageNumber="565" unit="mm" value="4.96">4.96 mm</quantity>
; width
<quantity id="694CA7BBFF86FFA1773AFC4CFD62FBCD" box="[546,649,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="6" pageNumber="565" unit="mm" value="1.16">1.16 mm</quantity>
. Membrane without setae. R and R
<subScript id="3230081BFF86FFA1710AFBB1FBF1FBC7" attach="left" box="[1042,1050,1034,1048]" fontSize="6" pageId="6" pageNumber="565">1</subScript>
with 21–22 setae; R
<quantity id="694CA7BBFF86FFA17019FBB1FACDFBCE" box="[1281,1318,1015,1048]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="565" unit="in" value="1.0">
<subScript id="3230081BFF86FFA17019FBB1FAE2FBC7" attach="left" box="[1281,1289,1034,1048]" fontSize="6" pageId="6" pageNumber="565">1</subScript>
in
</quantity>
distal part without setae; R
<subScript id="3230081BFF86FFA17456FB96FE83FBE4" attach="left" box="[334,360,1069,1083]" fontSize="6" pageId="6" pageNumber="565">4+5</subScript>
without setae. Costa extension 72 μm long. RM length/MCu length 3.25. Anal lobe developed, rounded and slightly protrude. Squama with 36–44 setae in 2–3 rows. Alula without setae. VR 0.84.
</paragraph>
<paragraph id="AE0B0A5EFF86FFA175DFFBDFFAF1FB7E" blockId="6.[151,1437,295,1186]" pageId="6" pageNumber="565">
Legs. Spur of fore tibia 128 μm long; spurs of mid tibia 92 μm and 100 μm long; spurs of hind tibia 136 μm and 92 μm long. Hind tibial comb with 12–13 setae. Lengths and proportions of leg segments as in
<tableCitation id="E3363FE5FF86FFA171D8FB3CFAFDFB7E" box="[1216,1302,1159,1185]" captionStart="TABLE 2" captionStartId="6.[151,239,1233,1257]" captionTargetPageId="6" captionText="TABLE 2. Lengths (in µm) and proportions of leg segments of Pagastia (P.) hanseni sp. nov., male (n=1)." pageId="6" pageNumber="565">Table 2</tableCitation>
.
</paragraph>
<caption id="FACB5AD6FF86FFA1758FFB6AFAE1FB36" box="[151,1290,1233,1257]" pageId="6" pageNumber="565" startId="6.[151,239,1233,1257]" targetBox="[166,1385,1273,1424]" targetIsTable="true" targetPageId="6">
<paragraph id="AE0B0A5EFF86FFA1758FFB6AFAE1FB36" blockId="6.[151,1290,1233,1257]" box="[151,1290,1233,1257]" pageId="6" pageNumber="565">
<emphasis id="9CC0D64CFF86FFA1758FFB6AFEE1FB36" bold="true" box="[151,266,1233,1257]" pageId="6" pageNumber="565">TABLE 2.</emphasis>
Lengths (in µm) and proportions of leg segments of
<taxonomicName id="69B471DDFF86FFA17627FB69FBCFFB36" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2021" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2021" box="[831,1060,1233,1257]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="hanseni" status="sp. nov." subGenus="Pagastia">
<emphasis id="9CC0D64CFF86FFA17627FB69FBCFFB36" box="[831,1060,1233,1257]" italics="true" pageId="6" pageNumber="565">Pagastia (P.) hanseni</emphasis>
</taxonomicName>
<emphasis id="9CC0D64CFF86FFA17132FB6AFB6BFB36" bold="true" box="[1066,1152,1233,1257]" pageId="6" pageNumber="565">
<taxonomicNameLabel id="87F36B37FF86FFA17132FB6AFB6BFB36" box="[1066,1152,1233,1257]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (n=1).
</paragraph>
</caption>
<paragraph id="AE0B0A5EFF86FFA175E7FB42FA8CFA55" pageId="6" pageNumber="565">
<table id="DCB4F8FEFF86005875BEFB42FA82FA4F" box="[166,1385,1273,1424]" colsContinueFrom="3.[166,1383,1514,1665]" gridcols="12" gridrows="4" pageId="6" pageNumber="565">
<tr id="1084081CFF86005875BEFB42FA82FAC7" box="[166,1385,1273,1304]" gridrow="0" pageId="6" pageNumber="565" rowspan-0="1">
<th id="53556160FF86005875E7FB42FED9FAC7" box="[255,306,1273,1304]" gridcol="1" gridrow="0" pageId="6" pageNumber="565">fe</th>
<th id="53556160FF8600587465FB42FE5BFAC7" box="[381,432,1273,1304]" gridcol="2" gridrow="0" pageId="6" pageNumber="565">ti</th>
<th id="53556160FF86005874E3FB42FDC5FAC7" box="[507,558,1273,1304]" gridcol="3" gridrow="0" pageId="6" pageNumber="565">ta1</th>
<th id="53556160FF8600587760FB42FD75FAC7" box="[632,670,1273,1304]" gridcol="4" gridrow="0" pageId="6" pageNumber="565">ta2</th>
<th id="53556160FF86005877C7FB42FCEEFAC7" box="[735,773,1273,1304]" gridcol="5" gridrow="0" pageId="6" pageNumber="565">ta3</th>
<th id="53556160FF860058765DFB42FC80FAC7" box="[837,875,1273,1304]" gridcol="6" gridrow="0" pageId="6" pageNumber="565">ta4</th>
<th id="53556160FF86005876B4FB42FC39FAC7" box="[940,978,1273,1304]" gridcol="7" gridrow="0" pageId="6" pageNumber="565">ta 5</th>
<th id="53556160FF860058710BFB42FBD4FAC7" box="[1043,1087,1273,1304]" gridcol="8" gridrow="0" pageId="6" pageNumber="565">LR</th>
<th id="53556160FF8600587161FB42FB4EFAC7" box="[1145,1189,1273,1304]" gridcol="9" gridrow="0" pageId="6" pageNumber="565">BV</th>
<th id="53556160FF86005871F8FB42FAE7FAC7" box="[1248,1292,1273,1304]" gridcol="10" gridrow="0" pageId="6" pageNumber="565">SV</th>
<th id="53556160FF860058705FFB42FA82FAC7" box="[1351,1385,1273,1304]" gridcol="11" gridrow="0" pageId="6" pageNumber="565">BR</th>
</tr>
<tr id="1084081CFF86005875BEFA9AFA82FAE0" box="[166,1385,1313,1343]" gridrow="1" pageId="6" pageNumber="565">
<th id="53556160FF86005875BEFA9AFF50FAE0" box="[166,187,1313,1343]" gridcol="0" gridrow="1" pageId="6" pageNumber="565">P1</th>
<td id="53556160FF86005875E7FA9AFED9FAE0" box="[255,306,1313,1343]" gridcol="1" gridrow="1" pageId="6" pageNumber="565">1440</td>
<td id="53556160FF8600587465FA9AFE5BFAE0" box="[381,432,1313,1343]" gridcol="2" gridrow="1" pageId="6" pageNumber="565">1560</td>
<td id="53556160FF86005874E3FA9AFDC5FAE0" box="[507,558,1313,1343]" gridcol="3" gridrow="1" pageId="6" pageNumber="565">1120</td>
<td id="53556160FF8600587760FA9AFD75FAE0" box="[632,670,1313,1343]" gridcol="4" gridrow="1" pageId="6" pageNumber="565">640</td>
<td id="53556160FF86005877C7FA9AFCEEFAE0" box="[735,773,1313,1343]" gridcol="5" gridrow="1" pageId="6" pageNumber="565">400</td>
<td id="53556160FF860058765DFA9AFC80FAE0" box="[837,875,1313,1343]" gridcol="6" gridrow="1" pageId="6" pageNumber="565">240</td>
<td id="53556160FF86005876B4FA9AFC39FAE0" box="[940,978,1313,1343]" gridcol="7" gridrow="1" pageId="6" pageNumber="565">240</td>
<td id="53556160FF860058710BFA9AFBD4FAE0" box="[1043,1087,1313,1343]" gridcol="8" gridrow="1" pageId="6" pageNumber="565">0.72</td>
<td id="53556160FF8600587161FA9AFB4EFAE0" box="[1145,1189,1313,1343]" gridcol="9" gridrow="1" pageId="6" pageNumber="565">2.72</td>
<td id="53556160FF86005871F8FA9AFAE7FAE0" box="[1248,1292,1313,1343]" gridcol="10" gridrow="1" pageId="6" pageNumber="565">2.68</td>
<td id="53556160FF860058705FFA9AFA82FAE0" box="[1351,1385,1313,1343]" gridcol="11" gridrow="1" pageId="6" pageNumber="565">2.8</td>
</tr>
<tr id="1084081CFF86005875BEFAF2FA82FAB8" box="[166,1385,1353,1383]" gridrow="2" pageId="6" pageNumber="565">
<th id="53556160FF86005875BEFAF2FF50FAB8" box="[166,187,1353,1383]" gridcol="0" gridrow="2" pageId="6" pageNumber="565">P2</th>
<td id="53556160FF86005875E7FAF2FED9FAB8" box="[255,306,1353,1383]" gridcol="1" gridrow="2" pageId="6" pageNumber="565">1560</td>
<td id="53556160FF8600587465FAF2FE5BFAB8" box="[381,432,1353,1383]" gridcol="2" gridrow="2" pageId="6" pageNumber="565">1440</td>
<td id="53556160FF86005874E3FAF2FDC5FAB8" box="[507,558,1353,1383]" gridcol="3" gridrow="2" pageId="6" pageNumber="565">740</td>
<td id="53556160FF8600587760FAF2FD75FAB8" box="[632,670,1353,1383]" gridcol="4" gridrow="2" pageId="6" pageNumber="565">460</td>
<td id="53556160FF86005877C7FAF2FCEEFAB8" box="[735,773,1353,1383]" gridcol="5" gridrow="2" pageId="6" pageNumber="565">320</td>
<td id="53556160FF860058765DFAF2FC80FAB8" box="[837,875,1353,1383]" gridcol="6" gridrow="2" pageId="6" pageNumber="565">200</td>
<td id="53556160FF86005876B4FAF2FC39FAB8" box="[940,978,1353,1383]" gridcol="7" gridrow="2" pageId="6" pageNumber="565">240</td>
<td id="53556160FF860058710BFAF2FBD4FAB8" box="[1043,1087,1353,1383]" gridcol="8" gridrow="2" pageId="6" pageNumber="565">0.51</td>
<td id="53556160FF8600587161FAF2FB4EFAB8" box="[1145,1189,1353,1383]" gridcol="9" gridrow="2" pageId="6" pageNumber="565">3.07</td>
<td id="53556160FF86005871F8FAF2FAE7FAB8" box="[1248,1292,1353,1383]" gridcol="10" gridrow="2" pageId="6" pageNumber="565">4.05</td>
<td id="53556160FF860058705FFAF2FA82FAB8" box="[1351,1385,1353,1383]" gridcol="11" gridrow="2" pageId="6" pageNumber="565">2.4</td>
</tr>
<tr id="1084081CFF86005875BEFAC9FA82FA4F" box="[166,1385,1394,1424]" gridrow="3" pageId="6" pageNumber="565">
<th id="53556160FF86005875BEFAC9FF50FA4F" box="[166,187,1394,1424]" gridcol="0" gridrow="3" pageId="6" pageNumber="565">P 3</th>
<td id="53556160FF86005875E7FAC9FED9FA4F" box="[255,306,1394,1424]" gridcol="1" gridrow="3" pageId="6" pageNumber="565">1900</td>
<td id="53556160FF8600587465FAC9FE5BFA4F" box="[381,432,1394,1424]" gridcol="2" gridrow="3" pageId="6" pageNumber="565">2080</td>
<td id="53556160FF86005874E3FAC9FDC5FA4F" box="[507,558,1394,1424]" gridcol="3" gridrow="3" pageId="6" pageNumber="565">1200</td>
<td id="53556160FF8600587760FAC9FD75FA4F" box="[632,670,1394,1424]" gridcol="4" gridrow="3" pageId="6" pageNumber="565">700</td>
<td id="53556160FF86005877C7FAC9FCEEFA4F" box="[735,773,1394,1424]" gridcol="5" gridrow="3" pageId="6" pageNumber="565">400</td>
<td id="53556160FF860058765DFAC9FC80FA4F" box="[837,875,1394,1424]" gridcol="6" gridrow="3" pageId="6" pageNumber="565">280</td>
<td id="53556160FF86005876B4FAC9FC39FA4F" box="[940,978,1394,1424]" gridcol="7" gridrow="3" pageId="6" pageNumber="565">260</td>
<td id="53556160FF860058710BFAC9FBD4FA4F" box="[1043,1087,1394,1424]" gridcol="8" gridrow="3" pageId="6" pageNumber="565">0.58</td>
<td id="53556160FF8600587161FAC9FB4EFA4F" box="[1145,1189,1394,1424]" gridcol="9" gridrow="3" pageId="6" pageNumber="565">5.51</td>
<td id="53556160FF86005871F8FAC9FAE7FA4F" box="[1248,1292,1394,1424]" gridcol="10" gridrow="3" pageId="6" pageNumber="565">3.32</td>
<td id="53556160FF860058705FFAC9FA82FA4F" box="[1351,1385,1394,1424]" gridcol="11" gridrow="3" pageId="6" pageNumber="565">4.5</td>
</tr>
</table>
</paragraph>
<paragraph id="AE0B0A5EFF86FFA175DFFA02FE1EF90F" blockId="6.[151,1443,1465,2032]" pageId="6" pageNumber="565">
Hypopygium (
<figureCitation id="368F16DBFF86FFA17473FA02FE3CFA0B" box="[363,471,1465,1492]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 1–7. Pagastia (P.) caelestomontana sp. nov. (1–4) and P. (P.) hanseni sp. nov. (5–7), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 – 200 µm, for figures 2, 4, 7 – 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="6" pageNumber="565">Figs. 5, 7</figureCitation>
,
<figureCitation id="368F16DBFF86FFA174FBFA01FDC7FA0B" box="[483,556,1466,1492]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1704,1728]" captionTargetBox="[151,1435,429,1666]" captionTargetId="figure-134@5.[151,1435,417,1680]" captionTargetPageId="5" captionText="FIGURES 8–11. Pagastia (P.) caelestomontana sp. nov. (8–9) and P. (P.) hanseni sp. nov. (10–11), males. 8, 10, hypopygium in dorsal view; 9, gonostylus; 11, anal point." figureDoi="http://doi.org/10.5281/zenodo.4668491" httpUri="https://zenodo.org/record/4668491/files/figure.png" pageId="6" pageNumber="565">10–11</figureCitation>
). Tergite IX with 29–30 setae on each side and with an anal point 88 μm long, apically with peg 28 μm long. Tergite IX laterally from anal point on each side with rounded small lobe (
<figureCitation id="368F16DBFF86FFA17024FA66FA78FA27" box="[1340,1427,1501,1528]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1704,1728]" captionTargetBox="[151,1435,429,1666]" captionTargetId="figure-134@5.[151,1435,417,1680]" captionTargetPageId="5" captionText="FIGURES 8–11. Pagastia (P.) caelestomontana sp. nov. (8–9) and P. (P.) hanseni sp. nov. (10–11), males. 8, 10, hypopygium in dorsal view; 9, gonostylus; 11, anal point." figureDoi="http://doi.org/10.5281/zenodo.4668491" httpUri="https://zenodo.org/record/4668491/files/figure.png" pageId="6" pageNumber="565">Fig. 11</figureCitation>
). Laterosternite IX with 11–12 setae. Transverse sternapodeme trapezoidal, 212 μm long. Phallapodeme 48–52 μm long. Gonocoxite 384–392 μm long; basal lobe of medium size, rounded, with long setae along inner margin and with 4 setae at inner part, 28–32 μm long (
<figureCitation id="368F16DBFF86FFA1776BF9F2FD29F9BB" box="[627,706,1609,1636]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 1–7. Pagastia (P.) caelestomontana sp. nov. (1–4) and P. (P.) hanseni sp. nov. (5–7), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 – 200 µm, for figures 2, 4, 7 – 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="6" pageNumber="565">Figs. 5</figureCitation>
,
<figureCitation id="368F16DBFF86FFA177D6F9F1FD06F9BB" box="[718,749,1610,1636]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1704,1728]" captionTargetBox="[151,1435,429,1666]" captionTargetId="figure-134@5.[151,1435,417,1680]" captionTargetPageId="5" captionText="FIGURES 8–11. Pagastia (P.) caelestomontana sp. nov. (8–9) and P. (P.) hanseni sp. nov. (10–11), males. 8, 10, hypopygium in dorsal view; 9, gonostylus; 11, anal point." figureDoi="http://doi.org/10.5281/zenodo.4668491" httpUri="https://zenodo.org/record/4668491/files/figure.png" pageId="6" pageNumber="565">10</figureCitation>
). Median aedeagal lobe 72 μm long, wedge-shaped, with fin- gerlike apex, 72 μm long; lateral aedeagal lobe narrow, 132–140 μm long (
<figureCitation id="368F16DBFF86FFA176D7F9D6FBF9F957" box="[975,1042,1645,1672]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 1–7. Pagastia (P.) caelestomontana sp. nov. (1–4) and P. (P.) hanseni sp. nov. (5–7), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 – 200 µm, for figures 2, 4, 7 – 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="6" pageNumber="565">Fig. 7</figureCitation>
). Gonostylus 280 μm long, slender, almost straight, of same thickness, sometimes slightly widened at the base, slightly curved apically with megaseta 16–20 μm long. HR 1.37–1.40.
</paragraph>
<paragraph id="AE0B0A5EFF86FFA175DFF961FE90F92C" blockId="6.[151,1443,1465,2032]" box="[199,379,1753,1780]" pageId="6" pageNumber="565">
<emphasis id="9CC0D64CFF86FFA175DFF961FEEFF92B" bold="true" box="[199,260,1754,1780]" italics="true" pageId="6" pageNumber="565">Pupa</emphasis>
unknown.
</paragraph>
<paragraph id="AE0B0A5EFF86FFA175DFF945FF26F8E4" blockId="6.[151,1443,1465,2032]" pageId="6" pageNumber="565">
<emphasis id="9CC0D64CFF86FFA175DFF945FEE0F8C7" bold="true" box="[199,267,1790,1816]" italics="true" pageId="6" pageNumber="565">Larva</emphasis>
not separable from
<emphasis id="9CC0D64CFF86FFA174E2F945FDFBF8C8" box="[506,528,1790,1815]" italics="true" pageId="6" pageNumber="565">P.</emphasis>
<taxonomicName id="69B471DDFF86FFA1770FF945FCE3F8C8" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2021" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2021" box="[535,776,1789,1816]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="caelestomontana" status="sp. nov." subGenus="Pagastia">
(
<emphasis id="9CC0D64CFF86FFA17739F945FDD9F8C8" box="[545,562,1790,1815]" italics="true" pageId="6" pageNumber="565">P</emphasis>
.)
<emphasis id="9CC0D64CFF86FFA17752F946FCE3F8C8" box="[586,776,1789,1815]" italics="true" pageId="6" pageNumber="565">caelestomontana</emphasis>
</taxonomicName>
<emphasis id="9CC0D64CFF86FFA17609F945FC98F8C7" bold="true" box="[785,883,1790,1816]" pageId="6" pageNumber="565">
<taxonomicNameLabel id="87F36B37FF86FFA17609F945FC86F8C7" box="[785,877,1790,1816]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
can be distinguished only by molecular genetic data.
</paragraph>
</subSubSection>
<subSubSection id="E6AE59D5FF86FFA175DFF8FEFC36F85C" pageId="6" pageNumber="565" type="diagnosis">
<paragraph id="AE0B0A5EFF86FFA175DFF8FEFC36F85C" blockId="6.[151,1443,1465,2032]" pageId="6" pageNumber="565">
<emphasis id="9CC0D64CFF86FFA175DFF8FEFEA9F880" bold="true" box="[199,322,1861,1887]" pageId="6" pageNumber="565">Diagnosis.</emphasis>
The male of this new species is most closely related to
<emphasis id="9CC0D64CFF86FFA176D3F8FDFC0BF880" box="[971,992,1862,1887]" italics="true" pageId="6" pageNumber="565">P.</emphasis>
<taxonomicName id="69B471DDFF86FFA176F1F8FDFB30F880" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2021" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2021" box="[1001,1243,1861,1888]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="caelestomontana" status="sp. nov." subGenus="Pagastia">
(
<emphasis id="9CC0D64CFF86FFA176EAF8FDFBE8F880" box="[1010,1027,1862,1887]" italics="true" pageId="6" pageNumber="565">P</emphasis>
.)
<emphasis id="9CC0D64CFF86FFA17105F8FEFB30F880" box="[1053,1243,1861,1887]" italics="true" pageId="6" pageNumber="565">caelestomontana</emphasis>
</taxonomicName>
<emphasis id="9CC0D64CFF86FFA171FCF8FDFAD4F8BF" bold="true" box="[1252,1343,1862,1888]" pageId="6" pageNumber="565">
<taxonomicNameLabel id="87F36B37FF86FFA171FCF8FDFAD4F8BF" box="[1252,1343,1862,1888]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
See the diagnosis of this species for details, the key and results of DNA barcoding.
</paragraph>
</subSubSection>
<subSubSection id="E6AE59D5FF86FFA175DFF835FA8BF813" pageId="6" pageNumber="565" type="biology_ecology">
<paragraph id="AE0B0A5EFF86FFA175DFF835FA8BF813" blockId="6.[151,1443,1465,2032]" pageId="6" pageNumber="565">
<emphasis id="9CC0D64CFF86FFA175DFF835FEC7F877" bold="true" box="[199,300,1934,1960]" pageId="6" pageNumber="565">Ecology.</emphasis>
Larvae were collected from stones and bowlders sometimes covered with algae and moss in mountain rivers, located at an altitude of
<quantity id="694CA7BBFF86FFA174E2F80AFD75F813" box="[506,670,1969,1996]" metricMagnitude="3" metricUnit="m" metricValue="2.8665" metricValueMax="3.717" metricValueMin="2.016" pageId="6" pageNumber="565" unit="m" value="2866.5" valueMax="3717.0" valueMin="2016.0">2016–3717 m</quantity>
, at a flow rate of 0.4–1.0 m/s, with water temperature
<emphasis id="9CC0D64CFF86FFA171E6F809FAF3F814" box="[1278,1304,1970,1995]" italics="true" pageId="6" pageNumber="565">ca</emphasis>
12°C.
</paragraph>
</subSubSection>
<subSubSection id="E6AE59D5FF86FFA175DFF86EFB23F82F" box="[199,1224,2005,2032]" pageId="6" pageNumber="565" type="distribution">
<paragraph id="AE0B0A5EFF86FFA175DFF86EFB23F82F" blockId="6.[151,1443,1465,2032]" box="[199,1224,2005,2032]" pageId="6" pageNumber="565">
<emphasis id="9CC0D64CFF86FFA175DFF86EFE8AF830" bold="true" box="[199,353,2005,2031]" pageId="6" pageNumber="565">Distribution.</emphasis>
Known only from the
<typeStatus id="710FB4FCFF86FFA17778F86DFD7BF82F" box="[608,656,2006,2032]" pageId="6" pageNumber="565">type</typeStatus>
locality in Pamir Mountains (
<collectingCountry id="D6A34ACEFF86FFA176C7F86EFBBAF82F" box="[991,1105,2005,2032]" name="Tajikistan" pageId="6" pageNumber="565">Tajikistan</collectingCountry>
) (
<figureCitation id="368F16DBFF86FFA17170F86EFB50F82F" box="[1128,1211,2005,2032]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1495,1519]" captionTargetBox="[151,1435,194,1457]" captionTargetId="figure-18@7.[151,1435,181,1471]" captionTargetPageId="7" captionText="FIGURES 12–15. Type localities of Pagastia (P.) caelestomontana sp. nov. (12), P. (P.) hanseni sp. nov. (13), and P. (P.) aff. lanceolata (Tokunaga) (14–15). 12, Otmek River, Talas district, Kirgizstan (photo by E.S. Chertoprud); 13, Soktosh River, Gorno-Badakhshan Autonomous Region, Roshtqal’a District, Tajikistan (photo by D.M. Palatov); 14, Tuzumtaikul River, Gorno- Badakhshan Autonomous Region, Shughnon District, Tajikistan (photo by M.V. Vinarski); 15, spring in front of the mouth of the Bartang River, Gorno-Badakhshan Autonomous Region, Rushanskiy District, Tajikistan (photo by M.V. Vinarski)." figureDoi="http://doi.org/10.5281/zenodo.4668493" httpUri="https://zenodo.org/record/4668493/files/figure.png" pageId="6" pageNumber="565">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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