treatments-xml/data/C0/33/D7/C033D7104F59FFCA4EA4FE53FB7CF9CB.xml

<document id="70E301B0088C53CB9FF9F16FA1F29A2D" ID="10.11646/zootaxa.4222.1.1" ID-CLB-Dataset="37268" ID-DOI="10.5281/zenodo.257769" ID-GBIF-Dataset="9ea59a17-588e-4af1-8c0d-ebcd50ad0395" ID-ISSN="1175-5326" ID-Zenodo-Dep="257769" ID-ZooBank="F0EC0A87-B1EE-4B5C-8F53-77A7EEA75F3A" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1485156926626" checkinUser="plazi" docAuthor="Lima, Flávio C. T." docDate="2017" docId="C033D7104F59FFCA4EA4FE53FB7CF9CB" docLanguage="en" docName="zootaxa.4222.1.1.pdf" docOrigin="Zootaxa 4222 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Brycon howesi Lima, 2017, new species" docType="treatment" docVersion="8" lastPageNumber="51" masterDocId="3C0AAF684F77FFF84E33FFC7FFD2FFE7" masterDocTitle="A revision of the cis-andean species of the genus Brycon Müller &amp; Troschel (Characiformes: Characidae)" masterLastPageNumber="189" masterPageNumber="1" pageNumber="47" updateTime="1698436136809" updateUser="ExternalLinkService">
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<mods:title id="6F56D0B24DEFD7300B2C1D95E248B678">A revision of the cis-andean species of the genus Brycon Müller &amp; Troschel (Characiformes: Characidae)</mods:title>
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<mods:namePart id="38DF1939CDD94DFAA5DD4FF1045D9278">Lima, Flávio C. T.</mods:namePart>
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<mods:date id="23428E1EA88505768FA941F9882A500B">2017</mods:date>
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<paragraph id="482566064F59FFD64EA4FE53FE32FE49" blockId="46.[151,480,404,463]" box="[151,480,404,430]" pageId="46" pageNumber="47">
<heading id="136DD16A4F59FFD64EA4FE53FE32FE49" bold="true" box="[151,480,404,430]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
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<taxonomicName id="8F9A1D854F59FFD64EA4FE53FE9BFE49" box="[151,329,404,430]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="howesi" status="sp. nov.">
<emphasis id="7AEEBA144F59FFD64EA4FE53FE9BFE49" bold="true" box="[151,329,404,430]" italics="true" pageId="46" pageNumber="47">Brycon howesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61DD076F4F59FFD64F63FE53FE32FE49" box="[336,480,404,430]" pageId="46" pageNumber="47" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="482566064F59FFD64EA4FE71FEFCFE28" blockId="46.[151,480,404,463]" box="[151,302,438,463]" pageId="46" pageNumber="47">
(
<figureCitation id="D0A17A834F59FFD64EACFE71FEF4FE28" box="[159,294,438,463]" captionStart-0="FIGURE 28" captionStart-1="FIGURE 29" captionStartId-0="46.[151,250,1948,1970]" captionStartId-1="47.[151,250,602,624]" captionTargetBox-0="[151,1436,1486,1927]" captionTargetBox-1="[151,1436,163,580]" captionTargetId-0="figure@46.[151,1436,1486,1927]" captionTargetId-1="figure@47.[151,1436,163,580]" captionTargetPageId-0="46" captionTargetPageId-1="47" captionText-0="FIGURE 28. Brycon howesi, new species, holotype, MZUSP 53803, 234.8 mm SL: Brazil, Minas Gerais, rio Jequitinhonha." captionText-1="FIGURE 29. Brycon howesi, new species, paratype, MZUSP 101506, 167.1 mm SL: Brazil, Minas Gerais, Rio Preto." httpUri-0="https://zenodo.org/record/257797/files/figure.png" httpUri-1="https://zenodo.org/record/257798/files/figure.png" pageId="46" pageNumber="47">Figs. 28–29</figureCitation>
)
</paragraph>
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<subSubSection id="0080358D4F59FFD64EA4FE39FAD5FDF3" box="[151,1287,510,532]" pageId="46" pageNumber="47" type="description">
<paragraph id="482566064F59FFD64EA4FE39FAD5FDF3" blockId="46.[151,1436,510,717]" box="[151,1287,510,532]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="688A41284F59FFD64EA4FE39FAD5FDF3" box="[151,1287,510,532]" pageId="46" pageNumber="47">
<taxonomicName id="8F9A1D854F59FFD64EA4FE39FE80FDF3" box="[151,338,510,532]" class="Actinopterygii" family="Characidae" genus="Chalceus" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F59FFD64EA4FE39FE80FDF3" box="[151,338,510,532]" italics="true" pageId="46" pageNumber="47">Chalceus opalinus</emphasis>
</taxonomicName>
(non Cuvier): Valenciennes, in 
<treatmentCitation id="C93B40174F59FFD64CA7FE39FC23FDF3" author="Cuvier" box="[660,1009,510,532]" page="246" pageId="46" pageNumber="47" year="1850">
<bibRefCitation id="2C0B1BF74F59FFD64CA7FE39FC13FDF3" author="Cuvier" box="[660,961,510,532]" pageId="46" pageNumber="47" refString="Cuvier, G. &amp; Valenciennes, A. (1850) Histoire naturelle des poissons. Tome vingt-deucieme. P. Bertrand editeur, Paris, i - xx, 1 - 532 + 1 - 91, pls. 634 - 650." type="book chapter" year="1850">Cuvier &amp; Valenciennes, 1850</bibRefCitation>
: 246
</treatmentCitation>
(part; “ 
<collectingRegion id="8A5EA8E44F59FFD64A0CFE39FBB7FDF3" box="[1087,1125,510,532]" country="Nigeria" name="Rivers" pageId="46" pageNumber="47">Rio</collectingRegion>
Tiquilelonha”).
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</paragraph>
</subSubSection>
<subSubSection id="0080358D4F59FFD64EA4FDD9FBFFFD97" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="482566064F59FFD64EA4FDD9FE2CFDB5" blockId="46.[151,1436,510,717]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="688A41284F59FFD64EA4FDD9FE2CFDB5" pageId="46" pageNumber="47">
<taxonomicName id="8F9A1D854F59FFD64EA4FDD9FD86FDD4" ID-CoL="NH7J" authority="Steindachner, 1877: 587" authorityName="Steindachner" authorityPageNumber="587" authorityYear="1877" box="[151,596,541,563]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F59FFD64EA4FDD9FEEEFDD4" box="[151,316,542,563]" italics="true" pageId="46" pageNumber="47">Brycon insignis</emphasis>
: 
<treatmentCitation id="C93B40174F59FFD64F62FDDAFD86FDD4" author="Steindachner" box="[337,596,541,563]" page="587" pageId="46" pageNumber="47" year="1877">
<bibRefCitation id="2C0B1BF74F59FFD64F62FDDAFDCFFDD4" author="Steindachner" box="[337,541,541,563]" pageId="46" pageNumber="47" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner, 1877</bibRefCitation>
: 587
</treatmentCitation>
</taxonomicName>
–591 (part; “ 
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Jequitinhonha”); 
<treatmentCitation id="C93B40174F59FFD64DFEFDDAFB22FDD4" author="Godinho" box="[973,1264,541,563]" page="415" pageId="46" pageNumber="47" year="1999">
<bibRefCitation id="2C0B1BF74F59FFD64DFEFDDAFB65FDD4" author="Godinho" box="[973,1207,541,563]" pageId="46" pageNumber="47" refString="Godinho, H. P., Godinho, A. L. &amp; Vono, V. (1999) Peixes da bacia do rio Jequitinhonha. In: R. H. Lowe-McConnell, Estudos ecologicos de comunidades de peixes tropicais. EDUSP, Sao Paulo, pp. 414 - 423." type="book chapter" year="1999">
Godinho 
<emphasis id="7AEEBA144F59FFD64A00FDD9FBBFFDD4" box="[1075,1133,541,563]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
, 1999
</bibRefCitation>
: 415
</treatmentCitation>
, 418, 421 (
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Jequitinhonha basin; ecology).
</treatmentCitationGroup>
</paragraph>
<paragraph id="482566064F59FFD64EA4FD9CFBFFFD97" blockId="46.[151,1436,510,717]" box="[151,1069,602,624]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="688A41284F59FFD64EA4FD9CFBFFFD97" box="[151,1069,602,624]" pageId="46" pageNumber="47">
<taxonomicName id="8F9A1D854F59FFD64EA4FD9CFEEFFD97" ID-CoL="NH83" box="[151,317,602,624]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F59FFD64EA4FD9CFEEFFD97" box="[151,317,602,624]" italics="true" pageId="46" pageNumber="47">Brycon opalinus</emphasis>
</taxonomicName>
(non Cuvier): 
<treatmentCitation id="C93B40174F59FFD64FE0FD9DFD3FFD97" author="Gery" box="[467,749,602,624]" page="813" pageId="46" pageNumber="47" year="1992">
<bibRefCitation id="2C0B1BF74F59FFD64FE0FD9DFD6DFD97" author="Gery" box="[467,703,602,624]" pageId="46" pageNumber="47" refString="Gery, J. &amp; Mahnert, V. (1992) Notes sur quelques Brycon des bassins de l'Amazone, du Parana-Paraguay et du Sud-Est bresilien (Pisces, Characiformes, Characidae). Revue Suisse de Zoologie, 99 (4), 793 - 819." type="journal article" year="1992">Géry &amp; Mahnert, 1992</bibRefCitation>
: 813
</treatmentCitation>
–814 (part; 
<collectingRegion id="8A5EA8E44F59FFD64D57FD9DFC5AFD97" box="[868,904,602,624]" country="Nigeria" name="Rivers" pageId="46" pageNumber="47">Rio</collectingRegion>
Jequitinhonha).
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</paragraph>
</subSubSection>
<subSubSection id="0080358D4F59FFD64EA4FDBDFBC6FD2B" pageId="46" pageNumber="47" type="description">
<paragraph id="482566064F59FFD64EA4FDBDFAD3FD49" blockId="46.[151,1436,510,717]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="688A41284F59FFD64EA4FDBDFAD3FD49" bridged="true" pageId="46" pageNumber="47">
<taxonomicName id="8F9A1D854F59FFD64EA4FDBDFEE0FD68" ID-CoL="NH77" box="[151,306,633,655]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="devillei">
<emphasis id="7AEEBA144F59FFD64EA4FDBDFEE0FD68" box="[151,306,633,655]" italics="true" pageId="46" pageNumber="47">Brycon devillei</emphasis>
</taxonomicName>
(non Castelnau): Vieira 
<emphasis id="7AEEBA144F59FFD64C1FFDBDFDB2FD68" box="[556,608,633,655]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
, 2008: 47–48 (part; 
<collectingRegion id="8A5EA8E44F59FFD64D02FDBEFC84FD68" box="[817,854,633,655]" country="Nigeria" name="Rivers" pageId="46" pageNumber="47">Rio</collectingRegion>
Jequitinhonha basin; ecology, conservation); 
<treatmentCitation id="C93B40174F59FFD64B1AFDBEFE83FD49" author="Azevedo" page="807" pageId="46" pageNumber="47" year="2011">
<bibRefCitation id="2C0B1BF74F59FFD64B1AFDBEFEF0FD49" author="Azevedo" pageId="46" pageNumber="47" refString="Azevedo, P. G., Melo, R. M. C. &amp; Young, R. J. (2011) Feeding and social behavior of the piabanha, Brycon devillei (Castelnau, 1855) (Characidae: Bryconinae) in the wild, with a note on following behavior. Neotropical Ichthyology, 9 (4), 807 - 814. http: // dx. doi. org / 10.1590 / S 1679 - 62252011005000046" type="journal article" year="2011">
Azevedo 
<emphasis id="7AEEBA144F59FFD64BB9FDBDFF33FD49" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
, 2011
</bibRefCitation>
: 807
</treatmentCitation>
–814 (feeding ecology and social behavior, rio Preto, rio Jequitinhonha basin, 
<collectingRegion id="8A5EA8E44F59FFD64A59FD5FFB25FD49" box="[1130,1271,664,686]" country="Brazil" name="Minas Gerais" pageId="46" pageNumber="47">Minas Gerais</collectingRegion>
).
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</paragraph>
<paragraph id="482566064F59FFD64EA4FD7FFBC6FD2B" blockId="46.[151,1436,510,717]" box="[151,1044,694,717]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="688A41284F59FFD64EA4FD7FFBC6FD2B" box="[151,1044,694,717]" pageId="46" pageNumber="47">
<taxonomicName id="8F9A1D854F59FFD64EA4FD7FFF33FD2A" ID-CoL="3DKB" box="[151,225,696,717]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F59FFD64EA4FD7FFF33FD2A" box="[151,225,696,717]" italics="true" pageId="46" pageNumber="47">Brycon</emphasis>
</taxonomicName>
sp.: 
<treatmentCitation id="C93B40174F59FFD64F23FD71FDDAFD2B" author="Pugedo" box="[272,520,694,717]" page="345" pageId="46" pageNumber="47" year="2016">
<bibRefCitation id="2C0B1BF74F59FFD64F23FD71FE0BFD2B" author="Pugedo" box="[272,473,694,717]" pageId="46" pageNumber="47" refString="Pugedo, M. L., Andrade Neto, F. R., Pessali, T. C., Birindelli, J. L. O. &amp; Carvalho, D. C. (2016) Integrative taxonomy supports new candidate fish species in a poorly studied neotropical region: the Jequitinhonha River basin. Genetica, 144, 341 - 349. http: // dx. doi. org / 10.1007 / s 10709 - 016 - 9903 - 4" type="journal article" year="2016">
Pugedo 
<emphasis id="7AEEBA144F59FFD64F50FD7FFE44FD2B" box="[355,406,694,717]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
, 2016
</bibRefCitation>
: 345
</treatmentCitation>
(photo), 346 (table) (barcoding, rio Jequitinhonha).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F59FFD64EA4FD3EFDCBFA7D" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="482566064F59FFD64EA4FD3EFDCBFA7D" blockId="46.[151,1436,761,1434]" pageId="46" pageNumber="47">
<emphasis id="7AEEBA144F59FFD64EA4FD3EFEC3FCF5" bold="true" box="[151,273,761,786]" pageId="46" pageNumber="47">Diagnosis.</emphasis>
<taxonomicName id="8F9A1D854F59FFD64F24FD3DFE6EFCF5" box="[279,444,761,786]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F59FFD64F24FD3DFE6EFCF5" box="[279,444,761,786]" italics="true" pageId="46" pageNumber="47">Brycon howesi</emphasis>
</taxonomicName>
can be diagnosed from all remaining cis-andean 
<taxonomicName id="8F9A1D854F59FFD64DEEFD3DFBFCFCF6" box="[989,1070,762,785]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F59FFD64DEEFD3DFBFCFCF6" box="[989,1070,762,785]" italics="true" pageId="46" pageNumber="47">Brycon</emphasis>
</taxonomicName>
species, with the exception of 
<taxonomicName id="8F9A1D854F59FFD64BB7FD3DFEC2FCD1" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="stolzmanni">
<emphasis id="7AEEBA144F59FFD64BB7FD3DFEC2FCD1" italics="true" pageId="46" pageNumber="47">B. stolzmanni</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F59FFD64F12FCD9FE59FCD2" box="[289,395,798,821]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="coxeyi">
<emphasis id="7AEEBA144F59FFD64F12FCD9FE59FCD2" box="[289,395,798,821]" italics="true" pageId="46" pageNumber="47">B. coxeyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F59FFD64FAFFCD9FDE0FCD2" box="[412,562,798,821]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="coquenani">
<emphasis id="7AEEBA144F59FFD64FAFFCD9FDE0FCD2" box="[412,562,798,821]" italics="true" pageId="46" pageNumber="47">B. coquenani</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F59FFD64C77FCD9FD1CFCD1" box="[580,718,797,822]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F59FFD64C77FCD9FD1CFCD1" box="[580,718,797,822]" italics="true" pageId="46" pageNumber="47">B. vermelha</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F59FFD64CECFCD9FC8BFCD2" box="[735,857,798,821]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F59FFD64CECFCD9FC8BFCD2" box="[735,857,798,821]" italics="true" pageId="46" pageNumber="47">B. insignis</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F59FFD64D59FCD9FC1DFCD1" box="[874,975,797,822]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="dulcis">
<emphasis id="7AEEBA144F59FFD64D59FCD9FC1DFCD1" box="[874,975,797,822]" italics="true" pageId="46" pageNumber="47">B. dulcis</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F59FFD64DD3FCD9FBE9FCD1" box="[992,1083,798,822]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="ferox">
<emphasis id="7AEEBA144F59FFD64DD3FCD9FBE9FCD1" box="[992,1083,798,822]" italics="true" pageId="46" pageNumber="47">B. ferox</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F59FFD64A7FFCD9FB7EFCD2" box="[1100,1196,798,821]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="vonoi">
<emphasis id="7AEEBA144F59FFD64A7FFCD9FB7EFCD2" box="[1100,1196,798,821]" italics="true" pageId="46" pageNumber="47">B. vonoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F59FFD64A8EFCD9FA92FCD1" box="[1213,1344,797,822]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F59FFD64A8EFCD9FA92FCD1" box="[1213,1344,797,822]" italics="true" pageId="46" pageNumber="47">B. opalinus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="8F9A1D854F59FFD64BB7FCD9FF2AFCBE" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="7AEEBA144F59FFD64BB7FCD9FF2AFCBE" italics="true" pageId="46" pageNumber="47">B. nattereri</emphasis>
</taxonomicName>
by possessing a color pattern consisting in a humeral blotch and a caudal peduncle blotch, without body stripes or other obvious color markings on caudal and anal-fins (vs. body stripes and caudal/anal fin color markings present; see 
<figureCitation id="D0A17A834F59FFD64F1BFC4EFEBDFC45" box="[296,367,905,930]" captionStart="FIGURE 5" captionStartId="11.[151,250,1476,1498]" captionTargetBox="[164,1419,611,1450]" captionTargetId="figure@11.[151,1436,589,1454]" captionTargetPageId="11" captionText="FIGURE 5. Diagrammatic representation of the patterns of dark colour markings found among cis-andean Brycon species, represented by selected species. A. Brycon stolzmanni. B. Brycon coxeyi. C. Brycon insignis. D. Brycon nattereri. E. Brycon hilarii. F. Brycon whitei. G. Brycon amazonicus. H. Brycon gouldingi. I. Brycon melanopterus. J-L. Brycon falcatus." httpUri="https://zenodo.org/record/257774/files/figure.png" pageId="46" pageNumber="47">Fig. 5</figureCitation>
). 
<taxonomicName id="8F9A1D854F59FFD64FBBFC4DFDE0FC45" box="[392,562,905,930]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F59FFD64FBBFC4DFDE0FC45" box="[392,562,905,930]" italics="true" pageId="46" pageNumber="47">Brycon howesi</emphasis>
</taxonomicName>
can be diagnosed from these species, with the exception of 
<taxonomicName id="8F9A1D854F59FFD64ACBFC4DFAA0FC46" box="[1272,1394,906,929]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F59FFD64ACBFC4DFAA0FC46" box="[1272,1394,906,929]" italics="true" pageId="46" pageNumber="47">B. insignis</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F59FFD64BB7FC4DFF2DFC21" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F59FFD64BB7FC4DFF2DFC21" italics="true" pageId="46" pageNumber="47">B. vermelha</emphasis>
</taxonomicName>
, and 
<taxonomicName id="8F9A1D854F59FFD64F7AFC69FE31FC22" box="[329,483,942,965]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="coquenani">
<emphasis id="7AEEBA144F59FFD64F7AFC69FE31FC22" box="[329,483,942,965]" italics="true" pageId="46" pageNumber="47">B. coquenani</emphasis>
</taxonomicName>
, by possessing a fifth infraorbital bone considerably wider than high (vs. fifth infraorbital bone about as wide as high; see 
<figureCitation id="D0A17A834F59FFD64C99FC16FD21FC0D" box="[682,755,977,1002]" captionStart="FIGURE 6" captionStartId="12.[151,250,1849,1871]" captionTargetBox="[419,1174,200,1801]" captionTargetId="figure@12.[358,1217,171,1822]" captionTargetPageId="12" captionText="FIGURE 6. Diagrammatic representation of the head profile, maxillary and infraorbitals showing the distinct morphologies found among cis-andean Brycon species, represented by selected species. A. Brycon amazonicus, MZUSP 3815, 152.3 mm SL. B. Brycon insignis, MCZ 21112, 232.2 mm SL. C. Brycon dulcis, MZUSP 1531, 146.6 mm SL. D. Brycon falcatus, MZUSP 3845, 195.1 mm SL. E. Brycon opalinus, MCZ 21123, 210.5 mm SL. F. Brycon orbignyanus, MZUSP 3900, 108.9 mm HL. G. Brycon ferox, MCZ 60934, 310.3 mm SL. H. Brycon orthotaenia, MZUSP 2007, 188.7 mm SL. I. Brycon polylepis, MZUSP 70413, 203.9 mm SL. J. Brycon stolzmanni, UMMZ 185306, 120.0 mm SL." httpUri="https://zenodo.org/record/257775/files/figure.png" pageId="46" pageNumber="47">Fig. 6</figureCitation>
). 
<taxonomicName id="8F9A1D854F59FFD64D3FFC15FC65FC0D" box="[780,951,977,1002]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F59FFD64D3FFC15FC65FC0D" box="[780,951,977,1002]" italics="true" pageId="46" pageNumber="47">Brycon howesi</emphasis>
</taxonomicName>
can be diagnosed from 
<taxonomicName id="8F9A1D854F59FFD64AEFFC15FAA6FC0E" box="[1244,1396,978,1001]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="coquenani">
<emphasis id="7AEEBA144F59FFD64AEFFC15FAA6FC0E" box="[1244,1396,978,1001]" italics="true" pageId="46" pageNumber="47">B. coquenani</emphasis>
</taxonomicName>
by possessing a higher head (65.6–76.2 % of head length, mean 70.6, vs. 59.8–64.9 %, mean 61.4, in 
<taxonomicName id="8F9A1D854F59FFD64AC4FC31FA59FBEA" box="[1271,1419,1014,1037]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="coquenani">
<emphasis id="7AEEBA144F59FFD64AC4FC31FA59FBEA" box="[1271,1419,1014,1037]" italics="true" pageId="46" pageNumber="47">B. coquenani</emphasis>
</taxonomicName>
), by possessing pointed tri- to pentacuspidate dentary teeth, with moderately developed lateral cusps (vs. pointed, trito tetracuspidate dentary teeth, with lateral cusps poorly developed), and by mature males displaying hooks only in anal and pelvic-fins (vs. mature males displaying hooks in all rayed fins, including caudal fin). 
<taxonomicName id="8F9A1D854F59FFD64AFBFBA5FABFFB9D" box="[1224,1389,1121,1146]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F59FFD64AFBFBA5FABFFB9D" box="[1224,1389,1121,1146]" italics="true" pageId="46" pageNumber="47">Brycon howesi</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="8F9A1D854F59FFD64F95FB41FDE6FB79" box="[422,564,1157,1182]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F59FFD64F95FB41FDE6FB79" box="[422,564,1157,1182]" italics="true" pageId="46" pageNumber="47">B. vermelha</emphasis>
</taxonomicName>
by presenting darkened dorsal, caudal, and anal-fins in both living and preserved specimens (vs. dorsal, caudal, and anal-fins reddish in living, clear in preserved 
<taxonomicName id="8F9A1D854F59FFD64ABAFB6DFAC2FB25" box="[1161,1296,1193,1218]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F59FFD64ABAFB6DFAC2FB25" box="[1161,1296,1193,1218]" italics="true" pageId="46" pageNumber="47">B. vermelha</emphasis>
</taxonomicName>
specimens), 7–9, modally 8 horizontal scale rows between the dorsal-fin origin and the lateral line (vs. 10–12, modally 10, in 
<taxonomicName id="8F9A1D854F59FFD64BB7FB09FF2DFAED" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F59FFD64BB7FB09FF2DFAED" italics="true" pageId="46" pageNumber="47">B. vermelha</emphasis>
</taxonomicName>
). 
<taxonomicName id="8F9A1D854F59FFD64F25FB35FE6EFAED" box="[278,444,1265,1290]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F59FFD64F25FB35FE6EFAED" box="[278,444,1265,1290]" italics="true" pageId="46" pageNumber="47">Brycon howesi</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="8F9A1D854F59FFD64CDFFB35FC49FAEE" box="[748,923,1266,1289]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F59FFD64CDFFB35FC49FAEE" box="[748,923,1266,1289]" italics="true" pageId="46" pageNumber="47">Brycon insignis</emphasis>
</taxonomicName>
by possessing 46–50, modally 49 lateral line scales (vs. 51–65, modally 
<quantity id="8F62CBE34F59FFD64FFCFAD2FDD8FAC9" box="[463,522,1301,1326]" metricMagnitude="0" metricUnit="m" metricValue="1.4731999999999998" pageId="46" pageNumber="47" unit="in" value="58.0">58 in</quantity>
<taxonomicName id="8F9A1D854F59FFD64C21FAD1FD58FACA" box="[530,650,1302,1325]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F59FFD64C21FAD1FD58FACA" box="[530,650,1302,1325]" italics="true" pageId="46" pageNumber="47">B. insignis</emphasis>
</taxonomicName>
), 7–9, modally 8 horizontal scale rows between the dorsal-fin origin and the lateral line (vs. 9–11, modally 
<quantity id="8F62CBE34F59FFD64C65FAFDFD68FAB5" box="[598,698,1337,1362]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="46" pageNumber="47" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10–11 in</quantity>
<taxonomicName id="8F9A1D854F59FFD64CF0FAFDFCE9FAB6" box="[707,827,1338,1361]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F59FFD64CF0FAFDFCE9FAB6" box="[707,827,1338,1361]" italics="true" pageId="46" pageNumber="47">B. insignis</emphasis>
</taxonomicName>
), 3–5, modally 4 horizontal scales rows between the lateral line and pelvic (vs. 4–7, modally 
<quantity id="8F62CBE34F59FFD64C5AFA9AFD44FA91" box="[617,662,1373,1398]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="46" pageNumber="47" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="8F9A1D854F59FFD64CADFA99FCC5FA92" box="[670,791,1374,1397]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F59FFD64CADFA99FCC5FA92" box="[670,791,1374,1397]" italics="true" pageId="46" pageNumber="47">B. insignis</emphasis>
</taxonomicName>
) and a relatively rounder snout profile in 
<taxonomicName id="8F9A1D854F59FFD64AC8FA99FAB8FA91" box="[1275,1386,1373,1398]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F59FFD64AC8FA99FAB8FA91" box="[1275,1386,1373,1398]" italics="true" pageId="46" pageNumber="47">B. howesi</emphasis>
</taxonomicName>
(vs. snout more pointed in 
<taxonomicName id="8F9A1D854F59FFD64FA0FA45FDDBFA7E" box="[403,521,1410,1433]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F59FFD64FA0FA45FDDBFA7E" box="[403,521,1410,1433]" italics="true" pageId="46" pageNumber="47">B. insignis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="1CE5368E4F59FFD64EA4F85BFA5BF855" box="[151,1417,1948,1971]" httpUri="https://zenodo.org/record/257797/files/figure.png" pageId="46" pageNumber="47" targetBox="[151,1436,1486,1927]" targetPageId="46">
<paragraph id="482566064F59FFD64EA4F85BFA5BF855" blockId="46.[151,1417,1948,1971]" box="[151,1417,1948,1971]" pageId="46" pageNumber="47">
<emphasis id="7AEEBA144F59FFD64EA4F85BFEF2F855" bold="true" box="[151,288,1948,1970]" pageId="46" pageNumber="47">FIGURE 28.</emphasis>
<taxonomicName id="8F9A1D854F59FFD64F16F859FE69F855" box="[293,443,1948,1971]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="howesi" status="sp. nov.">
<emphasis id="7AEEBA144F59FFD64F16F859FE69F855" box="[293,443,1948,1971]" italics="true" pageId="46" pageNumber="47">Brycon howesi</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="61DD076F4F59FFD64FFBF85AFD96F855" box="[456,580,1948,1970]" pageId="46" pageNumber="47" rank="species">new species</taxonomicNameLabel>
, holotype, MZUSP 53803, 234.8 mm SL: Brazil, Minas Gerais, rio Jequitinhonha.
</paragraph>
</caption>
<caption id="1CE5368E4F58FFD74EA4FD9DFA91FD97" box="[151,1347,602,624]" httpUri="https://zenodo.org/record/257798/files/figure.png" pageId="47" pageNumber="48" targetBox="[151,1436,163,580]" targetPageId="47">
<paragraph id="482566064F58FFD74EA4FD9DFA91FD97" blockId="47.[151,1347,602,624]" box="[151,1347,602,624]" pageId="47" pageNumber="48">
<emphasis id="7AEEBA144F58FFD74EA4FD9DFECDFD97" bold="true" box="[151,287,602,624]" pageId="47" pageNumber="48">FIGURE 29.</emphasis>
<taxonomicName id="8F9A1D854F58FFD74F16FD9CFE69FD97" box="[293,443,602,624]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="howesi" status="sp. nov.">
<emphasis id="7AEEBA144F58FFD74F16FD9CFE69FD97" box="[293,443,602,624]" italics="true" pageId="47" pageNumber="48">Brycon howesi</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="61DD076F4F58FFD74FFBFD9CFD96FD97" box="[456,580,602,624]" pageId="47" pageNumber="48" rank="species">new species</taxonomicNameLabel>
, paratype, MZUSP 101506, 167.1 mm SL: Brazil, Minas Gerais, Rio Preto.
</paragraph>
</caption>
<subSubSection id="0080358D4F58FFC94EF4FD50FC0FFD62" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48" type="description">
<paragraph id="482566064F58FFD74EF4FD50FCCBFC4C" blockId="47.[151,1436,663,940]" pageId="47" pageNumber="48">
<emphasis id="7AEEBA144F58FFD74EF4FD50FE8BFD57" bold="true" box="[199,345,663,688]" pageId="47" pageNumber="48">Description.</emphasis>
Morphometric data are presented in 
<tableCitation id="051853BD4F58FFD74D29FD50FCA4FD48" box="[794,886,663,688]" captionStart="TABLE 7" captionStartId="47.[151,239,983,1006]" captionTargetBox="[159,1313,1034,2006]" captionTargetPageId="47" captionText="TABLE 7. Morphometric data of Brycon howesi (A: holotype, MZUSP 103075)." httpUri="http://table.plazi.org/id/1CE5368E4F58FFD74EA4FC10FC28FC09" pageId="47" pageNumber="48" tableUuid="1CE5368E4F58FFD74EA4FC10FC28FC09">Table 7</tableCitation>
. Large-sized species, largest alcohol-preserved examined specimen 284.0 mm SL. Body moderately slender to moderately high. Largest body height slightly ahead of dorsal-fin origin. Dorsal body profile slightly convex from upper lip margin to vertical through anterior naris, slightly convex from latter point to basis of supraoccipital process, moderately convex from latter point to dorsal-fin origin, straight along dorsal-fin basis, and straight to slightly convex from dorsal-fin basis to adipose-fin origin. Dorsal profile of caudal peduncle slightly concave. Ventral profile slightly convex from lower lip to pelvicfin insertion, straight to slightly convex from this point to anal-fin origin and approximately straight along anal-fin base. Ventral profile of caudal peduncle slightly concave.
</paragraph>
<caption id="1CE5368E4F58FFD74EA4FC10FC28FC09" ID-Table-UUID="1CE5368E4F58FFD74EA4FC10FC28FC09" box="[151,1018,982,1006]" httpUri="http://table.plazi.org/id/1CE5368E4F58FFD74EA4FC10FC28FC09" pageId="47" pageNumber="48" targetBox="[159,1313,1034,2006]" targetIsTable="true" targetPageId="47">
<paragraph id="482566064F58FFD74EA4FC10FC28FC09" blockId="47.[151,1018,982,1006]" box="[151,1018,982,1006]" pageId="47" pageNumber="48">
<emphasis id="7AEEBA144F58FFD74EA4FC10FEDBFC09" bold="true" box="[151,265,983,1006]" pageId="47" pageNumber="48">TABLE 7.</emphasis>
Morphometric data of 
<taxonomicName id="8F9A1D854F58FFD74C32FC10FD4DFC09" box="[513,671,982,1006]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F58FFD74C32FC10FD4DFC09" box="[513,671,982,1006]" italics="true" pageId="47" pageNumber="48">Brycon howesi</emphasis>
</taxonomicName>
(A: holotype, MZUSP 103075).
</paragraph>
</caption>
<paragraph id="482566064F58FFD74D29FBCDFAC1F831" pageId="47" pageNumber="48">
<table id="3A9A94A64F5800074EACFBCDFAF3F831" box="[159,1313,1034,2006]" gridcols="5" gridrows="24" pageId="47" pageNumber="48">
<tr id="F6AA64444F5800074EACFBCDFAF3FBC7" box="[159,1313,1034,1056]" gridrow="0" pageId="47" pageNumber="48" rowspan-0="1">
<th id="B57B0D384F5800074D29FBCDFC81FBC7" box="[794,851,1034,1056]" gridcol="1" gridrow="0" pageId="47" pageNumber="48">A</th>
<th id="B57B0D384F5800074DA1FBCDFC7EFBC7" box="[914,940,1034,1056]" gridcol="2" gridrow="0" pageId="47" pageNumber="48">n</th>
<th id="B57B0D384F5800074A37FBCDFBA4FBC7" box="[1028,1142,1034,1056]" gridcol="3" gridrow="0" pageId="47" pageNumber="48">Range</th>
<th id="B57B0D384F5800074AD4FBCDFAF3FBC7" box="[1255,1313,1034,1056]" gridcol="4" gridrow="0" pageId="47" pageNumber="48">Mean</th>
</tr>
<tr id="F6AA64444F5800074EACFBF4FAF3FBAE" box="[159,1313,1075,1097]" gridrow="1" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFBF4FDAFFBAE" box="[159,637,1075,1097]" gridcol="0" gridrow="1" pageId="47" pageNumber="48">Standard length (SL)</th>
<td id="B57B0D384F5800074D29FBF4FC81FBAE" box="[794,851,1075,1097]" gridcol="1" gridrow="1" pageId="47" pageNumber="48">230.6</td>
<td id="B57B0D384F5800074DA1FBF4FC7EFBAE" box="[914,940,1075,1097]" gridcol="2" gridrow="1" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FBF4FBA4FBAE" box="[1028,1142,1075,1097]" gridcol="3" gridrow="1" pageId="47" pageNumber="48">71.6–284.0</td>
<td id="B57B0D384F5800074AD4FBF4FAF3FBAE" box="[1255,1313,1075,1097]" gridcol="4" gridrow="1" pageId="47" pageNumber="48">-</td>
</tr>
<tr id="F6AA64444F5800074EACFB9BFAF3FB95" box="[159,1313,1116,1138]" gridrow="2" pageId="47" pageNumber="48" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="B57B0D384F5800074EACFB9BFDAFFB95" box="[159,637,1116,1138]" gridcol="0" gridrow="2" pageId="47" pageNumber="48">Percentages of standard length</th>
</tr>
<tr id="F6AA64444F5800074EACFB42FAF3FB7C" box="[159,1313,1157,1179]" gridrow="3" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFB42FDAFFB7C" box="[159,637,1157,1179]" gridcol="0" gridrow="3" pageId="47" pageNumber="48">Depth at dorsal-fin origin</th>
<td id="B57B0D384F5800074D29FB42FC81FB7C" box="[794,851,1157,1179]" gridcol="1" gridrow="3" pageId="47" pageNumber="48">28.6</td>
<td id="B57B0D384F5800074DA1FB42FC7EFB7C" box="[914,940,1157,1179]" gridcol="2" gridrow="3" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FB42FBA4FB7C" box="[1028,1142,1157,1179]" gridcol="3" gridrow="3" pageId="47" pageNumber="48">25.7–33.2</td>
<td id="B57B0D384F5800074AD4FB42FAF3FB7C" box="[1255,1313,1157,1179]" gridcol="4" gridrow="3" pageId="47" pageNumber="48">29.3</td>
</tr>
<tr id="F6AA64444F5800074EACFB68FAF3FB22" box="[159,1313,1199,1221]" gridrow="4" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFB68FDAFFB22" box="[159,637,1199,1221]" gridcol="0" gridrow="4" pageId="47" pageNumber="48">Snout to dorsal-fin origin</th>
<td id="B57B0D384F5800074D29FB68FC81FB22" box="[794,851,1199,1221]" gridcol="1" gridrow="4" pageId="47" pageNumber="48">52.9</td>
<td id="B57B0D384F5800074DA1FB68FC7EFB22" box="[914,940,1199,1221]" gridcol="2" gridrow="4" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FB68FBA4FB22" box="[1028,1142,1199,1221]" gridcol="3" gridrow="4" pageId="47" pageNumber="48">46.8–56.0</td>
<td id="B57B0D384F5800074AD4FB68FAF3FB22" box="[1255,1313,1199,1221]" gridcol="4" gridrow="4" pageId="47" pageNumber="48">52.6</td>
</tr>
<tr id="F6AA64444F5800074EACFB1FFAF3FB09" box="[159,1313,1240,1262]" gridrow="5" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFB1FFDAFFB09" box="[159,637,1240,1262]" gridcol="0" gridrow="5" pageId="47" pageNumber="48">Dorsal-fin base length</th>
<td id="B57B0D384F5800074D29FB1FFC81FB09" box="[794,851,1240,1262]" gridcol="1" gridrow="5" pageId="47" pageNumber="48">11.8</td>
<td id="B57B0D384F5800074DA1FB1FFC7EFB09" box="[914,940,1240,1262]" gridcol="2" gridrow="5" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FB1FFBA4FB09" box="[1028,1142,1240,1262]" gridcol="3" gridrow="5" pageId="47" pageNumber="48">10.1–13.4</td>
<td id="B57B0D384F5800074AD4FB1FFAF3FB09" box="[1255,1313,1240,1262]" gridcol="4" gridrow="5" pageId="47" pageNumber="48">12.1</td>
</tr>
<tr id="F6AA64444F5800074EACFAC5FAF3FAFF" box="[159,1313,1282,1304]" gridrow="6" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFAC5FDAFFAFF" box="[159,637,1282,1304]" gridcol="0" gridrow="6" pageId="47" pageNumber="48">Posterior terminus of dorsal fin to adipose fin</th>
<td id="B57B0D384F5800074D29FAC5FC81FAFF" box="[794,851,1282,1304]" gridcol="1" gridrow="6" pageId="47" pageNumber="48">23.8</td>
<td id="B57B0D384F5800074DA1FAC5FC7EFAFF" box="[914,940,1282,1304]" gridcol="2" gridrow="6" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FAC5FBA4FAFF" box="[1028,1142,1282,1304]" gridcol="3" gridrow="6" pageId="47" pageNumber="48">21.5–25.0</td>
<td id="B57B0D384F5800074AD4FAC5FAF3FAFF" box="[1255,1313,1282,1304]" gridcol="4" gridrow="6" pageId="47" pageNumber="48">23.3</td>
</tr>
<tr id="F6AA64444F5800074EACFAECFAF3FAA6" box="[159,1313,1323,1345]" gridrow="7" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFAECFDAFFAA6" box="[159,637,1323,1345]" gridcol="0" gridrow="7" pageId="47" pageNumber="48">Posterior terminus of dorsal fin to hypural joint</th>
<td id="B57B0D384F5800074D29FAECFC81FAA6" box="[794,851,1323,1345]" gridcol="1" gridrow="7" pageId="47" pageNumber="48">35.6</td>
<td id="B57B0D384F5800074DA1FAECFC7EFAA6" box="[914,940,1323,1345]" gridcol="2" gridrow="7" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FAECFBA4FAA6" box="[1028,1142,1323,1345]" gridcol="3" gridrow="7" pageId="47" pageNumber="48">34.0–39.5</td>
<td id="B57B0D384F5800074AD4FAECFAF3FAA6" box="[1255,1313,1323,1345]" gridcol="4" gridrow="7" pageId="47" pageNumber="48">37.0</td>
</tr>
<tr id="F6AA64444F5800074EACFA93FAF3FA8D" box="[159,1313,1364,1386]" gridrow="8" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFA93FDAFFA8D" box="[159,637,1364,1386]" gridcol="0" gridrow="8" pageId="47" pageNumber="48">Snout to pelvic-fin insertion</th>
<td id="B57B0D384F5800074D29FA93FC81FA8D" box="[794,851,1364,1386]" gridcol="1" gridrow="8" pageId="47" pageNumber="48">47.5</td>
<td id="B57B0D384F5800074DA1FA93FC7EFA8D" box="[914,940,1364,1386]" gridcol="2" gridrow="8" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FA93FBA4FA8D" box="[1028,1142,1364,1386]" gridcol="3" gridrow="8" pageId="47" pageNumber="48">47.5–53.6</td>
<td id="B57B0D384F5800074AD4FA93FAF3FA8D" box="[1255,1313,1364,1386]" gridcol="4" gridrow="8" pageId="47" pageNumber="48">49.9</td>
</tr>
<tr id="F6AA64444F5800074EACFABAFAF3FA74" box="[159,1313,1405,1427]" gridrow="9" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFABAFDAFFA74" box="[159,637,1405,1427]" gridcol="0" gridrow="9" pageId="47" pageNumber="48">Snout to anal-fin origin</th>
<td id="B57B0D384F5800074D29FABAFC81FA74" box="[794,851,1405,1427]" gridcol="1" gridrow="9" pageId="47" pageNumber="48">64.6</td>
<td id="B57B0D384F5800074DA1FABAFC7EFA74" box="[914,940,1405,1427]" gridcol="2" gridrow="9" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FABAFBA4FA74" box="[1028,1142,1405,1427]" gridcol="3" gridrow="9" pageId="47" pageNumber="48">64.4–71.7</td>
<td id="B57B0D384F5800074AD4FABAFAF3FA74" box="[1255,1313,1405,1427]" gridcol="4" gridrow="9" pageId="47" pageNumber="48">67.7</td>
</tr>
<tr id="F6AA64444F5800074EACFA60FAF3FA5A" box="[159,1313,1447,1469]" gridrow="10" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFA60FDAFFA5A" box="[159,637,1447,1469]" gridcol="0" gridrow="10" pageId="47" pageNumber="48">Anal-fin base length</th>
<td id="B57B0D384F5800074D29FA60FC81FA5A" box="[794,851,1447,1469]" gridcol="1" gridrow="10" pageId="47" pageNumber="48">23.7</td>
<td id="B57B0D384F5800074DA1FA60FC7EFA5A" box="[914,940,1447,1469]" gridcol="2" gridrow="10" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FA60FBA4FA5A" box="[1028,1142,1447,1469]" gridcol="3" gridrow="10" pageId="47" pageNumber="48">19.1–26.4</td>
<td id="B57B0D384F5800074AD4FA60FAF3FA5A" box="[1255,1313,1447,1469]" gridcol="4" gridrow="10" pageId="47" pageNumber="48">23.8</td>
</tr>
<tr id="F6AA64444F5800074EACFA17FAF3FA01" box="[159,1313,1488,1510]" gridrow="11" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFA17FDAFFA01" box="[159,637,1488,1510]" gridcol="0" gridrow="11" pageId="47" pageNumber="48">Caudal peduncle length</th>
<td id="B57B0D384F5800074D29FA17FC81FA01" box="[794,851,1488,1510]" gridcol="1" gridrow="11" pageId="47" pageNumber="48">15.8</td>
<td id="B57B0D384F5800074DA1FA17FC7EFA01" box="[914,940,1488,1510]" gridcol="2" gridrow="11" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FA17FBA4FA01" box="[1028,1142,1488,1510]" gridcol="3" gridrow="11" pageId="47" pageNumber="48">11.9–15.8</td>
<td id="B57B0D384F5800074AD4FA17FAF3FA01" box="[1255,1313,1488,1510]" gridcol="4" gridrow="11" pageId="47" pageNumber="48">13.4</td>
</tr>
<tr id="F6AA64444F5800074EACFA3DFAF3F9F7" box="[159,1313,1530,1552]" gridrow="12" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACFA3DFDAFF9F7" box="[159,637,1530,1552]" gridcol="0" gridrow="12" pageId="47" pageNumber="48">Dorsal-fin height</th>
<td id="B57B0D384F5800074D29FA3DFC81F9F7" box="[794,851,1530,1552]" gridcol="1" gridrow="12" pageId="47" pageNumber="48">19.2</td>
<td id="B57B0D384F5800074DA1FA3DFC7EF9F7" box="[914,940,1530,1552]" gridcol="2" gridrow="12" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37FA3DFBA4F9F7" box="[1028,1142,1530,1552]" gridcol="3" gridrow="12" pageId="47" pageNumber="48">17.9–21.6</td>
<td id="B57B0D384F5800074AD4FA3DFAF3F9F7" box="[1255,1313,1530,1552]" gridcol="4" gridrow="12" pageId="47" pageNumber="48">20.0</td>
</tr>
<tr id="F6AA64444F5800074EACF9E4FAF3F9DE" box="[159,1313,1571,1593]" gridrow="13" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACF9E4FDAFF9DE" box="[159,637,1571,1593]" gridcol="0" gridrow="13" pageId="47" pageNumber="48">Pectoral-fin length</th>
<td id="B57B0D384F5800074D29F9E4FC81F9DE" box="[794,851,1571,1593]" gridcol="1" gridrow="13" pageId="47" pageNumber="48">17.9</td>
<td id="B57B0D384F5800074DA1F9E4FC7EF9DE" box="[914,940,1571,1593]" gridcol="2" gridrow="13" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37F9E4FBA4F9DE" box="[1028,1142,1571,1593]" gridcol="3" gridrow="13" pageId="47" pageNumber="48">16.1–20.9</td>
<td id="B57B0D384F5800074AD4F9E4FAF3F9DE" box="[1255,1313,1571,1593]" gridcol="4" gridrow="13" pageId="47" pageNumber="48">19.3</td>
</tr>
<tr id="F6AA64444F5800074EACF98BFAF3F985" box="[159,1313,1612,1634]" gridrow="14" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACF98BFDAFF985" box="[159,637,1612,1634]" gridcol="0" gridrow="14" pageId="47" pageNumber="48">Pelvic-fin length</th>
<td id="B57B0D384F5800074D29F98BFC81F985" box="[794,851,1612,1634]" gridcol="1" gridrow="14" pageId="47" pageNumber="48">14.1</td>
<td id="B57B0D384F5800074DA1F98BFC7EF985" box="[914,940,1612,1634]" gridcol="2" gridrow="14" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37F98BFBA4F985" box="[1028,1142,1612,1634]" gridcol="3" gridrow="14" pageId="47" pageNumber="48">14.1–16.8</td>
<td id="B57B0D384F5800074AD4F98BFAF3F985" box="[1255,1313,1612,1634]" gridcol="4" gridrow="14" pageId="47" pageNumber="48">15.2</td>
</tr>
<tr id="F6AA64444F5800074EACF9B2FAF3F96C" box="[159,1313,1653,1675]" gridrow="15" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACF9B2FDAFF96C" box="[159,637,1653,1675]" gridcol="0" gridrow="15" pageId="47" pageNumber="48">Caudal peduncle depth</th>
<td id="B57B0D384F5800074D29F9B2FC81F96C" box="[794,851,1653,1675]" gridcol="1" gridrow="15" pageId="47" pageNumber="48">8.5</td>
<td id="B57B0D384F5800074DA1F9B2FC7EF96C" box="[914,940,1653,1675]" gridcol="2" gridrow="15" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37F9B2FBA4F96C" box="[1028,1142,1653,1675]" gridcol="3" gridrow="15" pageId="47" pageNumber="48">8.1–9.3</td>
<td id="B57B0D384F5800074AD4F9B2FAF3F96C" box="[1255,1313,1653,1675]" gridcol="4" gridrow="15" pageId="47" pageNumber="48">8.7</td>
</tr>
<tr id="F6AA64444F5800074EACF958FAF3F952" box="[159,1313,1695,1717]" gridrow="16" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACF958FDAFF952" box="[159,637,1695,1717]" gridcol="0" gridrow="16" pageId="47" pageNumber="48">Head length</th>
<td id="B57B0D384F5800074D29F958FC81F952" box="[794,851,1695,1717]" gridcol="1" gridrow="16" pageId="47" pageNumber="48">26.9</td>
<td id="B57B0D384F5800074DA1F958FC7EF952" box="[914,940,1695,1717]" gridcol="2" gridrow="16" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37F958FBA4F952" box="[1028,1142,1695,1717]" gridcol="3" gridrow="16" pageId="47" pageNumber="48">25.3–31.7</td>
<td id="B57B0D384F5800074AD4F958FAF3F952" box="[1255,1313,1695,1717]" gridcol="4" gridrow="16" pageId="47" pageNumber="48">28.6</td>
</tr>
<tr id="F6AA64444F5800074EACF90FFAF3F939" box="[159,1313,1736,1758]" gridrow="17" pageId="47" pageNumber="48" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="B57B0D384F5800074EACF90FFDAFF939" box="[159,637,1736,1758]" gridcol="0" gridrow="17" pageId="47" pageNumber="48">Percentages of head length</th>
</tr>
<tr id="F6AA64444F5800074EACF935FAF3F8EF" box="[159,1313,1778,1800]" gridrow="18" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACF935FDAFF8EF" box="[159,637,1778,1800]" gridcol="0" gridrow="18" pageId="47" pageNumber="48">Head height</th>
<td id="B57B0D384F5800074D29F935FC81F8EF" box="[794,851,1778,1800]" gridcol="1" gridrow="18" pageId="47" pageNumber="48">70.9</td>
<td id="B57B0D384F5800074DA1F935FC7EF8EF" box="[914,940,1778,1800]" gridcol="2" gridrow="18" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37F935FBA4F8EF" box="[1028,1142,1778,1800]" gridcol="3" gridrow="18" pageId="47" pageNumber="48">65.6–76.9</td>
<td id="B57B0D384F5800074AD4F935FAF3F8EF" box="[1255,1313,1778,1800]" gridcol="4" gridrow="18" pageId="47" pageNumber="48">71.3</td>
</tr>
<tr id="F6AA64444F5800074EACF8DCFAF3F8D6" box="[159,1313,1819,1841]" gridrow="19" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACF8DCFDAFF8D6" box="[159,637,1819,1841]" gridcol="0" gridrow="19" pageId="47" pageNumber="48">Snout length</th>
<td id="B57B0D384F5800074D29F8DCFC81F8D6" box="[794,851,1819,1841]" gridcol="1" gridrow="19" pageId="47" pageNumber="48">30.8</td>
<td id="B57B0D384F5800074DA1F8DCFC7EF8D6" box="[914,940,1819,1841]" gridcol="2" gridrow="19" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37F8DCFBA4F8D6" box="[1028,1142,1819,1841]" gridcol="3" gridrow="19" pageId="47" pageNumber="48">26.8–32.3</td>
<td id="B57B0D384F5800074AD4F8DCFAF3F8D6" box="[1255,1313,1819,1841]" gridcol="4" gridrow="19" pageId="47" pageNumber="48">29.9</td>
</tr>
<tr id="F6AA64444F5800074EACF883FAF3F8BD" box="[159,1313,1860,1882]" gridrow="20" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACF883FDAFF8BD" box="[159,637,1860,1882]" gridcol="0" gridrow="20" pageId="47" pageNumber="48">Upper jaw length</th>
<td id="B57B0D384F5800074D29F883FC81F8BD" box="[794,851,1860,1882]" gridcol="1" gridrow="20" pageId="47" pageNumber="48">44.0</td>
<td id="B57B0D384F5800074DA1F883FC7EF8BD" box="[914,940,1860,1882]" gridcol="2" gridrow="20" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37F883FBA4F8BD" box="[1028,1142,1860,1882]" gridcol="3" gridrow="20" pageId="47" pageNumber="48">42.7–50.2</td>
<td id="B57B0D384F5800074AD4F883FAF3F8BD" box="[1255,1313,1860,1882]" gridcol="4" gridrow="20" pageId="47" pageNumber="48">45.9</td>
</tr>
<tr id="F6AA64444F5800074EACF8A9FAF3F863" box="[159,1313,1902,1924]" gridrow="21" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACF8A9FDAFF863" box="[159,637,1902,1924]" gridcol="0" gridrow="21" pageId="47" pageNumber="48">Horizontal eye diameter</th>
<td id="B57B0D384F5800074D29F8A9FC81F863" box="[794,851,1902,1924]" gridcol="1" gridrow="21" pageId="47" pageNumber="48">21.4</td>
<td id="B57B0D384F5800074DA1F8A9FC7EF863" box="[914,940,1902,1924]" gridcol="2" gridrow="21" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37F8A9FBA4F863" box="[1028,1142,1902,1924]" gridcol="3" gridrow="21" pageId="47" pageNumber="48">19.0–27.3</td>
<td id="B57B0D384F5800074AD4F8A9FAF3F863" box="[1255,1313,1902,1924]" gridcol="4" gridrow="21" pageId="47" pageNumber="48">22.7</td>
</tr>
<tr id="F6AA64444F5800074EACF850FAF3F84A" box="[159,1313,1943,1965]" gridrow="22" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACF850FDAFF84A" box="[159,637,1943,1965]" gridcol="0" gridrow="22" pageId="47" pageNumber="48">Post-orbital length</th>
<td id="B57B0D384F5800074D29F850FC81F84A" box="[794,851,1943,1965]" gridcol="1" gridrow="22" pageId="47" pageNumber="48">48.6</td>
<td id="B57B0D384F5800074DA1F850FC7EF84A" box="[914,940,1943,1965]" gridcol="2" gridrow="22" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37F850FBA4F84A" box="[1028,1142,1943,1965]" gridcol="3" gridrow="22" pageId="47" pageNumber="48">45.8–53.2</td>
<td id="B57B0D384F5800074AD4F850FAF3F84A" box="[1255,1313,1943,1965]" gridcol="4" gridrow="22" pageId="47" pageNumber="48">50.3</td>
</tr>
<tr id="F6AA64444F5800074EACF807FAF3F831" box="[159,1313,1984,2006]" gridrow="23" pageId="47" pageNumber="48">
<th id="B57B0D384F5800074EACF807FDAFF831" box="[159,637,1984,2006]" gridcol="0" gridrow="23" pageId="47" pageNumber="48">Least interorbital width</th>
<td id="B57B0D384F5800074D29F807FC81F831" box="[794,851,1984,2006]" gridcol="1" gridrow="23" pageId="47" pageNumber="48">34.5</td>
<td id="B57B0D384F5800074DA1F807FC7EF831" box="[914,940,1984,2006]" gridcol="2" gridrow="23" pageId="47" pageNumber="48">36</td>
<td id="B57B0D384F5800074A37F807FBA4F831" box="[1028,1142,1984,2006]" gridcol="3" gridrow="23" pageId="47" pageNumber="48">28.6–37.1</td>
<td id="B57B0D384F5800074AD4F807FAF3F831" box="[1255,1313,1984,2006]" gridcol="4" gridrow="23" pageId="47" pageNumber="48">33.0</td>
</tr>
</table>
</paragraph>
<paragraph id="482566064F47FFC84EF4FF50FC67FD08" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="49">
Head profile considerably acute anteriorly, mouth terminal. Jaws slightly anisognathous, premaxillary projecting slightly relative to dentary, outer row of premaxillary teeth exposed when mouth is closed in some specimens. Maxillary long, extending posteriorly to middle of pupil. Adipose eyelid well developed. Premaxillary teeth in three rows; teeth of third row largest. Five (1), 6 (1), 7 (3), 8 (10), 9 (9), 10 (8), 11 (4), or 12(1) tricuspidate teeth in outer series. Four (4), 5 (18), 6 (11), or 7 (3) tri- to tetracuspidate teeth in second, inner premaxillary row, plus 1(2), 2 (15), 3 (17), or 4 (2) tricuspidate teeth between the first and third rows. Two teeth in third premaxillary row, medial teeth largest, symphyseal teeth smaller, slightly tilted towards each other, both pentacuspidate. Maxillary with distal portion considerably expanded and rounded in profile. Seventeen to 33 maxillary teeth, slightly smaller than teeth of first premaxillary row, anterior teeth tricuspidate, posterior teeth unicuspidate. Dentary with 8(1), 10 (3), 11 (5), 12 (8), 13 (3), 15 (2), 16 (1), or 17 (1) teeth in main series. Anterior four dentary teeth assymetrical, considerably larger and bulkier than remaining teeth, tetra- to pentacuspidate, each with central cusp distinctly larger than remaining cusps. Remaining dentary teeth progressivelly smaller, tetra, tri- to unicuspidate. Main dentary present gaps (diastemas) between teeth in small specimens (MZUSP 5139, 6, 
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SL). Inner (lingual) series consisting of a small, single unicuspid symphyseal tooth, situated immediately posterior to symphyseal dentary teeth of main series, plus row of 23–25 small, unicuspidate teeth, originating on lingual crest of dentary replacement trench at level of the fifth to sixth teeth of main series. Inner symphyseal teeth present in all specimens with relatively intact symphyseal dentary area.
</paragraph>
<paragraph id="482566064F47FFC84EF4FD3BFCF9FC0B" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="49">
Scales cycloid. Lateral line complete, from supracleithrum to caudal-fin base. Forty-four (1), 45 (4), 46 (6), 47 (7), 48 (10), 49 (7), or 50 (2) scales in lateral line series. Laterosensory tube simple in relatively small (&lt;
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SL) specimens, deflected upwards in the first 5–6 scales, downwards in the remaining lateral-line scales. Larger specimens (&gt; 
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SL) with some scales with 2–3 branches, most lateral scales with a single branch. Tubules short, generally not extending beyond scale anterior third. Horizontal scale rows between dorsal-fin origin and lateral line 7 (2), 8 (31), or 9 (4). Horizontal scale rows between lateral line and pelvic-fin 3 (5), 4 (30), or 5 (2). Circumpeduncular scales 14(3), 15 (16), 16 (15), or 17 (3).
</paragraph>
<paragraph id="482566064F47FFC84EF4FC30FC2CFA27" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="49">
Dorsal-fin rays ii, 9. Dorsal fin origin at, or slightly ahead of, middle of SL. First dorsal-fin pterygiophore inserting behind neural spine of 12th (2) or 13th (1) vertebra. Anal-fin rays iii (not including first, small unbranched ray only visible in the cs specimen), 19(1), 20 (1), 21 (1), 22 (4), 23 (9), 24 (9), 25 (8), or 26 (1). First anal-fin pterygiophore inserting behind haemal spine of 22th (1), 23th (1), or 24th (1) vertebrae. Last unbranched and anterior 3–4 branched anal-fin rays longer, remaining rays progressively shorter towards anal-fin end. Anal fin displaying numerous (c. 20 per fin-ray main branch) middle-sized hooks on last unbranched and posterior main branch of branched rays 17–23, associated with dense, gelatinous tissue in six specimens (MZUSP 
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SL; MZUSP 
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, 3, 189.2–219.0 mm SL; MZUSP 
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, 
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SL; ZUEC 12869, 245.0 mm SL). A single hook per ray segment. Sheath of scales covering basis of anal-fin rays composed of two scale rows, lower scale row formed by 20–25 rectangular scales. Pectoral-fin rays i, 11 (1), 12 (11), 13 (18), 14 (4), or 15 (1). Pelvicfin rays i,7, two specimens i, 8. A few small hooks on posterior branch of branched pelvic-fin rays in five specimens (MZUSP 
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SL; MZUSP 
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, 3, 189.2–219.0 mm SL; MZUSP 
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, 
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SL). Main caudal-fin rays 10/9. Caudal fin forked, lobes slightly pointed.
</paragraph>
<paragraph id="482566064F47FFC84EF4FA0AFCD0F9CB" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="49">Four branchiostegal rays, three on anterior ceratohyal and one on posterior ceratohyal. First branchial arch with 11 (2), 12 (1), 13 (1), 14 (5), 15 (1), or 16 (1) lower, 1 at angle, and 10 (3), 11 (4), or 12 (4) upper gill rakers. Vertebrae 42 (1) or 43 (2). Supraneurals 9 (1) or 10 (2).</paragraph>
<paragraph id="482566064F47FFC94EF4F9F0FBC0FF32" blockId="48.[151,1437,151,2013]" lastBlockId="49.[151,1437,151,2013]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<emphasis id="7AEEBA144F47FFC84EF4F9F0FE04F9B7" bold="true" box="[199,470,1591,1616]" pageId="48" pageNumber="49">Coloration in alcohol.</emphasis>
Overall color pattern clear in specimens from most localities, but very dark in specimens from the rio Preto (MZUSP 
<date id="3C2440C64F47FFC84CB6F99BFD0EF992" box="[645,732,1628,1653]" pageId="48" pageNumber="49" value="06-10-15">101506</date>
, MZUSP 
<date id="3C2440C64F47FFC84D6EF99BFC66F992" box="[861,948,1628,1653]" pageId="48" pageNumber="49" value="07-10-15">101507</date>
, MZUSP 
<date id="3C2440C64F47FFC84A05F99BFB5DF992" box="[1078,1167,1628,1653]" pageId="48" pageNumber="49" value="08-10-15">101508</date>
). Top of head, snout, supraorbital, sixth infraorbital, and dorsal portion of body brown to dark-brown. Remaining infraorbitals, opercle, dentary, maxillary, lateral and ventral surfaces of body light-brown to brown. Specimens retaining guanine (e.g., MZUSP 55963, ZUEC 12869) with infraorbitals, opercle, and lateral surfaces of body with a silvery hue. Humeral blotch present, little to moderately conspicuous, situated at level of lateral line, extending longitudinally from second to fourth, and vertically to scale row situated immediately above lateral line. Caudal peduncle blotch moderately to highly conspicuous, aproximately rounded in shape, extending along 6–7 last lateral line scales. Pectoral, pelvic, dorsal, and anal-fins clear, with a moderate concentration of dark chromatophores at the interradial membranes in the clear-colored specimens; intensely dark-pigmented in the specimens from rio Preto (MZUSP 
<date id="3C2440C64F47FFC84EA4F867FF3CF85E" box="[151,238,1952,1977]" pageId="48" pageNumber="49" value="06-10-15">101506</date>
, MZUSP 
<date id="3C2440C64F47FFC84F6EF867FE66F85E" box="[349,436,1952,1977]" pageId="48" pageNumber="49" value="07-10-15">101507</date>
, MZUSP 
<date id="3C2440C64F47FFC84C17F867FDAFF85E" box="[548,637,1952,1977]" pageId="48" pageNumber="49" value="08-10-15">101508</date>
). Caudal fin with outer caudal fin rays with dark pigmentation on outer rays and basis of middle cudal-fin rays continuous with caudal peduncle blotch, forming a roughly V-shaped blotch. Adipose fin light- to dark-brown. Juveniles (MZUSP 5139, 
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SL) presenting a conspicuous humeral blotch and caudal blotch extending into 3–4 innermost caudal-fin rays.
</paragraph>
<paragraph id="482566064F46FFC94EF4FF18FA5BFE28" blockId="49.[151,1437,151,2013]" pageId="49" pageNumber="50">
<emphasis id="7AEEBA144F46FFC94EF4FF18FEB0FF1F" bold="true" box="[199,354,223,248]" pageId="49" pageNumber="50">Color in life.</emphasis>
Based on pictures of freshly collected or living specimens from the rio Preto, 
<collectingRegion id="8A5EA8E44F46FFC94ADDFF18FAC5FF1F" box="[1262,1303,223,248]" country="Nigeria" name="Rivers" pageId="49" pageNumber="50">Rio</collectingRegion>
Preto State Park, taken by P. Azevedo, a specimen collected at the rio Vacaria, a tributary of the rio Jequitinhonha, by F.R. Andrade Neto, and several specimens from the rio Itacambiruçu, collected by T.C. Pessali. Overall color pattern clear-colored to dark-brown; clear-colored specimens with a silvery hue on opercle and infraorbital bones and lateral surfaces of body. Caudal peduncle blotch and dark pigmentantion on outer caudal-fin rays very conspicuous. Adipose-fin cream-colored to orangish. Specimens from the clear to turbid water rio Vacaria and rio Itacambiruçu possess an overall color pattern much more clear when compared with specimens from the black-water rio Preto.
</paragraph>
<paragraph id="482566064F46FFC94EF4FE1BFC0FFD62" blockId="49.[151,1437,151,2013]" pageId="49" pageNumber="50">
<emphasis id="7AEEBA144F46FFC94EF4FE1BFE6BFE12" bold="true" box="[199,441,476,501]" pageId="49" pageNumber="50">Sexual dimorphism.</emphasis>
Five specimens (MZUSP 
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SL; MZUSP 
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, 3, 189.2–219.0 mm SL; MZUSP 
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, 
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SL) were recorded as possessing anal- and pelvic-fin hooks, a single specimen (ZUEC 12869, 245.0 mm SL) presented only anal-fin hooks. One specimen (MZUSP 
<date id="3C2440C64F46FFC94A47FDE3FB19FDDA" box="[1140,1227,548,573]" pageId="49" pageNumber="50" value="07-10-15">101507</date>
, 
<quantity id="8F62CBE34F46FFC94AEBFDE3FAB8FDDB" box="[1240,1386,548,573]" metricMagnitude="0" metricUnit="m" metricValue="1.1861" pageId="49" pageNumber="50" unit="mm" value="1186.1">1, 186.1 mm</quantity>
SL) proved to be a male after dissection, with relatively well-developed testes. There is an indication in the literature that females may grown larger than males (see “Ecological notes”, below).
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F46FFC94EF4FD48FB70FCDF" pageId="49" pageNumber="50" type="etymology">
<paragraph id="482566064F46FFC94EF4FD48FB79FD2A" blockId="49.[151,1437,151,2013]" pageId="49" pageNumber="50">
<emphasis id="7AEEBA144F46FFC94EF4FD48FE9CFD4F" bold="true" box="[199,334,655,680]" pageId="49" pageNumber="50">Etymology.</emphasis>
Named in honour of Gordon J. Howes (
<date id="3C2440C64F46FFC94D29FD57FC4FFD4F" box="[794,925,656,680]" pageId="49" pageNumber="50" value="1938" valueMax="2013">1938–2013</date>
), in recognition to his important contribution to the knowledge of the taxonomy of the genus 
<taxonomicName id="8F9A1D854F46FFC94C9EFD72FD2CFD2B" box="[685,766,693,716]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F46FFC94C9EFD72FD2CFD2B" box="[685,766,693,716]" italics="true" pageId="49" pageNumber="50">Brycon</emphasis>
</taxonomicName>
as well as for ichthyology as a whole.
</paragraph>
<paragraph id="482566064F46FFC94EF4FD10FB70FCDF" blockId="49.[151,1437,151,2013]" pageId="49" pageNumber="50">
<emphasis id="7AEEBA144F46FFC94EF4FD10FE41FD17" bold="true" box="[199,403,727,752]" pageId="49" pageNumber="50">Common names.</emphasis>
“ 
<taxonomicName id="8F9A1D854F46FFC94F95FD10FDC3FD17" box="[422,529,727,752]" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="184" phylum="Chordata" rank="species" species="piabanha">Piabanha</taxonomicName>
” (Godinho 
<emphasis id="7AEEBA144F46FFC94CA9FD1EFD07FD17" box="[666,725,727,752]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
, 1998: 415; V. Vono, P. Azevedo, T.C. Pessali, pers. comm.). Valenciennes (in 
<bibRefCitation id="2C0B1BF74F46FFC94F6EFD3BFD63FCF2" author="Cuvier" box="[349,689,764,789]" pageId="49" pageNumber="50" refString="Cuvier, G. &amp; Valenciennes, A. (1850) Histoire naturelle des poissons. Tome vingt-deucieme. P. Bertrand editeur, Paris, i - xx, 1 - 532 + 1 - 91, pls. 634 - 650." type="book chapter" year="1850">Cuvier &amp; Valenciennes, 1850</bibRefCitation>
: 246) mentions the common name “pirabanha” for the species, a name apparently heard by 
<collectingRegion id="8A5EA8E44F46FFC94FF3FCE7FE05FCDF" box="[448,471,800,824]" country="Nigeria" name="Rivers" pageId="49" pageNumber="50">A</collectingRegion>
. de Saint-Hillaire, the collector of the specimen studied by him.
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F46FFC94EF4FC83FB5DFB0F" pageId="49" pageNumber="50" type="distribution">
<paragraph id="482566064F46FFC94EF4FC83FE22FC67" blockId="49.[151,1437,151,2013]" pageId="49" pageNumber="50">
<emphasis id="7AEEBA144F46FFC94EF4FC83FEB2FCBA" bold="true" box="[199,352,836,861]" pageId="49" pageNumber="50">Distribution.</emphasis>
Endemic from the rio Jequitinhonha basin, an independent coastal river system at the state of 
<collectingRegion id="8A5EA8E44F46FFC94EA4FCA0FEE2FC67" box="[151,304,871,896]" country="Brazil" name="Minas Gerais" pageId="49" pageNumber="50">Minas Gerais</collectingRegion>
, 
<collectingCountry id="308D26964F46FFC94F08FCA0FEADFC67" box="[315,383,871,896]" name="Brazil" pageId="49" pageNumber="50">Brazil</collectingCountry>
(
<figureCitation id="D0A17A834F46FFC94FBCFCA0FE31FC67" box="[399,483,871,896]" captionStart="FIGURE 25" captionStartId="39.[151,250,1679,1701]" captionTargetBox="[170,1417,392,1639]" captionTargetId="figure@39.[151,1436,373,1658]" captionTargetPageId="39" captionText="FIGURE 25. Map of eastern Brazil, showing known localities of Brycon insignis (red dots), B. vermelha (grey dots), and B. howesi (black dots; dark star indicates type-locality)." httpUri="https://zenodo.org/record/257794/files/figure.png" pageId="49" pageNumber="50">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="482566064F46FFC94EF4FC4BFB5DFB0F" blockId="49.[151,1437,151,2013]" pageId="49" pageNumber="50">
<emphasis id="7AEEBA144F46FFC94EF4FC4BFE40FC42" bold="true" box="[199,402,908,933]" pageId="49" pageNumber="50">Ecological notes.</emphasis>
Feeding and social behavior of the species was quantitavely studied by 
<bibRefCitation id="2C0B1BF74F46FFC94AC1FC4BFF08FC2F" author="Azevedo" pageId="49" pageNumber="50" refString="Azevedo, P. G., Melo, R. M. C. &amp; Young, R. J. (2011) Feeding and social behavior of the piabanha, Brycon devillei (Castelnau, 1855) (Characidae: Bryconinae) in the wild, with a note on following behavior. Neotropical Ichthyology, 9 (4), 807 - 814. http: // dx. doi. org / 10.1590 / S 1679 - 62252011005000046" type="journal article" year="2011">
Azevedo 
<emphasis id="7AEEBA144F46FFC94B53FC4AFA4EFC42" box="[1376,1436,908,933]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
(2011
</bibRefCitation>
; as 
<taxonomicName id="8F9A1D854F46FFC94F38FC76FEADFC2F" box="[267,383,943,968]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="devillei">
<emphasis id="7AEEBA144F46FFC94F38FC76FEADFC2F" box="[267,383,943,968]" italics="true" pageId="49" pageNumber="50">B. devillei</emphasis>
</taxonomicName>
) at a dark water tributary of the rio Araçuaí (a tributary of the rio Jequitinhonha). 
<taxonomicName id="8F9A1D854F46FFC94B78FC76FF37FC0A" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F46FFC94B78FC76FF37FC0A" italics="true" pageId="49" pageNumber="50">Brycon howesi</emphasis>
</taxonomicName>
adopted mainly surface-picking as a foraging tactic, though “digging” and “active-hunting” were also observed (
<bibRefCitation id="2C0B1BF74F46FFC94F3FFC30FE2CFBF7" author="Azevedo" box="[268,510,1015,1040]" pageId="49" pageNumber="50" refString="Azevedo, P. G., Melo, R. M. C. &amp; Young, R. J. (2011) Feeding and social behavior of the piabanha, Brycon devillei (Castelnau, 1855) (Characidae: Bryconinae) in the wild, with a note on following behavior. Neotropical Ichthyology, 9 (4), 807 - 814. http: // dx. doi. org / 10.1590 / S 1679 - 62252011005000046" type="journal article" year="2011">
Azevedo 
<emphasis id="7AEEBA144F46FFC94F49FC3EFE66FBF7" box="[378,436,1015,1040]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
, 2011
</bibRefCitation>
). Surface-picking was more common during the rainy season, whereas “digging” was more frequent during the dry season (
<bibRefCitation id="2C0B1BF74F46FFC94CA0FBDBFC43FBD3" author="Azevedo" box="[659,913,1052,1077]" pageId="49" pageNumber="50" refString="Azevedo, P. G., Melo, R. M. C. &amp; Young, R. J. (2011) Feeding and social behavior of the piabanha, Brycon devillei (Castelnau, 1855) (Characidae: Bryconinae) in the wild, with a note on following behavior. Neotropical Ichthyology, 9 (4), 807 - 814. http: // dx. doi. org / 10.1590 / S 1679 - 62252011005000046" type="journal article" year="2011">
Azevedo 
<emphasis id="7AEEBA144F46FFC94D37FBDAFC90FBD2" box="[772,834,1052,1077]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
, 2011
</bibRefCitation>
). An elaborate cooperative hunting of small characids, and a feeding association of the kind “nuclear-follower behavior” with 
<emphasis id="7AEEBA144F46FFC94A78FB86FAFEFBBF" box="[1099,1324,1089,1112]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="8F9A1D854F46FFC94A78FB86FAFAFBBF" box="[1099,1320,1089,1112]" class="Actinopterygii" family="Anostomidae" genus="Leporinus" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="garmani">Leporinus garmani</taxonomicName>
,
</emphasis>
a bottom feeder, were also documented for the species (
<bibRefCitation id="2C0B1BF74F46FFC94CAAFBA3FC57FB9B" author="Azevedo" box="[665,901,1124,1149]" pageId="49" pageNumber="50" refString="Azevedo, P. G., Melo, R. M. C. &amp; Young, R. J. (2011) Feeding and social behavior of the piabanha, Brycon devillei (Castelnau, 1855) (Characidae: Bryconinae) in the wild, with a note on following behavior. Neotropical Ichthyology, 9 (4), 807 - 814. http: // dx. doi. org / 10.1590 / S 1679 - 62252011005000046" type="journal article" year="2011">
Azevedo 
<emphasis id="7AEEBA144F46FFC94D37FBA2FCEEFB9A" box="[772,828,1124,1149]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
, 2011
</bibRefCitation>
). 
<taxonomicName id="8F9A1D854F46FFC94DA4FBA2FBEFFB9A" box="[919,1085,1124,1149]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F46FFC94DA4FBA2FBEFFB9A" box="[919,1085,1124,1149]" italics="true" pageId="49" pageNumber="50">Brycon howesi</emphasis>
</taxonomicName>
was reported by Godinho 
<emphasis id="7AEEBA144F46FFC94B57FBA2FA4EFB9A" box="[1380,1436,1124,1149]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
(1998; as 
<taxonomicName id="8F9A1D854F46FFC94F39FB4EFE50FB47" box="[266,386,1161,1184]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F46FFC94F39FB4EFE50FB47" box="[266,386,1161,1184]" italics="true" pageId="49" pageNumber="50">B. insignis</emphasis>
</taxonomicName>
) as being mainly insectivore. Largest recorded standard lengths reported for the species by these authors were 254.0 mm SL for males and 342.0 mm SL for females. The species occurs both in clear- (e.g., rio Itacambiruçu) and dark-water (e.g., rio Preto) tributaries of the rio Jequitinhonha basin.
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F46FFC94EF4FB33FDEEF9EF" pageId="49" pageNumber="50" type="description">
<paragraph id="482566064F46FFC94EF4FB33FDEEF9EF" blockId="49.[151,1437,151,2013]" pageId="49" pageNumber="50">
<emphasis id="7AEEBA144F46FFC94EF4FB33FEBCFAEA" bold="true" box="[199,366,1268,1293]" pageId="49" pageNumber="50">Conservation.</emphasis>
The rio Jequitinhonha basin has a long history of anthropogenic disturbances that includes intensive mining, deforestation, and siltation. Probably as a result of these impacts, 
<taxonomicName id="8F9A1D854F46FFC94A0EFADEFB31FAD7" box="[1085,1251,1303,1328]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F46FFC94A0EFADEFB31FAD7" box="[1085,1251,1303,1328]" italics="true" pageId="49" pageNumber="50">Brycon howesi</emphasis>
</taxonomicName>
was reported by fishermen to be becoming increasingly scarcer in the middle rio Jequitinhonha basin (Godinho 
<emphasis id="7AEEBA144F46FFC94ADAFAFAFAF6FAB2" box="[1257,1316,1340,1365]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
, 1998). 
<collectingRegion id="8A5EA8E44F46FFC94BBBFAFBFA4EFAB3" box="[1416,1436,1340,1364]" country="Nigeria" name="Rivers" pageId="49" pageNumber="50">A</collectingRegion>
large hydroelectric dam, the Irapé dam, was build in 2006 a little above the area where most of the records of 
<taxonomicName id="8F9A1D854F46FFC94EA4FA42FE92FA7A" box="[151,320,1412,1437]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F46FFC94EA4FA42FE92FA7A" box="[151,320,1412,1437]" italics="true" pageId="49" pageNumber="50">Brycon howesi</emphasis>
</taxonomicName>
come from, in the middle rio Jequitinhonha basin, and the extent of its presumable deleterious effects on the populations of the species are still unknown. Fortunately, an apparently healthy population of the species occurs at the 
<collectingRegion id="8A5EA8E44F46FFC94FA7FA0BFE6EFA02" box="[404,444,1484,1509]" country="Nigeria" name="Rivers" pageId="49" pageNumber="50">Rio</collectingRegion>
Preto State Park, at the headwaters of the rio Araçuaí, the main tributary of the rio Jequitinhonha (
<bibRefCitation id="2C0B1BF74F46FFC94F71FA28FDE2F9EF" author="Azevedo" box="[322,560,1519,1544]" pageId="49" pageNumber="50" refString="Azevedo, P. G., Melo, R. M. C. &amp; Young, R. J. (2011) Feeding and social behavior of the piabanha, Brycon devillei (Castelnau, 1855) (Characidae: Bryconinae) in the wild, with a note on following behavior. Neotropical Ichthyology, 9 (4), 807 - 814. http: // dx. doi. org / 10.1590 / S 1679 - 62252011005000046" type="journal article" year="2011">
Azevedo 
<emphasis id="7AEEBA144F46FFC94F9CFA36FE35F9EF" box="[431,487,1519,1544]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
, 2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F46FFCA4EF4F9D3FBF7FE4A" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="discussion">
<paragraph id="482566064F46FFC94EF4F9D3FA4EF8AB" blockId="49.[151,1437,151,2013]" pageId="49" pageNumber="50">
<emphasis id="7AEEBA144F46FFC94EF4F9D3FEE9F9CA" bold="true" box="[199,315,1556,1581]" pageId="49" pageNumber="50">Remarks.</emphasis>
There are two references to specimens of 
<taxonomicName id="8F9A1D854F46FFC94D71F9D2FC3FF9CA" box="[834,1005,1556,1581]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F46FFC94D71F9D2FC3FF9CA" box="[834,1005,1556,1581]" italics="true" pageId="49" pageNumber="50">Brycon howesi</emphasis>
</taxonomicName>
in the taxonomic literature, both as misidentifications. The first report for the species was by Valenciennes (in 
<bibRefCitation id="2C0B1BF74F46FFC94DCDF9F0FA86F9B7" author="Cuvier" box="[1022,1364,1591,1616]" pageId="49" pageNumber="50" refString="Cuvier, G. &amp; Valenciennes, A. (1850) Histoire naturelle des poissons. Tome vingt-deucieme. P. Bertrand editeur, Paris, i - xx, 1 - 532 + 1 - 91, pls. 634 - 650." type="book chapter" year="1850">Cuvier &amp; Valenciennes, 1850</bibRefCitation>
: 246), which identified it as 
<taxonomicName id="8F9A1D854F46FFC94FBBF99BFD85F992" box="[392,599,1628,1653]" class="Actinopterygii" family="Characidae" genus="Chalceus" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F46FFC94FBBF99BFD85F992" box="[392,599,1628,1653]" italics="true" pageId="49" pageNumber="50">Chalceus opalinus</emphasis>
</taxonomicName>
(now 
<taxonomicName id="8F9A1D854F46FFC94CACF99AFC84F992" box="[671,854,1628,1653]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F46FFC94CACF99AFC84F992" box="[671,854,1628,1653]" italics="true" pageId="49" pageNumber="50">Brycon opalinus</emphasis>
</taxonomicName>
; see under this species). This record was based on a single, stuffed specimen (MNHN 
<collectingRegion id="8A5EA8E44F46FFC94C0AF9B8FD9DF97F" box="[569,591,1663,1688]" country="Nigeria" name="Rivers" pageId="49" pageNumber="50">A</collectingRegion>
.8612), collected by the botanist Auguste de Saint-Hilaire at the rio “Tiquilelonha” (an evident misspelling of Jequitinhonha), a region surveyed by him between 
<date id="3C2440C64F46FFC94AD2F963FA4EF95B" box="[1249,1436,1700,1725]" pageId="49" pageNumber="50" value="1817-04" valueMax="1817-07-31" valueMin="1817-04-00">April–July 1817</date>
(
<bibRefCitation id="2C0B1BF74F46FFC94EADF90FFE85F907" author="Papavero" box="[158,343,1736,1760]" pageId="49" pageNumber="50" refString="Papavero, N. (1971) Essays on the history of neotropical dipterology. Vol. I. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, 216 pp." type="book" year="1971">Papavero, 1971</bibRefCitation>
). This specimen certainly does not belong to the type-series of 
<taxonomicName id="8F9A1D854F46FFC94A62F900FAF7F907" box="[1105,1317,1735,1760]" class="Actinopterygii" family="Characidae" genus="Chalceus" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F46FFC94A62F900FAF7F907" box="[1105,1317,1735,1760]" italics="true" pageId="49" pageNumber="50">Chalceus opalinus</emphasis>
</taxonomicName>
, which is composed only of the 
<typeStatus id="9721D8A44F46FFC94FA0F92BFE2AF8E2" box="[403,504,1772,1797]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
, MNHN 
<collectingRegion id="8A5EA8E44F46FFC94C52F92BFDA4F8E2" box="[609,630,1772,1797]" country="Nigeria" name="Rivers" pageId="49" pageNumber="50">A</collectingRegion>
.8613 (see item “Remarks” of 
<taxonomicName id="8F9A1D854F46FFC94DFDF92AFB55F8E2" box="[974,1159,1772,1797]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F46FFC94DFDF92AFB55F8E2" box="[974,1159,1772,1797]" italics="true" pageId="49" pageNumber="50">Brycon opalinus</emphasis>
</taxonomicName>
, below). Although 
<bibRefCitation id="2C0B1BF74F46FFC94B57F92BFE73F8CF" author="Gery" pageId="49" pageNumber="50" refString="Gery, J. &amp; Mahnert, V. (1992) Notes sur quelques Brycon des bassins de l'Amazone, du Parana-Paraguay et du Sud-Est bresilien (Pisces, Characiformes, Characidae). Revue Suisse de Zoologie, 99 (4), 793 - 819." type="journal article" year="1992">Géry &amp; Mahnert (1992: 813)</bibRefCitation>
considered the specimen MNHN 
<collectingRegion id="8A5EA8E44F46FFC94D15F8C8FCE9F8CF" box="[806,827,1807,1832]" country="Nigeria" name="Rivers" pageId="49" pageNumber="50">A</collectingRegion>
.8612 to be probably conspecific with the 
<typeStatus id="9721D8A44F46FFC94B29F8C8FAAEF8CF" box="[1306,1404,1807,1832]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="8F9A1D854F46FFC94EA4F8F3FEB4F8AA" box="[151,358,1844,1869]" class="Actinopterygii" family="Characidae" genus="Chalceus" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F46FFC94EA4F8F3FEB4F8AA" box="[151,358,1844,1869]" italics="true" pageId="49" pageNumber="50">Chalceus opalinus</emphasis>
</taxonomicName>
, our examination of this same specimen leaves no doubt that it in fact represents 
<taxonomicName id="8F9A1D854F46FFC94AC3F8F2FA47F8AA" box="[1264,1429,1844,1869]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F46FFC94AC3F8F2FA47F8AA" box="[1264,1429,1844,1869]" italics="true" pageId="49" pageNumber="50">Brycon howesi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="482566064F46FFCA4EF4F890FBACFEFB" blockId="49.[151,1437,151,2013]" lastBlockId="50.[151,1437,151,1581]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<bibRefCitation id="2C0B1BF74F46FFC94EF4F890FE38F897" author="Steindachner" box="[199,490,1879,1904]" pageId="49" pageNumber="50" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner (1877: 591)</bibRefCitation>
reported “Rio Jequitinhonha” as being one of the syntypical localities of 
<taxonomicName id="8F9A1D854F46FFC94B78F89EFF3CF873" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F46FFC94B78F89EFF3CF873" italics="true" pageId="49" pageNumber="50">Brycon insignis</emphasis>
</taxonomicName>
, the remaining localities (“Mendez” and “Campos”) being situated at the rio Paraíba do Sul basin. A search in the NMW fish collection revealed a single specimen, NMW 62927 (1, 
<quantity id="8F62CBE34F46FFC94A32F867FBB8F85F" box="[1025,1130,1952,1977]" metricMagnitude="-2" metricUnit="m" metricValue="7.159999999999999" pageId="49" pageNumber="50" unit="mm" value="71.6">71.6 mm</quantity>
SL) collected at the “Rio Jequitinhonha”, which undoubtedly represents the material reported by 
<bibRefCitation id="2C0B1BF74F46FFC94D80F803FB4AF83B" author="Steindachner" box="[947,1176,1988,2013]" pageId="49" pageNumber="50" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner (1877)</bibRefCitation>
. The designation of the specimen NMW 62925 (
<quantity id="8F62CBE34F45FFCA4F9EFF5FFDF3FF48" box="[429,545,152,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.448" pageId="50" pageNumber="51" unit="mm" value="144.8">144.8 mm</quantity>
SL) as the lectotype of 
<taxonomicName id="8F9A1D854F45FFCA4D1CFF5EFC0DFF57" box="[815,991,153,176]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F45FFCA4D1CFF5EFC0DFF57" box="[815,991,153,176]" italics="true" pageId="50" pageNumber="51">Brycon insignis</emphasis>
</taxonomicName>
resulted in restricting this name for the species occurring in the rio Paraíba do Sul drainage (see “Remarks” of 
<taxonomicName id="8F9A1D854F45FFCA4D83FF7AFB8CFF33" box="[944,1118,189,212]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F45FFCA4D83FF7AFB8CFF33" box="[944,1118,189,212]" italics="true" pageId="50" pageNumber="51">Brycon insignis</emphasis>
</taxonomicName>
). Thus, the specimen NMW 62927, though a paralectotype of 
<taxonomicName id="8F9A1D854F45FFCA4C00FF26FD3BFF1F" box="[563,745,225,248]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F45FFCA4C00FF26FD3BFF1F" box="[563,745,225,248]" italics="true" pageId="50" pageNumber="51">Brycon insignis</emphasis>
</taxonomicName>
, is not conspecific with the lectotype and the remaining paralectotypes, representing instead the species which is herein named as 
<taxonomicName id="8F9A1D854F45FFCA4DE2FEC2FBA5FEFA" box="[977,1143,260,285]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F45FFCA4DE2FEC2FBA5FEFA" box="[977,1143,260,285]" italics="true" pageId="50" pageNumber="51">Brycon howesi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="482566064F45FFCA4EF4FEEFFBF7FE4A" blockId="50.[151,1437,151,1581]" pageId="50" pageNumber="51">
In fact, 
<taxonomicName id="8F9A1D854F45FFCA4F2FFEEEFE13FEA7" box="[284,449,295,320]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F45FFCA4F2FFEEEFE13FEA7" box="[284,449,295,320]" italics="true" pageId="50" pageNumber="51">Brycon howesi</emphasis>
</taxonomicName>
is more similar to 
<taxonomicName id="8F9A1D854F45FFCA4CA0FEEEFCDBFEA7" box="[659,777,297,320]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F45FFCA4CA0FEEEFCDBFEA7" box="[659,777,297,320]" italics="true" pageId="50" pageNumber="51">B. insignis</emphasis>
</taxonomicName>
than to any other 
<taxonomicName id="8F9A1D854F45FFCA4DE0FEEEFBF6FEA7" box="[979,1060,297,320]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F45FFCA4DE0FEEEFBF6FEA7" box="[979,1060,297,320]" italics="true" pageId="50" pageNumber="51">Brycon</emphasis>
</taxonomicName>
species. The V-shaped caudal-fin blotch formed by pigmentation concentrated on the outer caudal-fin rays is a feature uniquely shared between these two species. As noticed in the item “Diagnosis”, above, 
<taxonomicName id="8F9A1D854F45FFCA4D01FEB6FC09FE6F" box="[818,987,367,392]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F45FFCA4D01FEB6FC09FE6F" box="[818,987,367,392]" italics="true" pageId="50" pageNumber="51">Brycon howesi</emphasis>
</taxonomicName>
is clearly distinct from 
<taxonomicName id="8F9A1D854F45FFCA4AC8FEB6FAA7FE6F" box="[1275,1397,369,392]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F45FFCA4AC8FEB6FAA7FE6F" box="[1275,1397,369,392]" italics="true" pageId="50" pageNumber="51">B. insignis</emphasis>
</taxonomicName>
by possessing significantly lower scale counts and a distinctly rounder snout profile.
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F45FFCA4EF4FE70FCBFFB23" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="482566064F45FFCA4EF4FE70FD9FFDFF" blockId="50.[151,1437,151,1581]" pageId="50" pageNumber="51">
<materialsCitation id="F8F26C5B4F45FFCA4EF4FE70FB81FE13" ID-GBIF-Occurrence="1425222625" collectionCode="MZUSP" country="Brazil" county="Aracuai" latitude="-16.75" location="rio Aracuai" longLatPrecision="1281" longitude="-42.0" municipality="rio Jequitinhonha" pageId="50" pageNumber="51" specimenCode="MZUSP 103075" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
<emphasis id="7AEEBA144F45FFCA4EF4FE70FD92FE37" bold="true" box="[199,576,439,464]" pageId="50" pageNumber="51">
Material examined. 
<typeStatus id="9721D8A44F45FFCA4FF9FE70FDE9FE37" box="[458,571,439,464]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="183CCE7D4F45FFCA4C62FE7FFCC0FE37" box="[593,786,439,464]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4C62FE7FFD7CFE37" box="[593,686,440,464]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">MZUSP</collectionCode>
<date id="3C2440C64F45FFCA4C8DFE70FCC0FE37" box="[702,786,439,464]" pageId="50" pageNumber="51" value="75-10-30">103075</date>
</specimenCode>
(
<quantity id="8F62CBE34F45FFCA4D19FE70FC7AFE28" box="[810,936,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="2.306" pageId="50" pageNumber="51" unit="mm" value="230.6">230.6 mm</quantity>
SL): 
<collectingCountry id="308D26964F45FFCA4DC9FE70FB93FE37" box="[1018,1089,439,464]" name="Brazil" pageId="50" pageNumber="51">Brazil</collectingCountry>
, 
<collectingRegion id="8A5EA8E44F45FFCA4A65FE70FB2AFE37" box="[1110,1272,439,464]" country="Brazil" name="Minas Gerais" pageId="50" pageNumber="51">Minas Gerais</collectingRegion>
, 
<collectingCounty id="A1441E8A4F45FFCA4B3EFE7FFABAFE37" box="[1293,1384,440,464]" pageId="50" pageNumber="51">Araçuaí</collectingCounty>
, 
<collectingMunicipality id="A841FC7C4F45FFCA4B4EFE70FEE5FE12" pageId="50" pageNumber="51">rio Jequitinhonha</collectingMunicipality>
, confluence with 
<location id="4D4530DD4F45FFCA4C33FE1BFD51FE13" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4C33FE1BFD51FE13" box="[512,643,476,501]" country="Brazil" county="Aracuai" latitude="-16.75" longLatPrecision="1281" longitude="-42.0" municipality="rio Jequitinhonha" name="rio Aracuai" pageId="50" pageNumber="51" stateProvince="Minas Gerais">rio Araçuaí</location>
, below village of 
<location id="4D4530DD4F45FFCA4D7CFE1BFC53FE12" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4D7CFE1BFC53FE12" box="[847,897,476,501]" country="Brazil" county="Aracuai" latitude="-16.75" longLatPrecision="1281" longitude="-42.0" municipality="rio Jequitinhonha" name="Itira" pageId="50" pageNumber="51" stateProvince="Minas Gerais">Itira</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4DBEFE1BFC39FE12" box="[909,1003,476,501]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="925" value="-16.75">16°45’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4DCBFE1BFB81FE13" box="[1016,1107,476,500]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="925" value="-42.0">42°0’W</geoCoordinate>
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F45FFCA4A6CFE1BFD9FFDFF" ID-GBIF-Occurrence="1425222596" collectingDate="1989-02-19" collectorName="J. C. Garavello &amp; J. C. Soares" country="Brazil" location="A" pageId="50" pageNumber="51" specimenCount="1" stateProvince="A" typeStatus="holotype">
<collectorName id="E56F03D04F45FFCA4A6CFE1BFAD6FE12" box="[1119,1284,476,501]" pageId="50" pageNumber="51">J.C. Garavello</collectorName>
, 
<collectingRegion id="8A5EA8E44F45FFCA4B23FE1BFAFBFE13" box="[1296,1321,476,500]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
.S. Soares, 
<collectingRegion id="8A5EA8E44F45FFCA4EA4FDC7FF62FDFF" box="[151,176,512,536]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
.I. Alves &amp; 
<collectorName id="E56F03D04F45FFCA4F1DFDC7FE7CFDFF" box="[302,430,512,536]" pageId="50" pageNumber="51">J.C. Soares</collectorName>
, 
<collectingDate id="2C60B92E4F45FFCA4F89FDC7FD9BFDFF" box="[442,585,511,536]" pageId="50" pageNumber="51" value="1989-02-19">
<date id="3C2440C64F45FFCA4F89FDC7FD9BFDFF" box="[442,585,511,536]" pageId="50" pageNumber="51" value="1989-02-19">19 Feb 1989</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="482566064F45FFCA4EF4FDE3FCBFFB23" blockId="50.[151,1437,151,1581]" pageId="50" pageNumber="51">
<materialsCitation id="F8F26C5B4F45FFCA4EF4FDE3FF2FFD87" ID-GBIF-Occurrence="1425222617" collectionCode="MZUSP" country="Brazil" county="Aracuai" latitude="-16.75" location="rio Jequitinhonha" longLatPrecision="1281" longitude="-42.0" municipality="rio Jequitinhonha" pageId="50" pageNumber="51" specimenCode="MZUSP 53803" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<emphasis id="7AEEBA144F45FFCA4EF4FDE3FCAFFDDA" bold="true" box="[199,893,548,573]" pageId="50" pageNumber="51">
<typeStatus id="9721D8A44F45FFCA4EF4FDE3FE91FDDA" box="[199,323,548,573]" pageId="50" pageNumber="51" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="308D26964F45FFCA4F7CFDE3FE48FDDA" box="[335,410,548,573]" name="Brazil" pageId="50" pageNumber="51">Brazil</collectingCountry>
, 
<collectingRegion id="8A5EA8E44F45FFCA4F95FDE3FD98FDDA" box="[422,586,548,573]" country="Brazil" name="Minas Gerais" pageId="50" pageNumber="51">Minas Gerais</collectingRegion>
, 
<location id="4D4530DD4F45FFCA4C65FDE3FCFFFDDA" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4C65FDE3FCFFFDDA" box="[598,813,548,573]" country="Brazil" name="rio Jequitinhonha" pageId="50" pageNumber="51" stateProvince="Minas Gerais">rio Jequitinhonha</location>
basin:
</emphasis>
<specimenCode id="183CCE7D4F45FFCA4DB7FDE3FBFDFDDA" box="[900,1071,548,573]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4DB7FDE3FC33FDDB" box="[900,993,548,572]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">MZUSP</collectionCode>
53803
</specimenCode>
(
<quantity id="8F62CBE34F45FFCA4A72FDE3FB1CFDDB" box="[1089,1230,548,572]" metricMagnitude="0" metricUnit="m" metricValue="1.2348" pageId="50" pageNumber="51" unit="mm" value="1234.8">1, 234.8 mm</quantity>
SL): same data as holotype
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4F3BFD8FFECCFD63" ID-GBIF-Occurrence="1425222593" collectionCode="MZUSP" collectorName="G. B. Santos" country="Brazil" latitude="-17.1" location="rio Aracuai" longLatPrecision="1279" longitude="-42.65" municipality="Leme do Prado" pageId="50" pageNumber="51" specimenCode="MZUSP 53802" specimenCount="1" stateProvince="Rita" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4F3BFD8FFE61FD87" box="[264,435,583,608]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4F3BFD8FFEB7FD87" box="[264,357,584,608]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">MZUSP</collectionCode>
53802
</specimenCode>
(
<quantity id="8F62CBE34F45FFCA4FF4FD8FFD87FDB8" box="[455,597,584,608]" metricMagnitude="0" metricUnit="m" metricValue="1.2230999999999999" pageId="50" pageNumber="51" unit="mm" value="1223.1">1, 223.1 mm</quantity>
SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4CA4FD8FFC95FD87" box="[663,839,583,608]" pageId="50" pageNumber="51">Leme do Prado</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4D67FD80FC01FD87" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4D67FD80FC01FD87" box="[852,979,583,608]" country="Brazil" latitude="-17.1" longLatPrecision="1279" longitude="-42.65" municipality="Leme do Prado" name="rio Aracuai" pageId="50" pageNumber="51" stateProvince="Rita">rio Araçuaí</location>
at Santa 
<collectingRegion id="8A5EA8E44F45FFCA4A0CFD80FBBCFD87" box="[1087,1134,583,608]" country="Cameroon" name="East" pageId="50" pageNumber="51">Rita</collectingRegion>
village, 
<geoCoordinate id="2DAE00C14F45FFCA4AE2FD80FAF3FD87" box="[1233,1313,583,608]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="925" value="-17.1">17°6’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4B1DFD8FFA45FD87" box="[1326,1431,584,608]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="925" value="-42.65">42°39’W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4EA4FDABFECCFD63" box="[151,286,620,644]" pageId="50" pageNumber="51">G.B. Santos</collectorName>
</materialsCitation>
, 
<materialsCitation id="F8F26C5B4F45FFCA4F18FDABFB0DFD63" ID-GBIF-Occurrence="1425222615" box="[299,1247,620,645]" collectingDate="1987-10-18" collectionCode="MZUSP" country="Brazil" location="A" pageId="50" pageNumber="51" specimenCode="MZUSP 55963" specimenCount="1" stateProvince="A" typeStatus="paratype">
<collectingRegion id="8A5EA8E44F45FFCA4F18FDABFE96FD63" box="[299,324,620,644]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
.S. Soares &amp; 
<collectingRegion id="8A5EA8E44F45FFCA4FE5FDABFE22FD63" box="[470,496,620,644]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
.I.S. Alves, 
<collectingDate id="2C60B92E4F45FFCA4C41FDABFCD0FD63" box="[626,770,620,644]" pageId="50" pageNumber="51" value="1987-10-18">
<date id="3C2440C64F45FFCA4C41FDABFCD0FD63" box="[626,770,620,644]" pageId="50" pageNumber="51" value="1987-10-18">18 Oct 1987</date>
</collectingDate>
. 
<specimenCode id="183CCE7D4F45FFCA4D3DFDABFC68FD62" box="[782,954,620,645]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4D3DFDABFCB9FD63" box="[782,875,620,644]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">MZUSP</collectionCode>
55963
</specimenCode>
(6, 165.8– 
<quantity id="8F62CBE34F45FFCA4A06FDABFB78FD63" box="[1077,1194,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="2.4859999999999998" pageId="50" pageNumber="51" unit="mm" value="248.6">248.6 mm</quantity>
SL)
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F45FFCA4ADFFDABFAE0FD4F" ID-GBIF-Occurrence="1425222604" collectingDate="2010-08-06" collectionCode="ZUEC" collectorName="F. Andrade" country="Brazil" latitude="-16.625555" location="rio Jequitinhonha" longLatPrecision="21" longitude="-42.268333" municipality="Coronel Murta" pageId="50" pageNumber="51" specimenCode="ZUEC 6365" specimenCount="1" stateProvince="A" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4ADFFDABFAA7FD62" box="[1260,1397,620,645]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4ADFFDABFAE7FD63" box="[1260,1333,620,644]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
6365
</specimenCode>
(
<quantity id="8F62CBE34F45FFCA4BBAFDABFEDEFD40" metricMagnitude="0" metricUnit="m" metricValue="1.1649" pageId="50" pageNumber="51" unit="mm" value="1164.9">1, 164.9 mm</quantity>
SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4F61FD48FE2FFD40" box="[338,509,655,680]" pageId="50" pageNumber="51">Coronel Murta</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4C3FFD48FD07FD4F" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4C3FFD48FD07FD4F" box="[524,725,655,680]" country="Brazil" latitude="-16.625555" longLatPrecision="21" longitude="-42.268333" municipality="Coronel Murta" name="rio Jequitinhonha" pageId="50" pageNumber="51" stateProvince="A">rio Jequitinhonha</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4CD0FD48FCBCFD4F" box="[739,878,655,680]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.625555">16°37’32’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4D4EFD48FBD6FD4F" box="[893,1028,655,680]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.268333">42°16’6’’W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4A20FD57FB47FD4F" box="[1043,1173,655,680]" pageId="50" pageNumber="51">F. Andrade</collectorName>
, 
<collectingDate id="2C60B92E4F45FFCA4A90FD48FAE0FD4F" box="[1187,1330,655,680]" pageId="50" pageNumber="51" value="2010-08-06">
<date id="3C2440C64F45FFCA4A90FD48FAE0FD4F" box="[1187,1330,655,680]" pageId="50" pageNumber="51" value="2010-08-06">6 Aug 2010</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4B0CFD57FEE6FD17" ID-GBIF-Occurrence="1425222608" collectingDate="1966-06-25" collectionCode="MZUSP" collectorName="Exc. Departamento de Zoologia" country="Brazil" latitude="-16.566668" location="rio Jequitinhonha" longLatPrecision="1282" longitude="-41.483334" municipality="Itaobim" pageId="50" pageNumber="51" specimenCode="MZUSP 5139" specimenCount="1" stateProvince="A" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4B0CFD57FF1DFD2A" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4B0CFD57FA4EFD4F" box="[1343,1436,656,680]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">MZUSP</collectionCode>
5139
</specimenCode>
(6, 1 cs, 
<quantity id="8F62CBE34F45FFCA4F05FD73FE0BFD2B" box="[310,473,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="3.945" metricValueMax="4.76" metricValueMin="3.13" pageId="50" pageNumber="51" unit="mm" value="39.45" valueMax="47.6" valueMin="31.3">31.3–47.6 mm</quantity>
SL), 
<collectingMunicipality id="A841FC7C4F45FFCA4C2BFD73FDA0FD2A" box="[536,626,692,717]" pageId="50" pageNumber="51">Itaobim</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4C4DFD73FC96FD2A" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4C4DFD73FC96FD2A" box="[638,836,692,717]" country="Brazil" latitude="-16.566668" longLatPrecision="1282" longitude="-41.483334" municipality="Itaobim" name="rio Jequitinhonha" pageId="50" pageNumber="51" stateProvince="A">rio Jequitinhonha</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4D63FD73FC7CFD2A" box="[848,942,692,717]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="925" value="-16.566668">16°34’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4D89FD73FBF1FD2B" box="[954,1059,692,716]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="925" value="-41.483334">41°29’W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4A1CFD72FA4AFD2A" box="[1071,1432,692,717]" pageId="50" pageNumber="51">Exc. Departamento de Zoologia</collectorName>
, 
<collectingDate id="2C60B92E4F45FFCA4EA4FD10FEE6FD17" box="[151,308,727,752]" pageId="50" pageNumber="51" value="1966-06-25">
<date id="3C2440C64F45FFCA4EA4FD10FEE6FD17" box="[151,308,727,752]" pageId="50" pageNumber="51" value="1966-06-25">25 June 1966</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4F72FD1FFD53FCF3" ID-GBIF-Occurrence="1425222623" collectingDate="2011-05" collectionCode="ZUEC" collectorName="T. C. Pessali &amp; T. F. Teixeira" country="Brazil" latitude="-16.581112" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.873608" municipality="Grao Mogol" pageId="50" pageNumber="51" specimenCode="ZUEC 6939" specimenCount="1" stateProvince="A" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4F72FD1FFE19FD17" box="[321,459,727,752]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4F72FD1FFE58FD17" box="[321,394,728,752]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
6939
</specimenCode>
(3, 134.0–242.0 mm SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4D32FD1FFC42FD17" box="[769,912,727,752]" pageId="50" pageNumber="51">Grão Mogol</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4DAEFD10FB8EFD17" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4DAEFD10FB8EFD17" box="[925,1116,727,752]" country="Brazil" latitude="-16.581112" longLatPrecision="21" longitude="-42.873608" municipality="Grao Mogol" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçu</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4A5AFD10FB26FD17" box="[1129,1268,727,752]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.581112">16°34’52’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4B31FD10FA4AFD17" box="[1282,1432,727,752]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.873608">42°52’25’’W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4EA4FD3BFEF6FCF2" box="[151,292,764,789]" pageId="50" pageNumber="51">T.C. Pessali</collectorName>
&amp; 
<collectorName id="E56F03D04F45FFCA4F6BFD3AFE22FCF2" box="[344,496,764,789]" pageId="50" pageNumber="51">T.F. Teixeira</collectorName>
, 
<collectingDate id="2C60B92E4F45FFCA4C30FD3AFD53FCF3" box="[515,641,764,789]" pageId="50" pageNumber="51" value="2011-05">
<date id="3C2440C64F45FFCA4C30FD3AFD53FCF3" box="[515,641,764,789]" pageId="50" pageNumber="51" value="2011-05">May 2011</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4CA0FD3BFC51FCDF" ID-GBIF-Occurrence="1425222610" collectingDate="2011-05" collectionCode="ZUEC" collectorName="T. C. Pessali &amp; T. F. Teixeira" country="Brazil" latitude="-16.586668" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.854164" municipality="Grao Mogol" pageId="50" pageNumber="51" specimenCode="ZUEC 6940" specimenCount="1" stateProvince="A" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4CA0FD3BFCF6FCF2" box="[659,804,764,789]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4CA0FD3BFD0EFCF3" box="[659,732,764,788]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
6940
</specimenCode>
(
<quantity id="8F62CBE34F45FFCA4D0DFD3BFC09FCF3" box="[830,987,764,788]" metricMagnitude="0" metricUnit="m" metricValue="1.1437" pageId="50" pageNumber="51" unit="mm" value="1143.7">1, 143.7 mm</quantity>
SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4A18FD3BFB6DFCF2" box="[1067,1215,764,789]" pageId="50" pageNumber="51">Grão Mogol</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4AE0FD3BFA4AFCF2" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4AE0FD3BFA4AFCF2" box="[1235,1432,764,789]" country="Brazil" latitude="-16.586668" longLatPrecision="21" longitude="-42.854164" municipality="Grao Mogol" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçu</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4EA4FCD8FEF3FCDF" box="[151,289,799,824]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.586668">16°35’12’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4F1EFCD8FE11FCDF" box="[301,451,799,824]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.854164">42°51’15’’W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4FFDFCE7FD80FCDF" box="[462,594,799,824]" pageId="50" pageNumber="51">T.C. Pessali</collectorName>
&amp; 
<collectorName id="E56F03D04F45FFCA4C47FCE7FCD1FCDF" box="[628,771,799,824]" pageId="50" pageNumber="51">T.F. Teixeira</collectorName>
, 
<collectingDate id="2C60B92E4F45FFCA4D3DFCE7FC51FCDF" box="[782,899,800,824]" pageId="50" pageNumber="51" value="2011-05">
<date id="3C2440C64F45FFCA4D3DFCE7FC51FCDF" box="[782,899,800,824]" pageId="50" pageNumber="51" value="2011-05">May 2011</date>
</collectingDate>
</materialsCitation>
. ZUEC 7003 (1, 282.0 mm SL); MCP 48399 (1, 268.0 mm SL); 
<materialsCitation id="F8F26C5B4F45FFCA4F78FC83FEF3FC67" ID-GBIF-Occurrence="1425222630" collectionCode="ANSP" collectorName="T. C. Pessali" country="Brazil" latitude="-16.581112" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.873608" municipality="Grao Mogol" pageId="50" pageNumber="51" specimenCode="ANSP 200242" specimenCount="1" stateProvince="A" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4F78FC83FE3CFCBB" box="[331,494,836,860]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4F78FC83FE41FCBB" box="[331,403,836,860]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="50" pageNumber="51">ANSP</collectionCode>
<date id="3C2440C64F45FFCA4FA9FC83FE3CFCBB" box="[410,494,836,860]" pageId="50" pageNumber="51" value="42-02-20">200242</date>
</specimenCode>
(1, 232.0 mm SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4CE3FC83FC8FFCBA" box="[720,861,836,861]" pageId="50" pageNumber="51">Grão Mogol</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4D5AFC83FBF5FCBA" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4D5AFC83FBF5FCBA" box="[873,1063,836,861]" country="Brazil" latitude="-16.581112" longLatPrecision="21" longitude="-42.873608" municipality="Grao Mogol" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçú</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4A00FC83FB6CFCBA" box="[1075,1214,836,861]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.581112">16°34’52’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4AF9FC83FAB2FCBA" box="[1226,1376,836,861]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.873608">42°52’25’’W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4B5FFC83FF37FC67" pageId="50" pageNumber="51">T.C. Pessali</collectorName>
&amp; 
<collectingRegion id="8A5EA8E44F45FFCA4F39FCAFFEF3FC67" box="[266,289,872,896]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4F16FCAFFDE7FC67" ID-GBIF-Occurrence="1425222607" box="[293,565,871,896]" collectingDate="2011-02" country="Brazil" location="A" pageId="50" pageNumber="51" specimenCount="1" stateProvince="A" typeStatus="paratype">
<collectingRegion id="8A5EA8E44F45FFCA4F16FCAFFEEEFC67" box="[293,316,872,896]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
. Rodrigues, 
<collectingDate id="2C60B92E4F45FFCA4FFAFCA0FDE7FC67" box="[457,565,871,896]" pageId="50" pageNumber="51" value="2011-02">
<date id="3C2440C64F45FFCA4FFAFCA0FDE7FC67" box="[457,565,871,896]" pageId="50" pageNumber="51" value="2011-02">Feb 2011</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4C72FCAFFDCFFC43" ID-GBIF-Occurrence="1425222600" collectionCode="ZUEC" collectorName="T. C. Pessali" country="Brazil" latitude="-16.597223" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.839443" municipality="Grao Mogol" pageId="50" pageNumber="51" specimenCode="ZUEC 7000" specimenCount="1" stateProvince="A" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4C72FCAFFD1BFC67" box="[577,713,872,896]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4C72FCAFFD58FC67" box="[577,650,872,896]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
7000
</specimenCode>
(1, 208.0 mm SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4D99FCAFFBE5FC67" box="[938,1079,871,896]" pageId="50" pageNumber="51">Grão Mogol</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4A70FCA0FAD3FC67" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4A70FCA0FAD3FC67" box="[1091,1281,871,896]" country="Brazil" latitude="-16.597223" longLatPrecision="21" longitude="-42.839443" municipality="Grao Mogol" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçú</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4B3FFCA0FA45FC67" box="[1292,1431,871,896]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.597223">16°35’50’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4EA4FC4BFEFFFC42" box="[151,301,908,933]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.839443">42°50’22’’W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4F71FC4BFE02FC42" box="[322,464,908,933]" pageId="50" pageNumber="51">T.C. Pessali</collectorName>
&amp; 
<collectingRegion id="8A5EA8E44F45FFCA4C35FC4BFDCFFC43" box="[518,541,908,932]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4C11FC4BFC9CFC43" ID-GBIF-Occurrence="1425222603" box="[546,846,908,933]" collectingDate="2011-02" country="Brazil" location="A" pageId="50" pageNumber="51" specimenCount="1" stateProvince="A" typeStatus="paratype">
<collectingRegion id="8A5EA8E44F45FFCA4C11FC4BFDEBFC43" box="[546,569,908,932]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
. Rodrigues, 
<collectingDate id="2C60B92E4F45FFCA4CEBFC4BFC9CFC43" box="[728,846,908,933]" pageId="50" pageNumber="51" value="2011-02">
<date id="3C2440C64F45FFCA4CEBFC4BFC9CFC43" box="[728,846,908,933]" pageId="50" pageNumber="51" value="2011-02">Feb 2011</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4D52FC4BFAAFFC2F" ID-GBIF-Occurrence="1425222613" collectionCode="ZUEC" collectorName="Cachoeira Marias das Neves &amp; T. C. Pessali" country="Brazil" county="Grao Mogol" latitude="-16.58139" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.873886" municipality="Itacambirucu" pageId="50" pageNumber="51" specimenCode="ZUEC 12869, 1" specimenCount="1" stateProvince="A" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4D52FC4BFBF4FC43" box="[865,1062,908,933]" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4D52FC4BFC78FC43" box="[865,938,908,932]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
12869, 1
</specimenCode>
, 245.0 mm SL, 
<collectingCounty id="A1441E8A4F45FFCA4B31FC4BFA45FC42" box="[1282,1431,908,933]" pageId="50" pageNumber="51">Grão Mogol</collectingCounty>
, 
<collectingMunicipality id="A841FC7C4F45FFCA4EA4FC68FEFCFC2F" box="[151,302,943,968]" pageId="50" pageNumber="51">Itacambiruçu</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4F0AFC68FE25FC2F" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4F0AFC68FE25FC2F" box="[313,503,943,968]" country="Brazil" county="Grao Mogol" latitude="-16.58139" longLatPrecision="21" longitude="-42.873886" municipality="Itacambirucu" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçu</location>
, above 
<collectorName id="E56F03D04F45FFCA4C7EFC68FC43FC20" box="[589,913,943,968]" pageId="50" pageNumber="51">Cachoeira Marias das Neves</collectorName>
, 
<geoCoordinate id="2DAE00C14F45FFCA4DAEFC68FBC9FC2F" box="[925,1051,943,968]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.58139">16°34'53''S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4A14FC68FB62FC2F" box="[1063,1200,943,968]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.873886">42°52'26''W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4A88FC77FA93FC2F" box="[1211,1345,943,968]" pageId="50" pageNumber="51">T.C. Pessali</collectorName>
&amp; 
<collectingRegion id="8A5EA8E44F45FFCA4B55FC77FAAFFC2F" box="[1382,1405,944,968]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4BB2FC77FE67FC0B" ID-GBIF-Occurrence="1425222627" collectingDate="2011-12-25" country="Brazil" location="A" pageId="50" pageNumber="51" specimenCount="1" stateProvince="A" typeStatus="paratype">
<collectingRegion id="8A5EA8E44F45FFCA4BB2FC77FA4AFC2F" box="[1409,1432,944,968]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
. Rodrigues, 
<collectingDate id="2C60B92E4F45FFCA4F2FFC13FE67FC0B" box="[284,437,980,1005]" pageId="50" pageNumber="51" value="2011-12-25">
<date id="3C2440C64F45FFCA4F2FFC13FE67FC0B" box="[284,437,980,1005]" pageId="50" pageNumber="51" value="2011-12-25">25 Dec 2011</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4FF7FC13FC34FBF7" ID-GBIF-Occurrence="1425222614" collectionCode="ZUEC" collectorName="Cachoeira Marias das Neves &amp; T. C. Pessali" country="Brazil" county="Grao Mogol" latitude="-16.599445" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.926945" municipality="Itacambirucu" pageId="50" pageNumber="51" specimenCode="ZUEC 12870, 1" specimenCount="1" stateProvince="A" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4FF7FC13FDACFC0B" box="[452,638,980,1004]" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4FF7FC13FDDFFC0B" box="[452,525,980,1004]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
12870, 1
</specimenCode>
, 213.0 mm SL, 
<collectingCounty id="A1441E8A4F45FFCA4D77FC13FC06FC0A" box="[836,980,980,1005]" pageId="50" pageNumber="51">Grão Mogol</collectingCounty>
, 
<collectingMunicipality id="A841FC7C4F45FFCA4DD7FC13FBA9FC0A" box="[996,1147,980,1005]" pageId="50" pageNumber="51">Itacambiruçu</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4ABAFC13FA98FC0A" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4ABAFC13FA98FC0A" box="[1161,1354,980,1005]" country="Brazil" county="Grao Mogol" latitude="-16.599445" longLatPrecision="21" longitude="-42.926945" municipality="Itacambirucu" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçu</location>
, above 
<collectorName id="E56F03D04F45FFCA4EA4FC30FE34FBE8" box="[151,486,1015,1040]" pageId="50" pageNumber="51">Cachoeira Marias das Neves</collectorName>
, 
<geoCoordinate id="2DAE00C14F45FFCA4FC6FC30FDA1FBF7" box="[501,627,1015,1040]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.599445">16°35'58''S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4CB0FC30FCD9FBF7" box="[643,779,1015,1040]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.926945">42°55'37''W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4D28FC3FFC76FBF7" box="[795,932,1015,1040]" pageId="50" pageNumber="51">T.C. Pessali</collectorName>
&amp; 
<collectingRegion id="8A5EA8E44F45FFCA4DFCFC3FFC34FBF7" box="[975,998,1016,1040]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4DD8FC3FFAE3FBF7" ID-GBIF-Occurrence="1425222611" box="[1003,1329,1015,1040]" collectingDate="2011-12-28" country="Brazil" location="A" pageId="50" pageNumber="51" specimenCount="1" stateProvince="A" typeStatus="paratype">
<collectingRegion id="8A5EA8E44F45FFCA4DD8FC3FFBD0FBF7" box="[1003,1026,1016,1040]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
. Rodrigues, 
<collectingDate id="2C60B92E4F45FFCA4AA5FC3FFAE3FBF7" box="[1174,1329,1016,1040]" pageId="50" pageNumber="51" value="2011-12-28">
<date id="3C2440C64F45FFCA4AA5FC3FFAE3FBF7" box="[1174,1329,1016,1040]" pageId="50" pageNumber="51" value="2011-12-28">28 Dec 2011</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4B0CFC3FFB31FBD3" ID-GBIF-Occurrence="1425222597" collectionCode="MZUSP" country="Brazil" location="rio Aracuai" municipality="rio Preto" pageId="50" pageNumber="51" specimenCode="MZUSP 101506" specimenCount="1" stateProvince="Rio" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4B0CFC3FFF39FBD2" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4B0CFC3FFA4EFBF7" box="[1343,1436,1016,1040]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">MZUSP</collectionCode>
<date id="3C2440C64F45FFCA4EA4FBDBFF39FBD2" box="[151,235,1052,1077]" pageId="50" pageNumber="51" value="06-10-15">101506</date>
</specimenCode>
(4, 167.1–219.0 mm SL): São Gonçalo do 
<collectingRegion id="8A5EA8E44F45FFCA4CEEFBDBFCD7FBD2" box="[733,773,1052,1077]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">Rio</collectingRegion>
Preto, 
<collectingMunicipality id="A841FC7C4F45FFCA4D6BFBDBFC69FBD3" box="[856,955,1052,1077]" pageId="50" pageNumber="51">rio Preto</collectingMunicipality>
(tributary of 
<location id="4D4530DD4F45FFCA4A65FBDBFB09FBD3" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4A65FBDBFB09FBD3" box="[1110,1243,1052,1077]" country="Brazil" municipality="rio Preto" name="rio Aracuai" pageId="50" pageNumber="51" stateProvince="Rio">rio Araçuaí</location>
)
</materialsCitation>
, 
<materialsCitation id="F8F26C5B4F45FFCA4AC3FBDBFC9CFBBF" ID-GBIF-Occurrence="1425222616" collectingDate="2006-10-08" collectorName="P. G. Azevedo" country="Brazil" latitude="-18.113056" location="Rio" longLatPrecision="21" longitude="-43.344997" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Rio" typeStatus="paratype">
<collectingRegion id="8A5EA8E44F45FFCA4AC3FBDBFACAFBD2" box="[1264,1304,1052,1077]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">Rio</collectingRegion>
Preto State Park, 
<geoCoordinate id="2DAE00C14F45FFCA4EEDFBF8FE88FBBF" box="[222,346,1087,1112]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-18.113056">18°6’47’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4F5FFB87FDD0FBBF" box="[364,514,1088,1112]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-43.344997">43°20’42’’W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4C20FB87FD63FBBF" box="[531,689,1087,1112]" pageId="50" pageNumber="51">P.G. Azevedo</collectorName>
, 
<collectingDate id="2C60B92E4F45FFCA4CF1FB87FC9CFBBF" box="[706,846,1087,1112]" pageId="50" pageNumber="51" value="2006-10-08">
<date id="3C2440C64F45FFCA4CF1FB87FC9CFBBF" box="[706,846,1087,1112]" pageId="50" pageNumber="51" value="2006-10-08">8 Oct 2006</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4D6DFB87FE5BFB9A" ID-GBIF-Occurrence="1425222645" collectingDate="2006-10-08" collectionCode="MZUSP" collectorName="P. G. Azevedo" country="Brazil" latitude="-18.113056" location="Rio" longLatPrecision="21" longitude="-43.344997" pageId="50" pageNumber="51" specimenCode="MZUSP 101507" specimenCount="1" stateProvince="Rio" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4D6DFB87FBC9FBBF" box="[862,1051,1087,1112]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4D6DFB87FC69FBBF" box="[862,955,1088,1112]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">MZUSP</collectionCode>
<date id="3C2440C64F45FFCA4DF4FBF8FBC9FBBF" box="[967,1051,1087,1112]" pageId="50" pageNumber="51" value="07-10-15">101507</date>
</specimenCode>
(3, 186.1–243.0 mm SL): same locality and collector
</materialsCitation>
, 
<materialsCitation id="F8F26C5B4F45FFCA4FA5FBA3FA4BFB9A" ID-GBIF-Occurrence="1425222639" box="[406,1433,1124,1149]" collectingDate="2007-04-05" collectionCode="MZUSP" collectorName="P. G. Azevedo" country="Brazil" latitude="-18.113056" location="Rio" longLatPrecision="21" longitude="-43.344997" pageId="50" pageNumber="51" specimenCode="MZUSP 103685" specimenCount="1" stateProvince="Rio" typeStatus="paratype">
<collectingDate id="2C60B92E4F45FFCA4FA5FBA3FDFEFB9B" box="[406,556,1124,1149]" pageId="50" pageNumber="51" value="2007-04-05">
<date id="3C2440C64F45FFCA4FA5FBA3FDFEFB9B" box="[406,556,1124,1149]" pageId="50" pageNumber="51" value="2007-04-05">5 April 2007</date>
</collectingDate>
. 
<specimenCode id="183CCE7D4F45FFCA4C0BFBA3FD21FB9A" box="[568,755,1124,1149]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4C0BFBA3FD47FB9B" box="[568,661,1124,1148]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">MZUSP</collectionCode>
103685
</specimenCode>
(2 skel., 168.0–207.0 mm SL): same locality and collector
</materialsCitation>
, 
<date id="3C2440C64F45FFCA4EA4FB40FEDAFB47" box="[151,264,1159,1184]" pageId="50" pageNumber="51" value="2009-07">July 2009</date>
. 
<materialsCitation id="F8F26C5B4F45FFCA4F21FB4FFB4EFB47" ID-GBIF-Occurrence="1425222633" box="[274,1180,1159,1184]" collectingDate="2007-04-05" collectionCode="MZUSP" collectorName="F. Vieira &amp; G. B. Santos" country="Brazil" latitude="-18.113056" location="Rio" longLatPrecision="21" longitude="-43.344997" pageId="50" pageNumber="51" specimenCode="MZUSP 101508" specimenCount="1" stateProvince="Rio" typeStatus="paratype">
<specimenCode id="183CCE7D4F45FFCA4F21FB4FFE1BFB47" box="[274,457,1159,1184]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4F21FB4FFEBDFB47" box="[274,367,1160,1184]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="51">MZUSP</collectionCode>
<date id="3C2440C64F45FFCA4F46FB40FE1BFB47" box="[373,457,1159,1184]" pageId="50" pageNumber="51" value="08-10-15">101508</date>
</specimenCode>
(3, 244.0–284.0 mm SL): same locality; 
<collectorName id="E56F03D04F45FFCA4DA6FB4EFC26FB47" box="[917,1012,1159,1184]" pageId="50" pageNumber="51">F. Vieira</collectorName>
&amp; 
<collectorName id="E56F03D04F45FFCA4A24FB4FFB4EFB47" box="[1047,1180,1160,1184]" pageId="50" pageNumber="51">G.B. Santos</collectorName>
</materialsCitation>
, 
<materialsCitation id="F8F26C5B4F45FFCA4A9BFB4FFD0EFB22" ID-GBIF-Occurrence="1425222634" collectingDate="2006-01-10" collectionCode="MZUEL" collectorName="E. T. Grando" country="Brazil" latitude="-18.113056" location="Rio" longLatPrecision="21" longitude="-43.344997" pageId="50" pageNumber="51" specimenCode="MZUEL 3931" specimenCount="1" stateProvince="Rio" typeStatus="paratype">
<collectingDate id="2C60B92E4F45FFCA4A9BFB4FFAE0FB47" box="[1192,1330,1159,1184]" pageId="50" pageNumber="51" value="2006-01-10">
<date id="3C2440C64F45FFCA4A9BFB4FFAE0FB47" box="[1192,1330,1159,1184]" pageId="50" pageNumber="51" value="2006-01-10">
10 
<bibRefCitation id="2C0B1BF74F45FFCA4AF9FB4EFAE0FB47" author="Jan" box="[1226,1330,1159,1184]" pageId="50" pageNumber="51" refString="Bejarano, I., Blanco, M. P. &amp; Mojica, J. I. (2006) La comunidad ictica del rio Mesay durante el periodo de aguas altas (Caqueta, Amazonia colombiana). Caldasia, 28 (2), 359 - 370." type="journal article" year="2006">Jan 2006</bibRefCitation>
</date>
</collectingDate>
. 
<specimenCode id="183CCE7D4F45FFCA4B0FFB4EFF1DFB23" collectionCode="MZUEL" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4B0FFB4EFA4EFB47" box="[1340,1436,1161,1184]" pageId="50" pageNumber="51">MZUEL</collectionCode>
3931
</specimenCode>
(1, 130.0 mm SL): same locality; 
<collectorName id="E56F03D04F45FFCA4C63FB6AFD0EFB22" box="[592,732,1196,1221]" pageId="50" pageNumber="51">E.T. Grando</collectorName>
</materialsCitation>
, 
<date id="3C2440C64F45FFCA4CD4FB6BFCBBFB23" box="[743,873,1196,1221]" pageId="50" pageNumber="51" value="2004-02-02">2 Feb 2004</date>
.
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F45FFCA4EF4FB08FB7CF9CB" pageId="50" pageNumber="51" type="description">
<paragraph id="482566064F45FFCA4EF4FB08FB7CF9CB" blockId="50.[151,1437,151,1581]" pageId="50" pageNumber="51">
<materialsCitation id="F8F26C5B4F45FFCA4EF4FB08FC95FAEB" ID-GBIF-Occurrence="1425222632" collectingDate="1989-04" collectorName="V. Vono" county="Non" latitude="-16.616667" location="rio Jequitinhonha" longLatPrecision="1281" longitude="-42.166668" municipality="Coronel Murta" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
<emphasis id="7AEEBA144F45FFCA4EF4FB08FDD9FB0F" bold="true" box="[199,523,1231,1256]" pageId="50" pageNumber="51">
<collectingCounty id="A1441E8A4F45FFCA4EF4FB08FF2BFB0F" box="[199,249,1231,1256]" pageId="50" pageNumber="51">Non</collectingCounty>
<typeStatus id="9721D8A44F45FFCA4F30FB08FE91FB0F" box="[259,323,1231,1256]" pageId="50" pageNumber="51">types</typeStatus>
. 
<collectingRegion id="8A5EA8E44F45FFCA4F6EFB08FDD5FB0F" box="[349,519,1231,1256]" country="Brazil" name="Minas Gerais" pageId="50" pageNumber="51">Minas Gerais</collectingRegion>
:
</emphasis>
LISDEBE uncat. (2, 190.0–243.0 mm SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4A27FB08FB6DFB00" box="[1044,1215,1231,1256]" pageId="50" pageNumber="51">Coronel Murta</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4AFDFB08FA4AFB0F" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4AFDFB08FA4AFB0F" box="[1230,1432,1231,1256]" county="Non" latitude="-16.616667" longLatPrecision="1281" longitude="-42.166668" municipality="Coronel Murta" name="rio Jequitinhonha" pageId="50" pageNumber="51" stateProvince="Minas Gerais">rio Jequitinhonha</location>
, fazenda 
<location id="4D4530DD4F45FFCA4ECAFB33FE96FAEA" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4ECAFB33FE96FAEA" box="[249,324,1268,1293]" county="Non" latitude="-16.616667" longLatPrecision="1281" longitude="-42.166668" municipality="Coronel Murta" name="Jatoba" pageId="50" pageNumber="51" stateProvince="Minas Gerais">Jatobá</location>
, c. 
<geoCoordinate id="2DAE00C14F45FFCA4F43FB33FE1DFAEA" box="[368,463,1268,1293]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="925" value="-16.616667">16°37’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4FEDFB33FD9AFAEB" box="[478,584,1268,1292]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="925" value="-42.166668">42°10’W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4C64FB32FD65FAEB" box="[599,695,1269,1292]" pageId="50" pageNumber="51">V. Vono</collectorName>
, 
<collectingDate id="2C60B92E4F45FFCA4CF5FB33FC95FAEB" box="[710,839,1268,1293]" pageId="50" pageNumber="51" value="1989-04">
<date id="3C2440C64F45FFCA4CF5FB33FC95FAEB" box="[710,839,1268,1293]" pageId="50" pageNumber="51" value="1989-04">April 1989</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4D52FB33FCE4FAD7" ID-GBIF-Occurrence="1425222618" collectionCode="ZUEC" collectorName="T. C. Pessali" latitude="-16.586668" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.854164" municipality="Grao Mogol" pageId="50" pageNumber="51" specimenCode="ZUEC 6969" specimenCount="1" stateProvince="A" typeStatus="holotype">
<specimenCode id="183CCE7D4F45FFCA4D52FB33FC3FFAEA" box="[865,1005,1268,1293]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4D52FB33FC78FAEB" box="[865,938,1268,1292]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
6969
</specimenCode>
(
<quantity id="8F62CBE34F45FFCA4A30FB33FB44FAEB" box="[1027,1174,1268,1292]" metricMagnitude="0" metricUnit="m" metricValue="1.1017" pageId="50" pageNumber="51" unit="mm" value="1101.7">1, 101.7 mm</quantity>
SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4AEDFB33FABCFAEA" box="[1246,1390,1268,1293]" pageId="50" pageNumber="51">Grão Mogol</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4B4EFB33FEFCFAD7" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4B4EFB33FEFCFAD7" latitude="-16.586668" longLatPrecision="21" longitude="-42.854164" municipality="Grao Mogol" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçú</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4F09FAD0FE16FAD7" box="[314,452,1303,1328]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.586668">16°35’12’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4FE2FAD0FDB5FAD7" box="[465,615,1303,1328]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.854164">42°51’15’’W</geoCoordinate>
; 
<collectorName id="E56F03D04F45FFCA4C47FADFFD28FAD7" box="[628,762,1303,1328]" pageId="50" pageNumber="51">T.C. Pessali</collectorName>
&amp; 
<collectingRegion id="8A5EA8E44F45FFCA4D2CFADFFCE4FAD7" box="[799,822,1304,1328]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4D08FADFFC76FAB2" ID-GBIF-Occurrence="1425222649" collectingDate="2011-02" collectionCode="ZUEC" latitude="-16.586668" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.854164" municipality="Grao Mogol" pageId="50" pageNumber="51" specimenCode="ZUEC 6970" specimenCount="1" stateProvince="A" typeStatus="holotype">
<collectingRegion id="8A5EA8E44F45FFCA4D08FADFFC80FAD7" box="[827,850,1304,1328]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
. Rodrigues, 
<collectingDate id="2C60B92E4F45FFCA4DD3FAD0FB9EFAD7" box="[992,1100,1303,1328]" pageId="50" pageNumber="51" value="2011-02">
<date id="3C2440C64F45FFCA4DD3FAD0FB9EFAD7" box="[992,1100,1303,1328]" pageId="50" pageNumber="51" value="2011-02">Feb 2011</date>
</collectingDate>
. 
<specimenCode id="183CCE7D4F45FFCA4A6BFADFFB33FAD7" box="[1112,1249,1303,1328]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4A6BFADFFB73FAD7" box="[1112,1185,1304,1328]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
6970
</specimenCode>
(2, 109.1– 
<quantity id="8F62CBE34F45FFCA4B6FFADFFF11FAB3" metricMagnitude="-1" metricUnit="m" metricValue="1.384" pageId="50" pageNumber="51" unit="mm" value="138.4">138.4 mm</quantity>
SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4F3AFAFBFE4BFAB2" box="[265,409,1340,1365]" pageId="50" pageNumber="51">Grão Mogol</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4F95FAFBFDB5FAB2" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4F95FAFBFDB5FAB2" box="[422,615,1340,1365]" latitude="-16.586668" longLatPrecision="21" longitude="-42.854164" municipality="Grao Mogol" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçú</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4C46FAFBFD2DFAB2" box="[629,767,1340,1365]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.586668">16°35’12’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4D3DFAFBFC76FAB2" box="[782,932,1340,1365]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.854164">42°51’15’’W</geoCoordinate>
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F45FFCA4D80FAFBFBAEFAB3" ID-GBIF-Occurrence="1425222624" box="[947,1148,1340,1365]" collectorName="T. C. Pessali" location="A" pageId="50" pageNumber="51" specimenCount="1" stateProvince="A" typeStatus="holotype">
<collectorName id="E56F03D04F45FFCA4D80FAFBFBE9FAB2" box="[947,1083,1340,1365]" pageId="50" pageNumber="51">T.C. Pessali</collectorName>
&amp; 
<collectingRegion id="8A5EA8E44F45FFCA4A56FAFBFBAEFAB3" box="[1125,1148,1340,1364]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4AB3FAFBFB38FA9F" ID-GBIF-Occurrence="1425222646" collectingDate="2011-02" collectionCode="ZUEC" latitude="-16.581112" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.873608" municipality="Grao Mogol" pageId="50" pageNumber="51" specimenCode="ZUEC 6974" specimenCount="1" stateProvince="A" typeStatus="holotype">
<collectingRegion id="8A5EA8E44F45FFCA4AB3FAFBFB45FAB3" box="[1152,1175,1340,1364]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
. Rodrigues, 
<collectingDate id="2C60B92E4F45FFCA4B19FAFBFA4AFAB3" box="[1322,1432,1340,1365]" pageId="50" pageNumber="51" value="2011-02">
<date id="3C2440C64F45FFCA4B19FAFBFA4AFAB3" box="[1322,1432,1340,1365]" pageId="50" pageNumber="51" value="2011-02">Feb 2011</date>
</collectingDate>
. 
<specimenCode id="183CCE7D4F45FFCA4EA4FAA7FEF3FA9F" box="[151,289,1375,1400]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4EA4FAA7FF32FA9F" box="[151,224,1376,1400]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
6974
</specimenCode>
(3, 101.1– 
<quantity id="8F62CBE34F45FFCA4FADFA98FDC3FA90" box="[414,529,1375,1400]" metricMagnitude="-1" metricUnit="m" metricValue="1.7149999999999999" pageId="50" pageNumber="51" unit="mm" value="171.5">171.5 mm</quantity>
SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4C66FAA7FD30FA9F" box="[597,738,1375,1400]" pageId="50" pageNumber="51">Grão Mogol</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4CC3FA98FC7DFA9F" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4CC3FA98FC7DFA9F" box="[752,943,1375,1400]" latitude="-16.581112" longLatPrecision="21" longitude="-42.873608" municipality="Grao Mogol" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçú</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4D88FA98FB94FA9F" box="[955,1094,1375,1400]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.581112">16°34’52’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4A67FA98FB38FA9F" box="[1108,1258,1375,1400]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.873608">42°52’25’’W</geoCoordinate>
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F45FFCA4AC4FAA7FF7CFA7B" ID-GBIF-Occurrence="1425222621" collectorName="T. C. Pessali" location="A" pageId="50" pageNumber="51" specimenCount="1" stateProvince="A" typeStatus="holotype">
<collectorName id="E56F03D04F45FFCA4AC4FAA7FAAFFA9F" box="[1271,1405,1375,1400]" pageId="50" pageNumber="51">T.C. Pessali</collectorName>
&amp; 
<collectingRegion id="8A5EA8E44F45FFCA4EA4FA43FF7CFA7B" box="[151,174,1412,1436]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4E81FA43FEFEFA27" ID-GBIF-Occurrence="1425222637" collectingDate="2011-02" collectionCode="ZUEC" latitude="-16.573334" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.829445" municipality="Grao Mogol" pageId="50" pageNumber="51" specimenCode="ZUEC 6999" specimenCount="1" stateProvince="A" typeStatus="holotype">
<collectingRegion id="8A5EA8E44F45FFCA4E81FA43FF1BFA7B" box="[178,201,1412,1436]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
. Rodrigues, 
<collectingDate id="2C60B92E4F45FFCA4F54FA43FE09FA7B" box="[359,475,1412,1437]" pageId="50" pageNumber="51" value="2011-02">
<date id="3C2440C64F45FFCA4F54FA43FE09FA7B" box="[359,475,1412,1437]" pageId="50" pageNumber="51" value="2011-02">Feb 2011</date>
</collectingDate>
. 
<specimenCode id="183CCE7D4F45FFCA4FDCFA43FD52FA7A" box="[495,640,1412,1437]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4FDCFA43FDEAFA7B" box="[495,568,1412,1436]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
6999
</specimenCode>
(
<quantity id="8F62CBE34F45FFCA4CA8FA43FCEBFA7B" box="[667,825,1412,1436]" metricMagnitude="0" metricUnit="m" metricValue="1.1733" pageId="50" pageNumber="51" unit="mm" value="1173.3">1, 173.3 mm</quantity>
SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4DBAFA43FBCCFA7A" box="[905,1054,1412,1437]" pageId="50" pageNumber="51">Grão Mogol</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4A00FA43FB2BFA7A" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4A00FA43FB2BFA7A" box="[1075,1273,1412,1437]" latitude="-16.573334" longLatPrecision="21" longitude="-42.829445" municipality="Grao Mogol" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçú</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4B3FFA43FA45FA7A" box="[1292,1431,1412,1437]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.573334">16°34’24’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4EA4FA60FEFEFA27" box="[151,300,1447,1472]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.829445">42°49’46’’W</geoCoordinate>
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F45FFCA4F73FA6FFDC7FA27" ID-GBIF-Occurrence="1425222629" box="[320,533,1447,1472]" collectorName="T. C. Pessali" location="A" pageId="50" pageNumber="51" specimenCount="1" stateProvince="A" typeStatus="holotype">
<collectorName id="E56F03D04F45FFCA4F73FA6FFE1EFA27" box="[320,460,1447,1472]" pageId="50" pageNumber="51">T.C. Pessali</collectorName>
&amp; 
<collectingRegion id="8A5EA8E44F45FFCA4FCDFA6FFDC7FA27" box="[510,533,1448,1472]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4C29FA6FFD8FFA02" ID-GBIF-Occurrence="1425222631" collectingDate="2011-02" collectionCode="ZUEC" latitude="-16.59889" location="rio Itacambirucu" longLatPrecision="21" longitude="-42.835556" municipality="Grao Mogol" pageId="50" pageNumber="51" specimenCode="ZUEC 6367" specimenCount="1" stateProvince="A" typeStatus="holotype">
<collectingRegion id="8A5EA8E44F45FFCA4C29FA6FFDE3FA27" box="[538,561,1448,1472]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
. Rodrigues, 
<collectingDate id="2C60B92E4F45FFCA4CFFFA60FC92FA27" box="[716,832,1447,1472]" pageId="50" pageNumber="51" value="2011-02">
<date id="3C2440C64F45FFCA4CFFFA60FC92FA27" box="[716,832,1447,1472]" pageId="50" pageNumber="51" value="2011-02">Feb 2011</date>
</collectingDate>
. 
<specimenCode id="183CCE7D4F45FFCA4D62FA6FFC33FA27" box="[849,993,1447,1472]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">
<collectionCode id="2E8BFEC34F45FFCA4D62FA6FFC48FA27" box="[849,922,1448,1472]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="50" pageNumber="51">ZUEC</collectionCode>
6367
</specimenCode>
(1, 
<quantity id="8F62CBE34F45FFCA4A2FFA6FFB5AFA58" box="[1052,1160,1448,1472]" metricMagnitude="-2" metricUnit="m" metricValue="9.43" pageId="50" pageNumber="51" unit="mm" value="94.3">94.3 mm</quantity>
SL): 
<collectingMunicipality id="A841FC7C4F45FFCA4AE5FA6FFAB8FA27" box="[1238,1386,1447,1472]" pageId="50" pageNumber="51">Grão Mogol</collectingMunicipality>
, 
<location id="4D4530DD4F45FFCA4B4EFA60FEFCFA02" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4B4EFA60FEFCFA02" latitude="-16.59889" longLatPrecision="21" longitude="-42.835556" municipality="Grao Mogol" name="rio Itacambirucu" pageId="50" pageNumber="51" stateProvince="A">rio Itacambiruçu</location>
, 
<geoCoordinate id="2DAE00C14F45FFCA4F0FFA0BFE14FA02" box="[316,454,1484,1509]" direction="south" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="-16.59889">16°35’56’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F45FFCA4FE6FA0BFD8FFA02" box="[469,605,1484,1509]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-42.835556">42°50’8’’W</geoCoordinate>
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F45FFCA4C5FFA0AFEF9F9CA" ID-GBIF-Occurrence="1425222650" collectingDate="2010-11-22" collectionCode="NMW" collectorName="F. Andrade" location="Wertheimer" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Rio" typeStatus="paralectotype">
<collectorName id="E56F03D04F45FFCA4C5FFA0AFD3FFA02" box="[620,749,1484,1509]" pageId="50" pageNumber="51">F. Andrade</collectorName>
, 
<collectingDate id="2C60B92E4F45FFCA4CCFFA0BFC4AFA03" box="[764,920,1484,1508]" pageId="50" pageNumber="51" value="2010-11-22">
<date id="3C2440C64F45FFCA4CCFFA0BFC4AFA03" box="[764,920,1484,1508]" pageId="50" pageNumber="51" value="2010-11-22">22 Nov 2010</date>
</collectingDate>
. 
<emphasis id="7AEEBA144F45FFCA4D96FA0BFB44FA02" bold="true" box="[933,1174,1484,1509]" pageId="50" pageNumber="51">Imprecise localities:</emphasis>
<collectionCode id="2E8BFEC34F45FFCA4A93FA0AFB3AFA03" box="[1184,1256,1485,1508]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="50" pageNumber="51">NMW</collectionCode>
62927 (1, 
<quantity id="8F62CBE34F45FFCA4B58FA0BFF11F9E0" metricMagnitude="-2" metricUnit="m" metricValue="7.159999999999999" pageId="50" pageNumber="51" unit="mm" value="71.6">71.6 mm</quantity>
SL): “ 
<collectingRegion id="8A5EA8E44F45FFCA4F29FA28FE96F9EF" box="[282,324,1519,1544]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">Rio</collectingRegion>
Jequitinhonha” (no specific locality); 
<location id="4D4530DD4F45FFCA4D25FA28FC4EF9EF" LSID="urn:lsid:plazi:treatment:C033D7104F59FFCA4EA4FE53FB7CF9CB:4D4530DD4F45FFCA4D25FA28FC4EF9EF" box="[790,924,1519,1544]" name="Wertheimer" pageId="50" pageNumber="51" stateProvince="Rio">Wertheimer</location>
, no date. 
<typeStatus id="9721D8A44F45FFCA4A29FA28FB61F9EF" box="[1050,1203,1519,1544]" pageId="50" pageNumber="51" type="paralectotype">Paralectotype</typeStatus>
of 
<taxonomicName id="8F9A1D854F45FFCA4AD5FA36FEF9F9CA" authority="Steindachner." authorityName="Steindachner." class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F45FFCA4AD5FA36FA4EF9EF" box="[1254,1436,1521,1544]" italics="true" pageId="50" pageNumber="51">Brycon insignis</emphasis>
Steindachner
</taxonomicName>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4F06F9D2FC42F9CA" ID-GBIF-Occurrence="1425222635" box="[309,912,1556,1581]" collectionCode="MNHN" location="A" pageId="50" pageNumber="51" specimenCount="1" stateProvince="A" typeStatus="paralectotype">
<collectionCode id="2E8BFEC34F45FFCA4F06F9D2FE59F9CB" box="[309,395,1557,1580]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="50" pageNumber="51">MNHN</collectionCode>
<collectingRegion id="8A5EA8E44F45FFCA4FA2F9D3FE74F9CA" box="[401,422,1556,1581]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
.8612 (
<quantity id="8F62CBE34F45FFCA4FC4F9D3FD51F9CB" box="[503,643,1556,1581]" metricMagnitude="0" metricUnit="m" metricValue="1.2957999999999998" pageId="50" pageNumber="51" unit="mm" value="1295.8">1, 295.8 mm</quantity>
SL): “ 
<collectingRegion id="8A5EA8E44F45FFCA4CFEF9D3FD25F9CA" box="[717,759,1556,1581]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">Rio</collectingRegion>
Tiquilelonha
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F45FFCA4DA9F9D3FB7CF9CB" ID-GBIF-Occurrence="1425222653" box="[922,1198,1556,1581]" location="A" pageId="50" pageNumber="51" specimenCount="1" stateProvince="A" typeStatus="paralectotype">
<collectingRegion id="8A5EA8E44F45FFCA4DA9F9D3FC63F9CB" box="[922,945,1556,1580]" country="Nigeria" name="Rivers" pageId="50" pageNumber="51">A</collectingRegion>
. Saint- Hilaire, 1822”.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>