treatments-xml/data/61/2B/87/612B87FCACDC43F30B83F8CD8C0F07C8.xml

<document id="63A502CB92773761F4A36C8131763EC0" ID-DOI="10.11646/zootaxa.5483.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13209307" ID-ZooBank="AF0272DB-5588-411D-9EAE-DED4785BF170" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722786113536" checkinUser="plazi" docAuthor="WOOD, T. J." docDate="2024" docId="612B87FCACDC43F30B83F8CD8C0F07C8" docLanguage="en" docName="zootaxa.5483.1.1.pdf" docOrigin="Zootaxa 5483 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5483.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Andrena risha WOOD, 2024, spec. nov." docType="treatment" docVersion="2" lastPageNumber="137" masterDocId="9D12FF84AC59437B0B14FFCB8E650641" masterDocTitle="Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)" masterLastPageNumber="150" masterPageNumber="1" pageNumber="134" updateTime="1722787353532" updateUser="ExternalLinkService">
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<mods:title id="038E6C15918A4E3183D0B5B952A078B1">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<mods:date id="B88DE5071B4EEE39B926393B768D964E">2024</mods:date>
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<treatment id="612B87FCACDC43F30B83F8CD8C0F07C8" ID-DOI="http://doi.org/10.5281/zenodo.13209521" ID-Zenodo-Dep="13209521" LSID="urn:lsid:plazi:treatment:612B87FCACDC43F30B83F8CD8C0F07C8" httpUri="http://treatment.plazi.org/id/612B87FCACDC43F30B83F8CD8C0F07C8" lastPageId="136" lastPageNumber="137" pageId="133" pageNumber="134">
<subSubSection id="A1986561ACDC43FE0B83F8CD8CE10160" box="[151,644,1798,1825]" pageId="133" pageNumber="134" type="nomenclature">
<paragraph id="E93D36EAACDC43FE0B83F8CD8CE10160" blockId="133.[151,644,1798,1861]" box="[151,644,1798,1825]" pageId="133" pageNumber="134">
<heading id="B2758186ACDC43FE0B83F8CD8CE10160" bold="true" box="[151,644,1798,1825]" fontSize="11" level="2" pageId="133" pageNumber="134" reason="4">
<emphasis id="DBF6EAF8ACDC43FE0B83F8CD8CE10160" bold="true" box="[151,644,1798,1825]" pageId="133" pageNumber="134">
<taxonomicName id="2E824D69ACDC43FE0B83F8CD8C600160" authorityName="WOOD" authorityYear="2024" box="[151,517,1798,1825]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="risha" status="sp. nov.">
<emphasis id="DBF6EAF8ACDC43FE0B83F8CD8F640160" bold="true" box="[151,257,1798,1825]" italics="true" pageId="133" pageNumber="134">Andrena</emphasis>
(
<emphasis id="DBF6EAF8ACDC43FE0A06F8CC8FD30160" bold="true" box="[274,438,1798,1825]" italics="true" pageId="133" pageNumber="134">incertae sedis</emphasis>
) 
<emphasis id="DBF6EAF8ACDC43FE0AD3F8CD8C600160" bold="true" box="[455,517,1798,1825]" italics="true" pageId="133" pageNumber="134">risha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783ACDC43FE0918F8CC8CE10160" box="[524,644,1799,1825]" pageId="133" pageNumber="134" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACDC43FE0B83F8E08F290104" box="[151,332,1835,1861]" pageId="133" pageNumber="134" type="description">
<paragraph id="E93D36EAACDC43FE0B83F8E08F290104" blockId="133.[151,644,1798,1861]" box="[151,332,1835,1861]" pageId="133" pageNumber="134">
<figureCitation id="71B92A6FACDC43FE0B83F8E08F2D0104" box="[151,328,1835,1861]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="133" pageNumber="134">Figures 81A–H</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACDC43FE0B83F8B98B4901CE" box="[151,1324,1906,1935]" pageId="133" pageNumber="134" type="materials_examined">
<paragraph id="E93D36EAACDC43FE0B83F8B98B4901CE" blockId="133.[151,1436,1906,2041]" box="[151,1324,1906,1935]" pageId="133" pageNumber="134">
<materialsCitation id="59EA3CB7ACDC43FE0B83F8B98B4901CE" box="[151,1324,1906,1935]" collectingDate="1960-03-08" collectorName="J. Barbier" country="Algeria" location="Oran" pageId="133" pageNumber="134" specimenCount="1" specimenCount-female="1" stateProvince="Oran" typeStatus="holotype">
<emphasis id="DBF6EAF8ACDC43FE0B83F8B98FB401CD" bold="true" box="[151,465,1906,1932]" pageId="133" pageNumber="134">
<typeStatus id="36398848ACDC43FE0B83F8B98F5F01CD" box="[151,314,1906,1932]" pageId="133" pageNumber="134" type="holotype">HOLOTYPE</typeStatus>
: 
<collectingCountry id="9195767AACDC43FE0A51F8B98FA901CD" box="[325,460,1906,1932]" name="Algeria" pageId="133" pageNumber="134">ALGERIA</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808ACDC43FE0ACCF8B88C7501CC" box="[472,528,1907,1933]" country="Algeria" name="Oran" pageId="133" pageNumber="134">Oran</collectingRegion>
[inferred 35.7072 
<superScript id="1EF79BA2ACDC43FE09CFF8B88C9F01CC" attach="right" box="[731,762,1907,1933]" fontSize="6" pageId="133" pageNumber="134">oN</superScript>
, -0.6646 
<superScript id="1EF79BA2ACDC43FE084FF8B88D1D01CC" attach="right" box="[859,888,1907,1933]" fontSize="6" pageId="133" pageNumber="134">oE</superScript>
], 
<date id="9D3C102AACDC43FE0898F8B88D9F01CC" box="[908,1018,1907,1933]" pageId="133" pageNumber="134" value="1960-03-08">
<collectingDate id="8D78E9C2ACDC43FE0898F8B88D9F01CC" box="[908,1018,1907,1933]" pageId="133" pageNumber="134" value="1960-03-08">8.iii.1960</collectingDate>
</date>
, 
<specimenCount id="FF84FD63ACDC43FE0F11F8B88A7F01CC" box="[1029,1050,1907,1933]" count="1" pageId="133" pageNumber="134" type="female">♀</specimenCount>
, leg. 
<collectorName id="4477533CACDC43FE0F4CF8B88AA201CC" box="[1112,1223,1907,1933]" pageId="133" pageNumber="134">J. Barbier</collectorName>
, OÖLM.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACDC43FD0BD3F85D8FED0218" lastPageId="134" lastPageNumber="135" pageId="133" pageNumber="134" type="description">
<paragraph id="E93D36EAACDC43FD0BD3F85D8A230724" blockId="133.[151,1436,1906,2041]" lastBlockId="134.[151,1437,151,2013]" lastPageId="134" lastPageNumber="135" pageId="133" pageNumber="134">
<emphasis id="DBF6EAF8ACDC43FE0BD3F85D8FA701F0" bold="true" box="[199,450,1942,1969]" pageId="133" pageNumber="134">Description. Female.</emphasis>
Body length: 9.5 mm (
<figureCitation id="71B92A6FACDC43FE09CCF85C8D0701F0" box="[728,866,1943,1969]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="133" pageNumber="134">Figure 81A</figureCitation>
). 
<emphasis id="DBF6EAF8ACDC43FE086CF85C8DDB01F0" box="[888,958,1943,1969]" italics="true" pageId="133" pageNumber="134">Head:</emphasis>
Dark, 1.3 times wider than long (
<figureCitation id="71B92A6FACDC43FE0E45F85C8EAC0194" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="133" pageNumber="134">Figure 81B</figureCitation>
). Clypeus domed, densely but shallowly punctate, punctures separated by 0.5–1 puncture diameters, with hint of weak impunctate mid-line, underlying surface shagreened, weakly shining. Process of labrum rounded trapezoidal, slightly wider than long, surface shining (
<figureCitation id="71B92A6FACDF43FD0817FF5C8DEF06F0" box="[771,906,151,177]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="134" pageNumber="135">Figure 81C</figureCitation>
). Gena slightly exceeding width of compound eye; ocelloccipital distance 1.5 times diameter of lateral ocellus. Foveae moderately broad, occupying slightly more than malar areas space between compound eye and lateral ocellus, ventrally slightly passing lower margin of antennal insertions; foveae filled with dark brown hairs. Head covered with dark brow to light brown plumose hairs, hairs lightest around antennal insertions and on frons, none equalling length of scape. Antennae dark, A5–12 ventrally lightened by presence of grey scales; A3 exceeding A4, shorter than A4+5.
</paragraph>
<paragraph id="E93D36EAACDF43FD0BD3FEA48C4D0578" blockId="134.[151,1437,151,2013]" pageId="134" pageNumber="135">
<emphasis id="DBF6EAF8ACDF43FD0BD3FEA48F2D07C9" box="[199,328,367,392]" italics="true" pageId="134" pageNumber="135">
<taxonomicName id="2E824D69ACDF43FD0BD3FEA48F2707C9" box="[199,322,367,392]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="134" pageNumber="135" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
:
</emphasis>
Scutum and scutellum strongly microreticulate and predominantly dull, with obscure punctures disappearing into underlying sculpture (
<figureCitation id="71B92A6FACDF43FD0959FE588CB407EC" box="[589,721,403,429]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="134" pageNumber="135">Figure 81D</figureCitation>
). Pronotum with weak hint of humeral angle. Mesepisternum and dorsolateral part of propodeum with fine and uniform granular shagreen, weakly shining; propodeal triangle finely delineated by fine rim, internal surface with fine and uniform granular shagreen, basally with short raised rugae, propodeal triangle therefore poorly differentiated from remaining parts of propodeum (
<figureCitation id="71B92A6FACDF43FD0F4FFE348AB80458" box="[1115,1245,511,537]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="134" pageNumber="135">Figure 81E</figureCitation>
). Mesepisternum with long, densely plumose light brown hairs, becoming shorter and darker brown on scutum and scutellum (
<figureCitation id="71B92A6FACDF43FD0E45FDE88EA30420" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="134" pageNumber="135">Figure 81F</figureCitation>
). Propodeal corbicula complete, dorsal and anterior fringes composed of long densely plumose light brown hairs, internal surface with sparse, long, simple light brown hairs. Legs dark, apical tarsal segments lightened orange-brown; fore and mid femorae with long fringes of plumose light brown hairs, remaining legs with light to dark brown pubescence. Flocculus complete, composed of light brown plumose hairs; femoral scopae composed of mixture of light brown simple and plumose hairs (
<figureCitation id="71B92A6FACDF43FD09FEFD1C8D1504B0" box="[746,880,727,753]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="134" pageNumber="135">Figure 81G</figureCitation>
). Tibial scopae composed of simple hair, dorsally dark brown to black, ventrally light brown. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation orange, nervulus weakly antefurcal.
</paragraph>
<paragraph id="E93D36EAACDF43FD0BD3FC888FB50274" blockId="134.[151,1437,151,2013]" pageId="134" pageNumber="135">
<emphasis id="DBF6EAF8ACDF43FD0BD3FC888F20051D" box="[199,325,835,860]" italics="true" pageId="134" pageNumber="135">Metasoma:</emphasis>
Terga dark, T1–2 with marginal areas broadly lightened hyaline brown (
<figureCitation id="71B92A6FACDF43FD0F6FFC888B67051C" box="[1147,1282,835,861]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="134" pageNumber="135">Figure 81H</figureCitation>
). Tergal discs strongly shagreened, weakly shining; surface weakly punctate, punctures separated by 1–2 puncture diameters, disappearing into underlying sculpture. Marginal areas of T2–4 weakly but distinctly depressed. Discs of T1–4 with sparse brown hairs, longest on T1, becoming progressively shorter on subsequent discs; T1–4 with weak apical hairbands, not obscuring underlying surface. Apical fringe of T5 and hairs flanking pygidial plate dark brown. Pygidial plate rounded triangular, apical rim slightly raised, internal surface densely punctate, punctures confluent, interspaces slightly shining.
</paragraph>
<paragraph id="E93D36EAACDF43FD0BD3FBF48FED0218" blockId="134.[151,1437,151,2013]" box="[199,392,1087,1113]" pageId="134" pageNumber="135">
<emphasis id="DBF6EAF8ACDF43FD0BD3FBF48F6E0218" bold="true" box="[199,267,1087,1113]" pageId="134" pageNumber="135">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACDF43F30BD3FBA98D9C075C" lastPageId="136" lastPageNumber="137" pageId="134" pageNumber="135" type="diagnosis">
<paragraph id="E93D36EAACDF43FD0BD3FBA98AE50034" blockId="134.[151,1437,151,2013]" pageId="134" pageNumber="135">
<emphasis id="DBF6EAF8ACDF43FD0BD3FBA98F27023D" bold="true" box="[199,322,1122,1148]" pageId="134" pageNumber="135">Diagnosis.</emphasis>
<taxonomicName id="2E824D69ACDF43FD0A5CFBA88F8C023C" authorityName="WOOD" authorityYear="2024" box="[328,489,1123,1149]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="risha">
<emphasis id="DBF6EAF8ACDF43FD0A5CFBA88F8C023C" box="[328,489,1123,1149]" italics="true" pageId="134" pageNumber="135">Andrena risha</emphasis>
</taxonomicName>
presents a combination of characters that prevent confident placement in an existing subgenus. It is morphologically closest to 
<emphasis id="DBF6EAF8ACDF43FD0994FB4C8CFD02E0" box="[640,664,1159,1185]" italics="true" pageId="134" pageNumber="135">A.</emphasis>
(
<emphasis id="DBF6EAF8ACDF43FD09BFFB4C8D5102E0" box="[683,820,1159,1185]" italics="true" pageId="134" pageNumber="135">
? 
<taxonomicName id="2E824D69ACDF43FD09ADFB4C8D5102E0" authorityName="Hedicke" authorityYear="1933" box="[697,820,1159,1185]" pageId="134" pageNumber="150" rank="subGenus" subGenus="Euandrena">Euandrena</taxonomicName>
</emphasis>
) 
<taxonomicName id="2E824D69ACDF43FD0853FB4C8A5502E0" authority="Wood, 2022" authorityName="Wood" authorityYear="2022" box="[839,1072,1159,1185]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DBF6EAF8ACDF43FD0853FB4C8DFE02E1" box="[839,923,1159,1184]" italics="true" pageId="134" pageNumber="135">ramosa</emphasis>
Wood, 2022
</taxonomicName>
(
<collectingCountry id="9195767AACDF43FD0F55FB4C8AE702E0" box="[1089,1154,1159,1185]" name="Spain" pageId="134" pageNumber="135">Spain</collectingCountry>
and 
<collectingCountry id="9195767AACDF43FD0FABFB4C8B4502E0" box="[1215,1312,1159,1185]" name="Portugal" pageId="134" pageNumber="135">Portugal</collectingCountry>
) and more broadly falls into the broad 
<emphasis id="DBF6EAF8ACDF43FD0AF7FB608D1B0284" box="[483,894,1195,1221]" italics="true" pageId="134" pageNumber="135">
<taxonomicName id="2E824D69ACDF43FD0AF7FB608C3B0284" authorityName="Hedicke" authorityYear="1933" box="[483,606,1195,1221]" pageId="134" pageNumber="150" rank="subGenus" subGenus="Euandrena">Euandrena</taxonomicName>
-Margandrena-crocusella
</emphasis>
-group derived part of the 
<taxonomicName id="2E824D69ACDF43FD0FAFFB608B7F0284" box="[1211,1306,1195,1221]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8ACDF43FD0FAFFB608B7F0284" box="[1211,1306,1195,1221]" italics="true" pageId="134" pageNumber="135">Andrena</emphasis>
</taxonomicName>
phylogeny (
<bibRefCitation id="8D134B1BACDF43FD0B8BFB048FE102A8" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." box="[159,388,1231,1257]" pageId="134" pageNumber="135" refId="ref90124" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2022 b) Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution, 170, 107151. https: // doi. org / 10.1016 / j. ympev. 2021.107151" type="journal volume" year="2022">
Pisanty 
<emphasis id="DBF6EAF8ACDF43FD0BEFFB048F4802A8" box="[251,301,1231,1257]" italics="true" pageId="134" pageNumber="135">et al</emphasis>
. 2022b
</bibRefCitation>
) due to the pronotum with a very weak humeral angle; propodeum with almost no sculpture (
<figureCitation id="71B92A6FACDF43FD0B8BFB388F45034C" box="[159,288,1267,1293]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="134" pageNumber="135">Figure 81E</figureCitation>
), propodeal triangle thus subtly delineated; facial foveae relatively broad (occupying slightly more than ½ space between compound eye and lateral ocellus) and not narrowing ventrally; clypeus domed; A3 exceeding A 
<quantity id="2E7A9B0FACDF43FD0BB8FAF08EBC0314" box="[172,217,1339,1365]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="134" pageNumber="135" unit="in" value="4.0">4 in</quantity>
length, shorter than A4+5; pollen collecting hairs complex, flocculus composed of strongly plumose hairs (
<figureCitation id="71B92A6FACDF43FD0B8BFA948F420338" box="[159,295,1375,1401]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="134" pageNumber="135">Figure 81G</figureCitation>
); strongly plumose hairs present also on femoral scopae, ventral margins of fore and mid femorae, mesepisternum, and comprising propodeal corbiculae, propodeal corbiculae essentially complete; hind tibial scopae composed of simple hairs; and absence of other defining characters. The presence of abundant strongly plumose hairs that contrast the simple hairs of the tibial scopae (
<figureCitation id="71B92A6FACDF43FD0836FA008DB703A4" box="[802,978,1483,1509]" captionStart="FIGURE 80" captionStartId="132.[151,250,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-123@132.[151,1436,381,1823]" captionTargetPageId="132" captionText="FIGURE 80. Andrena (incertae sedis) hakkariensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Head, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209487" httpUri="https://zenodo.org/record/13209487/files/figure.png" pageId="134" pageNumber="135">Figures 80F–G</figureCitation>
) are strongly reminiscent of 
<taxonomicName id="2E824D69ACDF43FD0E32FA008BF903A5" authorityName="Wood" authorityYear="2022" box="[1318,1436,1483,1509]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DBF6EAF8ACDF43FD0E32FA008BF903A5" box="[1318,1436,1483,1509]" italics="true" pageId="134" pageNumber="135">A. ramosa</emphasis>
</taxonomicName>
which is uncertainly placed close to the subgenus 
<taxonomicName id="2E824D69ACDF43FD09CFFA248D330048" authorityName="Hedicke" authorityYear="1933" box="[731,854,1519,1545]" pageId="134" pageNumber="135" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8ACDF43FD09CFFA248D330048" box="[731,854,1519,1545]" italics="true" pageId="134" pageNumber="135">Euandrena</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="8D134B1BACDF43FD0882FA248A480048" author="Wood, T. J." box="[918,1069,1519,1545]" pageId="134" pageNumber="135" pagination="241 - 484" refId="ref92391" refString="Wood, T. J. (2023 b) The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera: Andrenidae). Journal of Hymenoptera Research, 96, 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood 2023b</bibRefCitation>
). The weak humeral angle, lack of sculpture on the propodeum, and relatively broad facial foveae that do not narrow ventrally make it difficult to place it in the subgenus 
<taxonomicName id="2E824D69ACDF43FD0AB9F9FC8C4D0010" authorityName="Hedicke" authorityYear="1933" box="[429,552,1591,1617]" pageId="134" pageNumber="135" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8ACDF43FD0AB9F9FC8C4D0010" box="[429,552,1591,1617]" italics="true" pageId="134" pageNumber="135">Euandrena</emphasis>
</taxonomicName>
with any confidence, as these characters are most closely associated with the 
<emphasis id="DBF6EAF8ACDF43FD0B83F9908F6E0034" box="[151,267,1627,1653]" italics="true" pageId="134" pageNumber="135">crocusella</emphasis>
-group. 
<taxonomicName id="2E824D69ACDF43FD0A77F9908C610034" authorityName="WOOD" authorityYear="2024" box="[355,516,1627,1653]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="risha">
<emphasis id="DBF6EAF8ACDF43FD0A77F9908C610034" box="[355,516,1627,1653]" italics="true" pageId="134" pageNumber="135">Andrena risha</emphasis>
</taxonomicName>
is therefore best left as 
<emphasis id="DBF6EAF8ACDF43FD081BF9908DCF0034" box="[783,938,1627,1653]" italics="true" pageId="134" pageNumber="135">incertae sedis</emphasis>
for the time being.
</paragraph>
<paragraph id="E93D36EAACDF43F30BD3F9B48D9C075C" blockId="134.[151,1437,151,2013]" lastBlockId="136.[151,1436,151,393]" lastPageId="136" lastPageNumber="137" pageId="134" pageNumber="135">
In direct comparison to 
<taxonomicName id="2E824D69ACDF43FD0ACEF9B48C2A00D9" authorityName="Wood" authorityYear="2022" box="[474,591,1663,1689]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DBF6EAF8ACDF43FD0ACEF9B48C2A00D9" box="[474,591,1663,1689]" italics="true" pageId="134" pageNumber="135">A. ramosa</emphasis>
</taxonomicName>
(characters of 
<taxonomicName id="2E824D69ACDF43FD09EFF9B48D1500D9" authorityName="Wood" authorityYear="2022" box="[763,880,1663,1689]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DBF6EAF8ACDF43FD09EFF9B48D1500D9" box="[763,880,1663,1689]" italics="true" pageId="134" pageNumber="135">A. ramosa</emphasis>
</taxonomicName>
illustrated in Wood 
<emphasis id="DBF6EAF8ACDF43FD0F4FF9B48AEB00D8" box="[1115,1166,1663,1689]" italics="true" pageId="134" pageNumber="135">et al</emphasis>
. 2022), 
<taxonomicName id="2E824D69ACDF43FD0FFBF9B48B2F00D8" authorityName="WOOD" authorityYear="2024" box="[1263,1354,1663,1689]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="risha">
<emphasis id="DBF6EAF8ACDF43FD0FFBF9B48B2F00D8" box="[1263,1354,1663,1689]" italics="true" pageId="134" pageNumber="135">A. risha</emphasis>
</taxonomicName>
can be separated due to the process of the labrum which is rounded trapezoidal, only slightly wider than long (
<figureCitation id="71B92A6FACDF43FD0E00F9688BFD00FC" box="[1300,1432,1699,1725]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="134" pageNumber="135">Figure 81C</figureCitation>
; in 
<taxonomicName id="2E824D69ACDF43FD0BA7F90C8F4000A1" authorityName="Wood" authorityYear="2022" box="[179,293,1735,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DBF6EAF8ACDF43FD0BA7F90C8F4000A1" box="[179,293,1735,1761]" italics="true" pageId="134" pageNumber="135">A. ramosa</emphasis>
</taxonomicName>
with the process of the labrum broad, 3 times wider than long), the clypeus which is evenly shagreened and weakly shining over its entire surface (in 
<taxonomicName id="2E824D69ACDF43FD0993F9208C9D0145" authorityName="Wood" authorityYear="2022" box="[647,760,1771,1797]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DBF6EAF8ACDF43FD0993F9208C9D0145" box="[647,760,1771,1797]" italics="true" pageId="134" pageNumber="135">A. ramosa</emphasis>
</taxonomicName>
the clypeus is shagreened laterally and basally, dull, shagreen becoming weaker and almost disappearing centrally, clypeus here almost smooth and shining), foveae occupying slightly more than ½ of the space between the compound eye and lateral ocellus (in 
<taxonomicName id="2E824D69ACDF43FD0F7FF8F88A84010D" authorityName="Wood" authorityYear="2022" box="[1131,1249,1843,1869]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DBF6EAF8ACDF43FD0F7FF8F88A84010D" box="[1131,1249,1843,1869]" italics="true" pageId="134" pageNumber="135">A. ramosa</emphasis>
</taxonomicName>
with the foveae occupying slightly less than ½ of the space between the compound eye and lateral ocellus), pronotum with weak humeral angle (in 
<taxonomicName id="2E824D69ACDF43FD0A77F8B08FB301D5" authorityName="Wood" authorityYear="2022" box="[355,470,1915,1941]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DBF6EAF8ACDF43FD0A77F8B08FB301D5" box="[355,470,1915,1941]" italics="true" pageId="134" pageNumber="135">A. ramosa</emphasis>
</taxonomicName>
with the pronotum rounded), scutum strongly microreticulate and predominantly dull, with obscure punctures disappearing into underlying sculpture (
<figureCitation id="71B92A6FACDF43FD0847F8548DB401F8" box="[851,977,1951,1977]" captionStart="FIGURE 78" captionStartId="127.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@127.[214,1373,181,1919]" captionTargetPageId="127" captionText="FIGURE 78. Andrena (incertae sedis) contraria spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209479" httpUri="https://zenodo.org/record/13209479/files/figure.png" pageId="134" pageNumber="135">Figure78D</figureCitation>
; in 
<taxonomicName id="2E824D69ACDF43FD08E1F8548A0301F9" authorityName="Wood" authorityYear="2022" box="[1013,1126,1951,1977]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DBF6EAF8ACDF43FD08E1F8548A0301F9" box="[1013,1126,1951,1977]" italics="true" pageId="134" pageNumber="135">A. ramosa</emphasis>
</taxonomicName>
with the scutum shagreened and dull to weakly shining but with clear shallow punctures, punctures separated by 1–3 puncture diameters), and the finely shagreened propodeum with minimal sculpture, propodeal triangle almost undifferentiated from dorsolateral parts of propodeum (
<figureCitation id="71B92A6FACD143F30A9EFF708C750694" box="[394,528,187,213]" captionStart="FIGURE 81" captionStartId="135.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@135.[214,1374,181,1919]" captionTargetPageId="135" captionText="FIGURE 81. Andrena (incertae sedis) risha spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Mesosoma, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="136" pageNumber="137">Figure 81E</figureCitation>
; in 
<taxonomicName id="2E824D69ACD143F3092AFF708CD10695" authorityName="Wood" authorityYear="2022" box="[574,692,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="DBF6EAF8ACD143F3092AFF708CD10695" box="[574,692,187,213]" italics="true" pageId="136" pageNumber="137">A. ramosa</emphasis>
</taxonomicName>
with clear different between propodeal triangle and dorsolateral parts of propodeum, these latter parts covered with granular microreticulation overlain by loose network of raised reticulation, sculpture thus contrasting smooth surface of propodeal triangle).
</paragraph>
</subSubSection>
<caption id="BDFD6662ACDE43FC0B83F85C8F1301B6" ID-DOI="http://doi.org/10.5281/zenodo.13209489" ID-Zenodo-Dep="13209489" httpUri="https://zenodo.org/record/13209489/files/figure.png" pageId="135" pageNumber="136" startId="135.[151,250,1943,1967]" targetBox="[214,1374,181,1919]" targetPageId="135" targetType="figure">
<paragraph id="E93D36EAACDE43FC0B83F85C8F1301B6" blockId="135.[151,1436,1943,2040]" pageId="135" pageNumber="136">
<emphasis id="DBF6EAF8ACDE43FC0B83F85C8F4701EE" bold="true" box="[151,290,1943,1967]" pageId="135" pageNumber="136">FIGURE 81.</emphasis>
<taxonomicName id="2E824D69ACDE43FC0A3EF85C8C0601EE" authorityName="WOOD" authorityYear="2024" box="[298,611,1943,1967]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="risha" status="sp. nov.">
<emphasis id="DBF6EAF8ACDE43FC0A3EF85C8FE501EE" box="[298,384,1943,1967]" italics="true" pageId="135" pageNumber="136">Andrena</emphasis>
(incertae sedis) 
<emphasis id="DBF6EAF8ACDE43FC093BF85C8C0601EE" box="[559,611,1943,1967]" italics="true" pageId="135" pageNumber="136">risha</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8ACDE43FC0978F8538CB001F1" bold="true" box="[620,725,1944,1968]" pageId="135" pageNumber="136">
<taxonomicNameLabel id="C0C55783ACDE43FC0978F8538CB001F1" box="[620,725,1944,1968]" pageId="135" pageNumber="136" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Labrum, ventral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. 
<taxonomicName id="2E824D69ACDE43FC0841F8778DA00192" box="[853,965,1980,2003]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="135" pageNumber="142" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
, lateral view; G. Hind leg, ventro-lateral view; H. Terga, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561ACD143F30BD3FEEC8C040724" pageId="136" pageNumber="137" type="etymology">
<paragraph id="E93D36EAACD143F30BD3FEEC8C040724" blockId="136.[151,1436,151,393]" pageId="136" pageNumber="137">
<emphasis id="DBF6EAF8ACD143F30BD3FEEC8F2B0700" bold="true" box="[199,334,295,321]" pageId="136" pageNumber="137">Etymology.</emphasis>
From the Arabic noun 
<taxonomicName id="2E824D69ACD143F30959FEEC8CF60700" authorityName="WOOD" authorityYear="2024" box="[589,659,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="risha">
“ 
<emphasis id="DBF6EAF8ACD143F3094DFEEC8CF60700" box="[601,659,295,321]" italics="true" pageId="136" pageNumber="137">risha</emphasis>
</taxonomicName>
” meaning feather, in reference to the plumose hairs covering the body of this species. It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD143F30BD3FEA58C0F07C8" box="[199,618,366,393]" pageId="136" pageNumber="137" type="distribution">
<paragraph id="E93D36EAACD143F30BD3FEA58C0F07C8" blockId="136.[151,1436,151,393]" box="[199,618,366,393]" pageId="136" pageNumber="137">
<emphasis id="DBF6EAF8ACD143F30BD3FEA58F0407C9" bold="true" box="[199,353,366,392]" pageId="136" pageNumber="137">Distribution.</emphasis>
North-western 
<collectingCountry id="9195767AACD143F3091BFEA48C0207C8" box="[527,615,367,393]" name="Algeria" pageId="136" pageNumber="137">Algeria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>