156 lines
18 KiB
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156 lines
18 KiB
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<document id="FFB3E0D4277693A9F1A6EEAF7316B630" ID-DOI="10.5281/zenodo.281856" ID-GBIF-Dataset="b174c7dc-9bd8-4a26-854b-33e5a6a9ee46" ID-ISSN="1175-5326" ID-Zenodo-Dep="281856" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460502285788" checkinUser="plazi" docAuthor="Parey, Sajad H. & Saini, Malkiat S." docDate="2012" docId="03EAF641B261FFD5FF40F999FB27FA97" docLanguage="en" docName="zt03403p068.pdf" docOrigin="Zootaxa 3403" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Eubasilissa asiatica Betten 1909" docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="FFD38E39B265FFD1FFD7F925FF84FF9D" masterDocTitle="Two new species of the genus Eubasilissa Martynov (Trichoptera: Phryganeidae) from the Indian Himalaya" masterLastPageNumber="68" masterPageNumber="61" pageNumber="65" updateTime="1698312062744" updateUser="plazi">
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<mods:title id="1810AFE5995A2182D701AF7859EF7F7C">Two new species of the genus Eubasilissa Martynov (Trichoptera: Phryganeidae) from the Indian Himalaya</mods:title>
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<mods:namePart id="E8B0C98EA3D30DF71409DF4EA8CE1E8E">Parey, Sajad H.</mods:namePart>
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<mods:roleTerm id="3949105C3784628949D877F164F76091">Author</mods:roleTerm>
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<mods:namePart id="0767735965F427168209F5A2D2BDA857">Saini, Malkiat S.</mods:namePart>
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<mods:typeOfResource id="0C7A17220EAA209F966121FFA64EA6FA">text</mods:typeOfResource>
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<mods:title id="CDD7185261DB5C9582147C1957957B73">Zootaxa</mods:title>
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<mods:part id="7ABC06F761BD5A63880FC99C544A574C">
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<mods:date id="9592E253D943420A522710448BFCFDB3">2012</mods:date>
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<mods:detail id="96D4838B5624ED41E49039BF487D4EFB" type="volume">
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<mods:number id="6D22A71117FDBECBA9D6397BB7BFAC23">3403</mods:number>
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<mods:start id="B44A95D0DE7A8366C8FBDE3B1FF7339A">61</mods:start>
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<mods:classification id="E6C962959C9158D6300DFD9BB63D1E94">journal article</mods:classification>
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<mods:identifier id="72DAF772239B86320A60EC43B690A666" type="DOI">10.5281/zenodo.281856</mods:identifier>
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<treatment id="03EAF641B261FFD5FF40F999FB27FA97" ID-DOI="http://doi.org/10.5281/zenodo.6170137" ID-GBIF-Taxon="119521535" ID-Zenodo-Dep="6170137" LSID="urn:lsid:plazi:treatment:03EAF641B261FFD5FF40F999FB27FA97" httpUri="http://treatment.plazi.org/id/03EAF641B261FFD5FF40F999FB27FA97" lastPageNumber="65" pageId="4" pageNumber="65">
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<subSubSection id="C35914DCB261FFD5FF40F999FE22FC8F" pageId="4" pageNumber="65" type="nomenclature">
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<paragraph id="8BFC4757B261FFD5FF40F999FDE0FF48" blockId="4.[151,612,187,246]" box="[151,612,187,214]" pageId="4" pageNumber="65">
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<heading id="D0B4F03BB261FFD5FF40F999FDE0FF48" bold="true" box="[151,612,187,214]" fontSize="11" level="1" pageId="4" pageNumber="65" reason="1">
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<treatmentCitationGroup id="AB536079B261FFD5FF40F999FDE0FF48" box="[151,612,187,214]" pageId="4" pageNumber="65">
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<taxonomicName id="4C433CD4B261FFD5FF40F999FDE0FF48" ID-CoL="3BNCZ" authority="Betten, 1909: 242" authorityName="Betten" authorityPageNumber="242" authorityYear="1909" box="[151,612,187,214]" class="Insecta" family="Phryganeidae" genus="Eubasilissa" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="asiatica">
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<emphasis id="B9379B45B261FFD5FF40F999FDE0FF48" bold="true" box="[151,612,187,214]" pageId="4" pageNumber="65">
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<emphasis id="B9379B45B261FFD5FF40F999FE02FF4B" bold="true" box="[151,390,188,214]" italics="true" pageId="4" pageNumber="65">Eubasilissa asiatica</emphasis>
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<treatmentCitation id="0AE26146B261FFD5FE5BF99EFDE0FF48" author="Betten" box="[396,612,187,213]" page="242" pageId="4" pageNumber="65" year="1909">
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<bibRefCitation id="EFD23AA6B261FFD5FE5BF99EFDAFFF48" author="Betten" box="[396,555,187,213]" pageId="4" pageNumber="65" refString="Betten, C. (1909) Notes on the Trichoptera in the Indian Museum. Records of the Indian Museum, 3, 231 - 242." type="journal article" year="1909">Betten, 1909</bibRefCitation>
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: 242
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</treatmentCitation>
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</emphasis>
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</taxonomicName>
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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<paragraph id="8BFC4757B261FFD5FF40F9FBFEDCFF6B" blockId="4.[151,612,187,246]" box="[151,344,222,246]" pageId="4" pageNumber="65">
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(
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<figureCitation id="13785BD2B261FFD5FF48F9FBFEA3FF6B" box="[159,295,222,246]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1921,1944]" captionTargetBox="[224,1374,220,1854]" captionTargetId="figure@5.[199,1385,193,1866]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Eubasilissa asiatica Betten. 15 – 18, Ə genitalia: 15, left lateral; 16, dorsal; 17, phallic apparatus, left lateral; 18, caudal. 19 – 20, Ƥ genitalia: 19, left lateral; 20, dorsal. 21 – 22, wings: 21, Ə right fore- & hind wings; 22, Ƥ right fore- & hind wings." httpUri="https://zenodo.org/record/281859/files/figure.png" pageId="4" pageNumber="65">Figs. 15–22</figureCitation>
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,
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<figureCitation id="13785BD2B261FFD5FEE5F9FBFECBFF6B" box="[306,335,222,246]" captionStart="FIGURES 23 – 25" captionStartId="6.[151,275,1964,1989]" captionTargetBox="[453,1156,180,1932]" captionTargetId="figure@6.[427,1179,161,1947]" captionTargetPageId="6" captionText="FIGURES 23 – 25. Collection sites of Eubasilissa spp. 23, E. sikkimensis sp. nov.; 24, E. schmidi sp. nov.; 25, E. asiatica Betten." httpUri="https://zenodo.org/record/281860/files/figure.png" pageId="4" pageNumber="65">25</figureCitation>
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)
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</paragraph>
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<paragraph id="8BFC4757B261FFD5FF40F803FEA6FEFF" blockId="4.[151,1437,294,1291]" pageId="4" pageNumber="65">
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Head and thorax brownish. Labrum brown, labial palpi yellowish. Colour pattern of male and female wings as in
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<figureCitation id="13785BD2B261FFD5FF40F86FFE9AFEFF" box="[151,286,330,354]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1921,1944]" captionTargetBox="[224,1374,220,1854]" captionTargetId="figure@5.[199,1385,193,1866]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Eubasilissa asiatica Betten. 15 – 18, Ə genitalia: 15, left lateral; 16, dorsal; 17, phallic apparatus, left lateral; 18, caudal. 19 – 20, Ƥ genitalia: 19, left lateral; 20, dorsal. 21 – 22, wings: 21, Ə right fore- & hind wings; 22, Ƥ right fore- & hind wings." httpUri="https://zenodo.org/record/281859/files/figure.png" pageId="4" pageNumber="65">Figs. 21–22</figureCitation>
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.
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</paragraph>
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<paragraph id="8BFC4757B261FFD5FF1EF84BFAA0FDA7" blockId="4.[151,1437,294,1291]" pageId="4" pageNumber="65">
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Male genitalia (
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<figureCitation id="13785BD2B261FFD5FE56F84BFD8BFE1B" box="[385,527,366,390]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1921,1944]" captionTargetBox="[224,1374,220,1854]" captionTargetId="figure@5.[199,1385,193,1866]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Eubasilissa asiatica Betten. 15 – 18, Ə genitalia: 15, left lateral; 16, dorsal; 17, phallic apparatus, left lateral; 18, caudal. 19 – 20, Ƥ genitalia: 19, left lateral; 20, dorsal. 21 – 22, wings: 21, Ə right fore- & hind wings; 22, Ƥ right fore- & hind wings." httpUri="https://zenodo.org/record/281859/files/figure.png" pageId="4" pageNumber="65">Figs. 15–18</figureCitation>
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): Segment IX subrectangular in lateral view; U-shaped, narrow at centre, and broadened near sides in dorsal view; posteroventral portion upturned and concave, produced into slightly dentate lobe. Segment X subtriangular in dorsal view, apically pointed. Preanal appendages reduced to pair of basolateral protrusions on segment X. Inferior appendages each elongate, subrectangular in lateral view, truncate apically, with subapicoventral notch with basal edge pointed, curved inwards. Phallotheca with slender, sclerotized posteroventral projection curved ventrad; membranous endotheca with pair of slender, sinuous sclerites.
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</paragraph>
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<paragraph id="8BFC4757B261FFD5FF1EFB63FE22FC8F" blockId="4.[151,1437,294,1291]" pageId="4" pageNumber="65">
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Female genitalia (
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<figureCitation id="13785BD2B261FFD5FE74FB63FDB2FDC3" box="[419,566,582,606]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1921,1944]" captionTargetBox="[224,1374,220,1854]" captionTargetId="figure@5.[199,1385,193,1866]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Eubasilissa asiatica Betten. 15 – 18, Ə genitalia: 15, left lateral; 16, dorsal; 17, phallic apparatus, left lateral; 18, caudal. 19 – 20, Ƥ genitalia: 19, left lateral; 20, dorsal. 21 – 22, wings: 21, Ə right fore- & hind wings; 22, Ƥ right fore- & hind wings." httpUri="https://zenodo.org/record/281859/files/figure.png" pageId="4" pageNumber="65">Figs. 19–20</figureCitation>
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): Subgenital plate with median lobe deeply divided, both ends closely approximate apically with small elliptical, median gap between them; lateral lobes narrow, triangular; median lobe appearing spoon-shaped, and lateral lobes slender in lateral view. Segment IX apicolaterally with club-shaped lobe. Vaginal sclerites highly modified, main sclerite extending into segment VIII, broad and with small opening near anterior end; pair of triangular submedial sclerites; and long transverse sclerite with rounded ends. Vaginal pouches long with rounded ends.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35914DCB261FFD5FF1EFA3BFE09FC3F" pageId="4" pageNumber="65" type="materials_examined">
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<paragraph id="8BFC4757B261FFD5FF1EFA3BFE09FC3F" blockId="4.[151,1437,294,1291]" pageId="4" pageNumber="65">
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<emphasis id="B9379B45B261FFD5FF1EFA3BFE36FCAA" bold="true" box="[201,434,798,823]" pageId="4" pageNumber="65">Material examined.</emphasis>
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<collectingCountry id="F35407C7B261FFD5FE6EFA3AFD89FCAA" box="[441,525,799,823]" name="India" pageId="4" pageNumber="65">INDIA</collectingCountry>
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: Jammu & Kashmir: Gurez,
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<quantity id="4CBBEAB2B261FFD5FC9AFA3BFC27FCAB" box="[845,931,798,822]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="65" unit="m" value="2400.0">2400 m</quantity>
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,
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<date id="FFFD6197B261FFD5FC78FA3BFBAAFCAB" box="[943,1070,798,822]" pageId="4" pageNumber="65" value="2010-09-05">05-ix-2010</date>
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, collectors Parey & Pandher, 13 1Ƥ; Apharwat,
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<quantity id="4CBBEAB2B261FFD5FE92FA67FE19FCC7" box="[325,413,834,858]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="65" unit="m" value="4000.0">4000 m</quantity>
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,
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<date id="FFFD6197B261FFD5FE7EFA67FDABFCC7" box="[425,559,834,858]" pageId="4" pageNumber="65" value="2010-07-30">30-vii-2010</date>
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, collector Rifat H Raina, 13.
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<collectingCountry id="F35407C7B261FFD5FCA8FA66FC57FCC6" box="[895,979,835,859]" name="India" pageId="4" pageNumber="65">INDIA</collectingCountry>
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: Himachal Pradesh: Punjpullah,
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<quantity id="4CBBEAB2B261FFD5FA97FA67FA13FCC7" box="[1344,1431,834,858]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="65" unit="m" value="1600.0">1600 m</quantity>
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,
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<date id="FFFD6197B261FFD5FF40FA43FE98FCE3" box="[151,284,870,894]" pageId="4" pageNumber="65" value="2010-07-12">12-07-2010</date>
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, collectors Parey, Muzamila A Dar & Vikram S Rathor, 13 1Ƥ; Kothi,
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<quantity id="4CBBEAB2B261FFD5FBEDFA43FB15FCE3" box="[1082,1169,870,894]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="65" unit="m" value="2300.0">2300 m</quantity>
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,
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<date id="FFFD6197B261FFD5FB4AFA43FAA0FCE3" box="[1181,1316,870,894]" pageId="4" pageNumber="65" value="2009-07-06">06-vii-2009</date>
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, collectors Parey & Pandher,
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<specimenCount id="9D458CDEB261FFD5FEB4FAAFFE09FC3F" box="[355,397,906,930]" pageId="4" pageNumber="65" type="generic">1Ƥ.</specimenCount>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35914DCB261FFD5FF1EFA8BFBF2FC5B" box="[201,1142,942,967]" pageId="4" pageNumber="65" type="distribution">
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<paragraph id="8BFC4757B261FFD5FF1EFA8BFBF2FC5B" blockId="4.[151,1437,294,1291]" box="[201,1142,942,967]" pageId="4" pageNumber="65">
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<emphasis id="B9379B45B261FFD5FF1EFA8BFEE6FC5A" bold="true" box="[201,354,942,967]" pageId="4" pageNumber="65">Distribution.</emphasis>
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<collectingCountry id="F35407C7B261FFD5FEBEFA8BFE71FC5B" box="[361,501,942,966]" name="Afghanistan" pageId="4" pageNumber="65">Afghanistan</collectingCountry>
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,
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<collectingCountry id="F35407C7B261FFD5FDD7FA8BFDE6FC5B" box="[512,610,942,966]" name="Pakistan" pageId="4" pageNumber="65">Pakistan</collectingCountry>
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,
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<collectingCountry id="F35407C7B261FFD5FDBAFA8BFD23FC5B" box="[621,679,942,966]" name="India" pageId="4" pageNumber="65">India</collectingCountry>
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(Jammu & Kashmir, Himachal Pradesh).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35914DCB261FFD5FF1EFAF7FB27FA97" pageId="4" pageNumber="65" type="discussion">
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<paragraph id="8BFC4757B261FFD5FF1EFAF7FB27FA97" blockId="4.[151,1437,294,1291]" pageId="4" pageNumber="65">
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<emphasis id="B9379B45B261FFD5FF1EFAF7FED5FC76" bold="true" box="[201,337,978,1003]" pageId="4" pageNumber="65">Comments.</emphasis>
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This species was described by
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<bibRefCitation id="EFD23AA6B261FFD5FD69FAF6FCDFFC77" author="Betten" box="[702,859,978,1002]" pageId="4" pageNumber="65" refString="Betten, C. (1909) Notes on the Trichoptera in the Indian Museum. Records of the Indian Museum, 3, 231 - 242." type="journal article" year="1909">Betten (1909)</bibRefCitation>
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from Sibsagar, Assam.
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<bibRefCitation id="EFD23AA6B261FFD5FBA7FAF7FA9DFC77" author="Schmid" box="[1136,1305,978,1002]" pageId="4" pageNumber="65" refString="Schmid, F. (1962) Le genre Eubasilissa Mart. en Inde. Bulletin de la Societe Vaudoise des Sciences Naturelles, 68 (309), 153 - 68." type="journal article" year="1962">Schmid (1962)</bibRefCitation>
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questioned that
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<taxonomicName id="4C433CD4B261FFD5FF1FFADDFEC4FB93" box="[200,320,1015,1039]" class="Insecta" family="Phryganeidae" genus="Eubasilissa" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="asiatica">
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<emphasis id="B9379B45B261FFD5FF1FFADDFEC4FB93" box="[200,320,1015,1039]" italics="true" pageId="4" pageNumber="65">E. asiatica</emphasis>
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</taxonomicName>
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or any phryganeid would live in the hot climate at the
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<quantity id="4CBBEAB2B261FFD5FC7FFAD3FC78FB93" box="[936,1020,1014,1038]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="4" pageNumber="65" unit="ft" value="1000.0">1000 ft</quantity>
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. elevation of the
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<typeStatus id="54F8F9F5B261FFD5FB6BFAD2FB68FB92" box="[1212,1260,1015,1039]" pageId="4" pageNumber="65">type</typeStatus>
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locality of Sibsagar in Assam, and concluded that the
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<typeStatus id="54F8F9F5B261FFD5FDB4FD3FFD41FBAF" box="[611,709,1050,1074]" pageId="4" pageNumber="65" type="holotype">holotype</typeStatus>
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was labeled incorrectly. Despite intensive collecting in Assam, Schmid could not find
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<taxonomicName id="4C433CD4B261FFD5FE76FD65FD9FFBCB" box="[417,539,1087,1111]" class="Insecta" family="Phryganeidae" genus="Eubasilissa" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="asiatica">
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<emphasis id="B9379B45B261FFD5FE76FD65FD9FFBCB" box="[417,539,1087,1111]" italics="true" pageId="4" pageNumber="65">E. asiatica</emphasis>
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</taxonomicName>
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in this region, but he did collect specimens in
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<collectingCountry id="F35407C7B261FFD5FBEAFD1BFB18FBCB" box="[1085,1180,1086,1110]" name="Pakistan" pageId="4" pageNumber="65">Pakistan</collectingCountry>
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and
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<collectingCountry id="F35407C7B261FFD5FB00FD1BFAE0FBCB" box="[1239,1380,1086,1110]" name="Afghanistan" pageId="4" pageNumber="65">Afghanistan</collectingCountry>
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. The present authors also could not find this species in Assam or its adjoining states Meghalaya, Sikkim, Nagaland, and West Bengal. However, we collected this species from Jammu & Kashmir and Himachal Pradesh. Male genitalia of
|
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<taxonomicName id="4C433CD4B261FFD5FF40FD89FE96FB5E" box="[151,274,1196,1219]" class="Insecta" family="Phryganeidae" genus="Eubasilissa" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="asiatica">
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<emphasis id="B9379B45B261FFD5FF40FD89FE96FB5E" box="[151,274,1196,1219]" italics="true" pageId="4" pageNumber="65">E. asiatica</emphasis>
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</taxonomicName>
|
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collected from Jammu & Kashmir and Himachal Pradesh are slightly different in morphology from those of the specimen described by
|
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<bibRefCitation id="EFD23AA6B261FFD5FD91FDEBFD7BFB7B" author="Wiggins" box="[582,767,1230,1254]" pageId="4" pageNumber="65" refString="Wiggins, G. B. (1998) The caddisfly family Phryganeidae (Trichoptera). University of Toronto Press, Toronto, ix + 306 pp." type="book" year="1998">Wiggins (1998)</bibRefCitation>
|
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|
. However, the phallic apparatus of our specimen almost matches that of
|
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|
<taxonomicName id="4C433CD4B261FFD5FE9EFDD1FE45FA96" box="[329,449,1268,1291]" class="Insecta" family="Phryganeidae" genus="Eubasilissa" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="asiatica">
|
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|
<emphasis id="B9379B45B261FFD5FE9EFDD1FE45FA96" box="[329,449,1268,1291]" italics="true" pageId="4" pageNumber="65">E. asiatica</emphasis>
|
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|
</taxonomicName>
|
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|
, so we consider this specimen to be a variant of the same species.
|
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|
</paragraph>
|
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|
</subSubSection>
|
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</treatment>
|
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</document>
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