treatments-xml/data/AF/37/BE/AF37BEF5BF775D25BA6B2D089170C748.xml

<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.7.90973" ID-Pensoft-Pub="2535-0730-1-51" ID-Pensoft-UUID="ABE1527AFB225DE3B1CDE6FE4E7BB38C" ID-ZooBank="15FBAE73903B4E14BDE3CCC2D65AAD81" ModsDocID="2535-0730-7-1-51" checkinTime="1672842668324" checkinUser="pensoft" docAuthor="Brahma, Shubhranil, Chatterjee, Somnath &amp; Hazra, Niladri" docDate="2023" docId="AF37BEF5BF775D25BA6B2D089170C748" docLanguage="en" docName="EvolutSyst 7(1): 51-66" docOrigin="Evolutionary Systematics 7 (1)" docPubDate="2023-01-03" docSource="http://dx.doi.org/10.3897/evolsyst.7.90973" docTitle="Dasyhelea (Sebessia) folia Brahma &amp; Chatterjee &amp; Hazra 2023, sp. nov." docType="treatment" docUuid="729C8E33-BF5A-43F8-906E-A41FE77F6A31" docUuidSource="ZooBank" docVersion="1" id="ABE1527AFB225DE3B1CDE6FE4E7BB38C" lastPageNumber="51" masterDocId="ABE1527AFB225DE3B1CDE6FE4E7BB38C" masterDocTitle="Five new Indian species of the genus Dasyhelea Kieffer (Diptera, Ceratopogonidae) with a key to the adult males" masterLastPageNumber="66" masterPageNumber="51" pageNumber="51" updateTime="1672842668324" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Five new Indian species of the genus Dasyhelea Kieffer (Diptera, Ceratopogonidae) with a key to the adult males</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brahma, Shubhranil</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5007-9704</mods:nameIdentifier>
<mods:affiliation>Iswar Chandra College, Department of Zoology, Belonia, Tripura, 799155, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chatterjee, Somnath</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8572-1779</mods:nameIdentifier>
<mods:affiliation>Entomology Research Unit, Department of Zoology, The University of Burdwan, Burdwan, 713104, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hazra, Niladri</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8149-9445</mods:nameIdentifier>
<mods:affiliation>Entomology Research Unit, Department of Zoology, The University of Burdwan, Burdwan, 713104, India</mods:affiliation>
<mods:nameIdentifier type="email">nhazra@zoo.buruniv.ac.in</mods:nameIdentifier>
</mods:name>
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<mods:titleInfo>
<mods:title>Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>7</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>51</mods:start>
<mods:end>66</mods:end>
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<mods:url>http://dx.doi.org/10.3897/evolsyst.7.90973</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.7.90973</mods:identifier>
<mods:identifier type="Pensoft-Pub">2535-0730-1-51</mods:identifier>
<mods:identifier type="ZooBank">15FBAE73903B4E14BDE3CCC2D65AAD81</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:729C8E33-BF5A-43F8-906E-A41FE77F6A31" httpUri="http://treatment.plazi.org/id/AF37BEF5BF775D25BA6B2D089170C748" lastPageNumber="51" pageId="0" pageNumber="51">
<subSubSection pageId="0" pageNumber="51" type="nomenclature">
<paragraph pageId="0" pageNumber="51">
<taxonomicName LSID="https://zoobank.org/729C8E33-BF5A-43F8-906E-A41FE77F6A31" authority="Brahma &amp; Chatterjee &amp; Hazra, 2023" authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Sebessia) folia" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="folia" status="sp. nov." subGenus="Sebessia">Dasyhelea (Sebessia) folia</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="51">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="type material">
<paragraph pageId="0" pageNumber="51">Type material.</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Holotype</emphasis>
♂, India, West Bengal, South 24 Parganas, Pakhiralaya [
<geoCoordinate degrees="22" direction="north" minutes="08" orientation="latitude" precision="1" seconds="13.0" value="22.136944">22°08'13.0&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="88" direction="east" minutes="49" orientation="longitude" precision="1" seconds="45.1" value="88.82919">88°49'45.1&quot;E</geoCoordinate>
], 25.V.2018, Coll. S. Brahma.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="diagnosis">
<paragraph pageId="0" pageNumber="51">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="51">
The only species in the subgenus 
<taxonomicName class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Sebessia)" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="subGenus" subGenus="Sebessia">Dasyhelea Sebessia</taxonomicName>
bearing a distinct oval shaped gonocoxite; one long seta near base of gonostylus; broad and leaf like basal arms of parameres; apicolateral process with one long seta at subapical notch.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="description">
<paragraph pageId="0" pageNumber="51">Description.</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Male adult</emphasis>
(n = 1) (Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7A-J</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-E Adult male genitalia structures. A Dasyhelea (Dasyhelea) incisura sp. nov.; B Dasyhelea (Dasyhelea) quasifulcillata sp. nov.; C Dasyhelea (Dasyhelea) trigona sp. nov.; D Dasyhelea (Sebessia) falxa sp. nov.; E Dasyhelea (Sebessia) folia sp. nov. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure8" httpUri="https://binary.pensoft.net/fig/790308" pageId="0" pageNumber="51">8E</figureCitation>
). Total body length 2.1 mm.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Head</emphasis>
. Light brown in colour. Eyes separated by about a diameter of an ommatidium. Frontal sclerite (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7A</figureCitation>
) 55 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 30 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, oval shaped. Flagellum (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7B</figureCitation>
) 0.6 mm long; flagellomere XIII without any prominent apical projection; length ratio of flagellomeres (I-XIII): 23: 13: 12: 12: 12: 13: 13: 13: 13: 30: 22: 20: 34; AR 1.07; flagellomeres X-XIII with sensilla basiconica. Clypeus as in figure 7C, with 3 pairs of setae. Palpus (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7D</figureCitation>
) pale brown; length ratio of palp segments (I-V): 10: 10: 16: 11: 12; PR 2.3; palp segment III with 1 capitate sensillum at its mid length.
</paragraph>
<caption doi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Figure 7.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="51">A-J</emphasis>
Adult male of 
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Sebessia) folia" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="folia" subGenus="Sebessia">Dasyhelea (Sebessia) folia</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="51">A</emphasis>
Frontal sclerite; 
<emphasis bold="true" pageId="0" pageNumber="51">B</emphasis>
Flagellum; 
<emphasis bold="true" pageId="0" pageNumber="51">C</emphasis>
Clypeus; 
<emphasis bold="true" pageId="0" pageNumber="51">D</emphasis>
Palpus; 
<emphasis bold="true" pageId="0" pageNumber="51">E</emphasis>
Thorax; 
<emphasis bold="true" pageId="0" pageNumber="51">F</emphasis>
Femora and tibiae (right to left) of fore, mid and hind legs; 
<emphasis bold="true" pageId="0" pageNumber="51">G</emphasis>
Hind tibial comb; 
<emphasis bold="true" pageId="0" pageNumber="51">H</emphasis>
Genitalia (left dorsal and right ventral view); 
<emphasis bold="true" pageId="0" pageNumber="51">I</emphasis>
Apicolateral process; 
<emphasis bold="true" pageId="0" pageNumber="51">J</emphasis>
Aedeagus. Scale bars: 0.05 mm.
</paragraph>
</caption>
<caption doi="10.3897/evolsyst.7.90973.figure8" httpUri="https://binary.pensoft.net/fig/790308" pageId="0" pageNumber="51" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Figure 8.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="51">A-E</emphasis>
Adult male genitalia structures. 
<emphasis bold="true" pageId="0" pageNumber="51">A</emphasis>
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Dasyhelea) incisura" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="incisura" subGenus="Dasyhelea">Dasyhelea (Dasyhelea) incisura</taxonomicName>
sp. nov.; 
<emphasis bold="true" pageId="0" pageNumber="51">B</emphasis>
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Dasyhelea) quasifulcillata" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="quasifulcillata" subGenus="Dasyhelea">Dasyhelea (Dasyhelea) quasifulcillata</taxonomicName>
sp. nov.; 
<emphasis bold="true" pageId="0" pageNumber="51">C</emphasis>
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Dasyhelea) trigona" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="trigona" subGenus="Dasyhelea">Dasyhelea (Dasyhelea) trigona</taxonomicName>
sp. nov.; 
<emphasis bold="true" pageId="0" pageNumber="51">D</emphasis>
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Sebessia) falxa" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="falxa" subGenus="Sebessia">Dasyhelea (Sebessia) falxa</taxonomicName>
sp. nov.; 
<emphasis bold="true" pageId="0" pageNumber="51">E</emphasis>
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Sebessia) folia" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="folia" subGenus="Sebessia">Dasyhelea (Sebessia) folia</taxonomicName>
sp. nov. Scale bars: 0.05 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Thorax</emphasis>
. Yellowish in colour.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Wing</emphasis>
(Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7E</figureCitation>
). Wing margin with short, prominent macrotrichia, macrotrichia in wing surface sparse. WL 0.91 mm long, WW 0.33 mm; CR 0.5. FCu proximal to costal extremity.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Legs</emphasis>
(Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7F</figureCitation>
). Pale brown in colour. Hind tibial comb (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7G</figureCitation>
) with 7 spines, third one longest. TRI 3.0, TRII 2.2 and TRIII 3.0.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Abdomen</emphasis>
. Light brown in colour.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Genitalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7H</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-E Adult male genitalia structures. A Dasyhelea (Dasyhelea) incisura sp. nov.; B Dasyhelea (Dasyhelea) quasifulcillata sp. nov.; C Dasyhelea (Dasyhelea) trigona sp. nov.; D Dasyhelea (Sebessia) falxa sp. nov.; E Dasyhelea (Sebessia) folia sp. nov. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure8" httpUri="https://binary.pensoft.net/fig/790308" pageId="0" pageNumber="51">8E</figureCitation>
). Tergite 9 broad apically, inverted dome shaped, 115 
<normalizedToken originalValue="μm">μm</normalizedToken>
long and 133.5 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide. Apicolateral processes (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7I</figureCitation>
) 27.5 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, base 7 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide and apex 2.3 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, a distinct notch at about mid length from base bearing one long seta; cercus bare. Sternite 9 (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7H</figureCitation>
) 1.93 
<normalizedToken originalValue="×">x</normalizedToken>
wider than length with bifurcated posteromedian margin. Gonocoxite oval shaped, 78 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, greatest width 48 
<normalizedToken originalValue="μm">μm</normalizedToken>
, having 3-4 stout, long setae on dorsal side. Gonostylus stumpy, 47 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 16 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide basally and 4.5 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide apically; less sclerotisation up to mid length; somewhat abruptly narrowed at about three fourth length from base; a single, distinct, dorsal, long seta present near base followed by another one, ventrally 4 small setae at middle region and one long seta at basal region. Each basal arm of parameres (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7H</figureCitation>
) 34.5 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, broad and leaf like, joined moderately with 30 
<normalizedToken originalValue="μm">μm</normalizedToken>
long posteromedian projection, extended beyond mid gonocoxite. Aedeagus (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-J Adult male of Dasyhelea (Sebessia) folia sp. nov. A Frontal sclerite; B Flagellum; C Clypeus; D Palpus; E Thorax; F Femora and tibiae (right to left) of fore, mid and hind legs; G Hind tibial comb; H Genitalia (left dorsal and right ventral view); I Apicolateral process; J Aedeagus. Scale bars: 0.05 mm." figureDoi="10.3897/evolsyst.7.90973.figure7" httpUri="https://binary.pensoft.net/fig/790307" pageId="0" pageNumber="51">7J</figureCitation>
) 
<normalizedToken originalValue="“H”">&quot;H&quot;</normalizedToken>
shaped, 32 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 43.5 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide; basal arm 18.5 
<normalizedToken originalValue="μm">μm</normalizedToken>
long enclosing 26.5 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide basal arch; posterolateral arm 9 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, stout; ventromedian projection absent.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Female adult.</emphasis>
Unknown.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Pupa and larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="etymology">
<paragraph pageId="0" pageNumber="51">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="51">
The name, &quot; 
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea folia" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="folia">
<emphasis italics="true" pageId="0" pageNumber="51">Dasyhelea folia</emphasis>
</taxonomicName>
&quot; derived from Latinised version of 
<emphasis italics="true" pageId="0" pageNumber="51">leaf</emphasis>
, referring to somewhat leaf shaped basal arms of parameres of male genitalia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="remarks">
<paragraph pageId="0" pageNumber="51">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (Sebessia) folia" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="folia" subGenus="Sebessia">Dasyhelea (Sebessia) folia</taxonomicName>
sp. nov. shows similarities with 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. dellapei" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="dellapei">
<emphasis italics="true" pageId="0" pageNumber="51">D. dellapei</emphasis>
</taxonomicName>
<normalizedToken originalValue="Díaz">Diaz</normalizedToken>
, Spinelli &amp; Ronderos, 2010 (Argentina), 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. fueguina" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="fueguina">
<emphasis italics="true" pageId="0" pageNumber="51">D. fueguina</emphasis>
</taxonomicName>
<normalizedToken originalValue="Díaz">Diaz</normalizedToken>
, Spinelli &amp; Ronderos, 2010 (Argentina) and 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. patagonica" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="patagonica">
<emphasis italics="true" pageId="0" pageNumber="51">D. patagonica</emphasis>
</taxonomicName>
Ingram &amp; Macfie, 1931 (Argentina, and Chile) in the shape of the basal arms of parameres, but the overall structure of parameres, gonostylus and aedeagus are markedly apart. 
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea folia" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="folia">
<emphasis italics="true" pageId="0" pageNumber="51">Dasyhelea folia</emphasis>
</taxonomicName>
has similarities with 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. (S.) holosericea" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="holosericea" subGenus="S.">D. (S.) holosericea</taxonomicName>
(Meigen, 1804) (Estonia, Finland, Poland, Lithuania, France, and Russia) in a similar shaped aedeagus and gonostylus, but the parameres, the apicolateral process and the gonocoxite are different. The new species is similar to 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. huertai" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="huertai">
<emphasis italics="true" pageId="0" pageNumber="51">D. huertai</emphasis>
</taxonomicName>
Grogan, Diaz, Spinelli &amp; Ronderos, 2016 (France) and 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. turnbowi" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="turnbowi">
<emphasis italics="true" pageId="0" pageNumber="51">D. turnbowi</emphasis>
</taxonomicName>
Grogan, Diaz, Spinelli &amp; Ronderos, 2016 (France) in respect to the parameres, but the venation pattern of the wing, the shape of tergite 9, the gonocoxite and gonostylus are opposing. The combination of characters stated in the diagnosis justifies 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="D. (Sebessia) folia" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="folia" subGenus="Sebessia">D. (Sebessia) folia</taxonomicName>
as a new member of this subgenus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="distribution">
<paragraph pageId="0" pageNumber="51">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="51">The material was collected from the Deltaic Proper of Gangetic West Bengal, at an altitude of 8 meters above sea level - new species from India.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="ecological notes">
<paragraph pageId="0" pageNumber="51">Ecological notes.</paragraph>
<paragraph pageId="0" pageNumber="51">
Adults of 
<taxonomicName authorityName="Brahma &amp; Chatterjee &amp; Hazra" authorityYear="2023" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyhelea (S.) folia" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="folia" subGenus="S.">Dasyhelea (S.) folia</taxonomicName>
sp. nov. were captured in proximity of a flower garden adjoining a brackish water river (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-E Photographs of collection sites where light traps were operated. A Collection site of Dasyhelea (Dasyhelea) incisura sp. nov.; B Collection site of Dasyhelea (D.) quasifulcillata sp. nov.; C Collection site of Dasyhelea (D.) trigona sp. nov. (adjacent fresh water river shown as different image at upper right corner); D Collection site of Dasyhelea (Sebessia) falxa sp. nov.; E. Collection site of Dasyhelea (S.) folia sp. nov. (adjacent brackish water river is same for both D and E, shown as different image at upper middle)." figureDoi="10.3897/evolsyst.7.90973.figure2" httpUri="https://binary.pensoft.net/fig/790302" pageId="0" pageNumber="51">2E</figureCitation>
), when the air temperature was 33-35 °C and the relative humidity was 80-82%.
</paragraph>
</subSubSection>
</treatment>
</document>