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<document id="745350579DE53462EB6BE4EE45844425" ID-CLB-Dataset="31142" ID-DOI="10.11646/zootaxa.4365.2.7" ID-GBIF-Dataset="0e9444d5-9938-46ed-8f44-5b775809d2bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="1117582" ID-ZooBank="AF2A21D6-88F3-4234-B5B1-E9E5FE84526C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513584024333" checkinUser="plazi" docAuthor="Pinheiro, Lívia R. &amp; Araujo Junior, Elias C." docDate="2017" docId="790587909705E36CF1F4FF0C02C7041D" docLanguage="en" docName="zootaxa.4365.2.7.pdf" docOrigin="Zootaxa 4365 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Episcepsis atlantica Pinheiro &amp; Araujo Junior 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="238" masterDocId="853CFFE89703E36BF163FF9B064E072A" masterDocTitle="Taxonomic notes and a new species of Episcepsis Butler, 1877 (Lepidoptera, Erebidae, Arctiinae, Arctiini, Ctenuchina)" masterLastPageNumber="240" masterPageNumber="231" pageNumber="237" updateTime="1698510196860" updateUser="ExternalLinkService">
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<mods:title id="92E2E6522AAD98438CB7A3BA39C253B7">Taxonomic notes and a new species of Episcepsis Butler, 1877 (Lepidoptera, Erebidae, Arctiinae, Arctiini, Ctenuchina)</mods:title>
</mods:titleInfo>
<mods:name id="6C8181027CA85580FE3B86C20E0CA5B9" type="personal">
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<mods:namePart id="CBD86235B544FAD47D982DA04D75A348">Pinheiro, Lívia R.</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="0E222090C4A12D063A5C99CA33ACF2A8">Araujo Junior, Elias C.</mods:namePart>
</mods:name>
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<mods:title id="0823622BEEFC81B5CB4AACE52D31DA60">Zootaxa</mods:title>
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<mods:part id="40702F53E41C6F56084B6C74C1822454">
<mods:date id="0D8F7CDC299F66FBDCF8ED87EDE57D1B">2017</mods:date>
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<mods:number id="4FE9CC69FFCC339713525A685341964E">2017-11-18</mods:number>
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<mods:detail id="DF8A99D7223CBDA4E6F9004CEBD4498A" type="volume">
<mods:number id="24FC81238BE6DAE5A8B1293E0404BCB5">4365</mods:number>
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<mods:classification id="DE5278B87243F148C481568CA34DD1D2">journal article</mods:classification>
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<mods:identifier id="F731238B7C2861EE5A28573B9950861A" type="DOI">10.11646/zootaxa.4365.2.7</mods:identifier>
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<mods:identifier id="0F656E142EA6F590B6784CA7C225AFA9" type="ISSN">1175-5326</mods:identifier>
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<treatment id="790587909705E36CF1F4FF0C02C7041D" ID-DOI="http://doi.org/10.5281/zenodo.6049820" ID-GBIF-Taxon="139334163" ID-Zenodo-Dep="6049820" LSID="urn:lsid:plazi:treatment:790587909705E36CF1F4FF0C02C7041D" httpUri="http://treatment.plazi.org/id/790587909705E36CF1F4FF0C02C7041D" lastPageId="7" lastPageNumber="238" pageId="6" pageNumber="237">
<subSubSection id="B9B6650D9705E36DF1F4FF0C071F07F9" pageId="6" pageNumber="237" type="nomenclature">
<paragraph id="F11336869705E36DF1F4FF0C05420798" blockId="6.[151,780,151,211]" box="[151,780,151,178]" pageId="6" pageNumber="237">
<heading id="AA5B81EA9705E36DF1F4FF0C05420798" bold="true" box="[151,780,151,178]" fontSize="11" level="1" pageId="6" pageNumber="237" reason="1">
<emphasis id="C3D8EA949705E36DF1F4FF0C05420798" bold="true" box="[151,780,151,178]" pageId="6" pageNumber="237">
<taxonomicName id="36AC4D059705E36DF1F4FF0C04EF0798" authority="Pinheiro &amp; Araujo Jr." authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[151,673,151,178]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF1F4FF0C07C8079B" bold="true" box="[151,390,151,177]" italics="true" pageId="6" pageNumber="237">Episcepsis atlantica</emphasis>
Pinheiro &amp; Araujo Jr.
</taxonomicName>
,
<taxonomicNameLabel id="D8EB57EF9705E36DF3CDFF0305420798" box="[686,780,152,178]" pageId="6" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F11336869705E36DF1F4FF21071F07F9" blockId="6.[151,780,151,211]" box="[151,337,186,211]" pageId="6" pageNumber="237">
(
<figureCitation id="69972A039705E36DF1FCFF21074F07F9" box="[159,257,186,211]" captionStart="FIGURE 1" captionStartId="3.[151,250,1351,1373]" captionTargetBox="[151,1436,195,1327]" captionTargetId="figure@3.[151,1436,195,1327]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of the species treated herein. A) E. aelia revis. stat., holotype; B) E. atlantica sp. nov., holotype; C) E. fumata, paralectotype;D) E.venata, holotype; E) E.coeruleocephala, lectotype. Scale bars: 1 cm." httpUri="https://zenodo.org/record/1117584/files/figure.png" pageId="6" pageNumber="237">Figs. 1B</figureCitation>
; 5A–I)
</paragraph>
</subSubSection>
<subSubSection id="B9B6650D9705E36DF1F4FE9903560649" pageId="6" pageNumber="237" type="materials_examined">
<paragraph id="F11336869705E36DF1F4FE9903560649" blockId="6.[151,1436,258,877]" pageId="6" pageNumber="237">
<materialsCitation id="41C43CDB9705E36DF1F4FE9903500631" ID-GBIF-Occurrence="1795578719" box="[151,1310,258,283]" collectingDate="1929-11" collectionCode="MZSP" collectorName="Dirings &amp; Two" country="Brazil" location="Timbo" pageId="6" pageNumber="237" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="holotype">
<typeStatus id="2E1788249705E36DF1F4FE9906B10631" box="[151,255,258,283]" pageId="6" pageNumber="237" type="holotype">Holotype</typeStatus>
<specimenCount id="E7AAFD0F9705E36DF065FE9807530630" box="[262,285,259,282]" pageId="6" pageNumber="237" type="male">♂</specimenCount>
,
<collectingCountry id="89BB76169705E36DF049FE9907DB0631" box="[298,405,258,283]" name="Brazil" pageId="6" pageNumber="237">
<emphasis id="C3D8EA949705E36DF049FE9907DB0631" bold="true" box="[298,405,258,283]" pageId="6" pageNumber="237">BRAZIL</emphasis>
</collectingCountry>
,
<collectingRegion id="3368F8649705E36DF0C7FE9804050631" box="[420,587,258,283]" country="Brazil" name="Santa Catarina" pageId="6" pageNumber="237">Santa Catarina</collectingRegion>
,
<location id="F473605D9705E36DF335FE9904ED0631" LSID="urn:lsid:plazi:treatment:790587909705E36CF1F4FF0C02C7041D:F473605D9705E36DF335FE9904ED0631" box="[598,675,258,283]" country="Brazil" name="Timbo" pageId="6" pageNumber="237" stateProvince="Santa Catarina">Timbó</location>
,
<collectingDate id="9556E9AE9705E36DF3CCFE9905480631" box="[687,774,258,283]" pageId="6" pageNumber="237" value="1929-11">
<date id="851210469705E36DF3CCFE9905480631" box="[687,774,258,283]" pageId="6" pageNumber="237" value="1929-11">xi.1929</date>
</collectingDate>
,
<collectorName id="5C5953509705E36DF271FE9905280631" box="[786,870,258,283]" pageId="6" pageNumber="237">Dirings</collectorName>
coll. (
<collectionCode id="97BDAE439705E36DF2CDFE9805B30631" box="[942,1021,259,283]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="237">MZSP</collectionCode>
).
<collectorName id="5C5953509705E36DF573FE98020F0630" box="[1040,1089,259,282]" pageId="6" pageNumber="237">Two</collectorName>
<typeStatus id="2E1788249705E36DF52AFE9802F90631" box="[1097,1207,259,283]" pageId="6" pageNumber="237" type="paratype">paratypes</typeStatus>
, females
</materialsCitation>
:
<materialsCitation id="41C43CDB9705E36DF449FE9902550615" ID-GBIF-Occurrence="1795578656" collectingDate="1958-01" collectionCode="MZSP" collectorName="Dirings" country="Brazil" location="Rio Vermelho" municipality="Sao Bento do Sul" pageId="6" pageNumber="237" specimenCount="1" stateProvince="Santa Catarina" typeStatus="holotype">
<collectingCountry id="89BB76169705E36DF449FE9903DB0631" box="[1322,1429,258,283]" name="Brazil" pageId="6" pageNumber="237">
<emphasis id="C3D8EA949705E36DF449FE9903DB0631" bold="true" box="[1322,1429,258,283]" pageId="6" pageNumber="237">BRAZIL</emphasis>
</collectingCountry>
,
<collectingRegion id="3368F8649705E36DF1F4FEBC07700615" box="[151,318,294,319]" country="Brazil" name="Santa Catarina" pageId="6" pageNumber="237">Santa Catarina</collectingRegion>
, [
<collectingMunicipality id="1177ACFC9705E36DF02CFEBC04570615" box="[335,537,294,319]" pageId="6" pageNumber="237">São Bento do Sul</collectingMunicipality>
],
<location id="F473605D9705E36DF348FEBD04840615" LSID="urn:lsid:plazi:treatment:790587909705E36CF1F4FF0C02C7041D:F473605D9705E36DF348FEBD04840615" box="[555,714,294,319]" country="Brazil" municipality="Sao Bento do Sul" name="Rio Vermelho" pageId="6" pageNumber="237" stateProvince="Santa Catarina">Rio Vermelho</location>
,
<collectingDate id="9556E9AE9705E36DF3B6FEBD05530615" box="[725,797,294,319]" pageId="6" pageNumber="237" value="1958-01">
<date id="851210469705E36DF3B6FEBD05530615" box="[725,797,294,319]" pageId="6" pageNumber="237" value="1958-01">i.1958</date>
</collectingDate>
,
<collectorName id="5C5953509705E36DF24AFEBD05330615" box="[809,893,294,319]" pageId="6" pageNumber="237">Dirings</collectorName>
coll. (
<collectionCode id="97BDAE439705E36DF2A0FEBC025C0615" box="[963,1042,295,319]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="237">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="41C43CDB9705E36DF545FEBD03560649" ID-GBIF-Occurrence="1795578721" collectingDate="2011-11-05" collectingDateMax="2011-11-20" collectingDateMin="2011-11-05" collectionCode="MZSP" collectorName="Simeao Moraes &amp; Livia Pinheiro &amp; Glaucia Marconato" country="Brazil" location="Serra Bonita" municipality="Camacan" pageId="6" pageNumber="237" specimenCode="MOL 479" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<collectingRegion id="3368F8649705E36DF545FEBD02240615" box="[1062,1130,294,319]" country="Brazil" name="Bahia" pageId="6" pageNumber="237">Bahia</collectingRegion>
,
<collectingMunicipality id="1177ACFC9705E36DF516FEBC02AE0615" box="[1141,1248,295,319]" pageId="6" pageNumber="237">Camacan</collectingMunicipality>
,
<location id="F473605D9705E36DF588FEBC03370615" LSID="urn:lsid:plazi:treatment:790587909705E36CF1F4FF0C02C7041D:F473605D9705E36DF588FEBC03370615" box="[1259,1401,294,319]" country="Brazil" municipality="Camacan" name="Serra Bonita" pageId="6" pageNumber="237" stateProvince="Bahia">Serra Bonita</location>
,
<collectingDate id="9556E9AE9705E36DF4E6FEBD075E0649" pageId="6" pageNumber="237" value="2011-11-05" valueMax="2011-11-20" valueMin="2011-11-05">
<date id="851210469705E36DF4E6FEBD075E0649" pageId="6" pageNumber="237" value="2011-11-05" valueMax="2011-11-20" valueMin="2011-11-05">5- 20.xi.2011</date>
</collectingDate>
,
<collectorName id="5C5953509705E36DF07FFED107830648" box="[284,461,330,355]" pageId="6" pageNumber="237">Simeão Moraes</collectorName>
,
<collectorName id="5C5953509705E36DF0BBFED104340649" box="[472,634,330,355]" pageId="6" pageNumber="237">Lívia Pinheiro</collectorName>
&amp;
<collectorName id="5C5953509705E36DF3FEFED1053A0648" box="[669,884,330,355]" pageId="6" pageNumber="237">Gláucia Marconato</collectorName>
leg., MZ 25.789,
<specimenCode id="A10A9EFD9705E36DF55CFED002E00649" box="[1087,1198,331,355]" collectionCode="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="6" pageNumber="237">MOL 479</specimenCode>
(
<collectionCode id="97BDAE439705E36DF5DEFED003420649" box="[1213,1292,331,355]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="237">MZSP</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B9B6650D9705E36DF1A4FEF504CF0446" pageId="6" pageNumber="237" type="diagnosis">
<paragraph id="F11336869705E36DF1A4FEF505A30575" blockId="6.[151,1436,258,877]" pageId="6" pageNumber="237">
<emphasis id="C3D8EA949705E36DF1A4FEF5070F06AD" bold="true" box="[199,321,366,391]" pageId="6" pageNumber="237">Diagnosis.</emphasis>
Externally indistinguishable from
<taxonomicName id="36AC4D059705E36DF382FEF4051F06AC" box="[737,849,367,390]" class="Insecta" family="Erebidae" genus="Heliura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="venata">
<emphasis id="C3D8EA949705E36DF382FEF4051F06AC" box="[737,849,367,390]" italics="true" pageId="6" pageNumber="237">E. venata</emphasis>
</taxonomicName>
and
<taxonomicName id="36AC4D059705E36DF2F7FEF405BC06AC" authorityName="Schaus" authorityYear="1889" box="[916,1010,366,390]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="aelia">
<emphasis id="C3D8EA949705E36DF2F7FEF405BC06AC" box="[916,1010,366,390]" italics="true" pageId="6" pageNumber="237">E. aelia</emphasis>
</taxonomicName>
. Differences in genitalia that allow recognition of
<taxonomicName id="36AC4D059705E36DF05FFE08078E0681" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[316,448,403,427]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF05FFE08078E0681" box="[316,448,403,427]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF0A4FE09046F0681" box="[455,545,402,427]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF0A4FE09046F0681" bold="true" box="[455,545,402,427]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
when compared to
<taxonomicName id="36AC4D059705E36DF39CFE0805190681" authorityName="Schaus" authorityYear="1889" box="[767,855,403,427]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="aelia">
<emphasis id="C3D8EA949705E36DF39CFE0805190681" box="[767,855,403,427]" italics="true" pageId="6" pageNumber="237">E. aelia</emphasis>
</taxonomicName>
are: cornuti decreasing dramatically in size distally in
<taxonomicName id="36AC4D059705E36DF1D7FE2C074206E4" authorityName="Schaus" authorityYear="1889" box="[180,268,438,462]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="aelia">
<emphasis id="C3D8EA949705E36DF1D7FE2C074206E4" box="[180,268,438,462]" italics="true" pageId="6" pageNumber="237">E. aelia</emphasis>
</taxonomicName>
, and only slightly in
<taxonomicName id="36AC4D059705E36DF363FE2C04CB06E4" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[512,645,438,462]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="C3D8EA949705E36DF363FE2C04CB06E4" box="[512,645,438,462]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
. Saccus more developed and markedly more produced in
<taxonomicName id="36AC4D059705E36DF47BFE2C03D206E4" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[1304,1436,438,462]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF47BFE2C03D206E4" box="[1304,1436,438,462]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF1F4FE4106BC06D9" box="[151,242,474,499]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF1F4FE4106BC06D9" bold="true" box="[151,242,474,499]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
Valvae with three lobes in
<taxonomicName id="36AC4D059705E36DF345FE40043306D9" authorityName="Schaus" authorityYear="1889" box="[550,637,475,499]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="aelia">
<emphasis id="C3D8EA949705E36DF345FE40043306D9" box="[550,637,475,499]" italics="true" pageId="6" pageNumber="237">E. aelia</emphasis>
</taxonomicName>
, but only two in
<taxonomicName id="36AC4D059705E36DF25CFE40058D06D9" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[831,963,475,499]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF25CFE40058D06D9" box="[831,963,475,499]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF2A9FE41026A06D9" box="[970,1060,474,499]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF2A9FE41026A06D9" bold="true" box="[970,1060,474,499]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="36AC4D059705E36DF548FE4002AF06D9" authorityName="Schaus" authorityYear="1889" box="[1067,1249,475,499]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="aelia">
<emphasis id="C3D8EA949705E36DF548FE4002AF06D9" box="[1067,1249,475,499]" italics="true" pageId="6" pageNumber="237">Episcepsis aelia</emphasis>
</taxonomicName>
has the cucullus shorter than valvula, while the opposite happens in
<taxonomicName id="36AC4D059705E36DF3BDFE64052C053C" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[734,866,510,534]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF3BDFE64052C053C" box="[734,866,510,534]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF20AFE65058A053D" box="[873,964,510,535]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF20AFE65058A053D" bold="true" box="[873,964,510,535]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
In female genitalia, both species differ in the following characters: corpus bursae not as twisted or corrugated in
<taxonomicName id="36AC4D059705E36DF2C8FDB802630511" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[939,1069,547,571]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF2C8FDB802630511" box="[939,1069,547,571]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF550FDB902C20511" box="[1075,1164,546,571]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF550FDB902C20511" bold="true" box="[1075,1164,546,571]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
as in
<taxonomicName id="36AC4D059705E36DF5AFFDB8036D0511" authorityName="Schaus" authorityYear="1889" box="[1228,1315,547,571]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="aelia">
<emphasis id="C3D8EA949705E36DF5AFFDB8036D0511" box="[1228,1315,547,571]" italics="true" pageId="6" pageNumber="237">E. aelia</emphasis>
</taxonomicName>
. The latter species lacks signa, while there are few ventral signa in
<taxonomicName id="36AC4D059705E36DF269FDDC05C30574" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[778,909,582,606]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF269FDDC05C30574" box="[778,909,582,606]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF2F7FDDD05A30575" box="[916,1005,582,607]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF2F7FDDD05A30575" bold="true" box="[916,1005,582,607]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="F11336869705E36DF1A4FDF104CF0446" blockId="6.[151,1436,258,877]" pageId="6" pageNumber="237">
When compared to
<taxonomicName id="36AC4D059705E36DF0C1FDF0044205A8" box="[418,524,619,642]" class="Insecta" family="Erebidae" genus="Heliura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="venata">
<emphasis id="C3D8EA949705E36DF0C1FDF0044205A8" box="[418,524,619,642]" italics="true" pageId="6" pageNumber="237">E. venata</emphasis>
</taxonomicName>
, the species differ in the following aspects: caecum slightly curved upwards in
<taxonomicName id="36AC4D059705E36DF4E7FDF006AC058E" class="Insecta" family="Erebidae" genus="Heliura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="venata">
<emphasis id="C3D8EA949705E36DF4E7FDF006AC058E" italics="true" pageId="6" pageNumber="237">E. venata</emphasis>
</taxonomicName>
, and straight in
<taxonomicName id="36AC4D059705E36DF0FAFD160453058E" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[409,541,652,676]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF0FAFD160453058E" box="[409,541,652,676]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF347FD170430058F" box="[548,638,652,677]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF347FD170430058F" bold="true" box="[548,638,652,677]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
Saccus developed and with apex almost sharp in
<taxonomicName id="36AC4D059705E36DF5D3FD16037D058E" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[1200,1331,652,676]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF5D3FD16037D058E" box="[1200,1331,652,676]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF458FD1703DB058F" box="[1339,1429,652,677]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF458FD1703DB058F" bold="true" box="[1339,1429,652,677]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
, and only weakly developed, with round apex and dorsally curved in
<taxonomicName id="36AC4D059705E36DF2DBFD35026B05EF" box="[952,1061,686,709]" class="Insecta" family="Erebidae" genus="Heliura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="venata">
<emphasis id="C3D8EA949705E36DF2DBFD35026B05EF" box="[952,1061,686,709]" italics="true" pageId="6" pageNumber="237">E. venata</emphasis>
</taxonomicName>
. Tegumen densely covered with scales only in
<taxonomicName id="36AC4D059705E36DF054FD5407F405CC" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[311,442,718,742]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF054FD5407F405CC" box="[311,442,718,742]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF0A2FD55045505CC" box="[449,539,718,743]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF0A2FD55045A05CD" bold="true" box="[449,532,718,743]" pageId="6" pageNumber="237">sp. nov</emphasis>
.
</taxonomicNameLabel>
<taxonomicName id="36AC4D059705E36DF341FD5404A505CC" authorityName="Butler" authorityYear="1877" box="[546,747,719,742]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="venata">
<emphasis id="C3D8EA949705E36DF341FD5404A505CC" box="[546,747,719,742]" italics="true" pageId="6" pageNumber="237">Episcepsis venata</emphasis>
</taxonomicName>
also has the cucullus shorter than valvula, while the opposite happens in
<taxonomicName id="36AC4D059705E36DF074FD6A07D40422" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[279,410,752,776]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF074FD6A07D40422" box="[279,410,752,776]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF0C2FD7407B50422" box="[417,507,751,776]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF0C2FD7407B50422" bold="true" box="[417,507,751,776]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
In female genitalia, we found the following differences: Dorsal surface of papillae anales with few setae in
<taxonomicName id="36AC4D059705E36DF0D5FC89046D0403" authorityName="Butler" authorityYear="1877" box="[438,547,786,809]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="venata">
<emphasis id="C3D8EA949705E36DF0D5FC89046D0403" box="[438,547,786,809]" italics="true" pageId="6" pageNumber="237">E. venata</emphasis>
</taxonomicName>
and lacking setae in
<taxonomicName id="36AC4D059705E36DF279FC8905EE0403" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[794,928,785,809]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF279FC8905EE0403" box="[794,928,785,809]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF2CAFC8A024B0400" box="[937,1029,785,810]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF2CAFC8A024B0400" bold="true" box="[937,1029,785,810]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
Antevaginal lamella with posterior margin sinuous only in
<taxonomicName id="36AC4D059705E36DF0C7FCA804410460" authorityName="Butler" authorityYear="1877" box="[420,527,819,842]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="venata">
<emphasis id="C3D8EA949705E36DF0C7FCA804410460" box="[420,527,819,842]" italics="true" pageId="6" pageNumber="237">E. venata</emphasis>
</taxonomicName>
. Ductus seminalis arising from ductus bursae on the left side in
<taxonomicName id="36AC4D059705E36DF58DFCA8033C0461" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[1262,1394,819,843]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF58DFCA8033C0461" box="[1262,1394,819,843]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF419FCA906860447" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF419FCA906860447" bold="true" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
and from accessory bursa in
<taxonomicName id="36AC4D059705E36DF373FCCE04340446" authorityName="Butler" authorityYear="1877" box="[528,634,853,876]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="venata">
<emphasis id="C3D8EA949705E36DF373FCCE04340446" box="[528,634,853,876]" italics="true" pageId="6" pageNumber="237">E. venata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="A5D3660E9705E36DF1F4F93704B501D5" httpUri="https://zenodo.org/record/1117592/files/figure.png" pageId="6" pageNumber="237" targetBox="[151,1436,895,1686]" targetPageId="6">
<paragraph id="F11336869705E36DF1F4F93704B501D5" blockId="6.[151,1435,1708,1791]" pageId="6" pageNumber="237">
<emphasis id="C3D8EA949705E36DF1F4F937075B01EB" bold="true" box="[151,277,1708,1730]" pageId="6" pageNumber="237">FIGURE 5.</emphasis>
Male and female genitalia of
<taxonomicName id="36AC4D059705E36DF331F936048401E8" authority="Pinheiro &amp; Araujo Junior, 2017" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[594,714,1708,1730]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
<emphasis id="C3D8EA949705E36DF331F936048401E8" box="[594,714,1708,1730]" italics="true" pageId="6" pageNumber="237">E. atlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8EB57EF9705E36DF3B0F937056B01EB" box="[723,805,1708,1729]" pageId="6" pageNumber="237" rank="species">
<emphasis id="C3D8EA949705E36DF3B0F937056B01EB" bold="true" box="[723,805,1708,1729]" pageId="6" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
A) Male abdominal pelt; B) Male genitalia, dorsal view; C) Same, lateral view; D) Same, posterior view; E) Aedeagus; F) Female abdominal pelt; G) Female genitalia, dorsal view; H) Same, lateral view; I) Same, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="B9B6650D9705E36CF1A4F8BD02AF05DA" lastPageId="7" lastPageNumber="238" pageId="6" pageNumber="237" type="description">
<paragraph id="F11336869705E36CF1A4F8BD02AF05DA" blockId="6.[151,1436,1830,2035]" lastBlockId="7.[151,1437,151,824]" lastPageId="7" lastPageNumber="238" pageId="6" pageNumber="237">
<emphasis id="C3D8EA949705E36DF1A4F8BD041A0015" bold="true" box="[199,596,1830,1855]" pageId="6" pageNumber="237">
Description
<specimenCount id="E7AAFD0F9705E36DF038F8BD073E0015" box="[347,368,1830,1855]" pageId="6" pageNumber="237" type="male">♂</specimenCount>
and
<specimenCount id="E7AAFD0F9705E36DF0D1F8BD07860015" box="[434,456,1830,1855]" pageId="6" pageNumber="237" type="female">♀</specimenCount>
. Abdomen.
</emphasis>
T8 of males almost rectangular, as sclerotized as previous tergites, with anterior margin heavily sclerotized, with two membranous projections shorter than the length of T8. S8 of males less sclerotized than previous sternites, except for laterals of anterior margin, heavily sclerotized. Intersegmental membrane 7-8 lacking coremata. T7 of females rectangular, twice longer than previous tergites, and as sclerotized as them. S7 also rectangular and as sclerotized as previous sternites, but only slightly longer than S6.
<emphasis id="C3D8EA949705E36DF43CF82D074B00D9" bold="true" pageId="6" pageNumber="237">Male genitalia.</emphasis>
Aedeagus much longer than wide, straight. Ejaculatory duct inserted dorsally. Caecum round. Vesica as long as aedeagus, dorsal surface with a longitudinal line of cornuti, proximal cornuti long, and distal cornuti shorter. Ventral surface with a longitudinal sclerotized plate. Saccus developed, slightly asymmetrical, apex almost sharp. Vinculum straight, very narrow. Tegumen longer than wide, densely covered with scales. Anterior margin of tegumen with a somewhat deep, V-shaped invagination. Dorsal surface of tegumen with a triangular sclerotization. Lateral posterior surfaces of tegumen densely scaled. Posterior margin of tegumen straight. Intersegmental membrane 9-10 visible only dorsally. Uncus with an inverted Y-shape. Base of uncus glabrous, and much wider than its lobe. Lobe of uncus somewhat cylindrical, finger-like, with a few tiny setae on dorsal surface and a sharp apex. Transtilla membranous. Juxta sclerotized, glabrous, wider than long. Valvae symmetrical, with two lobes, a cucullus weakily sclerotized and densely covered with scales, with round apex, and longer than valvula, and a valvula heavily sclerotized with a few setae and a sharp apex.
<emphasis id="C3D8EA949704E36CF22FFE47025606DF" bold="true" box="[844,1048,476,501]" pageId="7" pageNumber="238">Female genitalia.</emphasis>
T9 uniformly sclerotized. Papillae anales with short setae on most of their surface. Dorsal surface lacking setae. Pseudopapillae anales conspicuous, finger-like, with short setae. Anterior apophyses almost the same length of posterior apophyses. Pheromone glands undeveloped. Postvaginal lamella predominantly sclerotized. Ventral portion of posterior margin membranous, with a highly sclerotized tubercle below. Antevaginal lamella slightly sclerotized, lacking setae or spines. Ostium and antrum centralized. Ductus bursae heavily sclerotized and flattened, very short. Corpus bursae approximately spherical, somewhat twisted, most of its surface lacking signa. Dorsal distal portion of corpus bursae corrugated. Left ventral side with a pair of signa. Ductus seminalis arising from ductus bursae on the left side.
</paragraph>
</subSubSection>
<subSubSection id="B9B6650D9704E36CF1A4FD67023A043F" box="[199,1140,764,789]" pageId="7" pageNumber="238" type="distribution">
<paragraph id="F11336869704E36CF1A4FD67023A043F" blockId="7.[151,1437,151,824]" box="[199,1140,764,789]" pageId="7" pageNumber="238">
<emphasis id="C3D8EA949704E36CF1A4FD67072E043F" bold="true" box="[199,352,764,789]" pageId="7" pageNumber="238">Distribution.</emphasis>
<taxonomicName id="36AC4D059704E36CF004FD660406043E" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[359,584,764,788]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="C3D8EA949704E36CF004FD660406043E" box="[359,584,764,788]" italics="true" pageId="7" pageNumber="238">Episcepsis atlantica</emphasis>
</taxonomicName>
is known only from the Atlantic Forest of
<collectingCountry id="89BB76169704E36CF54BFD67023E043F" box="[1064,1136,764,789]" name="Brazil" pageId="7" pageNumber="238">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B9B6650D9704E36CF1A4FC8402C7041D" box="[199,1161,799,824]" pageId="7" pageNumber="238" type="etymology">
<paragraph id="F11336869704E36CF1A4FC8402C7041D" blockId="7.[151,1437,151,824]" box="[199,1161,799,824]" pageId="7" pageNumber="238">
<emphasis id="C3D8EA949704E36CF1A4FC8407000412" bold="true" box="[199,334,799,824]" pageId="7" pageNumber="238">Etymology.</emphasis>
The specific epithet
<taxonomicName id="36AC4D059704E36CF35BFCBB04D20412" authorityName="Pinheiro &amp; Araujo Junior" authorityYear="2017" box="[568,668,800,824]" class="Insecta" family="Erebidae" genus="Episcepsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="C3D8EA949704E36CF35BFCBB04D20412" box="[568,668,800,824]" italics="true" pageId="7" pageNumber="238">atlantica</emphasis>
</taxonomicName>
refers to the biome where this moth occurs.
</paragraph>
</subSubSection>
</treatment>
</document>
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