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<document id="6865748CFE802CF0C3BC13A4ACBB9699" ID-DOI="10.11646/zootaxa.3888.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10086415" ID-ZooBank="1B437AE1-CF28-4C1B-95B6-C31A295905A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623730057475" checkinUser="felipe" docAuthor="Schwarzhans, Werner" docDate="2014" docId="463A8F36FF8CFFC512979EFDE41AF826" docLanguage="en" docName="zt03888p073.pdf.imf" docOrigin="Zootaxa 3888 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hymenocephalus torvus Smith &amp; Radcliffe 1912" docType="treatment" docVersion="2" lastPageNumber="62" masterDocId="BA03F74EFFB0FFF812009814E16BFFF8" masterDocTitle="Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species" masterLastPageNumber="73" masterPageNumber="1" pageNumber="61" updateTime="1699488604623" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods id="2DC1A316FD1A88684FF4CA4639F4F93B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B86287AD472345DE70A3C5490A1148BF">Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="869FFD4FDCC6D5DB1A4973F842BC53C7">Schwarzhans, Werner</mods:namePart>
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<mods:typeOfResource id="A8E72125368E9C5AFD91846BF53CCA34">text</mods:typeOfResource>
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<mods:title id="FEA693310B8C476F59C33194A1EA0F45">Zootaxa</mods:title>
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<mods:part id="EB69EAA885F7FE37AE2C495C7AF3796F">
<mods:date id="5E89516647A9DD1E6B5275B81341EF25">2014</mods:date>
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<mods:number id="3683A9E9B1B3638EC533A8D77E42942E">2014-11-28</mods:number>
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<mods:detail id="CEB5EE35832F83E6F3E3F2B7E74EEDBD" type="volume">
<mods:number id="9AC27A325290B8285AEB1FD1E0626E47">3888</mods:number>
</mods:detail>
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<mods:number id="F48C5DEE54444D47E42AF4DB3610416E">1</mods:number>
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<mods:classification id="99A291B0C82BAA6B3C12F2D1176E8531">journal article</mods:classification>
<mods:identifier id="B08F0CB61396821F5AD251C8B9786D59" type="DOI">10.11646/zootaxa.3888.1.1</mods:identifier>
<mods:identifier id="74E8FC0844285FB41C296A0FD97FDB6F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="98EE69A0C6C283E6AF271C913D3572E2" type="Zenodo-Dep">10086415</mods:identifier>
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<treatment id="463A8F36FF8CFFC512979EFDE41AF826" LSID="urn:lsid:plazi:treatment:463A8F36FF8CFFC512979EFDE41AF826" httpUri="http://treatment.plazi.org/id/463A8F36FF8CFFC512979EFDE41AF826" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
<subSubSection id="86896DABFF8CFFC412979EFDE39BF8FB" box="[151,752,1769,1795]" pageId="60" pageNumber="61" type="nomenclature">
<paragraph id="CE2C3E20FF8CFFC412979EFDE39BF8FB" blockId="60.[151,752,1769,1829]" box="[151,752,1769,1795]" pageId="60" pageNumber="61">
<heading id="9564894CFF8CFFC412979EFDE39BF8FB" bold="true" box="[151,752,1769,1795]" fontSize="11" level="2" pageId="60" pageNumber="61" reason="4">
<taxonomicName id="099345A3FF8CFFC412979EFDE39BF8FB" ID-CoL="6MPB5" authority="Smith &amp; Radcliffe, 1912" authorityName="Smith &amp; Radcliffe" authorityYear="1912" box="[151,752,1769,1795]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="torvus">
<emphasis id="FCE7E232FF8CFFC412979EFDE39BF8FB" bold="true" box="[151,752,1769,1795]" pageId="60" pageNumber="61">
<emphasis id="FCE7E232FF8CFFC412979EFDE0D0F8FB" bold="true" box="[151,443,1769,1795]" italics="true" pageId="60" pageNumber="61">Hymenocephalus torvus</emphasis>
<bibRefCitation id="AA0243D1FF8CFFC413C29EFDE39BF8FB" author="Smith, H. M. &amp; Radcliffe, L." box="[450,752,1769,1795]" pageId="60" pageNumber="61" pagination="105 - 140" refId="ref52077" refString="Smith, H. M. &amp; Radcliffe, L. (1912) in Radcliffe. Descriptions of a new family, two new genera, and twenty-nine new species of anacanthine fishes from the Philippine Islands and contiguous waters. Proceedings of the United States National Museum, 43 (1924), 105 - 140. http: // dx. doi. org / 10.5479 / si. 00963801.43 - 1924.105" type="journal article" year="1912">Smith &amp; Radcliffe, 1912</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF8CFFC412979F18E025F8DD" box="[151,334,1804,1829]" pageId="60" pageNumber="61" type="description">
<paragraph id="CE2C3E20FF8CFFC412979F18E025F8DD" blockId="60.[151,752,1769,1829]" box="[151,334,1804,1829]" pageId="60" pageNumber="61">
<figureCitation id="56A822A5FF8CFFC412979F18E04CF8DD" box="[151,295,1804,1829]" captionStart="FIGURE 33" captionStartId="61.[151,250,1568,1590]" captionTargetBox="[178,1413,812,1538]" captionTargetId="figure-427@61.[170,1417,807,1543]" captionTargetPageId="61" captionText="FIGURE 33. Hymenocephalus torvus: A–B—USNM 149465, A—Lateral view of head, B—Dorsal view of head. C–G—MNHN 2005-0928, otoliths, C, D, F—Inner faces, E—Anterior view, G—Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10086485" httpUri="https://zenodo.org/record/10086485/files/figure.png" pageId="60" pageNumber="61">Figs. 33A–G</figureCitation>
,
<figureCitation id="56A822A5FF8CFFC413329F18E025F8DD" box="[306,334,1804,1829]" captionStart="FIGURE 37" captionStartId="66.[151,250,976,998]" captionTargetBox="[158,1430,272,949]" captionTargetId="figure-236@66.[152,1435,265,954]" captionTargetPageId="66" captionText="FIGURE 37. Geographical distribution of Hymenocephalus aeger, H. megalops, H. striatissimus, H. torvus, H. grimaldii, H. neglectissimus, H. semipellucidus, H. sp. 1 and H." figureDoi="http://doi.org/10.5281/zenodo.10086493" httpUri="https://zenodo.org/record/10086493/files/figure.png" pageId="60" pageNumber="61">37</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF8CFFC412979F40E22FF83E" pageId="60" pageNumber="61" type="reference_group">
<paragraph id="CE2C3E20FF8CFFC412979F40E01CF870" blockId="60.[151,1436,1876,1990]" pageId="60" pageNumber="61">
<treatmentCitationGroup id="EE83190EFF8CFFC412979F40E01CF870" pageId="60" pageNumber="61">
<treatmentCitation id="4F321831FF8CFFC412979F40E3D1F892" author="Smith, H. M. &amp; Radcliffe, L." box="[151,698,1876,1898]" page="110" pageId="60" pageNumber="61" year="1912">
<taxonomicName id="099345A3FF8CFFC412979F40E3D1F892" ID-CoL="6MPB5" authority="Smith &amp; Radcliffe, 1912: 110" authorityName="Smith &amp; Radcliffe" authorityPageNumber="110" authorityYear="1912" box="[151,698,1876,1898]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="torvus">
<emphasis id="FCE7E232FF8CFFC412979F40E0E1F891" box="[151,394,1876,1898]" italics="true" pageId="60" pageNumber="61">Hymenocephalus torvus</emphasis>
<bibRefCitation id="AA0243D1FF8CFFC413909F40E3D1F892" author="Smith, H. M. &amp; Radcliffe, L." box="[400,698,1876,1898]" pageId="60" pageNumber="61" pagination="105 - 140" refId="ref52077" refString="Smith, H. M. &amp; Radcliffe, L. (1912) in Radcliffe. Descriptions of a new family, two new genera, and twenty-nine new species of anacanthine fishes from the Philippine Islands and contiguous waters. Proceedings of the United States National Museum, 43 (1924), 105 - 140. http: // dx. doi. org / 10.5479 / si. 00963801.43 - 1924.105" type="journal article" year="1912">Smith &amp; Radcliffe, 1912: 110</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="11288082FF8CFFC410C79F40E39FF892" box="[711,756,1876,1898]" pageId="60" pageNumber="61">type</typeStatus>
locality: off Jolo,
<geoCoordinate id="ABA758E7FF8CFFC411AC9F40E56DF892" box="[940,1030,1876,1898]" degrees="06" direction="north" minutes="00" orientation="latitude" pageId="60" pageNumber="61" precision="925" value="6.0">06°00’N</geoCoordinate>
,
<geoCoordinate id="ABA758E7FF8CFFC416109F40E51FF892" box="[1040,1140,1876,1898]" degrees="120" direction="east" minutes="45" orientation="longitude" pageId="60" pageNumber="61" precision="925" value="120.75">120°45’E</geoCoordinate>
; location corrected by
<collectingCountry id="B6847EB0FF8CFFC417589F40E4F7F892" box="[1368,1436,1876,1898]" name="Jordan" pageId="60" pageNumber="61">Jordan</collectingCountry>
&amp;
<bibRefCitation id="AA0243D1FF8CFFC412E19F66E007F870" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[225,364,1906,1928]" pageId="60" pageNumber="61" pagination="369 - 588" refId="ref50763" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (1), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert, 1920</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="CE2C3E20FF8CFFC412979F85E39FF85F" blockId="60.[151,1436,1876,1990]" box="[151,756,1937,1959]" pageId="60" pageNumber="61">
<treatmentCitationGroup id="EE83190EFF8CFFC412979F85E39FF85F" box="[151,756,1937,1959]" pageId="60" pageNumber="61">
<treatmentCitation id="4F321831FF8CFFC412979F85E39BF85F" author="Iwamoto, T. &amp; McCosker, J. E." box="[151,752,1937,1959]" page="281" pageId="60" pageNumber="61" year="2014">
<taxonomicName id="099345A3FF8CFFC412979F85E39BF85F" ID-CoL="6MPB5" authority=": Iwamoto &amp; McCosker, 2014: 281" authorityName=": Iwamoto &amp; McCosker" authorityPageNumber="281" authorityYear="2014" box="[151,752,1937,1959]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="torvus">
<emphasis id="FCE7E232FF8CFFC412979F85E0E0F85F" box="[151,395,1937,1959]" italics="true" pageId="60" pageNumber="61">Hymenocephalus torvus</emphasis>
:
<bibRefCitation id="AA0243D1FF8CFFC413989F86E39BF85F" author="Iwamoto, T. &amp; McCosker, J. E." box="[408,752,1937,1959]" pageId="60" pageNumber="61" pagination="263 - 332" refId="ref51028" refString="Iwamoto, T. &amp; McCosker, J. E. (2014) Deep-water fishes of the 2011 Hearst Philippine biodiversity expedition of the California Academy of Sciences. In: Williams, G. C. &amp; Gosliner, T. M. (Eds.), The Coral Triangle. The 2011 Hearst Philippine Biodiversity Expedition. California Academy of Sciences, San Francisco, pp. 263 - 332." type="book chapter" year="2014">Iwamoto &amp; McCosker, 2014: 281</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CE2C3E20FF8CFFC412979FA4E22FF83E" blockId="60.[151,1436,1876,1990]" box="[151,836,1968,1990]" pageId="60" pageNumber="61">
<treatmentCitationGroup id="EE83190EFF8CFFC412979FA4E22FF83E" box="[151,836,1968,1990]" pageId="60" pageNumber="61">
<taxonomicName id="099345A3FF8CFFC412979FA4E37AF83E" ID-CoL="6MPB5" authorityName="Smith &amp; Radcliffe" authorityYear="1912" box="[151,529,1968,1990]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="60" pageNumber="61" phylum="Chordata" rank="subSpecies" species="striatissimus" subSpecies="torvus">
<emphasis id="FCE7E232FF8CFFC412979FA4E37AF83E" box="[151,529,1968,1990]" italics="true" pageId="60" pageNumber="61">Hymenocephalus striatissimus torvus</emphasis>
</taxonomicName>
:
<collectingCountry id="B6847EB0FF8CFFC4101E9FA4E309F83E" box="[542,610,1968,1990]" name="Jordan" pageId="60" pageNumber="61">Jordan</collectingCountry>
&amp;
<treatmentCitation id="4F321831FF8CFFC410829FA4E254F83E" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[642,831,1968,1990]" page="530" pageId="60" pageNumber="61" year="1920">
<bibRefCitation id="AA0243D1FF8CFFC410829FA4E254F83E" author="Gilbert, C. H. &amp; Hubbs, C. L." box="[642,831,1968,1990]" pageId="60" pageNumber="61" pagination="369 - 588" refId="ref50763" refString="Gilbert, C. H. &amp; Hubbs, C. L. (1920) The macrouroid fishes of the Philippine Islands and the East Indies. United States National Museum Bulletin, 100 (1), 369 - 588. http: // dx. doi. org / 10.5962 / bhl. title. 13637" type="journal article" year="1920">Gilbert, 1920: 530</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF8DFFC512979883E20AFEB8" pageId="61" pageNumber="62" type="materials_examined">
<paragraph id="CE2C3E20FF8DFFC512979883E20AFEB8" blockId="61.[151,1436,151,752]" pageId="61" pageNumber="62">
<emphasis id="FCE7E232FF8DFFC512979883E0EFFF48" bold="true" box="[151,388,151,176]" pageId="61" pageNumber="62">Material examined.</emphasis>
<specimenCount id="D895F5A9FF8DFFC5138E9883E346FF48" box="[398,557,151,176]" pageId="61" pageNumber="62" type="generic">15 specimens</specimenCount>
;
<materialsCitation id="7EFB347DFF8DFFC5103C9883E414FF57" box="[572,1407,151,176]" collectionCode="USNM" elevation="347" latitude="13.783334" longLatPrecision="1289" longitude="121.583336" pageId="61" pageNumber="62" specimenCode="USNM 148995" specimenCount="2">
<specimenCount id="D895F5A9FF8DFFC5103C9883E3A2FF48" box="[572,713,151,176]" pageId="61" pageNumber="62" type="generic">2 specimens</specimenCount>
(otoliths only)
<specimenCode id="9E35965BFF8DFFC511819883E558FF48" box="[897,1075,151,176]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="61" pageNumber="62" type="Museum">USNM 148995</specimenCode>
,
<geoCoordinate id="ABA758E7FF8DFFC516429883E5CEFF48" box="[1090,1189,151,176]" degrees="13" direction="north" minutes="47" orientation="latitude" pageId="61" pageNumber="62" precision="925" value="13.783334">13°47’N</geoCoordinate>
,
<geoCoordinate id="ABA758E7FF8DFFC516B49883E449FF48" box="[1204,1314,151,176]" degrees="121" direction="east" minutes="35" orientation="longitude" pageId="61" pageNumber="62" precision="925" value="121.583336">121°35’E</geoCoordinate>
,
<quantity id="096B93C5FF8DFFC517319883E414FF57" box="[1329,1407,151,176]" metricMagnitude="2" metricUnit="m" metricValue="3.47" pageId="61" pageNumber="62" unit="m" value="347.0">
<elevation id="45BED913FF8DFFC517319883E414FF57" box="[1329,1407,151,176]" metricMagnitude="2" metricUnit="m" metricValue="3.47" pageId="61" pageNumber="62" unit="m" value="347.0">347 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="7EFB347DFF8DFFC5178E9883E466FF2D" collectionCode="USNM" country="Philippines" location="Albatross" municipality="Expedition" pageId="61" pageNumber="62" specimenCode="USNM 149032, 96" specimenCount="5">
<specimenCount id="D895F5A9FF8DFFC5178E9883E067FF2D" pageId="61" pageNumber="62" type="generic">5 specimens</specimenCount>
<specimenCode id="9E35965BFF8DFFC5131598A8E0ADFF2D" box="[277,454,188,213]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="61" pageNumber="62" type="Museum">USNM 149032</specimenCode>
,
<specimenCode id="9E35965BFF8DFFC513D498A8E09AFF2D" box="[468,497,188,213]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="61" pageNumber="62" type="Museum">96</specimenCode>
+ –
<quantity id="096B93C5FF8DFFC5101898A8E31DFF2B" box="[536,630,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.43" pageId="61" pageNumber="62" unit="mm" value="143.0">143 mm</quantity>
TL,
<location id="CB4C68FBFF8DFFC510B298A8E275FF2D" LSID="urn:lsid:plazi:treatment:463A8F36FF8CFFC512979EFDE41AF826:CB4C68FBFF8DFFC510B298A8E275FF2D" box="[690,798,188,213]" country="Philippines" municipality="Expedition" name="Albatross" pageId="61" pageNumber="62">Albatross</location>
<collectingCountry id="B6847EB0FF8DFFC5112798A8E2CEFF2D" box="[807,933,188,213]" name="Philippines" pageId="61" pageNumber="62">Philippines</collectingCountry>
<collectingMunicipality id="2E48A45AFF8DFFC511AF98A8E541FF2D" box="[943,1066,188,213]" pageId="61" pageNumber="62">Expedition</collectingMunicipality>
(no further details)
</materialsCitation>
;
<materialsCitation id="7EFB347DFF8DFFC5171B98A8E25DFF0F" collectionCode="USNM" country="Philippines" elevation="402" latitude="8.616667" longLatPrecision="1301" longitude="124.6" pageId="61" pageNumber="62" specimenCount="1">
<specimenCount id="D895F5A9FF8DFFC5171B98A8E4F7FF2D" box="[1307,1436,188,213]" pageId="61" pageNumber="62" type="generic">1 specimen</specimenCount>
<collectionCode id="A882A6E5FF8DFFC5129798CBE183FF00" box="[151,232,223,248]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="61" pageNumber="62" type="Museum">USNM</collectionCode>
<quantity id="096B93C5FF8DFFC512F398CBE0D2FF0F" box="[243,441,223,248]" metricMagnitude="5" metricUnit="m" metricValue="1.4946515999999999" pageId="61" pageNumber="62" unit="mm" value="1.4946516E8">149465, 160 mm</quantity>
TL,
<geoCoordinate id="ABA758E7FF8DFFC513F898CBE330FF00" box="[504,603,223,248]" degrees="08" direction="north" minutes="37" orientation="latitude" pageId="61" pageNumber="62" precision="925" value="8.616667">08°37’N</geoCoordinate>
,
<geoCoordinate id="ABA758E7FF8DFFC5106B98CBE3B3FF00" box="[619,728,223,248]" degrees="124" direction="east" minutes="36" orientation="longitude" pageId="61" pageNumber="62" precision="925" value="124.6">124°36’E</geoCoordinate>
,
<quantity id="096B93C5FF8DFFC510E898CBE25DFF0F" box="[744,822,223,248]" metricMagnitude="2" metricUnit="m" metricValue="4.02" pageId="61" pageNumber="62" unit="m" value="402.0">
<elevation id="45BED913FF8DFFC510E898CBE25DFF0F" box="[744,822,223,248]" metricMagnitude="2" metricUnit="m" metricValue="4.02" pageId="61" pageNumber="62" unit="m" value="402.0">402 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="7EFB347DFF8DFFC5114698CBE009FEE3" collectionCode="USNM" country="Philippines" elevation="366" latitude="8.583333" longLatPrecision="1301" longitude="124.6" pageId="61" pageNumber="62" specimenCount="1">
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TL,
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,
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,
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;
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(otolith only)
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,
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,
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,
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;
<materialsCitation id="7EFB347DFF8DFFC516B39910E237FEB8" collectingDate="1914-09-03" collectionCode="ZMUC" country="Philippines" latitude="7.4166665" longLatPrecision="1303" longitude="123.23333" pageId="61" pageNumber="62" specimenCount="2">
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<collectionCode id="A882A6E5FF8DFFC5174B9910E4F7FEE5" box="[1355,1436,260,285]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="61" pageNumber="62">ZMUC</collectionCode>
unregistered
<date id="BA2D18E0FF8DFFC513289933E0C2FEB8" box="[296,425,295,320]" pageId="61" pageNumber="62" value="1914-09-03">
<collectingDate id="AA69E108FF8DFFC513289933E0C2FEB8" box="[296,425,295,320]" pageId="61" pageNumber="62" value="1914-09-03">09.03.1914</collectingDate>
</date>
, 117–
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TL,
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,
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.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF8DFFC512C79958E540FE0B" pageId="61" pageNumber="62" type="diagnosis">
<paragraph id="CE2C3E20FF8DFFC512C79958E3F7FE28" blockId="61.[151,1436,151,752]" pageId="61" pageNumber="62">
<emphasis id="FCE7E232FF8DFFC512C79958E02AFE9D" bold="true" box="[199,321,332,357]" pageId="61" pageNumber="62">Diagnosis.</emphasis>
Pelvic fin rays 7 (rarely 8 on one side); pectoral fin rays 12–14; gill rakers 19–22. Barbel short, 10–15% HL. Orbit 34–40% HL. Snout short, only slightly protruding, 15–20% HL. Ventral striae extending to periproct. Infraorbital supporter long, 80–90% OD. Otolith compressed, OL:OH = 0.8–0.9; colliculi fused; pseudocolliculum long, TCL:PCL = 1.35–1.5.
</paragraph>
<paragraph id="CE2C3E20FF8DFFC512C799C8E540FE0B" blockId="61.[151,1436,151,752]" box="[199,1067,476,501]" pageId="61" pageNumber="62">
<emphasis id="FCE7E232FF8DFFC512C799C8E009FE0D" bold="true" box="[199,354,476,501]" pageId="61" pageNumber="62">Comparison.</emphasis>
For differences see discussion to
<taxonomicName id="099345A3FF8DFFC510DC99C9E254FE0C" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[732,831,477,500]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="aeger">
<emphasis id="FCE7E232FF8DFFC510DC99C9E254FE0C" box="[732,831,477,500]" italics="true" pageId="61" pageNumber="62">H. aeger</emphasis>
</taxonomicName>
and
<taxonomicName id="099345A3FF8DFFC5117599C9E54FFE0C" authorityName=": Weber" authorityYear="1913" box="[885,1060,477,500]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="striatissimus">
<emphasis id="FCE7E232FF8DFFC5117599C9E54FFE0C" box="[885,1060,477,500]" italics="true" pageId="61" pageNumber="62">H. striatissimus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF8DFFC512C799EBE47CF8B6" pageId="61" pageNumber="62" type="description">
<paragraph id="CE2C3E20FF8DFFC512C799EBE292FD08" blockId="61.[151,1436,151,752]" pageId="61" pageNumber="62">
<emphasis id="FCE7E232FF8DFFC512C799EBE033FDE0" bold="true" box="[199,344,511,536]" pageId="61" pageNumber="62">Description.</emphasis>
Head morphology (n = 2) (
<figureCitation id="56A822A5FF8DFFC510A799EBE25DFDE0" box="[679,822,511,536]" captionStart="FIGURE 33" captionStartId="61.[151,250,1568,1590]" captionTargetBox="[178,1413,812,1538]" captionTargetId="figure-427@61.[170,1417,807,1543]" captionTargetPageId="61" captionText="FIGURE 33. Hymenocephalus torvus: A–B—USNM 149465, A—Lateral view of head, B—Dorsal view of head. C–G—MNHN 2005-0928, otoliths, C, D, F—Inner faces, E—Anterior view, G—Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10086485" httpUri="https://zenodo.org/record/10086485/files/figure.png" pageId="61" pageNumber="62">Fig. 33A–B</figureCitation>
): Snout short, slightly projecting, 15–20% HL, orbit diameter large, 34–40% HL, interorbital width 60–70% HW. Barbel short, 10–15% HL, not or just reaching vertical through anterior rim of orbit. Head canals well developed, infraorbital width 12–15% HL, supraorbital canal with 5 segments, width 13–15% HL, supratemporal canal above segment 4 of supraorbital canal, preopercular canal width 12–16% HL, postorbital-preopercular interspace 4–7% HL. Infranasal supporter moderately large; infraorbital supporter long, extended below almost entire length of orbit, 80–90% OD; preopercular supporter moderately long, 5–8% HL, with straight rear margin.
</paragraph>
<caption id="9AEC6EA8FF8DFFC512979E34E534F9AD" ID-DOI="http://doi.org/10.5281/zenodo.10086485" ID-Zenodo-Dep="10086485" httpUri="https://zenodo.org/record/10086485/files/figure.png" pageId="61" pageNumber="62" startId="61.[151,250,1568,1590]" targetBox="[178,1413,812,1538]" targetPageId="61">
<paragraph id="CE2C3E20FF8DFFC512979E34E534F9AD" blockId="61.[151,1437,1568,1621]" pageId="61" pageNumber="62">
<emphasis id="FCE7E232FF8DFFC512979E34E042F9CD" bold="true" box="[151,297,1568,1590]" pageId="61" pageNumber="62">FIGURE 33.</emphasis>
<taxonomicName id="099345A3FF8DFFC513399E34E35DF9CE" authorityName=": Iwamoto &amp; McCosker" authorityYear="2014" box="[313,566,1568,1590]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="torvus">
<emphasis id="FCE7E232FF8DFFC513399E34E35DF9CE" box="[313,566,1568,1590]" italics="true" pageId="61" pageNumber="62">Hymenocephalus torvus</emphasis>
</taxonomicName>
: A–B—USNM 149465, A—Lateral view of head, B—Dorsal view of head. C–G—MNHN 2005-0928, otoliths, C, D, F—Inner faces, E—Anterior view, G—Ventral view.
</paragraph>
</caption>
<paragraph id="CE2C3E20FF8DFFC512C79E95E47CF8B6" blockId="61.[151,1437,1665,2014]" pageId="61" pageNumber="62">
Otolith morphology (n = 10) (
<figureCitation id="56A822A5FF8DFFC510209E95E3C6F962" box="[544,685,1665,1690]" captionStart="FIGURE 33" captionStartId="61.[151,250,1568,1590]" captionTargetBox="[178,1413,812,1538]" captionTargetId="figure-427@61.[170,1417,807,1543]" captionTargetPageId="61" captionText="FIGURE 33. Hymenocephalus torvus: A–B—USNM 149465, A—Lateral view of head, B—Dorsal view of head. C–G—MNHN 2005-0928, otoliths, C, D, F—Inner faces, E—Anterior view, G—Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10086485" httpUri="https://zenodo.org/record/10086485/files/figure.png" pageId="61" pageNumber="62">Fig. 33D–G</figureCitation>
). Otolith large; OL:OH = 0.8–0.9; OH:OT = 3.5. Dorsal rim with very large, broad, undulating or crenulated predorsal lobe, distally marked by indentation; posterior tip rounded, positioned at about level of sulcus termination; ventral rim deep, regularly curved, smooth, deepest anterior of the middle; anterior rim high, nearly vertical. Inner face slightly convex, with median sulcus. Colliculi completely fused, terminating far from anterior and posterior tips of otolith; pseudocolliculum very long. TCL:PCL = 1.4–1. 5. Dorsal depression moderately large, moderately indistinct; ventral furrow distinct, close to ventral rim.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF8DFFC512C79F4DE349F841" pageId="61" pageNumber="62" type="discussion">
<paragraph id="CE2C3E20FF8DFFC512C79F4DE349F841" blockId="61.[151,1437,1665,2014]" pageId="61" pageNumber="62">
<emphasis id="FCE7E232FF8DFFC512C79F4DE02EF88A" bold="true" box="[199,325,1881,1906]" pageId="61" pageNumber="62">Discussion</emphasis>
. I follow
<bibRefCitation id="AA0243D1FF8DFFC513BA9F4EE25EF88A" author="Iwamoto, T. &amp; McCosker, J. E." box="[442,821,1881,1906]" pageId="61" pageNumber="62" pagination="263 - 332" refId="ref51028" refString="Iwamoto, T. &amp; McCosker, J. E. (2014) Deep-water fishes of the 2011 Hearst Philippine biodiversity expedition of the California Academy of Sciences. In: Williams, G. C. &amp; Gosliner, T. M. (Eds.), The Coral Triangle. The 2011 Hearst Philippine Biodiversity Expedition. California Academy of Sciences, San Francisco, pp. 263 - 332." type="book chapter" year="2014">Iwamoto and McCosker’s (2014)</bibRefCitation>
rationale in regarding
<taxonomicName id="099345A3FF8DFFC5163C9F4EE5CCF889" authorityName=": Iwamoto &amp; McCosker" authorityYear="2014" box="[1084,1191,1882,1905]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="torvus">
<emphasis id="FCE7E232FF8DFFC5163C9F4EE5CCF889" box="[1084,1191,1882,1905]" italics="true" pageId="61" pageNumber="62">H. torvus</emphasis>
</taxonomicName>
as a species separate from
<taxonomicName id="099345A3FF8DFFC512D49F6AE0EFF86D" authorityName=": Weber" authorityYear="1913" box="[212,388,1918,1941]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="striatissimus">
<emphasis id="FCE7E232FF8DFFC512D49F6AE0EFF86D" box="[212,388,1918,1941]" italics="true" pageId="61" pageNumber="62">H. striatissimus</emphasis>
</taxonomicName>
, with snout profile and pectoral fin counts probably adding further characters for distinction, while otolith morphology does not.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF8DFFC512C79FD1E41AF826" box="[199,1393,1989,2014]" pageId="61" pageNumber="62" type="distribution">
<paragraph id="CE2C3E20FF8DFFC512C79FD1E41AF826" blockId="61.[151,1437,1665,2014]" box="[199,1393,1989,2014]" pageId="61" pageNumber="62">
<emphasis id="FCE7E232FF8DFFC512C79FD1E032F826" bold="true" box="[199,345,1989,2014]" pageId="61" pageNumber="62">Distribution</emphasis>
(
<figureCitation id="56A822A5FF8DFFC513689FD1E0D7F826" box="[360,444,1989,2014]" captionStart="FIGURE 37" captionStartId="66.[151,250,976,998]" captionTargetBox="[158,1430,272,949]" captionTargetId="figure-236@66.[152,1435,265,954]" captionTargetPageId="66" captionText="FIGURE 37. Geographical distribution of Hymenocephalus aeger, H. megalops, H. striatissimus, H. torvus, H. grimaldii, H. neglectissimus, H. semipellucidus, H. sp. 1 and H." figureDoi="http://doi.org/10.5281/zenodo.10086493" httpUri="https://zenodo.org/record/10086493/files/figure.png" pageId="61" pageNumber="62">Fig. 37</figureCitation>
). The geographic distribution of
<taxonomicName id="099345A3FF8DFFC5112C9FD2E2FFF825" authorityName=": Iwamoto &amp; McCosker" authorityYear="2014" box="[812,916,1990,2013]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="torvus">
<emphasis id="FCE7E232FF8DFFC5112C9FD2E2FFF825" box="[812,916,1990,2013]" italics="true" pageId="61" pageNumber="62">H. torvus</emphasis>
</taxonomicName>
appears to be restricted to the
<collectingCountry id="B6847EB0FF8DFFC516EC9FD1E405F826" box="[1260,1390,1989,2014]" name="Philippines" pageId="61" pageNumber="62">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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