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<document id="5B4E1BEE65A24E8648149D8F1F9EC621" ID-DOI="10.5281/zenodo.10106680" ID-ISSN="2345-7600" ID-Zenodo-Dep="10106680" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630179122938" checkinUser="tatiana" docAuthor="Tan, Siong Kiat, Chan, Sow Yan &amp; Panha, Somsak" docDate="2011" docId="4B7C87FFFFE4FF9EFF29A53BFE8FB6EC" docLanguage="en" docName="59rbz039-046.pdf" docOrigin="Raffles Bulletin of Zoology 59 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Amphidromus (Amphidromus) atricallosus subsp. temasek Tan &amp; Chan &amp; Panha 2011, new subspecies" docType="treatment" docVersion="4" lastPageNumber="42" masterDocId="B745FF87FFE5FF9DFFA7A35BFF84B41B" masterDocTitle="A New Subspecies Of Amphidromus (Amphidromus) Atricallosus From Singapore (Mollusca: Gastropoda: Camaenidae)" masterLastPageNumber="46" masterPageNumber="39" pageNumber="40" updateTime="1700278366035" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="CC353F44DC4DAA2A7A2E0E68920B3001" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2B98A63246080C2A0502A2F4EF59DE5E">
<mods:title id="7B363C07F18D8F6D8AC74718DEC353B8">A New Subspecies Of Amphidromus (Amphidromus) Atricallosus From Singapore (Mollusca: Gastropoda: Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name id="A037ED877A16BA49E13247291E768712" type="personal">
<mods:role id="E20D6B7D399E4FCD9EB79EF998E30410">
<mods:roleTerm id="574344AADCDDDADEC6EB9816B2F61095">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E4BDC779FFBE7FBCCFA2665A23EF540B">Tan, Siong Kiat</mods:namePart>
</mods:name>
<mods:name id="D002E35E0F56D9CF188F6D2D221F09D3" type="personal">
<mods:role id="899FB0D38C97CCCB978AFB3731E0BD99">
<mods:roleTerm id="40B65E53314E398DEBB395616E5DED80">Author</mods:roleTerm>
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<mods:namePart id="054EFFC8671B784D42275D48B620B36E">Chan, Sow Yan</mods:namePart>
</mods:name>
<mods:name id="BAF02542F67B52F876F7BA9DBF82C3DD" type="personal">
<mods:role id="E1A12D08D20FFBCDA588320DBB56A5FD">
<mods:roleTerm id="4762FC3768B625A0EBA83FA1D6780619">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="089B288AF732F4527AEC1DC89D286878">Panha, Somsak</mods:namePart>
</mods:name>
<mods:typeOfResource id="E23CA516E855F55B34209D5DFD46503B">text</mods:typeOfResource>
<mods:relatedItem id="BC6CB2FB710BB183F0EC967466DA22EA" type="host">
<mods:titleInfo id="810A2E543BAAB2264FEE04BA5486AEAB">
<mods:title id="C56D4EFA245F05914FE495A557CC3E67">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="8B119BDBBEAB676346BE49D99F389446">
<mods:date id="5F271EF49A4132FEC99BCE64CEE6F84E">2011</mods:date>
<mods:detail id="6D3ADF2F03B62F68EB0D5DCAEDF9C60D" type="pubDate">
<mods:number id="AD5944C04BD31539CD384BFB1DE2899C">2011-02-28</mods:number>
</mods:detail>
<mods:detail id="327853B976816863E595DBAFCAA6C962" type="volume">
<mods:number id="C20097EB909CE10678749EB9057A0F5A">59</mods:number>
</mods:detail>
<mods:detail id="49B63457D224278CC2886E5AC558A1DC" type="issue">
<mods:number id="7D6FC7BA209219C2FF14FBA2DB7BA115">1</mods:number>
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<mods:start id="D1593DD95ABB987821A6F41F9BBC4275">39</mods:start>
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<mods:classification id="2FEF56CAC178C1C60C7EAA9564FC9CE2">journal article</mods:classification>
<mods:identifier id="E7255C67BB6830D200B430E284260DD7" type="DOI">10.5281/zenodo.10106680</mods:identifier>
<mods:identifier id="2FFBC4FB5D74479E37FFA8FB229EAE3C" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E362A74D258190D98BE1DE217DB8DB65" type="Zenodo-Dep">10106680</mods:identifier>
</mods:mods>
<treatment id="4B7C87FFFFE4FF9EFF29A53BFE8FB6EC" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC" httpUri="http://treatment.plazi.org/id/4B7C87FFFFE4FF9EFF29A53BFE8FB6EC" lastPageId="3" lastPageNumber="42" pageId="1" pageNumber="40">
<subSubSection id="8BCF6562FFE4FF9CFF29A53BFD83B283" pageId="1" pageNumber="40" type="nomenclature">
<paragraph id="C36A36E9FFE4FF9CFF29A53BFD83B283" blockId="1.[142,723,1632,1719]" pageId="1" pageNumber="40">
<heading id="98228185FFE4FF9CFF29A53BFD83B283" bold="true" centered="true" fontSize="10" level="2" pageId="1" pageNumber="40" reason="2">
<emphasis id="F1A1EAFBFFE4FF9CFF29A53BFD83B283" bold="true" pageId="1" pageNumber="40">
<taxonomicName id="04D54D6AFFE4FF9CFF29A53BFD48B263" ID-CoL="7GKQZ" authority="Tan &amp; Chan &amp; Panha, 2011" authorityName="Tan &amp; Chan &amp; Panha" authorityYear="2011" box="[142,716,1632,1656]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="40" phylum="Mollusca" rank="subSpecies" species="atricallosus" status="subsp. nov." subGenus="Amphidromus" subSpecies="temasek">
<emphasis id="F1A1EAFBFFE4FF9CFF29A53BFEAFB263" bold="true" box="[142,299,1632,1656]" italics="true" pageId="1" pageNumber="40">Amphidromus</emphasis>
(
<emphasis id="F1A1EAFBFFE4FF9CFE9AA53BFE5EB263" bold="true" box="[317,474,1632,1656]" italics="true" pageId="1" pageNumber="40">Amphidromus</emphasis>
)
<emphasis id="F1A1EAFBFFE4FF9CFE4CA53BFD48B263" bold="true" box="[491,716,1632,1656]" italics="true" pageId="1" pageNumber="40">atricallosus temasek</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="EA925780FFE4FF9CFEFDA5DBFD83B283" box="[346,519,1664,1688]" pageId="1" pageNumber="40" rank="subSpecies">new subspecies</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BCF6562FFE4FF9CFEEEA5FBFD9CB2AC" box="[329,536,1696,1719]" pageId="1" pageNumber="40" type="description">
<paragraph id="C36A36E9FFE4FF9CFEEEA5FBFD9CB2AC" blockId="1.[142,723,1632,1719]" box="[329,536,1696,1719]" pageId="1" pageNumber="40">
(
<figureCitation id="5BEE2A6CFFE4FF9CFEF6A5FBFE35B2AC" box="[337,433,1696,1719]" captionStart="Fig" captionStartId="2.[136,171,1948,1969]" captionTargetBox="[137,1474,565,1922]" captionTargetId="figure-125@2.[137,1475,565,1922]" captionTargetPageId="2" captionText="Fig. 1. A, Amphidromus atricallosus temasek, new subspecies: Holotype, SH 44 x SD 25.6 (ZRC.MOL.3058), Nee Soon Swamp Forest, Singapore; B–E Amphidromus atricallosus perakensis (Fulton, 1901): B, SH 45 x SD 26.4 (TSK 11041), Bukit Serdam, Pahang, West Malaysia; C, SH 47.1 x SD 25.3 (CBB), Tapah Hill, Perak, West Malaysia; D, SH 47.2 x SD 27.6 (ZRC.MOL.3064), Pulau Tulai, near Pulau Tioman, West Malaysia; E, SH 46.4 x SD 24.2 (ZRC.MOL.3062), Gunung Genting, Perak, West Malaysia; F, Amphidromus atricallosus atricallosus (Gould, 1843): SH 45 x SD 25.7 (CSY 409.4.4.16), Ranong, Thailand; G, Amphidromus atricallosus leucoxanthus (von Martens, 1864): SH 51 x SD 28.5 (CSY 409.3.49.1), Chantaburi, East Thailand." figureDoi="http://doi.org/10.5281/zenodo.10106684" httpUri="https://zenodo.org/record/10106684/files/figure.png" pageId="1" pageNumber="40">Figs. 1A</figureCitation>
,
<figureCitation id="5BEE2A6CFFE4FF9CFE18A5FBFE65B2AC" box="[447,481,1696,1719]" captionStart="Fig" captionStartId="3.[114,149,2035,2056]" captionTargetBox="[113,1452,1623,2009]" captionTargetId="figure-421@3.[113,1452,1623,2010]" captionTargetPageId="3" captionText="Fig. 2. A–C, Shells of juveniles: A, SH 31 x SD 19.7, juvenile shell of Amphidromus atricallosus temasek, new subspecies, showing the white columella; B–C, Amphidromus atricallosus perakensis (Fulton, 1901), juveniles, has a white zone bordering the columella: B, SH 33.9 x SD 22.4, Tapah Hill, Perak, West Malaysia; C, SH 34.7 x SD 25, Pulau Tulai, West Malaysia." figureDoi="http://doi.org/10.5281/zenodo.10106686" httpUri="https://zenodo.org/record/10106686/files/figure.png" pageId="1" pageNumber="40">2A</figureCitation>
,
<figureCitation id="5BEE2A6CFFE4FF9CFE48A5FBFD94B2AC" box="[495,528,1696,1719]" captionStart="Fig" captionStartId="4.[137,172,2063,2084]" captionTargetBox="[136,1474,229,2023]" captionTargetId="figure-43@4.[138,1474,229,1137]" captionTargetPageId="4" captionText="Fig. 3. A–B, Live animal colouration: A, Amphidromus atricallosus temasek, new subspecies, holotype (ZRC.MOL.3058); B, Amphidromus atricallosus perakensis (Fulton, 1901), Pulau Tulai, West Malaysia. (Photographs by Tan Heok Hui)" figureDoi="http://doi.org/10.5281/zenodo.10106692" httpUri="https://zenodo.org/record/10106692/files/figure.png" pageId="1" pageNumber="40">3A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8BCF6562FFE4FF9CFFD6A584FDD4B37C" pageId="1" pageNumber="40" type="reference_group">
<paragraph id="C36A36E9FFE4FF9CFFD6A584FD12B351" blockId="1.[113,752,1759,1895]" pageId="1" pageNumber="40">
<taxonomicName id="04D54D6AFFE4FF9CFFD6A584FE57B2EF" ID-CoL="7GL2Y" authorityName="Fulton" authorityYear="1901" box="[113,467,1759,1780]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="40" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="perakensis">
<emphasis id="F1A1EAFBFFE4FF9CFFD6A584FE57B2EF" box="[113,467,1759,1780]" italics="true" pageId="1" pageNumber="40">Amphidromus atricallosus perakensis</emphasis>
</taxonomicName>
– Ng &amp; Lim, 1992: 262;
<bibRefCitation id="A7444B18FFE4FF9CFD1BA584FEAFB30A" author="Chou, L" pageId="1" pageNumber="40" pagination="70 - 106" refId="ref6237" refString="Chou, L. M., P. K. L. Ng &amp; K. K. P. Lim, 1994. Animalia. In: Wee, Y. C. &amp; P. K. L. Ng (eds.), A First Look at the Biodiversity in Singapore. National Council on the Environment, Singapore. Pp 70 - 106." type="book chapter" year="1994">Chou et al., 1994: 73</bibRefCitation>
; Sutcharit &amp; Panha, 2006: 21–22, Fig. 4. F–G, Fig. 11. A–C, Fig. 12. C;
<bibRefCitation id="A7444B18FFE4FF9CFE3FA443FDF1B336" author="Prasankok, P &amp; Ota, M" box="[408,629,1816,1837]" pageId="1" pageNumber="40" pagination="189 - 197" refId="ref7006" refString="Prasankok, P., H. Ota, M. Toda &amp; S. Panha, 2007. Allozyme variation in the Camaenid tree snails Amphidromus atricallosus (Gould, 1843) and A. inversus (Muller, 1774). Zoological Science, 24: 189 - 197." type="journal article" year="2007">Prasankok et al., 2007</bibRefCitation>
;
<bibRefCitation id="A7444B18FFE4FF9CFD27A443FF6DB351" author="Sutcharit, C" pageId="1" pageNumber="40" pagination="661 - 672" refId="ref7181" refString="Sutcharit, C., T. Asami &amp; S. Panha, 2007. Evolution of whole-body enantiomorphy in the tree snail genus Amphidromus. Journal of Evolutionary Biology, 20 (2): 661 - 672." type="journal article" year="2007">Sutcharit et al, 2007</bibRefCitation>
; Chou &amp; Tan, 2008: 57; Lok &amp; Tan, 2008.
</paragraph>
<paragraph id="C36A36E9FFE4FF9CFFD6A409FDD4B37C" blockId="1.[113,752,1759,1895]" box="[113,592,1874,1895]" pageId="1" pageNumber="40">
<treatmentCitationGroup id="E3C511C7FFE4FF9CFFD6A409FDD4B37C" box="[113,592,1874,1895]" pageId="1" pageNumber="40">
<taxonomicName id="04D54D6AFFE4FF9CFFD6A409FEF4B37C" ID-CoL="D2K9" baseAuthorityName="Gould" baseAuthorityYear="1843" box="[113,368,1874,1895]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="40" phylum="Mollusca" rank="species" species="atricallosus">
<emphasis id="F1A1EAFBFFE4FF9CFFD6A409FEF4B37C" box="[113,368,1874,1895]" italics="true" pageId="1" pageNumber="40">Amphidromus atricallosus</emphasis>
</taxonomicName>
–
<treatmentCitation id="427410F8FFE4FF9CFE2CA409FDC8B37C" author="Ho, W" box="[395,588,1874,1895]" page="108" pageId="1" pageNumber="40" year="1995">
<bibRefCitation id="A7444B18FFE4FF9CFE2CA409FDC8B37C" author="Ho, W" box="[395,588,1874,1895]" pageId="1" pageNumber="40" pagination="91 - 113" refId="ref6652" refString="Ho, W. H., 1995. A Review of the Land-snail Fauna of Singapore. Raffles Bulletin of Zoology, 43 (1): 91 - 113." type="journal article" year="1995">Ho, 1995: 108–109</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8BCF6562FFE4FF9CFFD6A4D7FB9AB0E4" pageId="1" pageNumber="40" type="materials_examined">
<paragraph id="C36A36E9FFE4FF9CFFD6A4D7FBE1B69E" blockId="1.[113,753,1931,2097]" lastBlockId="1.[812,1452,192,645]" pageId="1" pageNumber="40">
Material examined. –
<materialsCitation id="73BD3CB4FFE4FF9CFEFCA4D0FDACB3C1" collectingDate="2010-10-03" collectionCode="ZRC" collectorName="Singapore &amp; S. K. Tan &amp; S. Y. Chan" country="Singapore" location="Nee Soon Swamp Forest" pageId="1" pageNumber="40" specimenCount="1" typeStatus="holotype">
<emphasis id="F1A1EAFBFFE4FF9CFEFCA4D0FE71B3BB" bold="true" box="[347,501,1931,1952]" pageId="1" pageNumber="40">
<collectingCountry id="BBC27679FFE4FF9CFEFCA4D0FE75B3BB" box="[347,497,1931,1952]" name="Singapore" pageId="1" pageNumber="40">SINGAPORE</collectingCountry>
:
</emphasis>
<typeStatus id="1C6E884BFFE4FF9CFE58A4D7FDDBB3BA" box="[511,607,1932,1953]" pageId="1" pageNumber="40" type="holotype">Holotype</typeStatus>
: 1 S SH 44 x SD 25.6 (
<collectionCode id="A5C4AE2CFFE4FF9CFF77A4F3FE87B3A6" box="[208,259,1960,1981]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
.MOL.3058),
<location id="C60A6032FFE4FF9CFE20A4F3FDF9B3A6" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFE20A4F3FDF9B3A6" box="[391,637,1960,1981]" country="Singapore" name="Nee Soon Swamp Forest" pageId="1" pageNumber="40">Nee Soon Swamp Forest</location>
,
<collectorName id="6E20533FFFE4FF9CFD2FA4F3FD68B3A6" box="[648,748,1960,1981]" pageId="1" pageNumber="40">Singapore</collectorName>
, coll.
<collectorName id="6E20533FFFE4FF9CFF03A49EFE82B3C1" box="[164,262,1989,2010]" pageId="1" pageNumber="40">S. K. Tan</collectorName>
&amp;
<collectorName id="6E20533FFFE4FF9CFE8BA49EFE1AB3C1" box="[300,414,1989,2010]" pageId="1" pageNumber="40">S. Y. Chan</collectorName>
,
<date id="B76B1029FFE4FF9CFE0CA49EFDACB3C1" box="[427,552,1989,2010]" pageId="1" pageNumber="40" value="2010-10-03">
<collectingDate id="A72FE9C1FFE4FF9CFE0CA49EFDACB3C1" box="[427,552,1989,2010]" pageId="1" pageNumber="40" value="2010-10-03">03 Oct.2010</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="73BD3CB4FFE4FF9CFD93A49EFD1DBC0F" collectionCode="ZRC" collectorName="Nee Soon &amp; E. Alfred" country="Singapore" pageId="1" pageNumber="40" specimenCount="1" typeStatus="paratype">
<typeStatus id="1C6E884BFFE4FF9CFD93A49EFD12B3C1" box="[564,662,1989,2010]" pageId="1" pageNumber="40" type="paratype">Paratypes</typeStatus>
: 2 D SH 42.7 x SD 26–SH 46.4 x SD 27.7, 2 S SH 40.1x SD 25–SH 40.8 x SD 25.1 (
<collectionCode id="A5C4AE2CFFE4FF9CFF43A4A4FE92BC0F" box="[228,278,2047,2068]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
.MOL.2992),
<collectorName id="6E20533FFFE4FF9CFE3CA4A4FE79BC0F" box="[411,509,2047,2068]" pageId="1" pageNumber="40">Nee Soon</collectorName>
, coll.
<collectorName id="6E20533FFFE4FF9CFD9DA4A4FD1DBC0F" box="[570,665,2047,2068]" pageId="1" pageNumber="40">E. Alfred</collectorName>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFD03A4A4FB00B4CE" collectingDate="1992-04-30" collectionCode="ZRC" collectorName="National Park Survey" country="Singapore" location="Nee Soon Swamp Forest" pageId="1" pageNumber="40" specimenCount="1" typeStatus="paratype">
1 D SH 46.9 x SD 28.3 (
<collectionCode id="A5C4AE2CFFE4FF9CFEB3AB47FEC2BC2A" box="[276,326,2076,2097]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
.1992.3163),
<location id="C60A6032FFE4FF9CFE61AB47FD3EBC2A" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFE61AB47FD3EBC2A" box="[454,698,2076,2097]" country="Singapore" name="Nee Soon Swamp Forest" pageId="1" pageNumber="40">Nee Soon Swamp Forest</location>
, coll.
<collectorName id="6E20533FFFE4FF9CFC8BA39BFC79B4CE" box="[812,1021,192,213]" pageId="1" pageNumber="40">National Park Survey</collectorName>
,
<date id="B76B1029FFE4FF9CFBAFA39BFB00B4CE" box="[1032,1156,192,213]" pageId="1" pageNumber="40" value="1992-04-30">
<collectingDate id="A72FE9C1FFE4FF9CFBAFA39BFB00B4CE" box="[1032,1156,192,213]" pageId="1" pageNumber="40" value="1992-04-30">30 Apr.1992</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFB29A39BFC76B514" collectionCode="ZRC, NS" collectorName="H. K. Lua" country="Singapore" location="Nee Soon" pageId="1" pageNumber="40" specimenCode="ZRC 1994.4118, NS 43" specimenCount="1" typeStatus="paratype">
1 D SH 45.0 x SD 26.4 (
<specimenCode id="93739E92FFE4FF9CFADCA39BFC12B4E9" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC 1994.4118</specimenCode>
), outside
<location id="C60A6032FFE4FF9CFC52A386FBD1B4E9" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFC52A386FBD1B4E9" box="[1013,1109,221,242]" country="Singapore" name="Nee Soon" pageId="1" pageNumber="40">Nee Soon</location>
range, Seletar Reservoir Park (
<specimenCode id="93739E92FFE4FF9CFA2CA386FCC3B514" collectionCode="NS" pageId="1" pageNumber="40">NS 43</specimenCode>
), coll.
<collectorName id="6E20533FFFE4FF9CFC2BA3A1FC76B514" box="[908,1010,250,271]" pageId="1" pageNumber="40">H. K. Lua</collectorName>
</materialsCitation>
,
<materialsCitation id="73BD3CB4FFE4FF9CFC5AA3A1FB21B530" collectingDate="1998-06-26" collectionCode="CUMZ" collectorName="S. Panha" country="Singapore" location="Nee Soon Nature Reserve" pageId="1" pageNumber="40" specimenCode="CUMZ 2133" specimenCount="1" typeStatus="paratype">
1994; 1 D (
<specimenCode id="93739E92FFE4FF9CFBC9A3A1FB74B514" box="[1134,1264,250,271]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="1" pageNumber="40">CUMZ 2133</specimenCode>
),
<location id="C60A6032FFE4FF9CFAA5A3A1FCF9B530" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFAA5A3A1FCF9B530" country="Singapore" name="Nee Soon Nature Reserve" pageId="1" pageNumber="40">Nee Soon Nature Reserve</location>
, coll.
<collectorName id="6E20533FFFE4FF9CFC18A24DFB9EB530" box="[959,1050,278,299]" pageId="1" pageNumber="40">S. Panha</collectorName>
,
<date id="B76B1029FFE4FF9CFB8FA24DFB21B530" box="[1064,1189,278,299]" pageId="1" pageNumber="40" value="1998-06-26">
<collectingDate id="A72FE9C1FFE4FF9CFB8FA24DFB21B530" box="[1064,1189,278,299]" pageId="1" pageNumber="40" value="1998-06-26">26 Jun.1998</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFB14A24DFA23B553" collectingDate="2000-02-19" collectionCode="CUMZ" collectorName="S. Panha &amp; Ms Loua" country="Singapore" location="Nee Soon Nature Reserve" pageId="1" pageNumber="40" specimenCode="CUMZ 2067" specimenCount="1" typeStatus="paratype">
2 D (
<specimenCode id="93739E92FFE4FF9CFB4DA24DFAEBB530" box="[1258,1391,278,299]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="1" pageNumber="40">CUMZ 2067</specimenCode>
),
<location id="C60A6032FFE4FF9CFA23A24DFB86B553" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFA23A24DFB86B553" country="Singapore" name="Nee Soon Nature Reserve" pageId="1" pageNumber="40">Nee Soon Nature Reserve</location>
, coll.
<collectorName id="6E20533FFFE4FF9CFBE4A268FB1EB553" box="[1091,1178,307,328]" pageId="1" pageNumber="40">S. Panha</collectorName>
&amp;
<collectorName id="6E20533FFFE4FF9CFB18A268FA9FB553" box="[1215,1307,307,328]" pageId="1" pageNumber="40">Ms Loua</collectorName>
,
<date id="B76B1029FFE4FF9CFA8FA268FA23B553" box="[1320,1447,307,328]" pageId="1" pageNumber="40" value="2000-02-19">
<collectingDate id="A72FE9C1FFE4FF9CFA8FA268FA23B553" box="[1320,1447,307,328]" pageId="1" pageNumber="40" value="2000-02-19">19 Feb.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFC8BA20BFBACB599" collectingDate="2001-02-19" collectionCode="CUMZ" collectorName="S. Panha &amp; C. Sutcharit" country="Singapore" location="Nee Soon Nature Reserve" pageId="1" pageNumber="40" specimenCode="CUMZ 2198" specimenCount="1" typeStatus="paratype">
10 D (
<specimenCode id="93739E92FFE4FF9CFCCCA20BFC69B57E" box="[875,1005,336,357]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="1" pageNumber="40">CUMZ 2198</specimenCode>
),
<location id="C60A6032FFE4FF9CFC58A20BFB7BB57E" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFC58A20BFB7BB57E" box="[1023,1279,336,357]" country="Singapore" name="Nee Soon Nature Reserve" pageId="1" pageNumber="40">Nee Soon Nature Reserve</location>
, coll.
<collectorName id="6E20533FFFE4FF9CFA9CA20BFA15B57E" box="[1339,1425,336,357]" pageId="1" pageNumber="40">S. Panha</collectorName>
&amp;
<collectorName id="6E20533FFFE4FF9CFC8BA236FC26B599" box="[812,930,365,386]" pageId="1" pageNumber="40">C. Sutcharit</collectorName>
,
<date id="B76B1029FFE4FF9CFC0AA236FBACB599" box="[941,1064,365,386]" pageId="1" pageNumber="40" value="2001-02-19">
<collectingDate id="A72FE9C1FFE4FF9CFC0AA236FBACB599" box="[941,1064,365,386]" pageId="1" pageNumber="40" value="2001-02-19">19 Feb.2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFB94A236FAB2B584" collectingDate="2004-03-04" collectionCode="CUMZ" collectorName="S. Panha &amp; P. Prasarnkok" country="Singapore" location="Nee Soon Nature Reserve" pageId="1" pageNumber="40" specimenCode="CUMZ 2633" specimenCount="1" typeStatus="paratype">
11 D (
<specimenCode id="93739E92FFE4FF9CFBD6A236FB76B599" box="[1137,1266,365,386]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="1" pageNumber="40">CUMZ 2633</specimenCode>
),
<location id="C60A6032FFE4FF9CFAA3A236FCF8B584" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFAA3A236FCF8B584" country="Singapore" name="Nee Soon Nature Reserve" pageId="1" pageNumber="40">Nee Soon Nature Reserve</location>
, coll.
<collectorName id="6E20533FFFE4FF9CFC12A2D1FB8DB584" box="[949,1033,394,415]" pageId="1" pageNumber="40">S. Panha</collectorName>
&amp;
<collectorName id="6E20533FFFE4FF9CFB80A2D1FB2BB584" box="[1063,1199,394,415]" pageId="1" pageNumber="40">P. Prasarnkok</collectorName>
,
<date id="B76B1029FFE4FF9CFB1FA2D1FAB2B584" box="[1208,1334,394,415]" pageId="1" pageNumber="40" value="2004-03-04">
<collectingDate id="A72FE9C1FFE4FF9CFB1FA2D1FAB2B584" box="[1208,1334,394,415]" pageId="1" pageNumber="40" value="2004-03-04">04 Mar.2004</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFAE7A2D1FB79B5C3" collectingDate="2008-10-13" collectionCode="ZRC" collectorName="H. H. Tan &amp; A. F. S. L. Lok" country="Singapore" location="Nee Soon Swamp Forest" pageId="1" pageNumber="40" specimenCount="1" typeStatus="paratype">
2 S (
<specimenCount id="D5D3FD60FFE4FF9CFAC9A2D1FA21B584" box="[1390,1445,394,415]" pageId="1" pageNumber="40" type="generic">1 juv.</specimenCount>
) SH 43.4 x SD 26.8 (
<collectionCode id="A5C4AE2CFFE4FF9CFC5EA2FDFBA8B5A0" box="[1017,1068,422,443]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
.MOL.2843),
<location id="C60A6032FFE4FF9CFB16A2FDFA23B5A0" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFB16A2FDFA23B5A0" box="[1201,1447,422,443]" country="Singapore" name="Nee Soon Swamp Forest" pageId="1" pageNumber="40">Nee Soon Swamp Forest</location>
, coll.
<collectorName id="6E20533FFFE4FF9CFCFAA298FC3BB5C3" box="[861,959,451,472]" pageId="1" pageNumber="40">H. H. Tan</collectorName>
&amp;
<collectorName id="6E20533FFFE4FF9CFC47A298FBF3B5C3" box="[992,1143,451,472]" pageId="1" pageNumber="40">A. F. S. L. Lok</collectorName>
,
<date id="B76B1029FFE4FF9CFB25A298FB79B5C3" box="[1154,1277,451,472]" pageId="1" pageNumber="40" value="2008-10-13">
<collectingDate id="A72FE9C1FFE4FF9CFB25A298FB79B5C3" box="[1154,1277,451,472]" pageId="1" pageNumber="40" value="2008-10-13">13 Oct.2008</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFAA0A298FA23B5EE" collectingDate="2010-09" collectionCode="ZRC" collectorName="Y. C. Ng" country="Singapore" location="Nee Soon Swamp Forest" pageId="1" pageNumber="40" specimenCount="1" typeStatus="paratype">
1 D (juv.) (
<collectionCode id="A5C4AE2CFFE4FF9CFAD2A298FA2CB5C3" box="[1397,1448,451,472]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
. MOL.3059),
<location id="C60A6032FFE4FF9CFC09A2BBFB20B5EE" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFC09A2BBFB20B5EE" box="[942,1188,480,501]" country="Singapore" name="Nee Soon Swamp Forest" pageId="1" pageNumber="40">Nee Soon Swamp Forest</location>
, coll.
<collectorName id="6E20533FFFE4FF9CFB45A2BBFABBB5EE" box="[1250,1343,480,501]" pageId="1" pageNumber="40">Y. C. Ng</collectorName>
,
<date id="B76B1029FFE4FF9CFAEDA2BBFA23B5EE" box="[1354,1447,480,501]" pageId="1" pageNumber="40" value="2010-09">
<collectingDate id="A72FE9C1FFE4FF9CFAEDA2BBFA23B5EE" box="[1354,1447,480,501]" pageId="1" pageNumber="40" value="2010-09">Sep.2010</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFC8BA2A6FAA0B634" collectingDate="2010-10-03" collectionCode="ZRC" collectorName="S. K. Tan &amp; S. Y. Chan" country="Singapore" location="Nee Soon Swamp Forest" pageId="1" pageNumber="40" specimenCount="1" typeStatus="paratype">
1 D SH 43.5 x SD 27.1 (
<collectionCode id="A5C4AE2CFFE4FF9CFB91A2A6FBEEB609" box="[1078,1130,509,530]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
.MOL.3057),
<location id="C60A6032FFE4FF9CFB53A2A6FCEEB634" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFB53A2A6FCEEB634" country="Singapore" name="Nee Soon Swamp Forest" pageId="1" pageNumber="40">Nee Soon Swamp Forest</location>
, coll.
<collectorName id="6E20533FFFE4FF9CFC0EA141FB8DB634" box="[937,1033,538,559]" pageId="1" pageNumber="40">S. K. Tan</collectorName>
&amp;
<collectorName id="6E20533FFFE4FF9CFB8BA141FB19B634" box="[1068,1181,538,559]" pageId="1" pageNumber="40">S. Y. Chan</collectorName>
,
<date id="B76B1029FFE4FF9CFB0FA141FAA0B634" box="[1192,1316,538,559]" pageId="1" pageNumber="40" value="2010-10-03">
<collectingDate id="A72FE9C1FFE4FF9CFB0FA141FAA0B634" box="[1192,1316,538,559]" pageId="1" pageNumber="40" value="2010-10-03">03 Oct.2010</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFA97A141FBE4B69E" collectingDate="2010-11-03" collectionCode="ZRC" collectorName="P. X. Ng &amp; B. Y. Q. Ng &amp; C. K. Yeo" country="Singapore" location="Nee Soon Swamp Forest" municipality="Nee Soon Range" pageId="1" pageNumber="40" specimenCount="1" typeStatus="paratype">
1 S SH 38.9 x SD 25.1 (
<collectionCode id="A5C4AE2CFFE4FF9CFC04A16DFC51B650" box="[931,981,566,587]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
.MOL.3060),
<location id="C60A6032FFE4FF9CFBFCA16DFAD0B650" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFBFCA16DFAD0B650" box="[1115,1364,566,587]" country="Singapore" municipality="Nee Soon Range" name="Nee Soon Swamp Forest" pageId="1" pageNumber="40">Nee Soon Swamp Forest</location>
, edge of swamp forest, behind
<collectingMunicipality id="230EAC93FFE4FF9CFBA0A108FB34B673" box="[1031,1200,595,616]" pageId="1" pageNumber="40">Nee Soon Range</collectingMunicipality>
, coll.
<collectorName id="6E20533FFFE4FF9CFB49A108FACEB673" box="[1262,1354,595,616]" pageId="1" pageNumber="40">P. X. Ng</collectorName>
,
<collectorName id="6E20533FFFE4FF9CFAF1A108FCCDB69E" pageId="1" pageNumber="40">B. Y. Q. Ng</collectorName>
&amp;
<collectorName id="6E20533FFFE4FF9CFCCBA12BFC57B69E" box="[876,979,624,645]" pageId="1" pageNumber="40">C. K. Yeo</collectorName>
,
<date id="B76B1029FFE4FF9CFC78A12BFBE4B69E" box="[991,1120,624,645]" pageId="1" pageNumber="40" value="2010-11-03">
<collectingDate id="A72FE9C1FFE4FF9CFC78A12BFBE4B69E" box="[991,1120,624,645]" pageId="1" pageNumber="40" value="2010-11-03">03 Nov.2010</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="C36A36E9FFE4FF9CFC8BA1F2FB9AB0E4" blockId="1.[812,1452,681,1279]" pageId="1" pageNumber="40">
<emphasis id="F1A1EAFBFFE4FF9CFC8BA1F2FC7EB6A5" bold="true" box="[812,1018,681,702]" pageId="1" pageNumber="40">
<emphasis id="F1A1EAFBFFE4FF9CFC8BA1F2FC63B6A5" bold="true" box="[812,999,681,702]" italics="true" pageId="1" pageNumber="40">Non-type material.</emphasis>
–
</emphasis>
<materialsCitation id="73BD3CB4FFE4FF9CFBA6A1F1FA23B70E" collectingDate="1969-09" collectionCode="ZRC" collectorName="Nee Soon" country="Singapore" location="SINGAPORE" pageId="1" pageNumber="40" specimenCode="ZRC 1989.1109, 1111, ZRC 1989.1135, 1144, ZRC 1990.1528" specimenCount="9">
3 D SH 42.1 x SD 24.9–SH 42.9 x SD 26.4 (
<specimenCode id="93739E92FFE4FF9CFC94A19DFC57B6C0" box="[819,979,710,731]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC 1989.1109</specimenCode>
–
<specimenCode id="93739E92FFE4FF9CFC47A19DFB8AB6C0" box="[992,1038,710,731]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">1111</specimenCode>
), no data; 10 D (
<specimenCount id="D5D3FD60FFE4FF9CFB0BA19DFB63B6C0" box="[1196,1255,710,731]" pageId="1" pageNumber="40" type="generic">9 juv.</specimenCount>
), SH 42.7 x SD 26.5 (
<specimenCode id="93739E92FFE4FF9CFC94A1B8FC56B6E3" box="[819,978,739,760]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC 1989.1135</specimenCode>
–
<specimenCode id="93739E92FFE4FF9CFC78A1B8FB8BB6E3" box="[991,1039,739,760]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">1144</specimenCode>
), no data.
<emphasis id="F1A1EAFBFFE4FF9CFBD4A1B8FA86B6E3" bold="true" box="[1139,1282,739,760]" pageId="1" pageNumber="40">
<collectingCountry id="BBC27679FFE4FF9CFBD4A1B8FA86B6E3" box="[1139,1282,739,760]" name="Singapore" pageId="1" pageNumber="40">SINGAPORE</collectingCountry>
</emphasis>
: 1 D (juv.) (
<specimenCode id="93739E92FFE4FF9CFADCA1B8FC12B70E" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC 1990.1528</specimenCode>
),
<collectorName id="6E20533FFFE4FF9CFC0FA05BFB8EB70E" box="[936,1034,768,789]" pageId="1" pageNumber="40">Nee Soon</collectorName>
, on tree along road, coll. CLM,
<date id="B76B1029FFE4FF9CFAEDA05BFA23B70E" box="[1354,1447,768,789]" pageId="1" pageNumber="40" value="1969-09">
<collectingDate id="A72FE9C1FFE4FF9CFAEDA05BFA23B70E" box="[1354,1447,768,789]" pageId="1" pageNumber="40" value="1969-09">Sep.1969</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFC8BA046FBE3B754" collectingDate="1988-09-09" collectionCode="ZRC" collectorName="C. M. Yang" country="Singapore" location="Nee Soon Swamp Forest" pageId="1" pageNumber="40" specimenCode="ZRC 1998.3373, 3377" specimenCount="1">
3 S (juv.) (
<specimenCode id="93739E92FFE4FF9CFC30A046FBBCB729" box="[919,1080,797,818]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC 1998.3373</specimenCode>
-
<specimenCode id="93739E92FFE4FF9CFB98A046FBF5B729" box="[1087,1137,797,818]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">3377</specimenCode>
),
<location id="C60A6032FFE4FF9CFB25A046FAF2B729" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFB25A046FAF2B729" box="[1154,1398,797,818]" country="Singapore" name="Nee Soon Swamp Forest" pageId="1" pageNumber="40">Nee Soon Swamp Forest</location>
, coll.
<collectorName id="6E20533FFFE4FF9CFC8BA061FC22B754" box="[812,934,826,847]" pageId="1" pageNumber="40">C. M. Yang</collectorName>
et al.,
<date id="B76B1029FFE4FF9CFC55A061FBE3B754" box="[1010,1127,826,847]" pageId="1" pageNumber="40" value="1988-09-09">
<collectingDate id="A72FE9C1FFE4FF9CFC55A061FBE3B754" box="[1010,1127,826,847]" pageId="1" pageNumber="40" value="1988-09-09">9 Sep.1988</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFBD3A061FB46B793" collectingDate="1990-05-09" collectionCode="ZRC" collectorName="K. Lim" country="Singapore" location="Nee Soon Swamp Forest" pageId="1" pageNumber="40" specimenCode="ZRC 1990.10741" specimenCount="1">
1 D SH 44.3 x SD 25.4 (
<specimenCode id="93739E92FFE4FF9CFADDA061FC26B770" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC 1990.10741</specimenCode>
),
<location id="C60A6032FFE4FF9CFC15A00DFB25B770" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFC15A00DFB25B770" box="[946,1185,854,875]" country="Singapore" name="Nee Soon Swamp Forest" pageId="1" pageNumber="40">Nee Soon Swamp Forest</location>
, NW sector, in leaf litter on open scrubland, coll.
<collectorName id="6E20533FFFE4FF9CFC5DA028FBC5B793" box="[1018,1089,883,904]" pageId="1" pageNumber="40">K. Lim</collectorName>
,
<date id="B76B1029FFE4FF9CFBECA028FB46B793" box="[1099,1218,883,904]" pageId="1" pageNumber="40" value="1990-05-09">
<collectingDate id="A72FE9C1FFE4FF9CFBECA028FB46B793" box="[1099,1218,883,904]" pageId="1" pageNumber="40" value="1990-05-09">9 May 1990</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFB6BA028FBCFB7D9" collectionCode="ZRC" collectorName="S. K. Tan" country="Singapore" location="Nee Soon Swamp Forest" pageId="1" pageNumber="40" specimenCount="1">
3 S (juv.), 2 D (broken, not measured) (
<collectionCode id="A5C4AE2CFFE4FF9CFC18A0CBFC75B7BE" box="[959,1009,912,933]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
.MOL.3061)
<location id="C60A6032FFE4FF9CFBCAA0CBFADDB7BE" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFBCAA0CBFADDB7BE" box="[1133,1369,912,933]" country="Singapore" name="Nee Soon Swamp Forest" pageId="1" pageNumber="40">Nee Soon Swamp Forest</location>
, dead on ground, coll.
<collectorName id="6E20533FFFE4FF9CFC09A0F6FB8AB7D9" box="[942,1038,941,962]" pageId="1" pageNumber="40">S. K. Tan</collectorName>
, 2009
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFBF1A0F6FC16B7E0" collectingDate="1993-12-25" collectionCode="CSY" collectorName="S. Y. Chan" country="Singapore" location="Pulau Tekong" pageId="1" pageNumber="40" specimenCount="1">
2 D (juv.) (
<collectionCode id="A5C4AE2CFFE4FF9CFB64A0F6FB77B7D9" box="[1219,1267,941,962]" pageId="1" pageNumber="40">CSY</collectionCode>
409.3.4.30),
<location id="C60A6032FFE4FF9CFAD3A0F6FCFCB7C4" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFAD3A0F6FCFCB7C4" country="Singapore" name="Pulau Tekong" pageId="1" pageNumber="40">Pulau Tekong</location>
, dead on ground among low shrubs, coll.
<collectorName id="6E20533FFFE4FF9CFABFA091FA03B7C4" box="[1304,1415,970,991]" pageId="1" pageNumber="40">S. Y. Chan</collectorName>
,
<date id="B76B1029FFE4FF9CFA34A091FC16B7E0" pageId="1" pageNumber="40" value="1993-12-25">
<collectingDate id="A72FE9C1FFE4FF9CFA34A091FC16B7E0" pageId="1" pageNumber="40" value="1993-12-25">25 Dec. 1993</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFC3BA0BDFA23B003" collectingDate="1994-10-25" collectionCode="CSY" collectorName="S. Y. Chan" country="Singapore" location="Dairy Farm" pageId="1" pageNumber="40" specimenCount="1">
1 S SH 43.7 x SD 26.2 (
<collectionCode id="A5C4AE2CFFE4FF9CFB22A0BDFB31B7E0" box="[1157,1205,998,1019]" pageId="1" pageNumber="40">CSY</collectionCode>
409.3.4.31),
<location id="C60A6032FFE4FF9CFA92A0BDFA23B7E0" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFA92A0BDFA23B7E0" box="[1333,1447,998,1019]" country="Singapore" name="Dairy Farm" pageId="1" pageNumber="40">Dairy Farm</location>
, dead on road path beside forest, coll.
<collectorName id="6E20533FFFE4FF9CFB01A758FA92B003" box="[1190,1302,1027,1048]" pageId="1" pageNumber="40">S. Y. Chan</collectorName>
,
<date id="B76B1029FFE4FF9CFA84A758FA23B003" box="[1315,1447,1027,1048]" pageId="1" pageNumber="40" value="1994-10-25">
<collectingDate id="A72FE9C1FFE4FF9CFA84A758FA23B003" box="[1315,1447,1027,1048]" pageId="1" pageNumber="40" value="1994-10-25">25 Oct. 1994</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFC8BA77BFC67B049" collectingDate="1995-04" collectorName="R. Koh" country="Singapore" location="Central Catchment area" pageId="1" pageNumber="40" specimenCount="1">
1 S SH 39.5 x SD 24.8,
<location id="C60A6032FFE4FF9CFBBAA77BFA81B02E" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFBBAA77BFA81B02E" box="[1053,1285,1056,1077]" country="Singapore" name="Central Catchment area" pageId="1" pageNumber="40">Central Catchment area</location>
, on artifact, coll,
<collectorName id="6E20533FFFE4FF9CFC8BA766FCF1B049" box="[812,885,1085,1106]" pageId="1" pageNumber="40">R. Koh</collectorName>
,
<date id="B76B1029FFE4FF9CFCD8A766FC67B049" box="[895,995,1085,1106]" pageId="1" pageNumber="40" value="1995-04">
<collectingDate id="A72FE9C1FFE4FF9CFCD8A766FC67B049" box="[895,995,1085,1106]" pageId="1" pageNumber="40" value="1995-04">Apr. 1995</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFC4AA766FA17B074" accessionNumber="TSK 11017" collectingDate="1999-04-04" collectionCode="TSK" collectorName="S. K. Tan" country="Singapore" location="Mandai Road Track" pageId="1" pageNumber="40" specimenCount="1">
1 D SH 37.4 x SD 25.1 (
<collectionCode id="A5C4AE2CFFE4FF9CFB7BA766FA8FB049" box="[1244,1291,1085,1106]" pageId="1" pageNumber="40">TSK</collectionCode>
11017),
<location id="C60A6032FFE4FF9CFAC5A766FC24B074" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFAC5A766FC24B074" country="Singapore" name="Mandai Road Track" pageId="1" pageNumber="40">Mandai Road Track</location>
7, on fishtail palm, coll.
<collectorName id="6E20533FFFE4FF9CFB05A701FA82B074" box="[1186,1286,1114,1135]" pageId="1" pageNumber="40">S. K. Tan</collectorName>
,
<date id="B76B1029FFE4FF9CFAB4A701FA17B074" box="[1299,1427,1114,1135]" pageId="1" pageNumber="40" value="1999-04-04">
<collectingDate id="A72FE9C1FFE4FF9CFAB4A701FA17B074" box="[1299,1427,1114,1135]" pageId="1" pageNumber="40" value="1999-04-04">04 Apr.1999</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="73BD3CB4FFE4FF9CFA38A701FA98B0B3" collectingDate="2010-07-23" collectionCode="ZRC" collectorName="A. F. S. L. Lok" country="Singapore" location="Lower Peirce Reservoir" pageId="1" pageNumber="40" specimenCount="1">
2 D (juv.), (
<collectionCode id="A5C4AE2CFFE4FF9CFC28A72DFC46B090" box="[911,962,1142,1163]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
.MOL.3065),
<location id="C60A6032FFE4FF9CFBE1A72DFAB5B090" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFBE1A72DFAB5B090" box="[1094,1329,1142,1163]" country="Singapore" name="Lower Peirce Reservoir" pageId="1" pageNumber="40">Lower Peirce Reservoir</location>
, arboreal on leaves, forest edge, coll.
<collectorName id="6E20533FFFE4FF9CFBB3A7C8FB27B0B3" box="[1044,1187,1171,1192]" pageId="1" pageNumber="40">A. F. S. L. Lok</collectorName>
,
<date id="B76B1029FFE4FF9CFB0CA7C8FA98B0B3" box="[1195,1308,1171,1192]" pageId="1" pageNumber="40" value="2010-07-23">
<collectingDate id="A72FE9C1FFE4FF9CFB0CA7C8FA98B0B3" box="[1195,1308,1171,1192]" pageId="1" pageNumber="40" value="2010-07-23">23 Jul.2010</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="73BD3CB4FFE4FF9CFA83A7C8FB9EB0E4" collectingDate="1996-10-13" collectionCode="CSY" collectorName="S. Y. Chan" country="Malaysia" location="Mount Ophir" municipality="Gunung Ledang" pageId="1" pageNumber="40" specimenCount="1" stateProvince="Johor">
<emphasis id="F1A1EAFBFFE4FF9CFA83A7C8FA20B0B3" bold="true" box="[1316,1444,1171,1192]" pageId="1" pageNumber="40">
<collectingCountry id="BBC27679FFE4FF9CFA83A7C8FA20B0B3" box="[1316,1444,1171,1192]" name="Malaysia" pageId="1" pageNumber="40">MALAYSIA</collectingCountry>
</emphasis>
: 1 D (broken, not measured) (
<collectionCode id="A5C4AE2CFFE4FF9CFBF4A7EBFB07B0DE" box="[1107,1155,1200,1221]" pageId="1" pageNumber="40">CSY</collectionCode>
409.3.4.14),
<collectingMunicipality id="230EAC93FFE4FF9CFAACA7EBFA2FB0DE" box="[1291,1451,1200,1221]" pageId="1" pageNumber="40">Gunung Ledang</collectingMunicipality>
(
<location id="C60A6032FFE4FF9CFC94A796FC33B0F9" LSID="urn:lsid:plazi:treatment:4B7C87FFFFE4FF9EFF29A53BFE8FB6EC:C60A6032FFE4FF9CFC94A796FC33B0F9" box="[819,951,1229,1250]" country="Malaysia" municipality="Gunung Ledang" name="Mount Ophir" pageId="1" pageNumber="40" stateProvince="Johor">Mount Ophir</location>
),
<collectingRegion id="0111F80BFFE4FF9CFC6FA796FB84B0F9" box="[968,1024,1229,1250]" country="Malaysia" name="Johor" pageId="1" pageNumber="40">Johor</collectingRegion>
, dead, among leaf litter in forest, coll.
<collectorName id="6E20533FFFE4FF9CFADDA796FCDBB0E4" pageId="1" pageNumber="40">S. Y. Chan</collectorName>
et al,
<date id="B76B1029FFE4FF9CFC38A7B1FB9EB0E4" box="[927,1050,1258,1279]" pageId="1" pageNumber="40" value="1996-10-13">
<collectingDate id="A72FE9C1FFE4FF9CFC38A7B1FB9EB0E4" box="[927,1050,1258,1279]" pageId="1" pageNumber="40" value="1996-10-13">13 Oct.1996</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BCF6562FFE4FF9CFC8BA673FBD8B364" pageId="1" pageNumber="40" type="diagnosis">
<paragraph id="C36A36E9FFE4FF9CFC8BA673FB9BB264" blockId="1.[812,1451,1320,1663]" pageId="1" pageNumber="40">
<emphasis id="F1A1EAFBFFE4FF9CFC8BA673FC37B15B" bold="true" box="[812,947,1320,1344]" italics="true" pageId="1" pageNumber="40">Diagnosis. –</emphasis>
Shell relatively large, ovate, chirally dimorphic coiling. Whorls 5½–6¾, slightly convex, surface generally smooth with indistinct axial striations or growth lines. Colour uniformly yellow with a thin white subsutural line (shells of living animals yellowish-green due to the dark colouration of soft body showing through), varix always absent. Columella simple, straight. Periostracum thin, transparent. Peristome in mature animals white, thickened and expanded except for parietal side, outer lip reflected but not adnate. Parietal callus usually inconspicuous, very rarely thickened to being whitish or translucent.
</paragraph>
<paragraph id="C36A36E9FFE4FF9CFC8BA5F3FBD8B364" blockId="1.[812,1451,1704,1919]" pageId="1" pageNumber="40">Animal body very pale greyish white in colour, reticulated, the recessed parts being darker, light brown around the head to greyish on the body. Mantle edge dark cream coloured to brown around the peristome. Upper and lower tentacles dark yellow, central part of the head between the upper tentacles occasionally pigmented with yellow, foot with longitudinal yellow stripe on either side.</paragraph>
</subSubSection>
<subSubSection id="8BCF6562FFE4FF9FFC8BA4F3FBAFB56C" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40" type="distribution">
<paragraph id="C36A36E9FFE4FF9FFC8BA4F3FBAFB56C" blockId="1.[812,1452,1960,2111]" lastBlockId="2.[835,1474,192,375]" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">
<emphasis id="F1A1EAFBFFE4FF9CFC8BA4F3FC49B3DB" bold="true" box="[812,973,1960,1984]" italics="true" pageId="1" pageNumber="40">Distribution. –</emphasis>
The distribution of
<taxonomicName id="04D54D6AFFE4FF9CFB0EA4F3FA20B3A4" authorityName="Tan &amp; Chan &amp; Panha" authorityYear="2011" box="[1193,1444,1960,1983]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="40" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="temasek">
<emphasis id="F1A1EAFBFFE4FF9CFB0EA4F3FA20B3A4" box="[1193,1444,1960,1983]" italics="true" pageId="1" pageNumber="40">A. atricallosus temasek</emphasis>
</taxonomicName>
, new subspecies, is largely confined to
<collectingCountry id="BBC27679FFE4FF9CFB5AA493FAF7B3C4" box="[1277,1395,1992,2015]" name="Singapore" pageId="1" pageNumber="40">Singapore</collectingCountry>
, and tentatively,
<collectingRegion id="0111F80BFFE4FF9CFC00A4B3FC60B3E4" box="[935,996,2024,2047]" country="Malaysia" name="Johor" pageId="1" pageNumber="40">Johor</collectingRegion>
, southern Peninsular
<collectingCountry id="BBC27679FFE4FF9CFB64A4B3FAA0B3E4" box="[1219,1316,2024,2047]" name="Malaysia" pageId="1" pageNumber="40">Malaysia</collectingCountry>
(Sutcharit &amp; Panha, 2006 [part];
<bibRefCitation id="A7444B18FFE4FF9CFC5AAB53FB63BC04" author="Prasankok, P &amp; Ota, M" box="[1021,1255,2056,2079]" pageId="1" pageNumber="40" pagination="189 - 197" refId="ref7006" refString="Prasankok, P., H. Ota, M. Toda &amp; S. Panha, 2007. Allozyme variation in the Camaenid tree snails Amphidromus atricallosus (Gould, 1843) and A. inversus (Muller, 1774). Zoological Science, 24: 189 - 197." type="journal article" year="2007">Prasankok et al., 2007</bibRefCitation>
[part]; this study). Known
<collectingCountry id="BBC27679FFE4FF9CFCD8AB73FC6DBC24" box="[895,1001,2088,2111]" name="Singapore" pageId="1" pageNumber="40">Singapore</collectingCountry>
localities include: Seletar (
<bibRefCitation id="A7444B18FFE4FF9CFB59AB73FA1BBC24" author="Hanitsch, R" box="[1278,1439,2088,2111]" pageId="1" pageNumber="40" refId="ref6619" refString="Hanitsch, R., 1908. Guide to the Zoological Collections of the Raffles Museum, Singapore. Straits Times Press, Limited, Singapore. 21 pls + 112 pp." type="book" year="1908">Hanitsch, 1908</bibRefCitation>
), Nee Soon Swamp Forest, Bukit Timah Nature Reserve, Mandai, and Pulau Tekong (Ng &amp; Lim, 1992; Chou &amp; Tan, 2008; Lok &amp; Tan, 2008). Although a specimen from
<collectingRegion id="0111F80BFFE7FF9FFF2FA27BFF47B52C" box="[136,195,288,311]" country="Malaysia" name="Johor" pageId="2" pageNumber="41">Johor</collectingRegion>
(CSY 409.3.4.14) is provisionally determined to be this subspecies in the material examined, identification was based on a weathered and broken specimen with only the body whorl intact. Verification with better preserved material would be required. Available evidence suggests
<taxonomicName id="04D54D6AFFE7FF9FFF2FA29BFE0EB5CC" authorityName="Tan &amp; Chan &amp; Panha" authorityYear="2011" box="[136,394,448,471]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="41" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="temasek">
<emphasis id="F1A1EAFBFFE7FF9FFF2FA29BFE0EB5CC" box="[136,394,448,471]" italics="true" pageId="2" pageNumber="41">A. atricallosus temasek</emphasis>
</taxonomicName>
to be endemic, but this has to be verified with greater sampling effort, particularly around the southern part of Peninsular
<collectingCountry id="BBC27679FFE7FF9FFBCFA39BFB4EB4CC" box="[1128,1226,192,215]" name="Malaysia" pageId="2" pageNumber="41">Malaysia</collectingCountry>
and the
<collectingRegion id="0111F80BFFE7FF9FFA84A39BFA21B4CC" box="[1315,1445,192,215]" country="Indonesia" name="Kepulauan Riau" pageId="2" pageNumber="41">Riau islands</collectingRegion>
of
<collectingCountry id="BBC27679FFE7FF9FFCE4A3BBFC2AB4EC" box="[835,942,224,247]" name="Indonesia" pageId="2" pageNumber="41">Indonesia</collectingCountry>
. In
<collectingCountry id="BBC27679FFE7FF9FFC73A3BBFBC7B4EC" box="[980,1091,224,247]" name="Singapore" pageId="2" pageNumber="41">Singapore</collectingCountry>
, these arboreal snails are apparently restricted to the remnant forests. Preference for particular tree species has not been observed or reported, and the animals can be found on a wide variety of plants and on man-made structures.
</paragraph>
</subSubSection>
<subSubSection id="8BCF6562FFE7FF9FFCE4A2FBFB31B5EC" pageId="2" pageNumber="41" type="etymology">
<paragraph id="C36A36E9FFE7FF9FFCE4A2FBFB31B5EC" blockId="2.[835,1475,416,503]" pageId="2" pageNumber="41">
<emphasis id="F1A1EAFBFFE7FF9FFCE4A2FBFC65B5A3" bold="true" box="[835,993,416,440]" italics="true" pageId="2" pageNumber="41">Etymology. –</emphasis>
The subspecies epithet is derived from Temasek, the historical name of the
<typeStatus id="1C6E884BFFE7FF9FFB63A29BFB76B5CC" box="[1220,1266,448,471]" pageId="2" pageNumber="41">type</typeStatus>
locality
<collectingCountry id="BBC27679FFE7FF9FFAE8A29BFA3BB5CC" box="[1359,1471,448,471]" name="Singapore" pageId="2" pageNumber="41">Singapore</collectingCountry>
. It is used as a noun in apposition.
</paragraph>
</subSubSection>
<caption id="97AA6661FFE7FF9FFF2FA4C7FC55BC5A" ID-DOI="http://doi.org/10.5281/zenodo.10106684" ID-Zenodo-Dep="10106684" httpUri="https://zenodo.org/record/10106684/files/figure.png" pageId="2" pageNumber="41" startId="2.[136,171,1948,1969]" targetBox="[137,1474,565,1922]" targetPageId="2">
<paragraph id="C36A36E9FFE7FF9FFF2FA4C7FC55BC5A" blockId="2.[136,1474,1948,2113]" pageId="2" pageNumber="41">
Fig. 1.
<emphasis id="F1A1EAFBFFE7FF9FFF68A4C7FF64B3AA" bold="true" box="[207,224,1948,1969]" pageId="2" pageNumber="41">A</emphasis>
,
<taxonomicName id="04D54D6AFFE7FF9FFF49A4C7FDF3B3AA" authority="Tan &amp; Chan &amp; Panha, 2011" authorityName=", new" authorityYear="2011" box="[238,631,1948,1969]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="39" phylum="Mollusca" rank="subSpecies" species="atricallosus" status="new" subSpecies="temasek">
<emphasis id="F1A1EAFBFFE7FF9FFF49A4C7FDC5B3AA" box="[238,577,1948,1969]" italics="true" pageId="2" pageNumber="41">Amphidromus atricallosus temasek</emphasis>
,
<taxonomicNameLabel id="EA925780FFE7FF9FFDE8A4C7FDF3B3AA" box="[591,631,1948,1969]" pageId="2" pageNumber="41" rank="subSpecies">new</taxonomicNameLabel>
</taxonomicName>
subspecies: Holotype, SH 44 x SD 25.6 (ZRC.MOL.3058), Nee Soon Swamp Forest, Singapore;
<emphasis id="F1A1EAFBFFE7FF9FFF5EA4E3FEA1B3D6" bold="true" box="[249,293,1976,1997]" pageId="2" pageNumber="41">B–E</emphasis>
<taxonomicName id="04D54D6AFFE7FF9FFE8AA4E2FCB1B3D5" authority="(Fulton, 1901)" baseAuthorityName="Fulton" baseAuthorityYear="1901" box="[301,821,1977,1998]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="41" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="perakensis">
<emphasis id="F1A1EAFBFFE7FF9FFE8AA4E2FD18B3D5" box="[301,668,1977,1998]" italics="true" pageId="2" pageNumber="41">Amphidromus atricallosus perakensis</emphasis>
(
<bibRefCitation id="A7444B18FFE7FF9FFD0CA4E2FCAAB3D5" author="Fulton, H" box="[683,814,1977,1998]" pageId="2" pageNumber="41" pagination="103 - 104" refId="ref6471" refString="Fulton, H., 1901. Descriptions of new species of Xesta, Amphidromus and Cyclostoma from Madagascar and Perak. Journal of Malacology, 8: 103 - 104, Pl. IX." type="journal article" year="1901">Fulton, 1901</bibRefCitation>
)
</taxonomicName>
:
<emphasis id="F1A1EAFBFFE7FF9FFCE6A4E3FCD5B3D6" bold="true" box="[833,849,1976,1997]" pageId="2" pageNumber="41">B</emphasis>
, SH 45 x SD 26.4 (TSK 11041), Bukit Serdam, Pahang, West Malaysia;
<emphasis id="F1A1EAFBFFE7FF9FFF49A48EFF7BB3F1" bold="true" box="[238,255,2005,2026]" pageId="2" pageNumber="41">C</emphasis>
, SH 47.1 x SD 25.3 (CBB), Tapah Hill, Perak, West Malaysia;
<emphasis id="F1A1EAFBFFE7FF9FFCDEA48EFC0EB3F1" bold="true" box="[889,906,2005,2026]" pageId="2" pageNumber="41">D</emphasis>
, SH 47.2 x SD 27.6 (ZRC.MOL.3064), Pulau Tulai, near Pulau Tioman, West Malaysia;
<emphasis id="F1A1EAFBFFE7FF9FFE6EA4A9FE5DBC1C" bold="true" box="[457,473,2034,2055]" pageId="2" pageNumber="41">E</emphasis>
, SH 46.4 x SD 24.2 (ZRC.MOL.3062), Gunung Genting, Perak, West Malaysia;
<emphasis id="F1A1EAFBFFE7FF9FFABBA4A9FAAFBC1C" bold="true" box="[1308,1323,2034,2055]" pageId="2" pageNumber="41">F</emphasis>
,
<taxonomicName id="04D54D6AFFE7FF9FFA9DA4A8FD85BC3F" authority="atricallosus (Gould, 1843)" authorityName="atricallosus (Gould" authorityYear="1843" baseAuthorityName="Gould" baseAuthorityYear="1843" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="41" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="atricallosus">
<emphasis id="F1A1EAFBFFE7FF9FFA9DA4A8FEF4BC3F" italics="true" pageId="2" pageNumber="41">Amphidromus atricallosus atricallosus</emphasis>
(
<bibRefCitation id="A7444B18FFE7FF9FFEDAAB54FE7EBC3F" author="Gould, A" box="[381,506,2063,2084]" pageId="2" pageNumber="41" pagination="139 - 141" refId="ref6507" refString="Gould, A. A., 1843. [S] hells not long since announced as having been received from Rev. Francis Mason, missionary at Tavoy, in British Burmah. In: Regular meeting - Dr. C. T Jackson, Vice-President, in the Chair. September 6 th, 1843. Proceedings of the Boston Society of Natural History, 1: 139 - 141." type="journal article" year="1843">Gould, 1843</bibRefCitation>
)
</taxonomicName>
: SH 45 x SD 25.7 (CSY 409.4.4.16), Ranong, Thailand;
<emphasis id="F1A1EAFBFFE7FF9FFB85AB54FBB1BC3F" bold="true" box="[1058,1077,2063,2084]" pageId="2" pageNumber="41">G</emphasis>
,
<taxonomicName id="04D54D6AFFE7FF9FFBE7AB54FED0BC5A" authority="(von Martens, 1864)" baseAuthorityName="von Martens" baseAuthorityYear="1864" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="41" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="leucoxanthus">
<emphasis id="F1A1EAFBFFE7FF9FFBE7AB54FA46BC3F" box="[1088,1474,2063,2084]" italics="true" pageId="2" pageNumber="41">Amphidromus atricallosus leucoxanthus</emphasis>
(von
<bibRefCitation id="A7444B18FFE7FF9FFF1BAB77FEC8BC5A" author="Martens, E" box="[188,332,2092,2113]" pageId="2" pageNumber="41" pagination="523 - 529" refId="ref6857" refString="Martens, E. von, 1864. Diagnosen neuer Heliceen aus dem ostasiatischen Archipel. Monatsberichte der Koniglichen Preussischen Akademien der Wissenschaften zu Berlin, 1861: 523 - 529." type="journal article" year="1864">Martens, 1864</bibRefCitation>
)
</taxonomicName>
: SH 51 x SD 28.5 (CSY 409.3.49.1), Chantaburi, East Thailand.
</paragraph>
</caption>
<subSubSection id="8BCF6562FFE6FF9EFFD6A39BFE8FB6EC" pageId="3" pageNumber="42" type="discussion">
<paragraph id="C36A36E9FFE6FF9EFFD6A39BFE8FB6EC" blockId="3.[113,752,192,759]" pageId="3" pageNumber="42">
<emphasis id="F1A1EAFBFFE6FF9EFFD6A39BFF74B4C3" bold="true" box="[113,240,192,216]" italics="true" pageId="3" pageNumber="42">Remarks. –</emphasis>
The thin white subsutural line and simple and straight columella that is never twisted is diagnostic. Shells of juvenile
<emphasis id="F1A1EAFBFFE6FF9EFF49A25BFF7CB50C" box="[238,248,256,279]" italics="true" pageId="3" pageNumber="42">s</emphasis>
can also be separated from
<taxonomicName id="04D54D6AFFE6FF9EFDEFA25BFEF1B52C" authority="Fulton, 1901" authorityName="Fulton" authorityYear="1901" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="42" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="perakensis">
<emphasis id="F1A1EAFBFFE6FF9EFDEFA25BFF67B52C" italics="true" pageId="3" pageNumber="42">A. atricallosus perakensis</emphasis>
<bibRefCitation id="A7444B18FFE6FF9EFF4DA27BFEF1B52C" author="Fulton, H" box="[234,373,288,311]" pageId="3" pageNumber="42" pagination="103 - 104" refId="ref6471" refString="Fulton, H., 1901. Descriptions of new species of Xesta, Amphidromus and Cyclostoma from Madagascar and Perak. Journal of Malacology, 8: 103 - 104, Pl. IX." type="journal article" year="1901">Fulton, 1901</bibRefCitation>
</taxonomicName>
, by lack of a white zone around the columella (see
<figureCitation id="5BEE2A6CFFE6FF9EFEB6A21BFED5B54C" box="[273,337,320,343]" captionStart="Fig" captionStartId="3.[114,149,2035,2056]" captionTargetBox="[113,1452,1623,2009]" captionTargetId="figure-421@3.[113,1452,1623,2010]" captionTargetPageId="3" captionText="Fig. 2. A–C, Shells of juveniles: A, SH 31 x SD 19.7, juvenile shell of Amphidromus atricallosus temasek, new subspecies, showing the white columella; B–C, Amphidromus atricallosus perakensis (Fulton, 1901), juveniles, has a white zone bordering the columella: B, SH 33.9 x SD 22.4, Tapah Hill, Perak, West Malaysia; C, SH 34.7 x SD 25, Pulau Tulai, West Malaysia." figureDoi="http://doi.org/10.5281/zenodo.10106686" httpUri="https://zenodo.org/record/10106686/files/figure.png" pageId="3" pageNumber="42">Fig. 2</figureCitation>
). Although the convexity of the whorls of
<taxonomicName id="04D54D6AFFE6FF9EFF2BA23BFE19B56C" authorityName="Fulton" authorityYear="1901" box="[140,413,352,375]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="42" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="perakensis">
<emphasis id="F1A1EAFBFFE6FF9EFF2BA23BFE19B56C" box="[140,413,352,375]" italics="true" pageId="3" pageNumber="42">A. atricallosus perakensis</emphasis>
</taxonomicName>
is evidently variable throughout West
<collectingCountry id="BBC27679FFE6FF9EFF16A2DBFE97B58C" box="[177,275,384,407]" name="Malaysia" pageId="3" pageNumber="42">Malaysia</collectingCountry>
populations, shell whorls of
<taxonomicName id="04D54D6AFFE6FF9EFDF5A2DBFF43B5AC" authorityName=", new" authorityYear="2011" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="42" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="temasek">
<emphasis id="F1A1EAFBFFE6FF9EFDF5A2DBFF43B5AC" italics="true" pageId="3" pageNumber="42">A. atricallosus temasek</emphasis>
</taxonomicName>
, new subspecies, seem to be generally more convex in profile and resulting in the suture appearing more sunken in comparison. Living
<taxonomicName id="04D54D6AFFE6FF9EFEFAA2BBFDD4B5EC" authorityName=", new" authorityYear="2011" box="[349,592,480,503]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="42" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="temasek">
<emphasis id="F1A1EAFBFFE6FF9EFEFAA2BBFDD4B5EC" box="[349,592,480,503]" italics="true" pageId="3" pageNumber="42">A. atricallosus temasek</emphasis>
</taxonomicName>
individuals are easily distinguished by the soft body colouration, and a pale yellowish-green shell with a thin white subsutural line (
<figureCitation id="5BEE2A6CFFE6FF9EFD62A17BFF22B64C" captionStart="Fig" captionStartId="4.[137,172,2063,2084]" captionTargetBox="[136,1474,229,2023]" captionTargetId="figure-43@4.[138,1474,229,1137]" captionTargetPageId="4" captionText="Fig. 3. A–B, Live animal colouration: A, Amphidromus atricallosus temasek, new subspecies, holotype (ZRC.MOL.3058); B, Amphidromus atricallosus perakensis (Fulton, 1901), Pulau Tulai, West Malaysia. (Photographs by Tan Heok Hui)" figureDoi="http://doi.org/10.5281/zenodo.10106692" httpUri="https://zenodo.org/record/10106692/files/figure.png" pageId="3" pageNumber="42">Fig. 3. A</figureCitation>
). The SEM images of the radula and genital system of
<taxonomicName id="04D54D6AFFE6FF9EFF35A13BFE16B66C" authorityName=", new" authorityYear="2011" box="[146,402,608,631]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="42" phylum="Mollusca" rank="subSpecies" species="atricallosus" subSpecies="temasek">
<emphasis id="F1A1EAFBFFE6FF9EFF35A13BFE16B66C" box="[146,402,608,631]" italics="true" pageId="3" pageNumber="42">A. atricallosus temasek</emphasis>
</taxonomicName>
, new subspecies, are described in detail and figured in Sutcharit &amp; Panha (2006, as
<emphasis id="F1A1EAFBFFE6FF9EFD0FA1DBFD44B68C" box="[680,704,640,663]" italics="true" pageId="3" pageNumber="42">A.</emphasis>
<taxonomicName id="04D54D6AFFE6FF9EFD60A1DBFF77B6AC" baseAuthorityName="Gould" baseAuthorityYear="1843" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="42" phylum="Mollusca" rank="species" species="atricallosus" subGenus="Amphidromus">
(
<emphasis id="F1A1EAFBFFE6FF9EFD77A1DBFD64B68C" box="[720,736,640,663]" italics="true" pageId="3" pageNumber="42">A</emphasis>
.)
<emphasis id="F1A1EAFBFFE6FF9EFFD6A1FBFF77B6AC" box="[113,243,672,695]" italics="true" pageId="3" pageNumber="42">atricallosus</emphasis>
</taxonomicName>
<taxonomicName id="04D54D6AFFE6FF9EFF59A1FBFEF0B6AC" authorityName="Fulton" authorityYear="1901" box="[254,372,672,695]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="42" phylum="Mollusca" rank="species" species="perakensis">
<emphasis id="F1A1EAFBFFE6FF9EFF59A1FBFEF0B6AC" box="[254,372,672,695]" italics="true" pageId="3" pageNumber="42">perakensis</emphasis>
</taxonomicName>
[CUMZ 2198; herein designated
<typeStatus id="1C6E884BFFE6FF9EFFD6A19BFF59B6CC" box="[113,221,704,727]" pageId="3" pageNumber="42" type="paratype">paratypes</typeStatus>
]; radula, pg. 16: Fig. 11A–C; genital system, pg. 17: Fig. 12C).
</paragraph>
</subSubSection>
</treatment>
</document>
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