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<document id="D431E1ED9CFFB4C260C19DAFD275001E" ID-DOI="10.1080/00222930150215399" ID-ISSN="1464-5262" ID-Zenodo-Dep="5276049" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630005110294" checkinUser="felipe" docAuthor="Brito, A. &amp; Miller, P. J." docDate="2001" docId="664ACA71FFED657D263C7CA4FE49FAA3" docLanguage="en" docName="JNatHist.35.2.253-277.pdf" docOrigin="Journal of Natural History 35 (2)" docSource="http://www.tandfonline.com/doi/abs/10.1080/00222930150215399" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Gnatholepis thompsoni Jordan 1904" docType="treatment" docVersion="3" lastPageNumber="259" masterDocId="9A73B209FFE96574275E782AFF9EFF8B" masterDocTitle="Gobiid ® shes from the Cape Verde Islands, including two new species of Gobius (Teleostei: Gobioidei)" masterLastPageNumber="277" masterPageNumber="253" pageNumber="255" updateTime="1699136295297" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="AA59F41252ED8C444C161F1085BB37BF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4B6DFDB8EE5848B354F5D85054B2758D">Gobiid ® shes from the Cape Verde Islands, including two new species of Gobius (Teleostei: Gobioidei)</mods:title>
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<mods:namePart id="D7AA7BCDE467C00C9FFF87592508BB81">Brito, A.</mods:namePart>
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<mods:namePart id="65D340F31E8BF5E969020A66A219C582">Miller, P. J.</mods:namePart>
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<mods:typeOfResource id="BB9E5953765606BC00C02741AEA901D7">text</mods:typeOfResource>
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<mods:title id="0DFED9708BC472FC6DA1428BEF650762">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="4BBF7947EB3557CA79D784BBFF6F5167">
<mods:date id="032E332843A84CADC78184192675083B">2001</mods:date>
<mods:detail id="D91D9B099268A4AE0EC25957148E96F4" type="pubDate">
<mods:number id="662A68E53F1FBCB53CD5BBC50F3EAF33">2010-12-03</mods:number>
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<mods:detail id="A1BC374620140DA6DE9673D2CD3A7437" type="volume">
<mods:number id="11FB9D9583A0E9363FECA5F8A835B27F">35</mods:number>
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<mods:number id="DA86BDC98A13699689DF369C08C26CB3">2</mods:number>
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<mods:start id="4847C50E708B3D10B77C600603B8A1CC">253</mods:start>
<mods:end id="A08895E8B1E4514DCB04366EC78F95D1">277</mods:end>
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<mods:url id="2512D62EE6001FB19122F2BFB74C9B81">http://www.tandfonline.com/doi/abs/10.1080/00222930150215399</mods:url>
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<mods:classification id="8E845D9535D3A6A20FBC9F5C7877B98B">journal article</mods:classification>
<mods:identifier id="469666F76B9F1BF9E2B149B92B35631F" type="DOI">10.1080/00222930150215399</mods:identifier>
<mods:identifier id="04379C0BF5B78D24BC3B42CAC42803CA" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="EEED27F636ED2BD703DE6EC7AE44789E" type="Zenodo-Dep">5276049</mods:identifier>
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<treatment id="664ACA71FFED657D263C7CA4FE49FAA3" LSID="urn:lsid:plazi:treatment:664ACA71FFED657D263C7CA4FE49FAA3" httpUri="http://treatment.plazi.org/id/664ACA71FFED657D263C7CA4FE49FAA3" lastPageId="9" lastPageNumber="259" pageId="4" pageNumber="255">
<subSubSection id="A6F928ECFFED6570263C7CA4FC95FB2D" box="[354,779,1166,1190]" pageId="4" pageNumber="255" type="nomenclature">
<paragraph id="EE5C7B67FFED6570263C7CA4FC95FB2D" blockId="4.[354,779,1166,1222]" box="[354,779,1166,1190]" pageId="4" pageNumber="255">
<heading id="B514CC0BFFED6570263C7CA4FC95FB2D" box="[354,779,1166,1190]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="255" reason="3">
<taxonomicName id="29E300E4FFED6570263C7CA4FC95FB2D" ID-CoL="3GLPG" authority="(Jordan, 1904)" authorityName="Jordan" authorityYear="1904" baseAuthorityName="Jordan" baseAuthorityYear="1904" box="[354,779,1166,1190]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="255" phylum="Chordata" rank="species" species="thompsoni">
<emphasis id="DC97A775FFED6570263C7CA4FDC7FB2D" bold="true" box="[354,601,1166,1190]" italics="true" pageId="4" pageNumber="255">Gnatholepis thompsoni</emphasis>
(
<collectingCountry id="96F43BF7FFED657025317CA4FD5CFB2D" box="[623,706,1166,1190]" name="Jordan" pageId="4" pageNumber="255">Jordan</collectingCountry>
, 1904)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A6F928ECFFED657D26AA7C84FE49FAA3" lastPageId="9" lastPageNumber="260" pageId="4" pageNumber="255" type="description">
<paragraph id="EE5C7B67FFED657026AA7C84FDE3FB4D" blockId="4.[354,779,1166,1222]" box="[500,637,1198,1222]" pageId="4" pageNumber="255">
(
<figureCitation id="76D867E2FFED6570255E7C84FDEBFB4D" box="[512,629,1198,1222]" captionStart="FIG" captionStartId="6.[92,108,1036,1057]" captionTargetBox="[95,1043,129,1009]" captionTargetId="figure-0@6.[95,1043,129,1009]" captionTargetPageId="6" captionText="FIG. 2. (A) Gnatholepis thompsoni, male, 43.01 13.0mm, SaÄo Vicente; (B) Gobius ateriformis sp. nov., holotype, male, 55.0 1 12.0mm (BMNH), SaÄo Tiago; (C) Mauligobius nigri, female, 70.0 1 16.0mm, SaÄo Tiago." figureDoi="http://doi.org/10.5281/zenodo.5276053" httpUri="https://zenodo.org/record/5276053/files/figure.png" pageId="4" pageNumber="255">Figure 2A</figureCitation>
)
</paragraph>
<paragraph id="EE5C7B67FFED657027DA7CCAFB89FABC" blockId="4.[92,1060,1247,1686]" pageId="4" pageNumber="255">
<emphasis id="DC97A775FFED657027DA7CCAFF78FB73" box="[132,230,1248,1272]" italics="true" pageId="4" pageNumber="255">Material</emphasis>
. Cape Verdes: 2 ll, 271 7 and 29.51
<quantity id="291BD682FFED657025ED7CCAFD6FFB73" box="[691,753,1248,1272]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="255" unit="mm" value="9.0">9 mm</quantity>
, Caleta de San Martinho, Ilha de SaÄo Tiago,
<date id="9A5D5DA7FFED657026767D2AFE6CFA93" box="[296,498,1280,1304]" pageId="4" pageNumber="255" value="1987-12-30">30 December 1987</date>
, A. Brito; 1, l43.01 13.0 mm,
<collectingCountry id="96F43BF7FFED657024617D2AFCF4FA93" box="[831,874,1280,1304]" name="Iceland" pageId="4" pageNumber="255">Ilha</collectingCountry>
de Sa Äo Vicente, 1993, U. Schliewen; 1,l611
<quantity id="291BD682FFED657026F17D35FE69FABC" box="[431,503,1311,1335]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="255" unit="mm" value="21.0">21 mm</quantity>
(RMNH), Baia das Gates (16 ss54¾N, 24ss55¾W),
</paragraph>
<paragraph id="EE5C7B67FFED657027027D15FC45FADC" blockId="4.[92,1060,1247,1686]" box="[92,987,1343,1368]" pageId="4" pageNumber="255">
Ilha de SaÄo Vicente,
<date id="9A5D5DA7FFED6570261C7D15FE51FADC" box="[322,463,1343,1367]" pageId="4" pageNumber="255" value="1982-07-22">22 July 1982</date>
, CANCAP;
<quantity id="291BD682FFED657025047D15FDE0FAD3" box="[602,638,1343,1368]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="255" unit="m" value="1.0">1 m</quantity>
, 40.51 12.0 mm, 1998, A. Brito.
</paragraph>
<paragraph id="EE5C7B67FFED657027DA7D75FB94FA1C" blockId="4.[92,1060,1247,1686]" pageId="4" pageNumber="255">
<emphasis id="DC97A775FFED657027DA7D75FEFCFAFC" box="[132,354,1375,1399]" italics="true" pageId="4" pageNumber="255">Associated material.</emphasis>
Western Atlantic: 10 ll, 27.0 1 8.0 to 41.5 1 12.0,
<quantity id="291BD682FFED657024907D75FB8EFAF3" box="[974,1040,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="255" unit="mm" value="10.0">10 mm</quantity>
, 28.5 to 42.5 1 13.0 mm, and 20 unsexed (ANSP 123774), Dowling’s Shoal,
<collectingCountry id="96F43BF7FFED657024EB7D55FB94FA1C" box="[949,1034,1407,1431]" name="Anguilla" pageId="4" pageNumber="255">Anguilla</collectingCountry>
</paragraph>
<paragraph id="EE5C7B67FFED657027027DB5FB88FA3C" blockId="4.[92,1060,1247,1686]" box="[92,1046,1439,1465]" pageId="4" pageNumber="255">
(18ss14¾N, 63ss05¾W); 15 ll, 23.5 1 4.8 to 32.0 1 11.0 mm and
<quantity id="291BD682FFED6570246D7DB5FCE8FA33" box="[819,886,1439,1464]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="255" unit="mm" value="13.0">13 mm</quantity>
, 17.0 1 4.5 to
</paragraph>
<paragraph id="EE5C7B67FFED657027027D95FB89FA5C" blockId="4.[92,1060,1247,1686]" box="[92,1047,1471,1495]" pageId="4" pageNumber="255">
37.0 1
<quantity id="291BD682FFED657027FB7D95FE9DFA5C" box="[165,259,1471,1495]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="4" pageNumber="255" unit="mm" value="10.7">10.7 mm</quantity>
(ANSP 124650), Bequia Island,
<collectingCountry id="96F43BF7FFED657025DF7D95FC9CFA5C" box="[641,770,1471,1495]" name="Saint Vincent and the Grenadines" pageId="4" pageNumber="255">Grenadines</collectingCountry>
of St Vincent (13 ss00¾N,
</paragraph>
<paragraph id="EE5C7B67FFED657027027DF5FB88FA7D" blockId="4.[92,1060,1247,1686]" box="[92,1046,1502,1528]" pageId="4" pageNumber="255">
61ss15¾W); 6 ll, 19.0 1 4.8 to 34.0 1
<quantity id="291BD682FFED657026AC7DF5FDCDFA7D" box="[498,595,1503,1527]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="255" unit="mm" value="11.5">11.5 mm</quantity>
, and
<quantity id="291BD682FFED657025D17DF5FD21FA7C" box="[655,703,1503,1527]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="255" unit="mm" value="2.0">2 mm</quantity>
, 29.0 1 8.5 and 43.0 1 14.0 mm
</paragraph>
<paragraph id="EE5C7B67FFED657027027DD5FB88F9BD" blockId="4.[92,1060,1247,1686]" pageId="4" pageNumber="255">
(ANSP 126767),
<collectingRegion id="2C27B585FFED657026747DD5FE41F99C" box="[298,479,1535,1559]" country="Bahamas" name="Crooked Island and Long Cay" pageId="4" pageNumber="255">Crooked Island</collectingRegion>
(22 ss45¾N, 74ss10¾W),
<collectingCountry id="96F43BF7FFED657025BA7DD5FCCFF99C" box="[740,849,1535,1559]" name="Bahamas" pageId="4" pageNumber="255">Bahamas</collectingCountry>
.
<collectingCountry id="96F43BF7FFED657024017DD5FC4BF99C" box="[863,981,1535,1559]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="4" pageNumber="255">Ascension</collectingCountry>
: 1, l 46 1
<quantity id="291BD682FFED657027CF7E35FF45F9BD" box="[145,219,1567,1591]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="255" unit="mm" value="15.0">15 mm</quantity>
(GCRL V76:15199); 4 ll, 22 1 d to 32.5 1
<quantity id="291BD682FFED657025817E35FCB5F9BD" box="[735,811,1567,1591]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="255" unit="mm" value="11.0">11 mm</quantity>
, and
<quantity id="291BD682FFED657024307E35FC3CF9BC" box="[878,930,1567,1591]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="255" unit="mm" value="3.0">3 mm</quantity>
, 26 1 7 to
</paragraph>
<paragraph id="EE5C7B67FFED657027027E15FB89F9DD" blockId="4.[92,1060,1247,1686]" box="[92,1047,1599,1623]" pageId="4" pageNumber="255">
30 1
<quantity id="291BD682FFED657027CF7E15FF50F9DD" box="[145,206,1599,1623]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="255" unit="mm" value="9.0">9 mm</quantity>
(BMNH 1979.1.5.168±174); 3 (BMNH 1979.1.5.179±181).
<collectingCountry id="96F43BF7FFED657024DE7E15FC4CF9DC" box="[896,978,1599,1623]" name="Ghana" pageId="4" pageNumber="255">Ghana</collectingCountry>
:
<quantity id="291BD682FFED657024837E15FB8EF9DC" box="[989,1040,1599,1623]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="255" unit="mm" value="2.0">2 mm</quantity>
,
</paragraph>
<paragraph id="EE5C7B67FFED657027027E75FB8FF9FC" blockId="4.[92,1060,1247,1686]" box="[92,1041,1631,1655]" pageId="4" pageNumber="255">
25.5 1 8 and 30.0 1 9.0 mm (BMNH), Vernon Bank, R. Lubbock.
<collectingRegion id="2C27B585FFED6570243F7E75FB93F9FC" box="[865,1037,1631,1655]" country="Spain" name="Canarias" pageId="4" pageNumber="255">Canary Islands</collectingRegion>
:
</paragraph>
<paragraph id="EE5C7B67FFED657027027E54FC8BF91D" blockId="4.[92,1060,1247,1686]" box="[92,789,1662,1686]" pageId="4" pageNumber="255">
<quantity id="291BD682FFED657027027E54FF1EF91D" box="[92,128,1662,1686]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="255" unit="m" value="1.0">1 m</quantity>
, 42.0 1
<quantity id="291BD682FFED657027847E54FEA5F91D" box="[218,315,1662,1686]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="255" unit="mm" value="12.5">12.5 mm</quantity>
, Boca Cangrejo, Tenerife, 1998, A. Brito.
</paragraph>
<caption id="BA9C2BEFFFEC657127027E05FDBBF9F1" ID-DOI="http://doi.org/10.5281/zenodo.5276051" ID-Zenodo-Dep="5276051" httpUri="https://zenodo.org/record/5276051/files/figure.png" pageId="5" pageNumber="256" startId="5.[92,108,1583,1604]" targetBox="[92,1048,129,1556]" targetPageId="5">
<paragraph id="EE5C7B67FFEC657127027E05FDBBF9F1" blockId="5.[92,1049,1583,1658]" pageId="5" pageNumber="256">
FIG. 1.
<taxonomicName id="29E300E4FFEC657127E97E05FEC1F9CE" authorityName="Van Tassell" authorityYear="1988" box="[183,351,1583,1605]" class="Actinopterygii" family="Gobiidae" genus="Didogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="256" phylum="Chordata" rank="species" species="kochi">
<emphasis id="DC97A775FFEC657127E97E05FEC1F9CE" box="[183,351,1583,1605]" italics="true" pageId="5" pageNumber="256">Didogobius kochi</emphasis>
</taxonomicName>
, dark phase; (B)
<taxonomicName id="29E300E4FFEC657125487E1AFDF2F9CE" authorityName="Van Tassell" authorityYear="1988" box="[534,620,1583,1605]" class="Actinopterygii" family="Gobiidae" genus="Didogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="256" phylum="Chordata" rank="species" species="kochi">
<emphasis id="DC97A775FFEC657125487E1AFDF2F9CE" box="[534,620,1583,1605]" italics="true" pageId="5" pageNumber="256">D. kochi</emphasis>
</taxonomicName>
, paler phase retaining prominent anterior vertical dark band; (C)
<taxonomicName id="29E300E4FFEC657126C87E63FDD6F9D4" authority="Brito &amp; Miller, 2001" authorityName="Brito &amp; Miller" authorityYear="2001" box="[406,584,1609,1631]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="256" phylum="Chordata" rank="species" species="ateriformis" status="sp. nov.">
<emphasis id="DC97A775FFEC657126C87E63FDD6F9D4" box="[406,584,1609,1631]" italics="true" pageId="5" pageNumber="256">Gobius ateriformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A41A0EFFEC657125087E60FD37F9D4" box="[598,681,1610,1631]" pageId="5" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
; (D)
<taxonomicName id="29E300E4FFEC657125817E63FC24F9D4" authority="Brito &amp; Miller, 2001" authorityName="Brito &amp; Miller" authorityYear="2001" box="[735,954,1609,1631]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="256" phylum="Chordata" rank="species" species="tetrophthalmus" status="sp. nov.">
<emphasis id="DC97A775FFEC657125817E63FC24F9D4" box="[735,954,1609,1631]" italics="true" pageId="5" pageNumber="256">Gobius tetrophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A41A0EFFEC657124987E60FB87F9D4" box="[966,1049,1610,1631]" pageId="5" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
All from Ilheu do Sal Rei, Boa Vista.
</paragraph>
</caption>
<caption id="BA9C2BEFFFEF657227027C26FD99FBDC" ID-DOI="http://doi.org/10.5281/zenodo.5276053" ID-Zenodo-Dep="5276053" httpUri="https://zenodo.org/record/5276053/files/figure.png" pageId="6" pageNumber="257" startId="6.[92,108,1036,1057]" targetBox="[95,1043,129,1009]" targetPageId="6">
<paragraph id="EE5C7B67FFEF657227027C26FD99FBDC" blockId="6.[92,1047,1035,1111]" pageId="6" pageNumber="257">
FIG. 2. (A)
<taxonomicName id="29E300E4FFEF657227BD7C21FE5EFBAA" authorityName="Jordan" authorityYear="1904" box="[227,448,1035,1057]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="257" phylum="Chordata" rank="species" species="thompsoni">
<emphasis id="DC97A775FFEF657227BD7C21FE5EFBAA" box="[227,448,1035,1057]" italics="true" pageId="6" pageNumber="257">Gnatholepis thompsoni</emphasis>
</taxonomicName>
, male, 43.01 13.0mm, SaÄo Vicente; (B)
<taxonomicName id="29E300E4FFEF657224017C21FB8EFBAA" authority="Brito &amp; Miller, 2001" authorityName="Brito &amp; Miller" authorityYear="2001" box="[863,1040,1035,1057]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="257" phylum="Chordata" rank="species" species="ateriformis" status="sp. nov.">
<emphasis id="DC97A775FFEF657224017C21FB8EFBAA" box="[863,1040,1035,1057]" italics="true" pageId="6" pageNumber="257">Gobius ateriformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A41A0EFFEF657227FD7C0DFF66FBB7" box="[163,248,1063,1084]" pageId="6" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male, 55.0 1 12.0 mm (BMNH), Sa Äo Tiago; (C)
<taxonomicName id="29E300E4FFEF657224017C0CFB92FBB7" baseAuthorityName="GuEnther" baseAuthorityYear="1861" box="[863,1036,1062,1084]" class="Actinopterygii" family="Gobiidae" genus="Mauligobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="257" phylum="Chordata" rank="species" species="nigri">
<emphasis id="DC97A775FFEF657224017C0CFB92FBB7" box="[863,1036,1062,1084]" italics="true" pageId="6" pageNumber="257">Mauligobius nigri</emphasis>
</taxonomicName>
, female, 70.0 1 16.0 mm, SaÄo Tiago.
</paragraph>
</caption>
<paragraph id="EE5C7B67FFEF657227DA7C8AFB91F91D" blockId="6.[92,1047,1184,1686]" pageId="6" pageNumber="257">
<emphasis id="DC97A775FFEF657227DA7C8AFE87FB33" box="[132,281,1184,1208]" italics="true" pageId="6" pageNumber="257">Generic identi</emphasis>
W
<emphasis id="DC97A775FFEF657226737C8BFEEEFB33" box="[301,368,1185,1208]" italics="true" pageId="6" pageNumber="257">cation</emphasis>
.
<taxonomicName id="29E300E4FFEF657226CF7C8AFD28FB33" authority="Bleeker, 1874" authorityName="Bleeker" authorityYear="1874" box="[401,694,1184,1208]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="257" phylum="Chordata" rank="genus">
<emphasis id="DC97A775FFEF657226CF7C8AFD92FB33" box="[401,524,1184,1208]" italics="true" pageId="6" pageNumber="257">Gnatholepis</emphasis>
Bleeker, 1874
</taxonomicName>
(
<typeStatus id="3158C5C5FFEF657225967C8BFD66FB32" box="[712,760,1185,1209]" pageId="6" pageNumber="257">type</typeStatus>
species
<taxonomicName id="29E300E4FFEF657224087C8AFF69FB53" authority="Bleeker, 1851" authorityName="Bleeker" authorityYear="1851" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="257" phylum="Chordata" rank="species" species="anjerensis">
<emphasis id="DC97A775FFEF657224087C8AFB90FB33" box="[854,1038,1184,1208]" italics="true" pageId="6" pageNumber="257">Gobius anjerensis</emphasis>
Bleeker, 1851
</taxonomicName>
, by original designation and monotypy) is a tropical Indo±West Paci®c and Atlantic gobionelline genus (
<emphasis id="DC97A775FFEF6572268B7CCBFD8EFB73" box="[469,528,1249,1272]" italics="true" pageId="6" pageNumber="257">sensu</emphasis>
<bibRefCitation id="8A720696FFEF657225497CCAFD2FFB73" author="MILLER, P. J." box="[535,689,1248,1272]" pageId="6" pageNumber="257" pagination="397" refId="ref13644" refString="MILLER, P. J., 1973 b, The osteology and adaptive features of Rhyacichthys aspro (Teleostei: Gobioidei) and the classi ® cation of gobioid ® shes, Journal of Zoology, London, 171, 397 ± 434." type="journal article" year="1973">Miller, 1973b</bibRefCitation>
). The Cape Verdean specimens listed above are placed in
<taxonomicName id="29E300E4FFEF657226DE7D2AFE65FA93" authorityName="Bleeker" authorityYear="1874" box="[384,507,1280,1304]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="257" phylum="Chordata" rank="genus">
<emphasis id="DC97A775FFEF657226DE7D2AFE65FA93" box="[384,507,1280,1304]" italics="true" pageId="6" pageNumber="257">Gnatholepis</emphasis>
</taxonomicName>
, by the possession of (i) head lateral-line canal system with anterior and posterior oculoscapular and preopercular canals; (ii) anterior oculoscapular canals separate in anterior interorbit, with paired pores
<emphasis id="DC97A775FFEF6572235D7D15FB8EFADC" bold="true" box="[1027,1040,1343,1367]" italics="true" pageId="6" pageNumber="257">l</emphasis>
, and extending to opposite anterior nostril (pores
<emphasis id="DC97A775FFEF657225C47D75FD36FAFC" bold="true" box="[666,680,1375,1399]" italics="true" pageId="6" pageNumber="257">S</emphasis>
and terminal
<emphasis id="DC97A775FFEF657224117D75FCC3FAFC" bold="true" box="[847,861,1375,1399]" italics="true" pageId="6" pageNumber="257">S</emphasis>
1); (iii) anterior oculoscapular pores
<emphasis id="DC97A775FFEF657226167D55FEC5FA1C" bold="true" box="[328,347,1407,1431]" italics="true" pageId="6" pageNumber="257">v</emphasis>
and
<emphasis id="DC97A775FFEF657226C87D55FE3AFA1C" bold="true" box="[406,420,1407,1431]" italics="true" pageId="6" pageNumber="257">B</emphasis>
absent; (iv) four transverse rows of suborbital sensory papillae before hyomandibular row
<emphasis id="DC97A775FFEF6572255F7DB5FD90FA3C" box="[513,526,1439,1463]" italics="true" pageId="6" pageNumber="257">b</emphasis>
; (v) three transverse rows above row
<emphasis id="DC97A775FFEF6572248E7DB5FC43FA3C" box="[976,989,1439,1463]" italics="true" pageId="6" pageNumber="257">b</emphasis>
, last near pore
<emphasis id="DC97A775FFEF6572278D7D95FF7EFA5C" bold="true" box="[211,224,1471,1495]" italics="true" pageId="6" pageNumber="257">a</emphasis>
; (vi) row
<emphasis id="DC97A775FFEF657226067D95FEF2FA5C" box="[344,364,1471,1495]" italics="true" pageId="6" pageNumber="257">6i</emphasis>
above level of rear end of hyomandibular row
<emphasis id="DC97A775FFEF657224CA7D95FC3FFA5C" box="[916,929,1471,1495]" italics="true" pageId="6" pageNumber="257">d</emphasis>
; (vii) row
<emphasis id="DC97A775FFEF657227027DF5FFF7FA7C" box="[92,105,1503,1527]" italics="true" pageId="6" pageNumber="257">b</emphasis>
short, not extending anteriorly beyond vertical of rear edge of pupil; (viii) row
<emphasis id="DC97A775FFEF657223547DF5FB89FA7C" box="[1034,1047,1503,1527]" italics="true" pageId="6" pageNumber="257">d</emphasis>
divided into two parts; (ix) head, opercle, and cheek covered with ctenoid scales, (x) snout short, convex in pro®le, (xi) anterior pelvic membrane (frenum) with villose free edge; (xii) lower lip with lateral dermal ¯ap more or less evident near angle of jaws; (xiii) jaw teeth with outer row enlarged, especially in lower jaw; (xiv) vertebrae 10 1 16 (including urostyle) 5 26; (xv) dorsal pterygiophore formula (3) 12210; (xvi)
</paragraph>
<paragraph id="EE5C7B67FFEE6573270278AFFB99FED3" blockId="7.[92,1047,132,1686]" pageId="7" pageNumber="258">
two anal ®n ptergiophores before ®rst haemal arch; and (xvii) caudal skeleton with two epurals (
<bibRefCitation id="8A720696FFEE657327AD7889FE1FFF30" author="MILLER, P. J." box="[243,385,163,187]" pageId="7" pageNumber="258" refId="ref13689" refString="MILLER, P. J., 1981, The gobiid genus Gnatholepis, its supposed congeners and its new-found relatives, Abstracts of the 1 st International Conference on the Systematics and Evolution of Indo ± Paci W c Fishes. (Sydney: Australian Museum)." type="proceedings paper" year="1981">Miller, 1981</bibRefCitation>
; Akihito
<emphasis id="DC97A775FFEE657326B3788EFD83FF30" box="[493,541,163,187]" italics="true" pageId="7" pageNumber="258">et al</emphasis>
., 1988;
<bibRefCitation id="8A720696FFEE657325277889FCF6FF30" author="BIRDSONG, R. S. &amp; MURDY, E. O. &amp; PEZOLD, F. L." box="[633,872,163,187]" pageId="7" pageNumber="258" pagination="174" refId="ref12870" refString="BIRDSONG, R. S., MURDY, E. O. and PEZOLD, F. L., 1988, A study of the vertebral column and median ® n osteology in gobioid ® shes with comments on gobioid relationships, Bulletin of Marine Science, 42, 174 ± 214." type="journal article" year="1988">
Birdsong
<emphasis id="DC97A775FFEE657325B6788EFC86FF30" box="[744,792,163,187]" italics="true" pageId="7" pageNumber="258">et al</emphasis>
., 1988
</bibRefCitation>
;
<bibRefCitation id="8A720696FFEE6573242D7889FB9BFF30" author="PEZOLD, F." box="[883,1029,163,187]" pageId="7" pageNumber="258" pagination="634" refId="ref14177" refString="PEZOLD, F., 1993, Evidence for a monophyletic Gobiinae, Copeia, 1993 (3), 634 ± 643." type="journal article" year="1993">Pezold, 1993</bibRefCitation>
).
<emphasis id="DC97A775FFEE657327DA78E9FF5EFF50" box="[132,192,195,219]" italics="true" pageId="7" pageNumber="258">Speci</emphasis>
W
<emphasis id="DC97A775FFEE6573279078EEFEBFFF50" box="[206,289,195,219]" italics="true" pageId="7" pageNumber="258">c identi</emphasis>
W
<emphasis id="DC97A775FFEE6573266F78EEFD8FFF50" box="[305,529,195,219]" italics="true" pageId="7" pageNumber="258">
cation.
<taxonomicName id="29E300E4FFEE657326C878E9FD8FFF50" authorityName="Bleeker" authorityYear="1874" box="[406,529,195,219]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="258" phylum="Chordata" rank="genus">Gnatholepis</taxonomicName>
</emphasis>
contains a number of nominal species, but only one is recognized for the Atlantic basin. This is the Goldspot Goby,
<emphasis id="DC97A775FFEE657324CA78C8FF4BFE92" italics="true" pageId="7" pageNumber="258">
<taxonomicName id="29E300E4FFEE657324CA78C8FF4FFE92" authorityName="Jordan" authorityYear="1904" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="258" phylum="Chordata" rank="species" species="thompsoni">Gnatholepis thompsoni</taxonomicName>
,
</emphasis>
originally described from the western Atlantic, where it occurs from Florida and
<collectingCountry id="96F43BF7FFEE657327B2790BFECDFEB2" box="[236,339,289,313]" name="Bermuda" pageId="7" pageNumber="258">Bermuda</collectingCountry>
through the Caribbean to northern South America (
<bibRefCitation id="8A720696FFEE657324EB790BFF0BFED3" author="RANDALL, J." pageId="7" pageNumber="258" refId="ref14201" refString="RANDALL, J., 1968, Caribbean Reef Fishes. (Neptune City, New Jersey: Tropical Fish Hobbyist), 318 pp." type="book" year="1968">Randall, 1968</bibRefCitation>
;
<bibRefCitation id="8A720696FFEE657327FF796AFECEFED3" author="CERVIGON, F." box="[161,336,320,344]" pageId="7" pageNumber="258" refId="ref13043" refString="CERVIGON, F., 1994, Los Peces Marinos de Venezuela. 2 nd ed. Vol. III. (Caracas: FundacioAn Cienti ® ca Los Roques), 295 pp." type="book" year="1994">Cervigon, 1994</bibRefCitation>
). More recently,
<taxonomicName id="29E300E4FFEE65732540796AFD07FED3" authorityName="Bleeker" authorityYear="1874" box="[542,665,320,344]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="258" phylum="Chordata" rank="genus">
<emphasis id="DC97A775FFEE65732540796AFD07FED3" box="[542,665,320,344]" italics="true" pageId="7" pageNumber="258">Gnatholepis</emphasis>
</taxonomicName>
populations have been reported
</paragraph>
<paragraph id="EE5C7B67FFEE657327027975FB90FD1B" blockId="7.[92,1047,132,1686]" pageId="7" pageNumber="258">
from the oceanic islands of
<collectingCountry id="96F43BF7FFEE657326C77975FD95FEFC" box="[409,523,351,375]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="7" pageNumber="258">Ascension</collectingCountry>
and St Helena (
<bibRefCitation id="8A720696FFEE657325947975FC73FEFC" author="EDWARDS, A. J. &amp; GLASS, C. W." box="[714,1005,351,375]" pageId="7" pageNumber="258" pagination="617" refId="ref13203" refString="EDWARDS, A. J. and GLASS, C. W., 1987, The ® shes of Saint Helena Island, South Atlantic Ocean. I. The shore ® shes, Journal of Natural History, 21, 617 ± 686." type="journal article" year="1987">Edwards and Glass, 1987</bibRefCitation>
) as well as from the West African coast (
<bibRefCitation id="8A720696FFEE657325417954FD30FE1D" author="MILLER, P. J." box="[543,686,382,406]" pageId="7" pageNumber="258" pagination="951" refId="ref13836" refString="MILLER, P. J., 1990, Gobiidae, in J. C. Quero, J. C. Hureau, C. Karrer, A. Post and L. Saldanha (eds), Check-list of the Fishes of the Eastern Tropical Atlantic. (Lisbon: JNICT, EIU, UNESCO), 2, 925 ± 951." type="journal article" year="1990">Miller, 1990</bibRefCitation>
) and, most recently, from the Canaries, around Tenerife (AB, personal observation). These records have been identi®ed as
<emphasis id="DC97A775FFEE657327A67997FE0DFE5E" box="[248,403,445,469]" italics="true" pageId="7" pageNumber="258">
<taxonomicName id="29E300E4FFEE657327A67997FE11FE5E" authorityName="Jordan" authorityYear="1904" box="[248,399,445,469]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="258" phylum="Chordata" rank="species" species="thompsoni">G. thompsoni</taxonomicName>
.
</emphasis>
Comparison of west Atlantic
<taxonomicName id="29E300E4FFEE657324537997FC3CFE5E" authorityName="Jordan" authorityYear="1904" box="[781,930,445,469]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="258" phylum="Chordata" rank="species" species="thompsoni">
<emphasis id="DC97A775FFEE657324537997FC3CFE5E" box="[781,930,445,469]" italics="true" pageId="7" pageNumber="258">G. thompsoni</emphasis>
</taxonomicName>
with the Indo±West Paci®c species
<taxonomicName id="29E300E4FFEE657326D679F6FD38FE7F" authority="Herre" authorityName="Herre" box="[392,678,476,500]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="258" phylum="Chordata" rank="species" species="scapulostigmata">
<emphasis id="DC97A775FFEE657326D679F6FDCCFE7F" box="[392,594,476,500]" italics="true" pageId="7" pageNumber="258">G. scapulostigmata</emphasis>
Herre
</taxonomicName>
(
<bibRefCitation id="8A720696FFEE657325E479F6FCD8FE7F" author="MILLER, P. J." box="[698,838,476,500]" pageId="7" pageNumber="258" refId="ref13689" refString="MILLER, P. J., 1981, The gobiid genus Gnatholepis, its supposed congeners and its new-found relatives, Abstracts of the 1 st International Conference on the Systematics and Evolution of Indo ± Paci W c Fishes. (Sydney: Australian Museum)." type="proceedings paper" year="1981">Miller, 1981</bibRefCitation>
and unpublished) indicates close similarity in possession of (i) a pronounced scapular spot, more or less ocelliform; (ii) anterior fork of suborbital dark bar faint or absent, and (iii) more or less evident longitudinal streaks on body, with thinner lines. The two are separable only by (i) modal P ray count, 16 (16±18) for
<taxonomicName id="29E300E4FFEE657325A37A73FC2CFDFA" authorityName="Herre" authorityYear="1953" box="[765,946,601,625]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="258" phylum="Chordata" rank="species" species="scapulostigma">
<emphasis id="DC97A775FFEE657325A37A73FC2CFDFA" box="[765,946,601,625]" italics="true" pageId="7" pageNumber="258">G. scapulostigma</emphasis>
</taxonomicName>
and 17 (16±18) for
<taxonomicName id="29E300E4FFEE657327BA7A52FEEEFD1B" authorityName="Jordan" authorityYear="1904" box="[228,368,632,656]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="258" phylum="Chordata" rank="species" species="thompsoni">
<emphasis id="DC97A775FFEE657327BA7A52FEEEFD1B" box="[228,368,632,656]" italics="true" pageId="7" pageNumber="258">G. thompsoni</emphasis>
</taxonomicName>
; and in coloration, with (ii) longitudinal streaks less evident
</paragraph>
<paragraph id="EE5C7B67FFEE657327027ABDFF3AFCE0" blockId="7.[92,1047,132,1686]" pageId="7" pageNumber="258">
along body, and (iii) thinner suborbital bar, in
<taxonomicName id="29E300E4FFEE657325227ABDFD73FD24" authorityName="Jordan" authorityYear="1904" box="[636,749,663,687]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="258" phylum="Chordata" rank="species" species="thompsoni">
<emphasis id="DC97A775FFEE657325227ABDFD73FD24" box="[636,749,663,687]" italics="true" pageId="7" pageNumber="258">thompsoni</emphasis>
</taxonomicName>
. In meristics, as indicated by the somewhat limited material, the eastern Atlantic material from the Cape Verdes and
<collectingCountry id="96F43BF7FFEE657327B47AFCFEA5FD65" box="[234,315,726,750]" name="Ghana" pageId="7" pageNumber="258">Ghana</collectingCountry>
, as well as that from
<collectingCountry id="96F43BF7FFEE657325167AFCFD23FD65" box="[584,701,726,750]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="7" pageNumber="258">Ascension</collectingCountry>
, corresponds in range to the larger samples of western Atlantic
<taxonomicName id="29E300E4FFEE657326A97ADFFD16FC86" authorityName="Jordan" authorityYear="1904" box="[503,648,757,781]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="258" phylum="Chordata" rank="species" species="thompsoni">
<emphasis id="DC97A775FFEE657326A97ADFFD16FC86" box="[503,648,757,781]" italics="true" pageId="7" pageNumber="258">G. thompsoni</emphasis>
</taxonomicName>
(table 2), but tend to have more frequent occurrence of 18 rather than 17 pectoral rays. Variants with D2 I /10 rather than D2 I/11 may be less frequent in the western Atlantic (table 2; and Cervignon, 1994).
</paragraph>
<paragraph id="EE5C7B67FFEE657327DA7B58FF24FBCD" blockId="7.[92,1047,132,1686]" pageId="7" pageNumber="258">
<emphasis id="DC97A775FFEE657327DA7B58FED2FC01" box="[132,332,882,906]" italics="true" pageId="7" pageNumber="258">General description</emphasis>
. See features noted under
<emphasis id="DC97A775FFEE657325C87B58FD75FC01" box="[662,747,882,906]" italics="true" pageId="7" pageNumber="258">Generic</emphasis>
and
<emphasis id="DC97A775FFEE6573247D7B58FCC1FC01" box="[803,863,882,906]" italics="true" pageId="7" pageNumber="258">Speci</emphasis>
W
<emphasis id="DC97A775FFEE657324337B59FC23FC01" box="[877,957,882,906]" italics="true" pageId="7" pageNumber="258">c identi</emphasis>
W
<emphasis id="DC97A775FFEE657324927B59FB91FC01" box="[972,1039,883,906]" italics="true" pageId="7" pageNumber="258">cation</emphasis>
. Body proportions as table 3. Anterior nostril a short tube, posterior nostril porelike. Angle of jaws below anterior edge of pupil; upper lip width about one quarter of preorbital width. Teeth caniniform, outer rows in both jaws enlarged and well spaced, especially in lower jaw, and two or three inner rows of much smaller teeth; tongue bi®d; branchiostegal membrane attached along entire lateral border of isthmus.
</paragraph>
<paragraph id="EE5C7B67FFEE657327DA7C67FB94FB0F" blockId="7.[92,1047,132,1686]" pageId="7" pageNumber="258">
<emphasis id="DC97A775FFEE657327DA7C67FF2DFBEF" box="[132,179,1101,1124]" italics="true" pageId="7" pageNumber="258">Fins</emphasis>
(see table 2). D1 VI (VI:4); D2 I /11 (10±11); A I /11 (10±11); P 17 (15±18); V I/5 1 I/5; C 17. In larger specimens, D1 IV and V to D2 1, D1 VI to D2
</paragraph>
<paragraph id="EE5C7B67FFEE657327027CA6FCB9FA8B" blockId="7.[92,1047,132,1686]" pageId="7" pageNumber="258">I when depressed; D2 and A rear tips over upper and lower origins of C. Interdorsal space membranous. Pectoral ®n with uppermost rays entirely within membrane. Pelvic disc complete, with deep anterior membrane having villose free edge (®gure 3D). Caudal ®n oblong, with bluntly pointed rear edge.</paragraph>
<paragraph id="EE5C7B67FFEE657327DA7D22FC51F91D" blockId="7.[92,1047,132,1686]" pageId="7" pageNumber="258">
<emphasis id="DC97A775FFEE657327DA7D22FF54FAAB" box="[132,202,1288,1312]" italics="true" pageId="7" pageNumber="258">Scales</emphasis>
. Body with large ctenoid scales,
<quantity id="291BD682FFEE6573253F7D22FD59FAAB" box="[609,711,1288,1312]" metricMagnitude="-1" metricUnit="m" metricValue="7.747" metricValueMax="7.874" metricValueMin="7.62" pageId="7" pageNumber="258" unit="in" value="30.5" valueMax="31.0" valueMin="30.0">30±31 in</quantity>
longitudinal series (table 2), nine±ten in transverse series; predorsal scales nine or ten in median series, breast six or seven in median series; scales at origin of C without enlarged lateral cteni. Nape, opercle, cheek to infraorbital row
<emphasis id="DC97A775FFEE657326B87D4FFE6DFAF6" box="[486,499,1381,1405]" italics="true" pageId="7" pageNumber="258">2</emphasis>
, and P base covered with large scales.
<emphasis id="DC97A775FFEE657327DA7DAFFF63FA16" box="[132,253,1413,1437]" italics="true" pageId="7" pageNumber="258">Coloration.</emphasis>
Preserved (®gure 2A): head and body pale brown, latter with several broad vertical dark bars, indistinct ®ne longitudinal striae, and most scales with dark mark at focus; intense dark blotch above upper origin of P typically with dense white emargination of upper border. Head with broad dark bar from eye to rear of jaw angle and extending ventrally across lower edge of cheek to branchiostegal membrane; interorbit with narrow dark transverse line; underside of head and breast densely stippled. D1 and D2 with several oblique rows of small dark marks; A uniformly dusky, lacking dark spots; C with scattered small dark marks; P with narrow horizontal dark streak across middle of base; V dark, with pale edge.
</paragraph>
<caption id="BA9C2BEFFFE1657C270278ABFC8EFE8A" pageId="8" pageNumber="259" startId="8.[92,149,129,151]">
<paragraph id="EE5C7B67FFE1657C270278ABFC8EFE8A" blockId="8.[92,1046,129,257]" pageId="8" pageNumber="259">
Table 2. Meristic variation in
<taxonomicName id="29E300E4FFE1657C26F478ABFD12FF1C" authorityName="Jordan" authorityYear="1904" box="[426,652,129,151]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="259" phylum="Chordata" rank="species" species="thompsoni">
<emphasis id="DC97A775FFE1657C26F478ABFD12FF1C" box="[426,652,129,151]" italics="true" pageId="8" pageNumber="259">Gnatholepis thompsoni</emphasis>
</taxonomicName>
. Counts of second dorsal (D2) and pectoral (P) rays, and scales in lateral (LL) and predorsal (PD) series, in samples from Western Atlantic (Bahamas, BA; Grenadines, GR; Anguilla, AN), Ascension, AS, Eastern Atlantic (Cape Verdes, CV; Ghana, GH);
<emphasis id="DC97A775FFE1657C25B878FBFD6CFF6D" box="[742,754,209,230]" italics="true" pageId="8" pageNumber="259">n</emphasis>
, number of observations;
<emphasis id="DC97A775FFE1657C235D78FBFB90FF6D" box="[1027,1038,208,230]" italics="true" pageId="8" pageNumber="259">xÅ</emphasis>
, mean;
<emphasis id="DC97A775FFE1657C27AF78C1FE8CFE8A" box="[241,274,235,257]" italics="true" pageId="8" pageNumber="259">SD</emphasis>
, standard deviation;
<emphasis id="DC97A775FFE1657C26B178C1FD90FE8A" box="[495,526,235,257]" italics="true" pageId="8" pageNumber="259">SE</emphasis>
, standard error of mean.
</paragraph>
</caption>
<paragraph id="EE5C7B67FFE1657C2702790BFD46FEBC" blockId="8.[92,730,288,470]" box="[92,728,288,311]" pageId="8" pageNumber="259">
D
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2 10 11
<emphasis id="DC97A775FFE1657C26F2790BFE26FEBD" box="[428,440,289,310]" italics="true" pageId="8" pageNumber="259">n</emphasis>
</geoCoordinate>
xÅ SD SE
</paragraph>
<paragraph id="EE5C7B67FFE1657C27027911FD51FEDA" blockId="8.[92,730,288,470]" box="[92,719,315,337]" pageId="8" pageNumber="259">BA ± 8811 ±±</paragraph>
<paragraph id="EE5C7B67FFE1657C2702797CFD44FEE7" blockId="8.[92,730,288,470]" box="[92,730,342,364]" pageId="8" pageNumber="259">GR 2 30 32 10.9 0.24 0.04</paragraph>
<paragraph id="EE5C7B67FFE1657C2702795AFD51FE0D" blockId="8.[92,730,288,470]" box="[92,719,368,390]" pageId="8" pageNumber="259">AN ± 7711 ±±</paragraph>
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<collectingCountry id="96F43BF7FFE1657C270279A1FFE5FE2A" box="[92,123,395,417]" name="American Samoa" pageId="8" pageNumber="259">AS</collectingCountry>
1 10 11 10.9 0.29 0.09
</paragraph>
<paragraph id="EE5C7B67FFE1657C2702798FFD44FE30" blockId="8.[92,730,288,470]" box="[92,730,421,443]" pageId="8" pageNumber="259">
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<specimenCode id="BE45D31CFFE1657C2702798FFE81FE30" box="[92,287,421,443]" collectionCode="CV" country="China" name="Municipal Museum of Chungking" pageId="8" pageNumber="259">CV 1</specimenCode>
</materialsCitation>
4 5 10.8 0.40 0.19
</paragraph>
<paragraph id="EE5C7B67FFE1657C270279EAFD51FE5D" blockId="8.[92,730,288,470]" box="[92,719,448,470]" pageId="8" pageNumber="259">GH ± 2211 ±±</paragraph>
<paragraph id="EE5C7B67FFE1657C270279C2FD46FE75" blockId="8.[92,731,487,669]" box="[92,728,487,510]" pageId="8" pageNumber="259">
A
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10 11
<emphasis id="DC97A775FFE1657C26F279C3FE26FE75" box="[428,440,489,510]" italics="true" pageId="8" pageNumber="259">n</emphasis>
</geoCoordinate>
xÅ SD SE
</paragraph>
<paragraph id="EE5C7B67FFE1657C27027A28FD44FD93" blockId="8.[92,731,487,669]" box="[92,730,514,536]" pageId="8" pageNumber="259">BA 1 7 8 10.9 0.33 0.12</paragraph>
<paragraph id="EE5C7B67FFE1657C27027A37FD44FDB8" blockId="8.[92,731,487,669]" box="[92,730,541,563]" pageId="8" pageNumber="259">GR 2 29 31 10.9 0.25 0.05</paragraph>
<paragraph id="EE5C7B67FFE1657C27027A12FD51FDC6" blockId="8.[92,731,487,669]" box="[92,719,568,590]" pageId="8" pageNumber="259">AN ± 5511 ±±</paragraph>
<paragraph id="EE5C7B67FFE1657C27027A78FD45FDE3" blockId="8.[92,731,487,669]" box="[92,731,594,616]" pageId="8" pageNumber="259">
<collectingCountry id="96F43BF7FFE1657C27027A78FFE5FDE3" box="[92,123,594,616]" name="American Samoa" pageId="8" pageNumber="259">AS</collectingCountry>
± 111111 ±±
</paragraph>
<paragraph id="EE5C7B67FFE1657C27027A47FD44FD08" blockId="8.[92,731,487,669]" box="[92,730,621,643]" pageId="8" pageNumber="259">
<materialsCitation id="5E8B713AFFE1657C27027A47FE81FD08" box="[92,287,621,643]" collectingCountry="As" collectionCode="CV" country="As" pageId="8" pageNumber="259" specimenCode="CV 1" specimenCount="1">
<specimenCode id="BE45D31CFFE1657C27027A47FE81FD08" box="[92,287,621,643]" collectionCode="CV" country="China" name="Municipal Museum of Chungking" pageId="8" pageNumber="259">CV 1</specimenCode>
</materialsCitation>
4 5 10.8 0.40 0.19
</paragraph>
<paragraph id="EE5C7B67FFE1657C27027AADFD51FD16" blockId="8.[92,731,487,669]" box="[92,719,647,669]" pageId="8" pageNumber="259">GH ± 2211 ±±</paragraph>
<paragraph id="EE5C7B67FFE1657C27027A9AFC06FCEE" blockId="8.[92,920,687,869]" pageId="8" pageNumber="259">
P 15
<materialsCitation id="5E8B713AFFE1657C26087A85FC06FCEE" collectionCode="CV" country="American Samoa" latitude="16.288332" location="AS" pageId="8" pageNumber="259" specimenCode="CV 1" specimenCount="1">
<geoCoordinate id="8BD71DA0FFE1657C26087A85FDF1FD4E" box="[342,623,687,709]" degrees="16" direction="north" minutes="17" orientation="latitude" pageId="8" pageNumber="259" precision="15" seconds="18" value="16.288332">
16 17 18
<emphasis id="DC97A775FFE1657C253D7A9AFDF1FD4E" box="[611,623,688,709]" italics="true" pageId="8" pageNumber="259">n</emphasis>
</geoCoordinate>
xÅ SD SE BA ± 4 10 2 16 16.9 0.60 0.15 GR ± 7 43 5 55 17.0 0.47 0.06 AN ± 5 41 6 52 17.0 0.46 0.06
<collectingCountry id="96F43BF7FFE1657C27027B30FFE5FCBB" box="[92,123,794,816]" name="American Samoa" pageId="8" pageNumber="259">AS</collectingCountry>
± 1 8 7 16 17.4 0.60 0.15
<specimenCode id="BE45D31CFFE1657C27027B1EFE81FCC1" box="[92,287,820,842]" collectionCode="CV" country="China" name="Municipal Museum of Chungking" pageId="8" pageNumber="259">CV 1</specimenCode>
± 4 4 9 17.2 0.92 0.31 GH ± ± 1 3 4 17.8 0.43 0.22
</materialsCitation>
</paragraph>
<paragraph id="EE5C7B67FFE1657C27027B5DFC73FB9A" blockId="8.[92,1016,886,1041]" pageId="8" pageNumber="259">
<materialsCitation id="5E8B713AFFE1657C27027B5DFCA6FB9A" collectionCode="CV" country="American Samoa" latitude="29.508612" location="AS" pageId="8" pageNumber="259" specimenCount="1">
LL 27 28
<geoCoordinate id="8BD71DA0FFE1657C26F87B5CFD51FC07" box="[422,719,886,908]" degrees="29" direction="north" minutes="30" orientation="latitude" pageId="8" pageNumber="259" precision="15" seconds="31" value="29.508612">
29 30 31
<emphasis id="DC97A775FFE1657C259D7B5DFD51FC07" box="[707,719,887,908]" italics="true" pageId="8" pageNumber="259">n</emphasis>
</geoCoordinate>
xÅ SD SE BA ± ± 2 5 1 8 29.9 0.60 0.21 AN ± 2 11 15 5 33 29.7 0.80 0.14
<collectingCountry id="96F43BF7FFE1657C27027BECFFE5FC57" box="[92,123,966,988]" name="American Samoa" pageId="8" pageNumber="259">AS</collectingCountry>
1 1 2 2 3 9 29.6 1.34 0.45
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± ± ± 3 3 6 30.5 0.50 0.20 GH ± 4 ± ± ± 4 28.0
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± ±
</paragraph>
<paragraph id="EE5C7B67FFE1657C27DA7C4AFCFDFBF3" blockId="8.[92,1041,1120,1686]" box="[132,867,1120,1145]" pageId="8" pageNumber="259">
<emphasis id="DC97A775FFE1657C27DA7C4AFEDBFBF3" box="[132,325,1120,1144]" italics="true" pageId="8" pageNumber="259">Lateral-line system</emphasis>
. As
<emphasis id="DC97A775FFE1657C26FC7C4AFDA9FBF3" box="[418,567,1120,1144]" italics="true" pageId="8" pageNumber="259">Generic identi</emphasis>
W
<emphasis id="DC97A775FFE1657C25127C4BFD0EFBF3" box="[588,656,1121,1144]" italics="true" pageId="8" pageNumber="259">cation</emphasis>
and ®gures 3A±C.
</paragraph>
<paragraph id="EE5C7B67FFE1657C27DA7CAAFB97F9BC" blockId="8.[92,1041,1120,1686]" pageId="8" pageNumber="259">
<emphasis id="DC97A775FFE1657C27DA7CAAFF47FB13" box="[132,217,1152,1176]" italics="true" pageId="8" pageNumber="259">Biology</emphasis>
. Distribution. Present throughout the Cape Verdes archipelago. In the eastern Atlantic, recorded from
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and from the islands of
<collectingCountry id="96F43BF7FFE1657C24127C8AFC20FB33" box="[844,958,1184,1208]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="8" pageNumber="259">Ascension</collectingCountry>
and St Helena (
<bibRefCitation id="8A720696FFE1657C279C7CEAFE70FB53" author="EDWARDS, A. J. &amp; GLASS, C. W." box="[194,494,1216,1240]" pageId="8" pageNumber="259" pagination="617" refId="ref13203" refString="EDWARDS, A. J. and GLASS, C. W., 1987, The ® shes of Saint Helena Island, South Atlantic Ocean. I. The shore ® shes, Journal of Natural History, 21, 617 ± 686." type="journal article" year="1987">Edwards and Glass, 1987</bibRefCitation>
;
<bibRefCitation id="8A720696FFE1657C26A27CEAFD17FB53" author="MILLER, P. J." box="[508,649,1216,1240]" pageId="8" pageNumber="259" pagination="951" refId="ref13836" refString="MILLER, P. J., 1990, Gobiidae, in J. C. Quero, J. C. Hureau, C. Karrer, A. Post and L. Saldanha (eds), Check-list of the Fishes of the Eastern Tropical Atlantic. (Lisbon: JNICT, EIU, UNESCO), 2, 925 ± 951." type="journal article" year="1990">Miller, 1990</bibRefCitation>
), and recently at localities on the east and south-east coast of Tenerife, Canary Islands (AB, personal observation). In the western Atlantic, it has been found from
<collectingCountry id="96F43BF7FFE1657C25DD7D2AFD74FA93" box="[643,746,1280,1304]" name="Bermuda" pageId="8" pageNumber="259">Bermuda</collectingCountry>
and Florida to the Lesser Antilles and coast of central America (
<bibRefCitation id="8A720696FFE1657C25517D35FD22FABC" author="CERVIGON, F." box="[527,700,1311,1335]" pageId="8" pageNumber="259" refId="ref13043" refString="CERVIGON, F., 1994, Los Peces Marinos de Venezuela. 2 nd ed. Vol. III. (Caracas: FundacioAn Cienti ® ca Los Roques), 295 pp." type="book" year="1994">Cervigon, 1994</bibRefCitation>
). Habitat. At the Cape Verdes the most common sublittoral goby, found from shore to
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on sand and stones. A similar habitat on sand near rocks and rubble at
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to at least
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is described by
<bibRefCitation id="8A720696FFE1657C27DC7D55FE27FA1C" author="EDWARDS, A. J. &amp; GLASS, C. W." box="[130,441,1407,1431]" pageId="8" pageNumber="259" pagination="617" refId="ref13203" refString="EDWARDS, A. J. and GLASS, C. W., 1987, The ® shes of Saint Helena Island, South Atlantic Ocean. I. The shore ® shes, Journal of Natural History, 21, 617 ± 686." type="journal article" year="1987">Edwards and Glass (1987)</bibRefCitation>
for St Helena. Tenerife specimens have been found at
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on stones and rock with sand. In the western Atlantic, at
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, this species has been reported as abundant, occurring from shore to
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, especially at less than
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, on substrata of algae-covered rocks, coral, rubble and sand, excavating burrows in sand (
<bibRefCitation id="8A720696FFE1657C26EF7DD5FDC9F99C" author="RANDALL, J." box="[433,599,1535,1559]" pageId="8" pageNumber="259" refId="ref14201" refString="RANDALL, J., 1968, Caribbean Reef Fishes. (Neptune City, New Jersey: Tropical Fish Hobbyist), 318 pp." type="book" year="1968">Randall, 1968</bibRefCitation>
; Bo Èhlke and Chaplin, 1968;
<bibRefCitation id="8A720696FFE1657C24F67DD5FF0CF9BC" author="CERVIGON, F." pageId="8" pageNumber="259" refId="ref13043" refString="CERVIGON, F., 1994, Los Peces Marinos de Venezuela. 2 nd ed. Vol. III. (Caracas: FundacioAn Cienti ® ca Los Roques), 295 pp." type="book" year="1994">Cervigon, 1994</bibRefCitation>
). In the Western Atlantic,
<taxonomicName id="29E300E4FFE1657C26887E35FDF6F9BC" authorityName="Jordan" authorityYear="1904" box="[470,616,1567,1591]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="259" phylum="Chordata" rank="species" species="thompsoni">
<emphasis id="DC97A775FFE1657C26887E35FDF6F9BC" box="[470,616,1567,1591]" italics="true" pageId="8" pageNumber="259">G. thompsoni</emphasis>
</taxonomicName>
is often associated with the Bridled
</paragraph>
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Goby,
<taxonomicName id="29E300E4FFE1657C27F17E15FE73F9DC" authorityName="Gill" authorityYear="1863" box="[175,493,1599,1623]" class="Actinopterygii" family="Gobiidae" genus="Coryphopterus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="259" phylum="Chordata" rank="species" species="glaucofraenum">
<emphasis id="DC97A775FFE1657C27F17E15FE73F9DC" box="[175,493,1599,1623]" italics="true" pageId="8" pageNumber="259">Coryphopterus glaucofraenum</emphasis>
</taxonomicName>
and
<taxonomicName id="29E300E4FFE1657C25617E15FC3EF9DC" authority="(Cervignon, 1994)" baseAuthorityName="Cervignon" baseAuthorityYear="1994" box="[575,928,1599,1623]" class="Actinopterygii" family="Gobiidae" genus="Coryphopterus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="259" phylum="Chordata" rank="species" species="tortugae">
<emphasis id="DC97A775FFE1657C25617E15FD5CF9DC" box="[575,706,1599,1623]" italics="true" pageId="8" pageNumber="259">C. tortugae</emphasis>
(Cervignon, 1994)
</taxonomicName>
, although neither of the latter has yet been found in the eastern Atlantic. Feeding appears to involve sorting of sand in the buccal cavity and ejection of the latter from the gill openings (
<bibRefCitation id="8A720696FFE0657D278A7CFAFD9FFB63" author="BOHLKE, J. E. &amp; CHAPLIN, C. G. C." box="[212,513,1232,1256]" pageId="9" pageNumber="260" refId="ref12925" refString="BOHLKE, J. E. and CHAPLIN, C. G. C., 1968, Fishes of the Bahamas and Adjacent Tropical Waters. (Wynnewood, PA.: Livingston), 771 pp." type="book" year="1968">Bohlke and Chaplin, 1968</bibRefCitation>
). Size. The largest Cape Verdean example is a male, 61 1
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; with maximum values of 58 and
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for standard and total length noted by
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.
</paragraph>
</subSubSection>
</treatment>
</document>
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