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<document id="DB5C14B79F907166FE2EBFC1C6D3B7C8" ID-DOI="10.11646/zootaxa.1161.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5058330" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625190197324" checkinUser="felipe" docAuthor="Hippa, Heikki" docDate="2006" docId="5D4C8785285EFFB55C1A0C42FE07F8AE" docLanguage="en" docName="zt01161p049.pdf" docOrigin="Zootaxa 1161 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1161.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Manota procera Hippa 2006, sp. n." docType="treatment" docUuid="642D243B-6F50-4117-9229-25B562FFFCBC" docUuidSource="ZooBank" docVersion="4" lastPageNumber="27" masterDocId="A175FFFD2844FFAF5D120E76FFC5FFE7" masterDocTitle="Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species" masterLastPageNumber="49" masterPageNumber="1" pageNumber="27" updateTime="1698983105622" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3EB8A5486512EE8497BE21E40EEBF38C">Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species</mods:title>
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<mods:namePart id="58D745232E273C036C294B58CBB570E8">Hippa, Heikki</mods:namePart>
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<mods:title id="DEE5D944BE61C94A02116E38119F27FA">Zootaxa</mods:title>
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<mods:date id="D77A0C62DEB5C674C79575777460D496">2006</mods:date>
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<mods:number id="C778C27E8F38936CEE1FDD4AE3E10ED6">2006-03-30</mods:number>
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<mods:number id="A1D71E7DEB19B3B73A0E3DD02D414CD2">1161</mods:number>
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<mods:url id="31B6CC7B7B3D0DB607D2C95ACFD22B52">https://biotaxa.org/Zootaxa/article/view/zootaxa.1161.1.1</mods:url>
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<mods:classification id="106D982CA2E00C1D4BFF0F7D11962891">journal article</mods:classification>
<mods:identifier id="D66C9EBAD56B995DCC29CCBD0A34E51F" type="DOI">10.11646/zootaxa.1161.1.1</mods:identifier>
<mods:identifier id="BFFCCE5224C8DDCE055BE0F941D935A1" type="ISSN">1175­5334</mods:identifier>
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<treatment id="5D4C8785285EFFB55C1A0C42FE07F8AE" ID-DOI="http://doi.org/10.5281/zenodo.5059310" ID-Zenodo-Dep="5059310" LSID="urn:lsid:zoobank.org:act:642D243B-6F50-4117-9229-25B562FFFCBC" httpUri="http://treatment.plazi.org/id/5D4C8785285EFFB55C1A0C42FE07F8AE" lastPageNumber="27" pageId="26" pageNumber="27">
<subSubSection id="9DFF6518285EFFB55C1A0C42FDD1FDAA" box="[264,532,563,590]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="D55A3693285EFFB55C1A0C42FDD1FDAA" blockId="26.[264,532,563,625]" box="[264,532,563,590]" pageId="26" pageNumber="27">
<heading id="8E1281FF285EFFB55C1A0C42FDD1FDAA" bold="true" box="[264,532,563,590]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="E791EA81285EFFB55C1A0C42FDD1FDAA" bold="true" box="[264,532,563,590]" pageId="26" pageNumber="27">
<taxonomicName id="12E54D10285EFFB55C1A0C42FE0CFDA9" ID-CoL="3XVY7" authority="Hippa, 2006" authorityName="Hippa" authorityYear="2006" box="[264,457,564,590]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="procera" status="sp. nov.">
<emphasis id="E791EA81285EFFB55C1A0C42FE0CFDA9" bold="true" box="[264,457,564,590]" italics="true" pageId="26" pageNumber="27">Manota procera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCA257FA285EFFB55CC00C45FDD1FDAA" box="[466,532,563,589]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9DFF6518285EFFB55C1A0C21FE07F8AE" pageId="26" pageNumber="27" type="description">
<paragraph id="D55A3693285EFFB55C1A0C21FE21FD96" blockId="26.[264,532,563,625]" box="[264,484,599,625]" pageId="26" pageNumber="27">
(
<figureCitation id="4DDE2A16285EFFB55C030C21FE5EFD96" box="[273,411,599,625]" captionStart="FIGURE 9" captionStartId="27.[264,367,1807,1831]" captionTargetBox="[277,1293,325,1765]" captionTargetId="figure-66@27.[277,1318,309,1796]" captionTargetPageId="27" captionText="FIGURE 9. Antennal flagellomere 4, lateral view (A and C); hypopygium, ventral (B and D) and dorsal view (E). A, B. Manota procera sp. n. (holotype). C, D, E. Manota transversa sp. n. (holotype). Scale for A and D 0.05 mm; for B, C, E, and F 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058360" httpUri="https://zenodo.org/record/5058360/files/figure.png" pageId="26" pageNumber="27">Figs. 9A, B</figureCitation>
;
<figureCitation id="4DDE2A16285EFFB55CBA0C21FE19FD96" box="[424,476,599,625]" captionStart="FIGURE 10" captionStartId="29.[264,368,829,853]" captionTargetBox="[292,1287,323,808]" captionTargetId="figure-251@29.[281,1306,311,825]" captionTargetPageId="29" captionText="FIGURE 10. Apical part of gonostylus, dorsal view. A. Manota procera sp. n. (holotype). B. Manota calcarata sp. n. (paratype). C. Manota transversa sp. n. (holotype). Scale 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5058362" httpUri="https://zenodo.org/record/5058362/files/figure.png" pageId="26" pageNumber="27">10A</figureCitation>
)
</paragraph>
<paragraph id="D55A3693285EFFB55C1A0CD1FDD6FD26" blockId="26.[264,1323,679,1505]" box="[264,531,679,705]" pageId="26" pageNumber="27">
A small­sized
<taxonomicName id="12E54D10285EFFB55CA30CD1FDC9FD27" authorityName="Williston" authorityYear="1896" box="[433,524,679,704]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="E791EA81285EFFB55CA30CD1FDC9FD27" box="[433,524,679,704]" italics="true" pageId="26" pageNumber="27">Manota</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D55A3693285EFFB55C2A0CB9FEA3FA5E" blockId="26.[264,1323,679,1505]" pageId="26" pageNumber="27">
Male.
<emphasis id="E791EA81285EFFB55C970CB8FE0BFD0F" bold="true" box="[389,462,718,744]" pageId="26" pageNumber="27">Head.</emphasis>
Flagellomere 4,
<figureCitation id="4DDE2A16285EFFB55F8F0CB9FD38FD0E" box="[669,765,719,745]" captionStart="FIGURE 9" captionStartId="27.[264,367,1807,1831]" captionTargetBox="[277,1293,325,1765]" captionTargetId="figure-66@27.[277,1318,309,1796]" captionTargetPageId="27" captionText="FIGURE 9. Antennal flagellomere 4, lateral view (A and C); hypopygium, ventral (B and D) and dorsal view (E). A, B. Manota procera sp. n. (holotype). C, D, E. Manota transversa sp. n. (holotype). Scale for A and D 0.05 mm; for B, C, E, and F 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058360" httpUri="https://zenodo.org/record/5058360/files/figure.png" pageId="26" pageNumber="27">Fig. 9A</figureCitation>
. Maxillary palpus with palpomere 3 bearing 5 apically expanded curved sensilla; palpomere 4 with parasegment.
<emphasis id="E791EA81285EFFB5590B0C80FBB8FCF7" bold="true" box="[1049,1149,758,784]" pageId="26" pageNumber="27">Thorax.</emphasis>
Anepisternum setose. Anterior basalare non­setose. Preepisternum 2 setose. Laterotergite non­setose. Episternum 3 setose.
<emphasis id="E791EA81285EFFB55F180D30FD8BFC87" bold="true" box="[522,590,838,864]" pageId="26" pageNumber="27">Wing</emphasis>
. Length
<quantity id="121D9B76285EFFB55FA90D31FCDFFC86" box="[699,794,839,865]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="27" unit="mm" value="1.8">1.8 mm</quantity>
. Wing membrane dorsally with few setae at posterior margin. Sc distally of h non­setose. Setae indicating A1 not observed with certainty.
<emphasis id="E791EA81285EFFB55C6E0DE0FDDBFC57" bold="true" box="[380,542,918,944]" pageId="26" pageNumber="27">Hypopygium</emphasis>
(
<figureCitation id="4DDE2A16285EFFB55F3C0DE1FD50FC56" box="[558,661,919,945]" captionStart="FIGURE 9" captionStartId="27.[264,367,1807,1831]" captionTargetBox="[277,1293,325,1765]" captionTargetId="figure-66@27.[277,1318,309,1796]" captionTargetPageId="27" captionText="FIGURE 9. Antennal flagellomere 4, lateral view (A and C); hypopygium, ventral (B and D) and dorsal view (E). A, B. Manota procera sp. n. (holotype). C, D, E. Manota transversa sp. n. (holotype). Scale for A and D 0.05 mm; for B, C, E, and F 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058360" httpUri="https://zenodo.org/record/5058360/files/figure.png" pageId="26" pageNumber="27">Figs. 9B</figureCitation>
,
<figureCitation id="4DDE2A16285EFFB55FB30DE1FD12FC56" box="[673,727,919,945]" captionStart="FIGURE 10" captionStartId="29.[264,368,829,853]" captionTargetBox="[292,1287,323,808]" captionTargetId="figure-251@29.[281,1306,311,825]" captionTargetPageId="29" captionText="FIGURE 10. Apical part of gonostylus, dorsal view. A. Manota procera sp. n. (holotype). B. Manota calcarata sp. n. (paratype). C. Manota transversa sp. n. (holotype). Scale 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5058362" httpUri="https://zenodo.org/record/5058362/files/figure.png" pageId="26" pageNumber="27">10A</figureCitation>
). Sternite 9 large, extending posteriorly to halfway along ventral length of gonocoxa, laterally sharply delimited, posterior margin indistinct, medially confluent with more posterior membranous part with transverse straight margin, setae similar to adjacent ventral setae of gonocoxa. Dorsomesial margin of gonocoxa simple, few stronger setae at apicomesial rounded corner, in position III 2 sigmoid or curved simple megasetae arising from common long basal body; gonocoxa apicolaterally with long, tapering, mesially setose, subtriangular lobe that extends posteriorly as far as apex of gonostylus. Gonostylus elongate­quadrangular in outline, slightly convex on lateral margin, concave on mesial margin, transversely truncated on apical margin, at apicomesial angle, on margin and on dorsal side group of strong setae arranged in 2 indistinct longitudinal rows (
<figureCitation id="4DDE2A16285EFFB55F730B51FD15FAA6" box="[609,720,1319,1345]" captionStart="FIGURE 10" captionStartId="29.[264,368,829,853]" captionTargetBox="[292,1287,323,808]" captionTargetId="figure-251@29.[281,1306,311,825]" captionTargetPageId="29" captionText="FIGURE 10. Apical part of gonostylus, dorsal view. A. Manota procera sp. n. (holotype). B. Manota calcarata sp. n. (paratype). C. Manota transversa sp. n. (holotype). Scale 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5058362" httpUri="https://zenodo.org/record/5058362/files/figure.png" pageId="26" pageNumber="27">Fig. 10A</figureCitation>
). Parastylar lobe present, directed anteriad, with 4 apical setae. Tegmen short and broad, subtriangular but with marked lateral shoulders. Setae on sternite
<quantity id="121D9B76285EFFB55CC40B01FDD6FA76" box="[470,531,1399,1425]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="26" pageNumber="27" unit="in" value="10.0">10 in</quantity>
indistinct double row on each half, close to sides of apical part of tegmen.
</paragraph>
<paragraph id="D55A3693285EFFB55C2A0BB1FCDBFA06" blockId="26.[264,1323,679,1505]" box="[312,798,1479,1505]" pageId="26" pageNumber="27">Female and preimaginal stages unknown.</paragraph>
<paragraph id="D55A3693285EFFB55C1A0861FE4DF9D7" blockId="26.[264,1322,1559,1665]" box="[264,392,1559,1584]" pageId="26" pageNumber="27">
<emphasis id="E791EA81285EFFB55C1A0861FE4DF9D7" box="[264,392,1559,1584]" italics="true" pageId="26" pageNumber="27">Discussion</emphasis>
</paragraph>
<paragraph id="D55A3693285EFFB55C2A0849FDD8F966" blockId="26.[264,1322,1559,1665]" pageId="26" pageNumber="27">
<taxonomicName id="12E54D10285EFFB55C2A0849FE3EF9BF" authorityName="Hippa" authorityYear="2006" box="[312,507,1599,1624]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="procera">
<emphasis id="E791EA81285EFFB55C2A0849FE3EF9BF" box="[312,507,1599,1624]" italics="true" pageId="26" pageNumber="27">Manota procera</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="12E54D10285EFFB55FBA0849FC95F9BF" authorityName="Hippa" authorityYear="2006" box="[680,848,1599,1624]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="transversa">
<emphasis id="E791EA81285EFFB55FBA0849FC95F9BF" box="[680,848,1599,1624]" italics="true" pageId="26" pageNumber="27">M. transversa</emphasis>
</taxonomicName>
. For the distinguishing characters, see under the latter species.
</paragraph>
<paragraph id="D55A3693285EFFB55C1A08C1FE8FF937" blockId="26.[264,1323,1719,1865]" box="[264,330,1719,1744]" pageId="26" pageNumber="27">
<emphasis id="E791EA81285EFFB55C1A08C1FE8FF937" box="[264,330,1719,1744]" italics="true" pageId="26" pageNumber="27">Types</emphasis>
</paragraph>
<paragraph id="D55A3693285EFFB55C2A08A9FE07F8AE" blockId="26.[264,1323,1719,1865]" pageId="26" pageNumber="27">
<materialsCitation id="658D3CCE285EFFB55C2A08A9FE07F8AE" collectingDate="1997-02-24" collectingDateMax="1997-03-21" collectingDateMin="1997-02-24" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund." country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
<emphasis id="E791EA81285EFFB55C2A08A9FE66F91F" box="[312,419,1759,1784]" italics="true" pageId="26" pageNumber="27">
<typeStatus id="0A5E8831285EFFB55C2A08A9FE66F91F" box="[312,419,1759,1784]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCounty id="3C3B4E1F285EFFB55CA608A9FE36F91E" box="[436,499,1759,1785]" pageId="26" pageNumber="27">Male</collectingCounty>
,
<collectingMunicipality id="353EACE9285EFFB55F1008A9FDF9F91E" box="[514,572,1759,1785]" pageId="26" pageNumber="27">West</collectingMunicipality>
<collectingCountry id="ADF27603285EFFB55F5708A9FD76F91E" box="[581,691,1759,1785]" name="Malaysia" pageId="26" pageNumber="27">Malaysia</collectingCountry>
,
<collectingRegion id="1721F871285EFFB55FD308A9FCEEF91E" box="[705,811,1759,1785]" country="Malaysia" name="Selangor" pageId="26" pageNumber="27">Selangor</collectingRegion>
,
<location id="D03A6048285EFFB55E2808A9FC12F91E" LSID="urn:lsid:plazi:treatment:5D4C8785285EFFB55C1A0C42FE07F8AE:D03A6048285EFFB55E2808A9FC12F91E" box="[826,983,1759,1785]" country="Malaysia" county="Male" municipality="West" name="Ulu Gombak" pageId="26" pageNumber="27" stateProvince="Selangor">Ulu Gombak</location>
,
<location id="D03A6048285EFFB55EF708A9FE60F8C6" LSID="urn:lsid:plazi:treatment:5D4C8785285EFFB55C1A0C42FE07F8AE:D03A6048285EFFB55EF708A9FE60F8C6" country="Malaysia" county="Male" municipality="West" name="University of Malaya Field Study Centre" pageId="26" pageNumber="27" stateProvince="Selangor">University of Malaya Field Study Centre</location>
,
<quantity id="121D9B76285EFFB55CA00971FE3EF8C6" box="[434,507,1799,1825]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="26" pageNumber="27" unit="ft" value="800.0">
<elevation id="5EC8D1A0285EFFB55CA00971FE3EF8C6" box="[434,507,1799,1825]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="26" pageNumber="27" unit="ft" value="800.0">800 ft</elevation>
</quantity>
, Malaise in jungle,
<date id="A15B1053285EFFB55FF60971FBD4F8C6" box="[740,1041,1799,1825]" pageId="26" pageNumber="27" value="1997-02-24" valueMax="1997-03-21" valueMin="1997-02-24">
<collectingDate id="B11FE9BB285EFFB55FF60971FBD4F8C6" box="[740,1041,1799,1825]" pageId="26" pageNumber="27" value="1997-02-24" valueMax="1997-03-21" valueMin="1997-02-24">24. Feb.–21. March 1997</collectingDate>
</date>
,
<collectorName id="78105345285EFFB5590D0971FB4BF8C6" box="[1055,1166,1799,1825]" pageId="26" pageNumber="27">H. Hippa</collectorName>
,
<collectorName id="78105345285EFFB559880971FAEFF8C6" box="[1178,1322,1799,1825]" pageId="26" pageNumber="27">M. Jaschhof</collectorName>
and
<collectorName id="78105345285EFFB55C280959FE07F8AE" box="[314,450,1839,1865]" pageId="26" pageNumber="27">B. Viklund.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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