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<document id="17178A7EAC5A89FE0CA7F2B72D1ED17A" ID-CLB-Dataset="45910" ID-DOI="10.5281/zenodo.201918" ID-GBIF-Dataset="790d3a4e-4192-43ad-92a0-19d1add50cc0" ID-ISSN="1175-5326" ID-Zenodo-Dep="201918" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460413596942" checkinUser="plazi" docAuthor="Carriol, René-Pierre, Cahuzac, Bruno &amp; Lesport, Jean-François" docDate="2011" docId="AA5C87A4F749FFB814DE868D089D5F6C" docLanguage="en" docName="zt03109p068.pdf" docOrigin="Zootaxa 3109" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acasta martillacensis Carriol, sp. nov." docType="treatment" docVersion="9" lastPageNumber="65" masterDocId="5665FFDCF74AFFBE144985300B325E53" masterDocTitle="New species of Acasta and Balanus (Balanoidea: Acastinae, Balaninae) from the Early Miocene (Burdigalian) of the Aquitaine Basin, France" masterLastPageNumber="68" masterPageNumber="60" pageNumber="63" updateTime="1698328255533" updateUser="ExternalLinkService">
<mods:mods id="DD5A20173A56FBAFE975C1966060B3CA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="57AB119BC518651F4A1E76397A06AF39">
<mods:title id="FDCE897E9B1F7D52ED8D1172F327055B">New species of Acasta and Balanus (Balanoidea: Acastinae, Balaninae) from the Early Miocene (Burdigalian) of the Aquitaine Basin, France</mods:title>
</mods:titleInfo>
<mods:name id="EAAFF5818D3C91ECD8472DBAFCCC39EE" type="personal">
<mods:role id="92B148587DF0AB31A21EFEF196A0F7F3">
<mods:roleTerm id="CCDD7F7700E715DBF24F597B3E75FC54">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB684694D9D643A1C652617E9062E2FF">Carriol, René-Pierre</mods:namePart>
</mods:name>
<mods:name id="598672B425066D9CACD727259E9091C2" type="personal">
<mods:role id="159BB6D2174CE5E1139D611ADEF56272">
<mods:roleTerm id="E82164758E4F8FF7242DAABB4C901C0C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="88DB42BF1D4A8A0E70A527A4C0EB40E2">Cahuzac, Bruno</mods:namePart>
</mods:name>
<mods:name id="3D3D86D7DDAFAB11DF15328C99E8360E" type="personal">
<mods:role id="FC12C0AE0E581BFDD72BC9B277314EB5">
<mods:roleTerm id="4BBF1F797730FD18B31DBF2A39CBD3D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B1CBC97B9A83B9F4DA164FEC7790B787">Lesport, Jean-François</mods:namePart>
</mods:name>
<mods:typeOfResource id="036F22DAF84E694EB796A86643AFC15A">text</mods:typeOfResource>
<mods:relatedItem id="4855C7AD0B545847F88B09EA8462A6A2" type="host">
<mods:titleInfo id="29E6AD7F32C3AA781411B8500D7BB120">
<mods:title id="1AAD58E26EB1CAE02B1B93915E7F4BA8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7AC453E4B0C443B8B77C453A326A0B1F">
<mods:date id="FF461A7985BA75A2580735FD06589058">2011</mods:date>
<mods:detail id="8C41822507BA90C344C913C05BA5ED8F" type="volume">
<mods:number id="81A23A845C9524B69B908CE9C14D107B">3109</mods:number>
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<mods:extent id="351164727E64967CC23209D39F901049" unit="page">
<mods:start id="F3BE65FAB02DCDB99166999D583024F3">60</mods:start>
<mods:end id="70961145CB7474FAD80AFBC98CC67BC3">68</mods:end>
</mods:extent>
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<mods:classification id="F453ED3386EB3ACF0C27AC5D440112F2">journal article</mods:classification>
<mods:identifier id="FE70EE5EF19F9BEAAC261C7135D344BA" type="CLB-Dataset">45910</mods:identifier>
<mods:identifier id="46EB37A3A15CE09CE7EBCD64BDDCB04D" type="DOI">10.5281/zenodo.201918</mods:identifier>
<mods:identifier id="3A5989817E1F2259A6D8D92AD299BD7A" type="GBIF-Dataset">790d3a4e-4192-43ad-92a0-19d1add50cc0</mods:identifier>
<mods:identifier id="1AC8DFDBA7D2601DE4ED8E59E71D0DFE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E95A5DF8AEADE2DFAE802AFECC6DCDCC" type="Zenodo-Dep">201918</mods:identifier>
</mods:mods>
<treatment id="AA5C87A4F749FFB814DE868D089D5F6C" ID-DOI="http://doi.org/10.5281/zenodo.6192586" ID-GBIF-Taxon="119424214" ID-Zenodo-Dep="6192586" LSID="urn:lsid:zoobank.org:act:599D92AE-51AB-4C76-9C28-C24EF2800A91" httpUri="http://treatment.plazi.org/id/AA5C87A4F749FFB814DE868D089D5F6C" lastPageId="6" lastPageNumber="65" pageId="3" pageNumber="63">
<subSubSection id="6AEF6539F749FFBD14DE868D0A075DA4" pageId="3" pageNumber="63" type="nomenclature">
<paragraph id="224A36B2F749FFBD14DE868D09535D84" blockId="3.[151,609,957,1015]" box="[151,609,957,983]" pageId="3" pageNumber="63">
<heading id="790281DEF749FFBD14DE868D09535D84" bold="true" box="[151,609,957,983]" fontSize="11" level="1" pageId="3" pageNumber="63" reason="1">
<emphasis id="1081EAA0F749FFBD14DE868D09535D84" bold="true" box="[151,609,957,983]" pageId="3" pageNumber="63">
<taxonomicName id="E5F54D31F749FFBD14DE868D0ACF5D84" authority="Carriol" authorityName="Carriol" box="[151,509,957,983]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="martillacensis" status="sp. nov.">
<emphasis id="1081EAA0F749FFBD14DE868D0AAA5D84" bold="true" box="[151,408,957,983]" italics="true" pageId="3" pageNumber="63">Acasta martillacensis</emphasis>
Carriol
</taxonomicName>
<taxonomicNameLabel id="0BB257DBF749FFBD164D868D09535D84" box="[516,609,957,983]" pageId="3" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="224A36B2F749FFBD14DE86EF0A075DA4" blockId="3.[151,609,957,1015]" box="[151,309,991,1015]" pageId="3" pageNumber="63">
<figureCitation id="BACE2A37F749FFBD14DE86EF0A2E5DA4" box="[151,284,991,1015]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,250,1913,1936]" captionTargetBox="[193,1378,193,1869]" captionTargetId="figure@4.[193,1380,193,1871]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Acasta martillacensis sp. nov., holotype (no. 85 – 12 – 68), level GA 39191, a shell (basis, plus wall plates) with right scutum which was inside of the shell: A, right side of shell; B, external side of basis; C – D, external and internal side of scutum. E – H, Balanus aculeatus sp. nov., holotype (no. 19 – 12 – 23), carina with a fragment of basis in connection: E, basis magnified showing septed tubes; F, external side; G, internal side; H, lateral view." httpUri="https://zenodo.org/record/201920/files/figure.png" pageId="3" pageNumber="63">Figs. 2A–D</figureCitation>
,
<figureCitation id="BACE2A37F749FFBD156E86EF0A075DA4" box="[295,309,991,1015]" captionStart="FIGURE 3 A – B" captionStartId="5.[151,250,1204,1227]" captionTargetBox="[361,1223,191,1124]" captionTargetId="figure@5.[336,1248,173,1153]" captionTargetPageId="5" captionText="FIGURE 3 A – B. Acasta martillacensis sp. nov., paratypes (no. 85 – 12 – 69 and no. 85 – 12 – 70, respectively): A, fragment of paries transversally cut; B, fragment of paries longitudinally cut at sheath level. p, paries; r, rib; sh, sheath; sp, septae; t, tube." httpUri="https://zenodo.org/record/201921/files/figure.png" pageId="3" pageNumber="63">3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6AEF6539F749FFBD14DE81180A685B0C" pageId="3" pageNumber="63" type="materials_examined">
<paragraph id="224A36B2F749FFBD14DE81180A175A37" blockId="3.[151,1437,1063,2024]" pageId="3" pageNumber="63">
<emphasis id="1081EAA0F749FFBD14DE81180A015A13" bold="true" box="[151,307,1063,1088]" pageId="3" pageNumber="63">
<typeStatus id="FD4E8810F749FFBD14DE81180BE25A13" box="[151,208,1064,1088]" pageId="3" pageNumber="63">Type</typeStatus>
locality.
</emphasis>
Martillac ditch section, near 'Couvent de la Solitude', Département Gironde, Aquitaine Basin, southwest
<collectingCountry id="5AE27622F749FFBD1498817D0A135A37" box="[209,289,1101,1124]" name="France" pageId="3" pageNumber="63">France</collectingCountry>
.
</paragraph>
<paragraph id="224A36B2F749FFBD148E81400AFB5AFF" blockId="3.[151,1437,1063,2024]" pageId="3" pageNumber="63">
<emphasis id="1081EAA0F749FFBD148E81400A5C5ADB" bold="true" box="[199,366,1136,1160]" pageId="3" pageNumber="63">
<typeStatus id="FD4E8810F749FFBD148E81400A325ADB" box="[199,256,1136,1160]" pageId="3" pageNumber="63">Type</typeStatus>
stratum.
</emphasis>
Early Burdigalian, fine grey-blue sands in the lower portion of the section exposed, level GA 39191, “Sables à Mactres”.
</paragraph>
<paragraph id="224A36B2F749FFBD148E8187082E5B4B" blockId="3.[151,1437,1063,2024]" pageId="3" pageNumber="63">
<emphasis id="1081EAA0F749FFBD148E81870A805A83" bold="true" box="[199,434,1207,1232]" pageId="3" pageNumber="63">Material examined.</emphasis>
<typeStatus id="FD4E8810F749FFBD15F3818709105A9C" box="[442,546,1207,1231]" pageId="3" pageNumber="63" type="holotype">Holotype</typeStatus>
(no. 85-12-68, Cahuzac Collection), a shell (basis with wall plates), retaining the right scutum.
<typeStatus id="FD4E8810F749FFBD151381ED0AF85AA6" box="[346,458,1245,1269]" pageId="3" pageNumber="63" type="paratype">Paratypes</typeStatus>
, (no. 85-12-69, Lesport Collection) a rostrum; (no. 85-12-70, Lesport Collection) half a basis with carina, a carinolateral, lateral and half rostrum.
</paragraph>
<paragraph id="224A36B2F749FFBD148E80130A685B0C" blockId="3.[151,1437,1063,2024]" pageId="3" pageNumber="63">
<emphasis id="1081EAA0F749FFBD148E80130A805B6F" bold="true" box="[199,434,1315,1340]" pageId="3" pageNumber="63">Additional material</emphasis>
(Lesport Collection). 2 rostra, 2 carinae, 1 basis, 1 fragmentary basis with a partial lateral and carinolateral.
</paragraph>
</subSubSection>
<subSubSection id="6AEF6539F749FFBD148E805C0988586B" pageId="3" pageNumber="63" type="description">
<paragraph id="224A36B2F749FFBD148E805C0988586B" blockId="3.[151,1437,1063,2024]" pageId="3" pageNumber="63">
<emphasis id="1081EAA0F749FFBD148E805C0A4F5BD7" bold="true" box="[199,381,1388,1412]" pageId="3" pageNumber="63">Measurements.</emphasis>
<typeStatus id="FD4E8810F749FFBD15CC805C0AC05BD7" box="[389,498,1388,1412]" pageId="3" pageNumber="63" type="holotype">Holotype</typeStatus>
: opercular aperture, carino-rostral diameter
<quantity id="E50D9B57F749FFBD17AE805C0F705BD7" box="[999,1090,1388,1412]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="63" unit="mm" value="3.5">3.5 mm</quantity>
; carina at mid-height
<quantity id="E50D9B57F749FFBD1174805C0EA55BD7" box="[1341,1431,1388,1412]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="63" unit="mm" value="4.4">4.4 mm</quantity>
; rostrum at mid-height
<quantity id="E50D9B57F749FFBD15D380A00AC75BFB" box="[410,501,1424,1448]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="63" unit="mm" value="3.9">3.9 mm</quantity>
, sheath height
<quantity id="E50D9B57F749FFBD16EC80A008325BFB" box="[677,768,1424,1448]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="63" unit="mm" value="1.9">1.9 mm</quantity>
; basis, carino-rostral diameter
<quantity id="E50D9B57F749FFBD102D80A00F8D5BFB" box="[1124,1215,1424,1448]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="63" unit="mm" value="4.1">4.1 mm</quantity>
; scutum, occludent margin
<quantity id="E50D9B57F749FFBD14A580840A775B9F" box="[236,325,1460,1484]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="63" unit="mm" value="3.1">3.1 mm</quantity>
, articular margin 2.0 mm, articular ridge
<quantity id="E50D9B57F749FFBD175B808408595B9F" box="[786,875,1460,1484]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="63" unit="mm" value="1.4">1.4 mm</quantity>
.
<typeStatus id="FD4E8810F749FFBD173F808508EA5B9E" box="[886,984,1461,1485]" pageId="3" pageNumber="63" type="paratype">Paratype</typeStatus>
no. 85-12-69: rostrum at mid-height
<quantity id="E50D9B57F749FFBD113080840BF45BA3" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="63" unit="mm" value="6.3">6.3 mm</quantity>
.
<typeStatus id="FD4E8810F749FFBD149B80E90A065BA2" box="[210,308,1497,1521]" pageId="3" pageNumber="63" type="paratype">Paratype</typeStatus>
no. 85-12-70: basis, carino-rostral diameter 7.0 mm, opercular aperture, carino-rostral diameter
<quantity id="E50D9B57F749FFBD113080E80BF45847" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="63" unit="mm" value="5.5">5.5 mm</quantity>
. Additional material: 2 rostra, at mid-height 5.1 and
<quantity id="E50D9B57F749FFBD175D80CC085F5847" box="[788,877,1532,1556]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="63" unit="mm" value="6.6">6.6 mm</quantity>
, respectively; 2 carinae, at mid-height 5.6 and
<quantity id="E50D9B57F749FFBD113080CC0BF7586B" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="3" pageNumber="63" unit="mm" value="7.1">7.1 mm</quantity>
, respectively; basis, larger diameter 6.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="6AEF6539F749FFBD148E8373091758F7" pageId="3" pageNumber="63" type="diagnosis">
<paragraph id="224A36B2F749FFBD148E8373091758F7" blockId="3.[151,1437,1063,2024]" pageId="3" pageNumber="63">
<emphasis id="1081EAA0F749FFBD148E83730A73580F" bold="true" box="[199,321,1603,1628]" pageId="3" pageNumber="63">Diagnosis.</emphasis>
Shell with internal longitudinal ribs; cellular space between sheath and wall; carinal sheath adhering to the wall in the centre of its basal edge; basis cup shaped; scutum externally with radial striae, and internally with adductor ridge, not prominent.
</paragraph>
</subSubSection>
<subSubSection id="6AEF6539F749FFBB148E839F0E155BB1" lastPageId="5" lastPageNumber="65" pageId="3" pageNumber="63" type="description">
<paragraph id="224A36B2F749FFBB148E839F0E155BB1" blockId="3.[151,1437,1063,2024]" lastBlockId="5.[151,1437,1302,2011]" lastPageId="5" lastPageNumber="65" pageId="3" pageNumber="63">
<emphasis id="1081EAA0F749FFBD148E839F0A6B589B" bold="true" box="[199,345,1711,1736]" pageId="3" pageNumber="63">Description.</emphasis>
Shell white, occasionally with pink hue towards apex and on radii (original coloration visible on
<typeStatus id="FD4E8810F749FFBD14DE83E40BCC58BF" box="[151,254,1748,1772]" pageId="3" pageNumber="63" type="holotype">holotype</typeStatus>
). Parietes internally with well-developed ribbing extending up to sheath or nearly so. Between ribs, in parts of plate where they are close, fine longitudinal spaces present. These tubes do not show an aperture between the ribs near basal margin of plates (see
<figureCitation id="BACE2A37F749FFBD162E822C09995967" box="[615,683,1820,1844]" captionStart="FIGURE 3 A – B" captionStartId="5.[151,250,1204,1227]" captionTargetBox="[361,1223,191,1124]" captionTargetId="figure@5.[336,1248,173,1153]" captionTargetPageId="5" captionText="FIGURE 3 A – B. Acasta martillacensis sp. nov., paratypes (no. 85 – 12 – 69 and no. 85 – 12 – 70, respectively): A, fragment of paries transversally cut; B, fragment of paries longitudinally cut at sheath level. p, paries; r, rib; sh, sheath; sp, septae; t, tube." httpUri="https://zenodo.org/record/201921/files/figure.png" pageId="3" pageNumber="63">Fig. 3</figureCitation>
); only in broken plates can tubes be demonstrated. Parietes externally with growth ridges crossed by very fine longitudinal ribs, and with, hollow, upward-curving spines sloping downwards, spines occasionally bifurcated. Radii and alae not very wide, upper edge oblique, sutural edge of radii with striations. Sheath broad, occupying about half length of compartment, transversally striated with basal edge free from wall except in carina, where central part of basal edge turns down, adhering to wall. Cellular space, secondarily calcified, between sheath and wall. Basis cup-shaped, calcareous, non porous, concentrically ringed outside but smooth inside, with margin bearing small pits for reception of teeth formed by ends of internal parietal ribs. Scutum wide, taller than wide, slightly bowed between apex and basal margin; occludent margin slightly concave, tergal margin straight, basal margin convex; exterior ornamented by growth increments crossed by incised radial striae; interior, with articular ridge not prominent barely extending over the tergal margin, and with adductor ridge not prominent, of same length as tergal edge of adductor muscle pit; lateral depressor muscle scar rounded, rather deep; adductor muscle pit shallow, rather wide with its posterior extremity not clearly discernible.
</paragraph>
</subSubSection>
<caption id="768A663AF74EFFBA14DE824908E059BF" httpUri="https://zenodo.org/record/201920/files/figure.png" pageId="4" pageNumber="64" targetBox="[193,1378,193,1869]" targetPageId="4">
<paragraph id="224A36B2F74EFFBA14DE824908E059BF" blockId="4.[151,1436,1913,2028]" pageId="4" pageNumber="64">
<emphasis id="1081EAA0F74EFFBA14DE82490A2659DC" bold="true" box="[151,276,1913,1936]" pageId="4" pageNumber="64">FIGURE 2.</emphasis>
A–D,
<taxonomicName id="E5F54D31F74EFFBA1512824B090A59C3" box="[347,568,1914,1936]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="4" pageNumber="64" phylum="Arthropoda" rank="species" species="martillacensis" status="sp. nov.">
<emphasis id="1081EAA0F74EFFBA1512824B090A59C3" box="[347,568,1914,1936]" italics="true" pageId="4" pageNumber="64">Acasta martillacensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BB257DBF74EFFBA1676824A09A359DC" box="[575,657,1914,1935]" pageId="4" pageNumber="64" rank="species">
<emphasis id="1081EAA0F74EFFBA1676824A09A359DC" bold="true" box="[575,657,1914,1935]" pageId="4" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype (no. 85–12–68), level GA 39191, a shell (basis, plus wall plates) with right scutum which was inside of the shell: A, right side of shell; B, external side of basis; C–D, external and internal side of scutum. E–H,
<taxonomicName id="E5F54D31F74EFFBA150C82870932599E" box="[325,512,1975,1997]" class="Maxillopoda" family="Balanidae" genus="Balanus" kingdom="Animalia" order="Sessilia" pageId="4" pageNumber="64" phylum="Arthropoda" rank="species" species="aculeatus" status="sp. nov.">
<emphasis id="1081EAA0F74EFFBA150C82870932599E" box="[325,512,1975,1997]" italics="true" pageId="4" pageNumber="64">Balanus aculeatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BB257DBF74EFFBA164E8288096A599E" box="[519,600,1976,1997]" pageId="4" pageNumber="64" rank="species">
<emphasis id="1081EAA0F74EFFBA164E8288096A599E" bold="true" box="[519,600,1976,1997]" pageId="4" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype (no. 19–12–23), carina with a fragment of basis in connection: E, basis magnified showing septed tubes; F, external side; G, internal side; H, lateral view.
</paragraph>
</caption>
<caption id="768A663AF74FFFBB14DE81840EB85ABA" httpUri="https://zenodo.org/record/201921/files/figure.png" pageId="5" pageNumber="65" targetBox="[361,1223,191,1124]" targetPageId="5">
<paragraph id="224A36B2F74FFFBB14DE81840EB85ABA" blockId="5.[151,1435,1204,1258]" pageId="5" pageNumber="65">
<emphasis id="1081EAA0F74FFFBB14DE81840A775A99" bold="true" box="[151,325,1204,1227]" pageId="5" pageNumber="65">FIGURE 3A–B.</emphasis>
<taxonomicName id="E5F54D31F74FFFBB15078186091E5A98" box="[334,556,1205,1227]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="65" phylum="Arthropoda" rank="species" species="martillacensis" status="sp. nov.">
<emphasis id="1081EAA0F74FFFBB15078186091E5A98" box="[334,556,1205,1227]" italics="true" pageId="5" pageNumber="65">Acasta martillacensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BB257DBF74FFFBB167C818509BA5A99" box="[565,648,1205,1226]" pageId="5" pageNumber="65" rank="species">
<emphasis id="1081EAA0F74FFFBB167C818509BA5A99" bold="true" box="[565,648,1205,1226]" pageId="5" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratypes (no. 85–12–69 and no. 85–12–70, respectively): A, fragment of paries transversally cut; B, fragment of paries longitudinally cut at sheath level. p, paries; r, rib; sh, sheath; sp, septae; t, tube.
</paragraph>
</caption>
<subSubSection id="6AEF6539F74FFFBB148E80DF0F3F5855" box="[199,1037,1518,1543]" pageId="5" pageNumber="65" type="etymology">
<paragraph id="224A36B2F74FFFBB148E80DF0F3F5855" blockId="5.[151,1437,1302,2011]" box="[199,1037,1518,1543]" pageId="5" pageNumber="65">
<emphasis id="1081EAA0F74FFFBB148E80DF0A7C5854" bold="true" box="[199,334,1519,1543]" pageId="5" pageNumber="65">Etymology.</emphasis>
In reference to Martillac, Aquitaine Basin (southwest
<collectingCountry id="5AE27622F74FFFBB17F980DE0F335855" box="[944,1025,1518,1542]" name="France" pageId="5" pageNumber="65">France</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="6AEF6539F74FFFB8148E8323089D5F6C" lastPageId="6" lastPageNumber="66" pageId="5" pageNumber="65" type="discussion">
<paragraph id="224A36B2F74FFFBB148E83230BD85975" blockId="5.[151,1437,1302,2011]" pageId="5" pageNumber="65">
<emphasis id="1081EAA0F74FFFBB148E83230A095878" bold="true" box="[199,315,1555,1579]" pageId="5" pageNumber="65">Remarks.</emphasis>
The specimens of
<taxonomicName id="E5F54D31F74FFFBB165D832208355879" box="[532,775,1554,1578]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="65" phylum="Arthropoda" rank="species" species="martillacensis" status="sp. nov.">
<emphasis id="1081EAA0F74FFFBB165D832208355879" box="[532,775,1554,1578]" italics="true" pageId="5" pageNumber="65">Acasta martillacensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BB257DBF74FFFBB1759832308595878" box="[784,875,1555,1579]" pageId="5" pageNumber="65" rank="species">
<emphasis id="1081EAA0F74FFFBB1759832308595878" bold="true" box="[784,875,1555,1579]" pageId="5" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
have ribbed parietal plates as do other numerous species of
<taxonomicName id="E5F54D31F74FFFBB154683060A69581D" box="[271,347,1590,1614]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="1081EAA0F74FFFBB154683060A69581D" box="[271,347,1590,1614]" italics="true" pageId="5" pageNumber="65">Acasta</emphasis>
</taxonomicName>
. No apertures of the tubes are visible between these ribs near the basal margin of the plate. It is along breaks of certain specimens that these tubes appear and these specimens, which seem rather old in view of their size, are only partially tubiferous. Tubes are present only in particular parts of the plates, in the parts where ribs are numerous and in contact, as if the ribs by contacting had closed a longitudinal space between them. These tubes are not the same that those present in
<emphasis id="1081EAA0F74FFFBB16C783F608B1588D" box="[654,899,1734,1758]" italics="true" pageId="5" pageNumber="65">Perforatus perforatus</emphasis>
and are different to those in
<taxonomicName id="E5F54D31F74FFFBB109883F60EA6588C" box="[1233,1428,1734,1759]" class="Maxillopoda" family="Balanidae" genus="Balanus" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="65" phylum="Arthropoda" rank="species" species="crenatus">
<emphasis id="1081EAA0F74FFFBB109883F60EA6588C" box="[1233,1428,1734,1759]" italics="true" pageId="5" pageNumber="65">Balanus crenatus</emphasis>
</taxonomicName>
. They are not tubes in the sense we usually understand and the presence of solid paries remains a characteristic of
<taxonomicName id="E5F54D31F74FFFBB14DE823E0BD15975" box="[151,227,1806,1830]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="1081EAA0F74FFFBB14DE823E0BD15975" box="[151,227,1806,1830]" italics="true" pageId="5" pageNumber="65">Acasta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="224A36B2F74FFFB8148E8202089D5F6C" blockId="5.[151,1437,1302,2011]" lastBlockId="6.[151,1437,151,319]" lastPageId="6" lastPageNumber="66" pageId="5" pageNumber="65">
<emphasis id="1081EAA0F74FFFBB148E82020A0E5918" bold="true" box="[199,316,1842,1867]" pageId="5" pageNumber="65">Affinities.</emphasis>
Among the 26 extant species retained in the genus
<taxonomicName id="E5F54D31F74FFFBB1735820208FA5919" box="[892,968,1842,1866]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="1081EAA0F74FFFBB1735820208FA5919" box="[892,968,1842,1866]" italics="true" pageId="5" pageNumber="65">Acasta</emphasis>
</taxonomicName>
by
<bibRefCitation id="46644B43F74FFFBB17BB82020F835919" author="Kolbasov" box="[1010,1201,1842,1866]" pageId="5" pageNumber="65" refString="Kolbasov, G. A. (1993) Revision of the genus Acasta Leach (Cirripedia: Balanoidea). Zoological Journal of the Linnean Society, 109, 395 - 427." type="journal article" year="1993">Kolbasov (1993)</bibRefCitation>
, one species has fossil representatives:
<taxonomicName id="E5F54D31F74FFFBB153D82660901593D" box="[372,563,1878,1902]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="65" phylum="Arthropoda" rank="species" species="spongites">
<emphasis id="1081EAA0F74FFFBB153D82660901593D" box="[372,563,1878,1902]" italics="true" pageId="5" pageNumber="65">Acasta spongites</emphasis>
</taxonomicName>
, the extant species from the Atlantic and the Mediterranean Sea, which has been described from the Pliocene of
<collectingCountry id="5AE27622F74FFFBB167A824B094D59C1" box="[563,639,1915,1938]" name="France" pageId="5" pageNumber="65">France</collectingCountry>
(
<collectingCountry id="5AE27622F74FFFBB16C7824A098259C1" box="[654,688,1914,1938]" name="South Georgia and the South Sandwich Islands" pageId="5" pageNumber="65">De</collectingCountry>
Alessandri 1907) and
<collectingCountry id="5AE27622F74FFFBB17E6824A08D059C1" box="[943,994,1914,1938]" name="Italy" pageId="5" pageNumber="65">Italy</collectingCountry>
(Moroni 1952). There are eight extinct species (
<bibRefCitation id="46644B43F74FFFBB14B082AE0AA359E5" author="Carriol" box="[249,401,1950,1974]" pageId="5" pageNumber="65" refString="Carriol, R. - P. (2008) New genus and new species of Cirripedia (Chthamalidae, Tetraclitidae, Archaeobalanidae and Balanidae) from the Middle Miocene of the faluns of Touraine (France). Zootaxa, 1675, 31 - 48." type="journal article" year="2008">Carriol 2008</bibRefCitation>
). The
<taxonomicName id="E5F54D31F74FFFBB159282AE091559E5" box="[475,551,1950,1974]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="1081EAA0F74FFFBB159282AE091559E5" box="[475,551,1950,1974]" italics="true" pageId="5" pageNumber="65">Acasta</emphasis>
</taxonomicName>
with fossil representatives, except
<taxonomicName id="E5F54D31F74FFFBB17F482AE0FC759E5" authority="Carriol, 2008" authorityName="Carriol" authorityYear="2008" box="[957,1269,1950,1974]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="65" phylum="Arthropoda" rank="species" species="vesiculosa">
<emphasis id="1081EAA0F74FFFBB17F482AE0F6659E5" box="[957,1108,1950,1974]" italics="true" pageId="5" pageNumber="65">A. vesiculosa</emphasis>
<bibRefCitation id="46644B43F74FFFBB101482AE0FC759E5" author="Carriol" box="[1117,1269,1950,1974]" pageId="5" pageNumber="65" refString="Carriol, R. - P. (2008) New genus and new species of Cirripedia (Chthamalidae, Tetraclitidae, Archaeobalanidae and Balanidae) from the Middle Miocene of the faluns of Touraine (France). Zootaxa, 1675, 31 - 48." type="journal article" year="2008">Carriol, 2008</bibRefCitation>
</taxonomicName>
(see complete description in
<bibRefCitation id="46644B43F74FFFBB157582F209045989" author="Schneider" box="[316,566,1986,2010]" pageId="5" pageNumber="65" refString="Schneider, S., Berning, B., Bitner, M. A., Carriol, R. - P., Jager, M., Kriwet, J., Kroh, A. &amp; Werner, W. (2009) A parautochthonous shallow marine fauna from the Late Burdigalian (early Ottnangian) of Gurlarn (Lower Bavaria, SE Germany): Macrofaunal inventory and paleoecology. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen, 254, 63 - 103." type="journal article" year="2009">
Schneider
<emphasis id="1081EAA0F74FFFBB15FC82F30AC25989" box="[437,496,1986,2010]" italics="true" pageId="5" pageNumber="65">et al.</emphasis>
2009
</bibRefCitation>
), may be distinguished from
<taxonomicName id="E5F54D31F74FFFBB17C182F20F4E5989" box="[904,1148,1986,2010]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="65" phylum="Arthropoda" rank="species" species="martillacensis" status="sp. nov.">
<emphasis id="1081EAA0F74FFFBB17C182F20F4E5989" box="[904,1148,1986,2010]" italics="true" pageId="5" pageNumber="65">Acasta martillacensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BB257DBF74FFFBB10CF82F30FD05988" box="[1158,1250,1987,2011]" pageId="5" pageNumber="65" rank="species">
<emphasis id="1081EAA0F74FFFBB10CF82F30FD05988" bold="true" box="[1158,1250,1987,2011]" pageId="5" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
in the lack of a vesicular sheath.
<taxonomicName id="E5F54D31F74CFFB8151E85A8097B5EE3" box="[343,585,152,176]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="66" phylum="Arthropoda" rank="species" species="martillacensis" status="sp. nov.">
<emphasis id="1081EAA0F74CFFB8151E85A8097B5EE3" box="[343,585,152,176]" italics="true" pageId="6" pageNumber="66">Acasta martillacensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BB257DBF74CFFB8160685A8099A5EE3" box="[591,680,152,176]" pageId="6" pageNumber="66" rank="species">
<emphasis id="1081EAA0F74CFFB8160685A8099A5EE3" bold="true" box="[591,680,152,176]" pageId="6" pageNumber="66">sp. nov.</emphasis>
</taxonomicNameLabel>
is close to
<taxonomicName id="E5F54D31F74CFFB8176D85A808855EE3" box="[804,951,152,176]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="66" phylum="Arthropoda" rank="species" species="vesiculosa">
<emphasis id="1081EAA0F74CFFB8176D85A808855EE3" box="[804,951,152,176]" italics="true" pageId="6" pageNumber="66">A. vesiculosa</emphasis>
</taxonomicName>
in having the vesicular sheath and a carinal sheath adhering to the wall in the centre of its basal edge. However, the former can be distinguished by the internal ribbing of the wall, its cup-shaped basis and its scutum, in which the adductor ridge is not prominent. The claim that
<taxonomicName id="E5F54D31F74CFFB8148184340A6E5F4F" box="[200,348,260,284]" class="Maxillopoda" family="Archaeobalanidae" genus="Acasta" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="66" phylum="Arthropoda" rank="species" species="vesiculosa">
<emphasis id="1081EAA0F74CFFB8148184340A6E5F4F" box="[200,348,260,284]" italics="true" pageId="6" pageNumber="66">A. vesiculosa</emphasis>
</taxonomicName>
from the middle Miocene of
<collectingCountry id="5AE27622F74CFFB816EF843509C05F4F" box="[678,754,261,284]" name="France" pageId="6" pageNumber="66">France</collectingCountry>
(
<bibRefCitation id="46644B43F74CFFB81749843408A15F4F" author="Carriol" box="[768,915,260,284]" pageId="6" pageNumber="66" refString="Carriol, R. - P. (2008) New genus and new species of Cirripedia (Chthamalidae, Tetraclitidae, Archaeobalanidae and Balanidae) from the Middle Miocene of the faluns of Touraine (France). Zootaxa, 1675, 31 - 48." type="journal article" year="2008">Carriol 2008</bibRefCitation>
) showed internal ribbing of the wall is erroneous, due to the poor preservation of a small set of specimens available.
</paragraph>
</subSubSection>
</treatment>
</document>
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